178 lines
21 KiB
XML
178 lines
21 KiB
XML
<document id="FFBCE85389BCB71756F3157FA4F52CBE" ID-CLB-Dataset="50962" ID-DOI="10.5281/zenodo.273298" ID-GBIF-Dataset="ef5f0237-93e1-45ca-8022-0022a946cd35" ID-ISSN="11755326" ID-Zenodo-Dep="273298" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459746912548" checkinUser="plazi" docAuthor="García, Alfonso N. & Aldrete" docDate="2005" docId="74065C270E721145FE99FEF162C0F980" docLanguage="en" docName="zt01077p060.pdf" docOrigin="Zootaxa 1077" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Incapsocus penai García & Aldrete, 2005, sp. n." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="883F245F0E701146FF91FFED6172FF8E" masterDocTitle="Three new monotypic genera of Epipsocidae (Psocoptera) from Peru and Brazil" masterLastPageNumber="60" masterPageNumber="51" pageNumber="53" updateTime="1698214001876" updateUser="ExternalLinkService">
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<mods:title id="73706B3F00029A17E336AE699687F8C3">Three new monotypic genera of Epipsocidae (Psocoptera) from Peru and Brazil</mods:title>
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<mods:namePart id="A86765090C6A2005BADA5141649DD998">García, Alfonso N.</mods:namePart>
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<mods:namePart id="9AFECF97E5F9F31456DBB0A94A8F7955">Aldrete</mods:namePart>
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<mods:title id="0792D1295A0A7987CE5AFDD0717AACEF">Zootaxa</mods:title>
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<mods:part id="6C153A1FF918B5138ABB6B227DC22887">
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<mods:date id="D819EBC611E44957844C0D3F700B8E79">2005</mods:date>
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<mods:number id="249D44D8131D2BFADB8EE6A640BA9616">1077</mods:number>
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<treatment id="74065C270E721145FE99FEF162C0F980" ID-DOI="http://doi.org/10.5281/zenodo.6269053" ID-GBIF-Taxon="119336611" ID-Zenodo-Dep="6269053" LSID="urn:lsid:plazi:treatment:74065C270E721145FE99FEF162C0F980" httpUri="http://treatment.plazi.org/id/74065C270E721145FE99FEF162C0F980" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
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<subSubSection id="B4B5BEBA0E721144FE99FEF16280FE77" pageId="2" pageNumber="53" type="nomenclature">
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<paragraph id="FC10ED310E721144FE99FEF163C7FEBB" blockId="2.[264,693,283,310]" box="[264,693,283,310]" pageId="2" pageNumber="53">
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<heading id="A7585A5D0E721144FE99FEF163C7FEBB" bold="true" box="[264,693,283,310]" fontSize="11" level="1" pageId="2" pageNumber="53" reason="1">
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<emphasis id="CEDB31230E721144FE99FEF163C7FEBB" bold="true" box="[264,693,283,310]" pageId="2" pageNumber="53">
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<emphasis id="CEDB31230E721144FE99FEF16090FEB8" bold="true" box="[264,482,284,310]" italics="true" pageId="2" pageNumber="53">
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<taxonomicName id="3BAF96B20E721144FE99FEF160AEFEB8" ID-CoL="3PL5T" box="[264,476,284,310]" class="Insecta" family="Epipsocidae" genus="Incapsocus" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="penai" status="sp. nov.">Incapsocus penai</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="D5E88C580E721144FE7BFEF6635EFEBB" box="[490,556,283,309]" pageId="2" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="6494F1B40E721144FDAAFEF663DEFEBB" box="[571,684,283,309]" captionStart="FIGURES 1 – 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[290,1316,554,1759]" captionTargetId="figure@2.[264,1323,542,1782]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Incapsocus penai. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3 – 5 to common scale." httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="2" pageNumber="53">Figs. 1–5</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="FC10ED310E721144FE99FE8B6280FE77" blockId="2.[264,1322,358,505]" pageId="2" pageNumber="53">
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<emphasis id="CEDB31230E721144FE99FE8B6022FE0E" bold="true" box="[264,336,358,384]" pageId="2" pageNumber="53">Color</emphasis>
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(in 80% alcohol). Body pale brown. Compound eyes black, ocelli hyaline, without pigmented centripetal crescents. Epicranial sulcus ochre, well defined. Wings almost hyaline, with a faint yellowish hue; veins pale brown, forewing pterostigma dark brown. Abdomen dirty white, with ochre, transverse subcuticular rings.
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</paragraph>
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</subSubSection>
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<caption id="A8D0BDB90E721144FE99F91660B6F8D6" httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="2" pageNumber="53" targetBox="[290,1316,554,1759]" targetPageId="2">
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<paragraph id="FC10ED310E721144FE99F91660B6F8D6" blockId="2.[264,1323,1787,1880]" pageId="2" pageNumber="53">
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<emphasis id="CEDB31230E721144FE99F91660C7F89D" bold="true" box="[264,437,1787,1811]" pageId="2" pageNumber="53">FIGURES 1–5.</emphasis>
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<taxonomicName id="3BAF96B20E721144FE2CF911630AF89D" box="[445,632,1788,1811]" class="Insecta" family="Epipsocidae" genus="Incapsocus" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="penai">
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<emphasis id="CEDB31230E721144FE2CF911630AF89D" box="[445,632,1788,1811]" italics="true" pageId="2" pageNumber="53">Incapsocus penai</emphasis>
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</taxonomicName>
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. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3–5 to common scale.
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</paragraph>
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</caption>
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<subSubSection id="B4B5BEBA0E731145FEA9FEF66556FC00" pageId="3" pageNumber="54" type="description">
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<paragraph id="FC10ED310E731145FEA9FEF6650EFCB0" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="54">
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<emphasis id="CEDB31230E731145FEA9FEF660AAFEBB" bold="true" box="[312,472,283,309]" pageId="3" pageNumber="54">Morphology.</emphasis>
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Epicranial sulcus with very short lateral arms. Outer cusp of lacinial tip broad, with 6 denticles (
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<figureCitation id="6494F1B40E731145FDB6FEA96303FED0" box="[551,625,324,350]" captionStart="FIGURES 1 – 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[290,1316,554,1759]" captionTargetId="figure@2.[264,1323,542,1782]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Incapsocus penai. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3 – 5 to common scale." httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="3" pageNumber="54">Fig. 2</figureCitation>
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). Five distal inner labral sensilla, of same size, in line: one central placoid and on each side of it one trichoid and one placoid. Forewing (
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<figureCitation id="6494F1B40E731145FB0EFE816596FE08" box="[1183,1252,364,390]" captionStart="FIGURES 1 – 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[290,1316,554,1759]" captionTargetId="figure@2.[264,1323,542,1782]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Incapsocus penai. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3 – 5 to common scale." httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="3" pageNumber="54">Fig. 1</figureCitation>
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) with pterostigma broadly triangular, narrow anteriorly and widening distally; Rs forked, each stem divided in two branches. Right forewing with M 5branched, left forewing with M 6 branched. Areola postica low, elongate. Hindwing (
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<figureCitation id="6494F1B40E731145FCE9FE0962B2FE70" box="[888,960,484,510]" captionStart="FIGURES 1 – 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[290,1316,554,1759]" captionTargetId="figure@2.[264,1323,542,1782]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Incapsocus penai. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3 – 5 to common scale." httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="3" pageNumber="54">Fig. 1</figureCitation>
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) with RsM joined by a long fusion, M forked. Hypandrium (
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<figureCitation id="6494F1B40E731145FD12FDE163B8FDA8" box="[643,714,524,550]" captionStart="FIGURES 1 – 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[290,1316,554,1759]" captionTargetId="figure@2.[264,1323,542,1782]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Incapsocus penai. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3 – 5 to common scale." httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="3" pageNumber="54">Fig. 4</figureCitation>
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) rounded distally, pigmented area concave, setae as illustrated, continuous along posterior border, without defined lateral fields. Phallosome (
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<figureCitation id="6494F1B40E731145FE80FDB1602AFDF8" box="[273,344,604,630]" captionStart="FIGURES 1 – 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[290,1316,554,1759]" captionTargetId="figure@2.[264,1323,542,1782]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Incapsocus penai. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3 – 5 to common scale." httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="3" pageNumber="54">Fig. 3</figureCitation>
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) simple, open anteriorly, aedeagal arch broadtipped; without external parameres, phallic sclerites absent. Clunium projected posteriorly in area over epiproct (
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<figureCitation id="6494F1B40E731145FB1FFD6965A4FD10" box="[1166,1238,644,670]" captionStart="FIGURES 1 – 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[290,1316,554,1759]" captionTargetId="figure@2.[264,1323,542,1782]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Incapsocus penai. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3 – 5 to common scale." httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="3" pageNumber="54">Fig. 5</figureCitation>
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). Paraprocts robust, with setae as illustrated, a field of microspines mesally on outer edge, and sensory field slightly elliptic, with 29–30 trichobothria issuing from basal rosettes (
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<figureCitation id="6494F1B40E731145FB45FD39646EFD60" box="[1236,1308,724,750]" captionStart="FIGURES 1 – 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[290,1316,554,1759]" captionTargetId="figure@2.[264,1323,542,1782]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Incapsocus penai. Male. 1. Fore and hindwing. 2. Lacinial tip. 3. Phallosome. 4. Hypandrium. 5. Posterior border of clunium, right paraproct and epiproct. Scales in mm. Figs. 3 – 5 to common scale." httpUri="https://zenodo.org/record/273299/files/figure.png" pageId="3" pageNumber="54">Fig. 5</figureCitation>
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). Epiproct trapeziform, with 3 setae and a field of microspines along posterior border, a field of posteromesal setae, and a group of 5 setae, on line, near anterior border.
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</paragraph>
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<paragraph id="FC10ED310E731145FEA9FCA16556FC00" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="54">
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<emphasis id="CEDB31230E731145FEA9FCA16085FCE8" bold="true" box="[312,503,844,870]" pageId="3" pageNumber="54">Measurements.</emphasis>
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FW: 4039, HW: 2920, F: 901, T: 1734, t1: 695, t2: 183, ctt1: 28, f1: 799, f2: 795, f3: 602: IO: 495, D: 302, d: 233, IO/D: 1.63, PO: 0.77.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4B5BEBA0E731145FEA9FC716227FB88" pageId="3" pageNumber="54" type="materials_examined">
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<paragraph id="FC10ED310E731145FEA9FC716227FB88" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="54">
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<emphasis id="CEDB31230E731145FEA9FC716366FC38" bold="true" box="[312,532,924,950]" pageId="3" pageNumber="54">Material studied.</emphasis>
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<typeStatus id="231453930E731145FDB3FC7163FDFC38" box="[546,655,924,950]" pageId="3" pageNumber="54" type="holotype">Holotype</typeStatus>
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♂.
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<collectingCountry id="84B8ADA10E731145FD52FC716260FC38" box="[707,786,924,950]" name="Peru" pageId="3" pageNumber="54">PERU</collectingCountry>
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. Department of Cuzco. Quincemil (
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<geoCoordinate id="999B8BF60E731145FB42FC716455FC38" box="[1235,1319,924,950]" direction="south" orientation="latitude" pageId="3" pageNumber="54" precision="5555" value="-13.2">13.2°S</geoCoordinate>
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:
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<geoCoordinate id="999B8BF60E731145FE99FC296006FC50" box="[264,372,964,990]" direction="west" orientation="longitude" pageId="3" pageNumber="54" precision="925" value="-71.28333">70°77’W</geoCoordinate>
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,
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<quantity id="3B5740D40E731145FE13FC2960A0FC50" box="[386,466,964,990]" metricMagnitude="2" metricUnit="m" metricValue="6.1899999999999995" pageId="3" pageNumber="54" unit="m" value="619.0">619 m</quantity>
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),
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<date id="8811CBF10E731145FE78FC29633FFC50" box="[489,589,964,990]" pageId="3" pageNumber="54" value="1962-09">IX.1962</date>
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, in tropical forest (Luis E. Peña), E. L. Mockford collection, Illinois State University, Normal, Illinois,
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<collectingCountry id="84B8ADA10E731145FD67FC016227FB88" box="[758,853,1004,1030]" name="United States of America" pageId="3" pageNumber="54">U. S. A.</collectingCountry>
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</paragraph>
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</subSubSection>
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<subSubSection id="B4B5BEBA0E731145FEA9FBF96529FBD8" pageId="3" pageNumber="54" type="etymology">
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<paragraph id="FC10ED310E731145FEA9FBF96529FBD8" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="54">
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<emphasis id="CEDB31230E731145FEA9FBF960B6FBA0" bold="true" box="[312,452,1044,1070]" pageId="3" pageNumber="54">Etymology.</emphasis>
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The genus name is a composite word from Inca, a dominant ethnic Peruvian group, and psocus= psocid. The species is dedicated to its collector.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4B5BEBA0E731145FEA9FB8962C0F980" pageId="3" pageNumber="54" type="discussion">
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<paragraph id="FC10ED310E731145FEA9FB8962C2FA30" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="54">
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<emphasis id="CEDB31230E731145FEA9FB8960C3FBF0" bold="true" box="[312,433,1124,1150]" pageId="3" pageNumber="54">Remarks.</emphasis>
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Preliminary results of a phylogenetic analysis of the 40 genera that comprise the infraorder Epipsocetae (Casasola González,
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<emphasis id="CEDB31230E731145FC18FB61657CFB2B" box="[905,1038,1164,1189]" italics="true" pageId="3" pageNumber="54">in progress</emphasis>
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), place
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<taxonomicName id="3BAF96B20E731145FBE1FB616586FB2B" box="[1136,1268,1164,1189]" class="Insecta" family="Epipsocidae" genus="Incapsocus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="CEDB31230E731145FBE1FB616586FB2B" box="[1136,1268,1164,1189]" italics="true" pageId="3" pageNumber="54">Incapsocus</emphasis>
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</taxonomicName>
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in a clade as sister group to an assemblage constituted by
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<emphasis id="CEDB31230E731145FC11FB596518FB43" box="[896,1130,1204,1229]" italics="true" pageId="3" pageNumber="54">
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<taxonomicName id="3BAF96B20E731145FC11FB596569FB43" box="[896,1051,1204,1229]" class="Insecta" family="Epipsocidae" genus="Cubitiglabra" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Cubitiglabra</taxonomicName>
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,
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<taxonomicName id="3BAF96B20E731145FBB6FB596517FB43" box="[1063,1125,1204,1229]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">Goja</taxonomicName>
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,
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</emphasis>
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and
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<taxonomicName id="3BAF96B20E731145FB32FB59605FFB7B" class="Insecta" family="Epipsocidae" genus="Ianthorntonia" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="CEDB31230E731145FB32FB59605FFB7B" italics="true" pageId="3" pageNumber="54">Ianthorntonia</emphasis>
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</taxonomicName>
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. The clade is supported by having wing venation supernumerary (=forewing with Rs more than 2branched, M more than 3branched, and hindwing with M forked).
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<taxonomicName id="3BAF96B20E731145FB55FAE9605DFACB" class="Insecta" family="Epipsocidae" genus="Incapsocus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="CEDB31230E731145FB55FAE9605DFACB" italics="true" pageId="3" pageNumber="54">Incapsocus</emphasis>
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</taxonomicName>
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stands apart from the above 3 genera on account principally of genital differences and wing venation details:
|
||
<taxonomicName id="3BAF96B20E731145FDA3FAB963BBFAE3" box="[562,713,1364,1389]" class="Insecta" family="Epipsocidae" genus="Cubitiglabra" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEDB31230E731145FDA3FAB963BBFAE3" box="[562,713,1364,1389]" italics="true" pageId="3" pageNumber="54">Cubitiglabra</emphasis>
|
||
</taxonomicName>
|
||
has a closed phallosome and distinct external parameres;
|
||
<taxonomicName id="3BAF96B20E731145FE02FA9160BCFA1B" box="[403,462,1404,1429]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEDB31230E731145FE02FA9160BCFA1B" box="[403,462,1404,1429]" italics="true" pageId="3" pageNumber="54">Goja</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BAF96B20E731145FD9CFA9163DDFA1B" box="[525,687,1404,1429]" class="Insecta" family="Epipsocidae" genus="Ianthorntonia" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEDB31230E731145FD9CFA9163DDFA1B" box="[525,687,1404,1429]" italics="true" pageId="3" pageNumber="54">Ianthorntonia</emphasis>
|
||
</taxonomicName>
|
||
are sister genera, presenting a complex phallosome, with external parameres and well defined phallic sclerites.
|
||
</paragraph>
|
||
<paragraph id="FC10ED310E731145FEA9FA2162C0F980" blockId="3.[264,1324,283,1550]" pageId="3" pageNumber="54">
|
||
Geographically,
|
||
<taxonomicName id="3BAF96B20E731145FE6FFA2163F0FA6B" box="[510,642,1484,1509]" class="Insecta" family="Epipsocidae" genus="Incapsocus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEDB31230E731145FE6FFA2163F0FA6B" box="[510,642,1484,1509]" italics="true" pageId="3" pageNumber="54">Incapsocus</emphasis>
|
||
</taxonomicName>
|
||
, some species of
|
||
<taxonomicName id="3BAF96B20E731145FCF5FA2162EDFA6B" box="[868,927,1484,1509]" class="Insecta" family="Epipsocidae" genus="Goja" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEDB31230E731145FCF5FA2162EDFA6B" box="[868,927,1484,1509]" italics="true" pageId="3" pageNumber="54">Goja</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3BAF96B20E731145FC78FA2165F9FA6B" box="[1001,1163,1484,1509]" class="Insecta" family="Epipsocidae" genus="Ianthorntonia" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEDB31230E731145FC78FA2165F9FA6B" box="[1001,1163,1484,1509]" italics="true" pageId="3" pageNumber="54">Ianthorntonia</emphasis>
|
||
</taxonomicName>
|
||
are Andean, while the two known species of
|
||
<taxonomicName id="3BAF96B20E731145FD13FA19626BF983" box="[642,793,1524,1549]" class="Insecta" family="Epipsocidae" genus="Cubitiglabra" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CEDB31230E731145FD13FA19626BF983" box="[642,793,1524,1549]" italics="true" pageId="3" pageNumber="54">Cubitiglabra</emphasis>
|
||
</taxonomicName>
|
||
are Chinese.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |