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<document id="206BCE94B96D032DB12FE11315488345" ID-DOI="10.17161/jom.v0i30.4698" ID-ISSN="2325-4467" ID-Zenodo-Dep="13736830" ID-ZooBank="urn:lsid:zoobank.org:pub:D3E9ABEE-FC97-4D2F-8834-E6D1F6C81D55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1722892979211" checkinUser="felipe" docAuthor="Engel, Michael S., Breitkreuz, Laura C. V. &amp; Ohl, Michael" docDate="2014" docId="737287F90E12FFE592627DD9FBDA6D1E" docLanguage="en" docName="JMelittology.2013.30.1-18.pdf" docOrigin="Journal of Melittology 2014 (30)" docSource="https://doi.org/10.17161/jom.v0i30.4698" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.4:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="4" docTitle="Mochlomelikertes Engel, Breitkreuz, &amp; Ohl 2014, new genus" docType="treatment" docUuid="D766BE9A-2AB7-4CF4-87FC-B78572B56C3B" docUuidSource="ZooBank" docVersion="4" lastPageNumber="9" masterDocId="8F4BFF810E16FFED93317878FF996F07" masterDocTitle="The first male of the extinct bee tribe Melikertini (Hymenoptera: Apidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="5" updateTime="1725892004978" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="44E4940E79516494A1EF5FA3AD03A630">The first male of the extinct bee tribe Melikertini (Hymenoptera: Apidae)</mods:title>
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<mods:namePart id="0D29D3A22E502C89AC4B690C41A49C19">Engel, Michael S.</mods:namePart>
<mods:affiliation id="7C09D2B5F4420E4F746E3509A12ECB77">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West &amp; Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 (msengel @ ku. edu; l-breitkreuz @ ku. edu).</mods:affiliation>
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<mods:namePart id="88F93BCB995036BDFED57A7CF414DFB8">Breitkreuz, Laura C. V.</mods:namePart>
<mods:affiliation id="AAC40362A7557B6A5639AEDC3F485292">Division of Entomology, Natural History Museum and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 (msengel @ ku. edu; l-breitkreuz @ ku. edu).</mods:affiliation>
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<mods:namePart id="99739A66C80AED65CEE809C521D210A8">Ohl, Michael</mods:namePart>
<mods:affiliation id="31561B9C267BDA3B52249DB951C8F75E">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West</mods:affiliation>
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<mods:title id="2B60CED2C58E9106E6E53F58F5973755">Journal of Melittology</mods:title>
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<mods:date id="B7E708E25D9ACD104B9A11B9E196B09C">2014</mods:date>
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<taxonomicName id="3CDB4D6C0E12FFE992627DD9FCCB6ABE" authority="Engel, Breitkreuz, &amp; Ohl" authorityName="Engel, Breitkreuz, &amp; Ohl" authorityYear="2014" box="[339,850,1441,1465]" class="Insecta" family="Apidae" genus="Mochlomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C9AFEAFD0E12FFE992627DD9FDB16ABE" bold="true" box="[339,552,1441,1465]" italics="true" pageId="4" pageNumber="5">Mochlomelikertes</emphasis>
Engel, Breitkreuz, &amp; Ohl
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,
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ZooBank:
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<paragraph id="FB6436EF0E12FFE993E17D87FB1C6911" blockId="4.[160,1168,1534,1782]" box="[208,1157,1534,1558]" pageId="4" pageNumber="5">
<materialsCitation id="4BB33CB20E12FFE993E17D87FB186911" box="[208,1153,1534,1558]" collectionCode="SPECIES" pageId="4" pageNumber="5" specimenCount="1" typeStatus="holotype">
<typeStatus id="2460884D0E12FFE993E17D87FE9F6911" box="[208,262,1535,1558]" pageId="4" pageNumber="5">TYPE</typeStatus>
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:
<taxonomicName id="3CDB4D6C0E12FFE9925D7D86FC7C6911" authority="Engel, Breitkreuz, &amp; Ohl" authorityName="Engel, Breitkreuz, &amp; Ohl" authorityYear="2014" box="[364,997,1534,1558]" class="Insecta" family="Apidae" genus="Mochlomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hoffeinsorum" status="sp. nov.">
<emphasis id="C9AFEAFD0E12FFE9925D7D86FD256911" box="[364,700,1534,1558]" italics="true" pageId="4" pageNumber="5">Mochlomelikertes hoffeinsorum</emphasis>
Engel, Breitkreuz, &amp; Ohl
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,
<taxonomicNameLabel id="D29C57860E12FFE990C07D87FB186911" box="[1009,1153,1534,1558]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
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.
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<subSubSection id="B3C165640E12FFE593E17E66FF436EBE" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="FB6436EF0E12FFE593E17E66FF436EBE" blockId="4.[160,1168,1534,1782]" lastBlockId="8.[160,1168,225,538]" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
DIAGNOSIS:
<specimenCount id="EDDDFD660E12FFE992677E67FEF3693E" box="[342,362,1567,1593]" count="1" pageId="4" pageNumber="5" type="male"></specimenCount>
: Medium-sized bee,
<emphasis id="C9AFEAFD0E12FFE9916F7E67FDEC6931" box="[606,629,1567,1590]" italics="true" pageId="4" pageNumber="5">ca</emphasis>
.
<quantity id="3C239B0A0E12FFE991B37E66FD5D6931" box="[642,708,1566,1590]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="mm" value="8.0">8 mm</quantity>
in length. Mandibles simple, elongate; axes of mandibular articulations converging anteriorly such that closed mandibles overlap at slightly obtuse angle (
<figureCitation id="63E02A6A0E12FFE991037E26FDEE6971" box="[562,631,1630,1654]" captionStart="Figures 78" captionStartId="6.[160,239,1399,1421]" captionTargetBox="[160,1168,224,1395]" captionTargetId="figure-134@6.[160,1168,224,1395]" captionTargetPageId="6" captionText="Figures 78. Photographs of head of holotype (CCHH 1766-1) male of Mochlomelikertes hoffeinso- rum, new genus and species. 7. Dorsal view. 8. Ventral apical oblique view showing mouthparts." figureDoi="http://doi.org/10.5281/zenodo.13736842" httpUri="https://zenodo.org/record/13736842/files/figure.png" pageId="4" pageNumber="5">Fig. 8</figureCitation>
); clypeus with apical margin pale, contrasting with brown integument of remainder of head (
<figureCitation id="63E02A6A0E12FFE990377E06FCCF6991" box="[774,854,1662,1686]" captionStart="Figures 56" captionStartId="5.[160,239,1557,1579]" captionTargetBox="[160,1168,224,1541]" captionTargetId="figure-70@5.[160,1168,224,1553]" captionTargetPageId="5" captionText="Figures 56. Photograph and illustration of head of holotype (CCHH 1766-1) male of Mochlomelikertes hoffeinsorum, new genus and species. 5. Left oblique view. 6. Illustration of head as preserved." figureDoi="http://doi.org/10.5281/zenodo.13736838" httpUri="https://zenodo.org/record/13736838/files/figure.png" pageId="4" pageNumber="5">Figs. 5</figureCitation>
,
<figureCitation id="63E02A6A0E12FFE990557E06FCEB6991" box="[868,882,1662,1686]" captionStart="Figures 78" captionStartId="6.[160,239,1399,1421]" captionTargetBox="[160,1168,224,1395]" captionTargetId="figure-134@6.[160,1168,224,1395]" captionTargetPageId="6" captionText="Figures 78. Photographs of head of holotype (CCHH 1766-1) male of Mochlomelikertes hoffeinso- rum, new genus and species. 7. Dorsal view. 8. Ventral apical oblique view showing mouthparts." figureDoi="http://doi.org/10.5281/zenodo.13736842" httpUri="https://zenodo.org/record/13736842/files/figure.png" pageId="4" pageNumber="5">8</figureCitation>
); clypeus without basal prominence or protrusion; epistomal sulcus forming broadly obtuse angle. Pronotal lobe densely setose; mesoscutum with anterior border low, broadly rounded; tegula elongate-ovoid; mesoscutellum with elongate medial extension of posterior margin (
<figureCitation id="63E02A6A0E13FFE893997E1DFE96697A" box="[168,271,1637,1661]" captionStart="Figures 34" captionStartId="3.[160,241,1404,1426]" captionTargetBox="[160,1168,224,1398]" captionTargetId="figure-123@3.[160,1168,224,1400]" captionTargetPageId="3" captionText="Figures 34. Photographs of holotype (CCHH 1766-1) male of Mochlomelikertes hoffeinsorum, new genus and species. 3. Left lateral view. 4. Right lateral oblique view." figureDoi="http://doi.org/10.5281/zenodo.13736836" httpUri="https://zenodo.org/record/13736836/files/figure.png" pageId="5" pageNumber="6">Figs. 3, 4</figureCitation>
,
<figureCitation id="63E02A6A0E13FFE892287E1DFECB697A" box="[281,338,1637,1661]" captionStart="Figures 911" captionStartId="7.[160,241,1601,1623]" captionTargetBox="[160,1168,224,1586]" captionTargetId="figure-8@7.[160,1168,224,1596]" captionTargetPageId="7" captionText="Figures 911. Holotype (CCHH 1766-1) male of Mochlomelikertes hoffeinsorum, new genus and species. 9. Photograph of dorsal view of mesosoma. 10. Photograph of left lateral view of mesoscutellum and its posterior extension. 11. Illustration of mesosomal dorsum." figureDoi="http://doi.org/10.5281/zenodo.13736844" httpUri="https://zenodo.org/record/13736844/files/figure.png" pageId="5" pageNumber="6">911</figureCitation>
), with process greatly elongate and projecting over metanotum, propodeum, and anterior portions of metasoma (at least extending beyond apical margin of second metasomal tergum), and with projection slightly longer than remainder of mesosoma and densely setose on dorsal surface (
<figureCitation id="63E02A6A0E13FFE891F77EBDFCA669DA" box="[710,831,1733,1757]" captionStart="Figures 911" captionStartId="7.[160,241,1601,1623]" captionTargetBox="[160,1168,224,1586]" captionTargetId="figure-8@7.[160,1168,224,1596]" captionTargetPageId="7" captionText="Figures 911. Holotype (CCHH 1766-1) male of Mochlomelikertes hoffeinsorum, new genus and species. 9. Photograph of dorsal view of mesosoma. 10. Photograph of left lateral view of mesoscutellum and its posterior extension. 11. Illustration of mesosomal dorsum." figureDoi="http://doi.org/10.5281/zenodo.13736844" httpUri="https://zenodo.org/record/13736844/files/figure.png" pageId="5" pageNumber="6">Figs. 911</figureCitation>
). Forewing with prominent pterostigma; two submarginal cells present (1rs-m absent); second submarginal cell greatly elongate, length 3.5 times width (resulting from absence of 1rs-m and thereby representing equivalent length of what would otherwise be the second and long third submarginal cells); first submarginal cell angled, horsehead-shaped (resulting from strongly angled second abscissa Rs); r-rs relatively long, meeting pterostigma near midpoint; 1m-cu short, relatively straight; basal vein basad cu-a. Mesobasitarsus elongate, nearly as long as mesotibia (
<figureCitation id="63E02A6A0E10FFEB91027E12FD1C6985" box="[563,645,1642,1666]" captionStart="Figures 1213" captionStartId="10.[160,241,1108,1130]" captionTargetBox="[160,1168,224,1102]" captionTargetId="figure-211@10.[160,1168,224,1103]" captionTargetPageId="10" captionText="Figures 1213. Photographs of legs of holotype (CCHH 1766-1) male of Mochlomelikertes hoffeinsorum, new genus and species. 12. Apex of mesotibia and entire mesotarsus and mesopretarsus. 13. Metafemur, metatibia, metatarsus, and metapretarsus (with portions of left mesotibia and mesotarsus in foreground)." figureDoi="http://doi.org/10.5281/zenodo.13736846" httpUri="https://zenodo.org/record/13736846/files/figure.png" pageId="6" pageNumber="7">Fig. 12</figureCitation>
), both podites slightly swollen with numerous elongate setae along outer and posterior surfaces, such setae minutely branched in apical quarters; metatibia without corbicula (true for all males), with outer surface slightly swollen, outer and posterior surfaces of metatibia and metabasitarsus with numerous, elongate setae (
<figureCitation id="63E02A6A0E10FFEB92D37E92FDAA6805" box="[482,563,1770,1794]" captionStart="Figures 1213" captionStartId="10.[160,241,1108,1130]" captionTargetBox="[160,1168,224,1102]" captionTargetId="figure-211@10.[160,1168,224,1103]" captionTargetPageId="10" captionText="Figures 1213. Photographs of legs of holotype (CCHH 1766-1) male of Mochlomelikertes hoffeinsorum, new genus and species. 12. Apex of mesotibia and entire mesotarsus and mesopretarsus. 13. Metafemur, metatibia, metatarsus, and metapretarsus (with portions of left mesotibia and mesotarsus in foreground)." figureDoi="http://doi.org/10.5281/zenodo.13736846" httpUri="https://zenodo.org/record/13736846/files/figure.png" pageId="6" pageNumber="7">Fig. 13</figureCitation>
), such setae minutely branched in apical thirds to quarters; inner surface of metatibia with keirotrichia; keirotrichiate zone not elevated, running from base to near apex, separated from posterior border by distance less than one-third width of keirotrichiate zone; single metatibial spur present, spur minutely serrate; metabasitarsus subquadrate, slightly longer than wide, inner surface with dense, stiff, simple setae forming loose comb rows (typical among higher corbiculate bees); pretarsal claws cleft. Metasoma sparsely setose, most setae minute, simple, and appressed to subappressed; visible metasomal sterna unmodified; gonostyli relatively broad, thumb-like, apically rounded, with a few distinct setae along outer apical borders.
</paragraph>
</subSubSection>
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<paragraph id="FB6436EF0E13FFE893917E6DFB17694F" blockId="5.[160,1166,1557,1608]" pageId="5" pageNumber="6">
<emphasis id="C9AFEAFD0E13FFE893917E6DFE85692D" bold="true" box="[160,284,1557,1579]" pageId="5" pageNumber="6">Figures 56.</emphasis>
Photograph and illustration of head of holotype (CCHH 1766-1) male of
<taxonomicName id="3CDB4D6C0E13FFE897347E6DFED06940" authorityName="Engel &amp; Breitkreuz &amp; Ohl" authorityYear="2014" class="Insecta" family="Apidae" genus="Mochlomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hoffeinsorum" status="new genus and species">
<emphasis id="C9AFEAFD0E13FFE897347E6DFED06940" italics="true" pageId="5" pageNumber="6">Mochlomelikertes hoffeinsorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D29C57860E13FFE892647E4AFDD9694F" box="[341,576,1586,1608]" pageId="5" pageNumber="6" rank="species">new genus and species</taxonomicNameLabel>
.
<emphasis id="C9AFEAFD0E13FFE8917F7E4AFDF96940" bold="true" box="[590,608,1586,1607]" pageId="5" pageNumber="6">5.</emphasis>
Left oblique view.
<emphasis id="C9AFEAFD0E13FFE890167E49FCA06941" bold="true" box="[807,825,1585,1606]" pageId="5" pageNumber="6">6.</emphasis>
Illustration of head as preserved.
</paragraph>
</caption>
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<paragraph id="FB6436EF0E10FFEB93917D0FFBE56AAD" blockId="6.[160,1148,1399,1450]" pageId="6" pageNumber="7">
<emphasis id="C9AFEAFD0E10FFEB93917D0FFE856A8B" bold="true" box="[160,284,1399,1421]" pageId="6" pageNumber="7">Figures 78.</emphasis>
Photographs of head of holotype (CCHH 1766-1) male of
<taxonomicName id="3CDB4D6C0E10FFEB905F7D0FFF516AAD" authorityName="Engel &amp; Breitkreuz &amp; Ohl" authorityYear="2014" class="Insecta" family="Apidae" genus="Mochlomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hoffeinsorum" status="new genus and species">
<emphasis id="C9AFEAFD0E10FFEB905F7D0FFF516AAD" italics="true" pageId="6" pageNumber="7">Mochlomelikertes hoffeinsorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D29C57860E10FFEB93E27DECFE266AAD" box="[211,447,1428,1450]" pageId="6" pageNumber="7" rank="species">new genus and species</taxonomicNameLabel>
.
<emphasis id="C9AFEAFD0E10FFEB92FD7DECFE476AAE" bold="true" box="[460,478,1428,1449]" pageId="6" pageNumber="7">7.</emphasis>
Dorsal view.
<emphasis id="C9AFEAFD0E10FFEB915E7DECFD196AAE" bold="true" box="[623,640,1428,1449]" pageId="6" pageNumber="7">8.</emphasis>
Ventral apical oblique view showing mouthparts.
</paragraph>
</caption>
<caption id="AFA466670E11FFEA93917E39FB9D6996" ID-DOI="http://doi.org/10.5281/zenodo.13736844" ID-Zenodo-Dep="13736844" httpUri="https://zenodo.org/record/13736844/files/figure.png" pageId="7" pageNumber="8" startId="7.[160,241,1601,1623]" targetBox="[160,1168,224,1586]" targetPageId="7" targetType="figure">
<paragraph id="FB6436EF0E11FFEA93917E39FB9D6996" blockId="7.[160,1150,1601,1681]" pageId="7" pageNumber="8">
<emphasis id="C9AFEAFD0E11FFEA93917E39FEB76951" bold="true" box="[160,302,1601,1623]" pageId="7" pageNumber="8">Figures 911.</emphasis>
Holotype (CCHH 1766-1) male of
<taxonomicName id="3CDB4D6C0E11FFEA91937E39FC496951" authorityName="Engel &amp; Breitkreuz &amp; Ohl" authorityYear="2014" box="[674,976,1601,1622]" class="Insecta" family="Apidae" genus="Mochlomelikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hoffeinsorum" status="new genus and species">
<emphasis id="C9AFEAFD0E11FFEA91937E39FC496951" box="[674,976,1601,1622]" italics="true" pageId="7" pageNumber="8">Mochlomelikertes hoffeinsorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D29C57860E11FFEA90ED7E39FF776973" pageId="7" pageNumber="8" rank="species">new genus and species</taxonomicNameLabel>
.
<emphasis id="C9AFEAFD0E11FFEA93CC7E26FE966974" bold="true" box="[253,271,1630,1651]" pageId="7" pageNumber="8">9.</emphasis>
Photograph of dorsal view of mesosoma.
<emphasis id="C9AFEAFD0E11FFEA91E57E26FD6B6974" bold="true" box="[724,754,1630,1651]" pageId="7" pageNumber="8">10.</emphasis>
Photograph of left lateral view of mesoscutellum and its posterior extension.
<emphasis id="C9AFEAFD0E11FFEA91407E02FD166988" bold="true" box="[625,655,1658,1679]" pageId="7" pageNumber="8">11.</emphasis>
Illustration of mesosomal dorsum.
</paragraph>
</caption>
<subSubSection id="B3C165640E1EFFE593E179B9FBDA6D1E" pageId="8" pageNumber="9" type="etymology">
<paragraph id="FB6436EF0E1EFFE593E179B9FBDA6D1E" blockId="8.[160,1168,225,538]" pageId="8" pageNumber="9">
ETYMOLOGY: The new genus-group name is a combination of the
<emphasis id="C9AFEAFD0E1EFFE590EB79B9FBA96EDE" box="[986,1072,449,473]" italics="true" pageId="8" pageNumber="9">mochlos</emphasis>
(Greek, “lever”, a reference to large mesoscutellar extension which resembles an extended lever) and
<taxonomicName id="3CDB4D6C0E1EFFE5923A7A79FEE56D1E" authorityName="Engel" authorityYear="1998" box="[267,380,513,537]" class="Insecta" family="Apidae" genus="Melikertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="C9AFEAFD0E1EFFE5923A7A79FEE56D1E" box="[267,380,513,537]" italics="true" pageId="8" pageNumber="9">Melikertes</emphasis>
</taxonomicName>
,
<typeStatus id="2460884D0E1EFFE592B87A7AFE246D1D" box="[393,445,514,538]" pageId="8" pageNumber="9">type</typeStatus>
genus of the tribe
<taxonomicName id="3CDB4D6C0E1EFFE591A97A79FC866D1E" box="[664,799,513,537]" class="Insecta" family="Apidae" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Melikertini">Melikertini</taxonomicName>
. The name is masculine.
</paragraph>
</subSubSection>
</treatment>
</document>