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<document id="BDCAA5930CBF1DC57A051F1617F71552" ID-DOI="10.11646/zootaxa.3919.1.5" ID-GBIF-Dataset="fb29460a-8754-44f1-bc88-f8715eda6a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="235217" ID-ZooBank="4100DA7B-62A7-4E15-BE30-C9331B9CE035" IM.bibliography_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460989802810" checkinUser="plazi" docAuthor="Katlav, Alihan, Hajiqanbar, Hamidreza &amp; Talebi, Ali Asghar" docDate="2015" docId="734F8766FFB9FFE281B70BE6FC6F912E" docLanguage="en" docName="zt03919p110.pdf" docOrigin="Zootaxa 3919 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar, sp. nov." docType="treatment" docVersion="8" lastPageNumber="108" masterDocId="8F76FF1EFFB8FFEA81200F42FF8C927C" masterDocTitle="Pseudopygmephorellus mazandaranicus sp. nov. (Acari: Heterostigmata: Pygmephoridae), phoretic on scarabaeid dung beetles (Coleoptera: Scarabaeidae) from Iran" masterLastPageNumber="110" masterPageNumber="100" pageNumber="101" updateTime="1698561533060" updateUser="plazi">
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<mods:title id="F87D0BF2952FB9B00D1F9736FC41F2A8">Pseudopygmephorellus mazandaranicus sp. nov. (Acari: Heterostigmata: Pygmephoridae), phoretic on scarabaeid dung beetles (Coleoptera: Scarabaeidae) from Iran</mods:title>
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<mods:name id="A65B98B232B215892E16CCB714F8EFEA" type="personal">
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<mods:namePart id="055AF875CA4578FB13CB3FF242549FBA">Katlav, Alihan</mods:namePart>
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<mods:name id="88C103A8A2080545CE2B2C79AF2BB10C" type="personal">
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<mods:namePart id="D28D0D451115DF25DACBF903E5D19F8B">Hajiqanbar, Hamidreza</mods:namePart>
</mods:name>
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<mods:namePart id="EA714D60F1A8117EE10B796356B6F221">Talebi, Ali Asghar</mods:namePart>
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<mods:title id="8CFF6F9100BB77E33D12B906BBE19084">Zootaxa</mods:title>
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<mods:part id="A3339BFEBEEA6355F20A7ADE19EDAA1A">
<mods:date id="00FB468F9D5172D49B831D4755346DF3">2015</mods:date>
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<mods:number id="C1721FD2C27361F1DF4DA660076DF316">3919</mods:number>
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<treatment id="734F8766FFB9FFE281B70BE6FC6F912E" ID-DOI="http://doi.org/10.5281/zenodo.6102471" ID-GBIF-Taxon="119596479" ID-Zenodo-Dep="6102471" LSID="urn:lsid:plazi:treatment:734F8766FFB9FFE281B70BE6FC6F912E" httpUri="http://treatment.plazi.org/id/734F8766FFB9FFE281B70BE6FC6F912E" lastPageId="8" lastPageNumber="108" pageId="1" pageNumber="101">
<subSubSection id="B3FC65FBFFB9FFEB81B70BE6FB9C96C2" box="[151,1040,1188,1214]" pageId="1" pageNumber="101" type="nomenclature">
<paragraph id="FB593670FFB9FFEB81B70BE6FB9C96C2" blockId="1.[151,1040,1188,1214]" box="[151,1040,1188,1214]" pageId="1" pageNumber="101">
<heading id="A011811CFFB9FFEB81B70BE6FB9C96C2" bold="true" box="[151,1040,1188,1214]" fontSize="11" level="1" pageId="1" pageNumber="101" reason="1">
<emphasis id="C992EA62FFB9FFEB81B70BE6FB9C96C2" bold="true" box="[151,1040,1188,1214]" pageId="1" pageNumber="101">
<taxonomicName id="3CE64DF3FFB9FFEB81B70BE6FC2696C2" authority="Katlav and Hajiqanbar" authorityName="Katlav and Hajiqanbar" box="[151,938,1188,1214]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="101" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFB9FFEB81B70BE6FDF296C2" bold="true" box="[151,638,1188,1214]" italics="true" pageId="1" pageNumber="101">Pseudopygmephorellus mazandaranicus</emphasis>
Katlav and Hajiqanbar
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFB9FFEB82920BE6FB9C96C2" box="[946,1040,1188,1214]" pageId="1" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B3FC65FBFFB9FFEB81B70BA8FAC99731" pageId="1" pageNumber="101" type="diagnosis">
<paragraph id="FB593670FFB9FFEB81B70BA8FAC99731" blockId="1.[151,1437,1258,2024]" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB81B70BA8FE9D977F" bold="true" box="[151,273,1258,1283]" pageId="1" pageNumber="101">Diagnosis.</emphasis>
The phoretic female of the new species is characterized by having: two claws on tarsus IV; posterior margin of the prodorsum nearly straight; setiform epimeral setae 1
<emphasis id="C992EA62FFB9FFEB82B20A4DFC2C975A" box="[914,928,1295,1318]" italics="true" pageId="1" pageNumber="101">a</emphasis>
, 1
<emphasis id="C992EA62FFB9FFEB829E0A4CFC40975A" box="[958,972,1294,1318]" italics="true" pageId="1" pageNumber="101">b</emphasis>
, 2
<emphasis id="C992EA62FFB9FFEB82CA0A4DFC74975A" box="[1002,1016,1295,1318]" italics="true" pageId="1" pageNumber="101">a</emphasis>
, 2
<emphasis id="C992EA62FFB9FFEB85360A4DFBAE975A" box="[1046,1058,1295,1318]" italics="true" pageId="1" pageNumber="101">c</emphasis>
; setae
<date id="8F5810B0FFB9FFEB85540A4DFB059757" box="[1140,1161,1295,1323]" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB85540A4DFB0C975A" box="[1140,1152,1295,1318]" italics="true" pageId="1" pageNumber="101">v</emphasis>
1
</date>
situated posterior to the level of setae
<date id="8F5810B0FFB9FFEB80110A74FECA972E" box="[305,326,1334,1362]" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB80110A74FEB19731" box="[305,317,1334,1357]" italics="true" pageId="1" pageNumber="101">v</emphasis>
2
</date>
; reduced apodemes 5; setiform TaII
<emphasis id="C992EA62FFB9FFEB83FA0A74FD789731" box="[730,756,1334,1357]" italics="true" pageId="1" pageNumber="101">tc'</emphasis>
(slender and tapering); setiform and pointed FeIV
<emphasis id="C992EA62FFB9FFEB84080A74FAB19731" box="[1320,1341,1334,1357]" italics="true" pageId="1" pageNumber="101">v</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="B3FC65FBFFB9FFEF81E70A1EFD1193AC" lastPageId="5" lastPageNumber="105" pageId="1" pageNumber="101" type="description">
<paragraph id="FB593670FFB9FFEB81E70A1EFE4697E5" blockId="1.[151,1437,1258,2024]" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB81E70A1EFED59709" bold="true" box="[199,345,1372,1397]" pageId="1" pageNumber="101">Description.</emphasis>
Female. Length of idiosoma 212 (194229), maximum width (measured on widest part of segment C) 128 (119135).
</paragraph>
<paragraph id="FB593670FFB9FFEB81E70AE6FC5D94C0" blockId="1.[151,1437,1258,2024]" pageId="1" pageNumber="101">
<taxonomicName id="3CE64DF3FFB9FFEB81E70AE6FEDA97C0" box="[199,342,1444,1468]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="1" pageNumber="101" phylum="Nematoda" rank="genus">
<emphasis id="C992EA62FFB9FFEB81E70AE6FEDA97C0" box="[199,342,1444,1468]" italics="true" pageId="1" pageNumber="101">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="63DD2AF5FFB9FFEB80460AE6FE5E97C1" box="[358,466,1444,1469]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,880,903]" captionTargetBox="[302,1285,384,856]" captionTargetId="figure@2.[302,1285,384,857]" captionTargetPageId="2" captionText="FIGURES 1 3. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 1. Gnathosoma, dorsal view; 2. Gnathosoma, ventral view; 3. Pharyngeal system. Scale bar 10 µm." httpUri="https://zenodo.org/record/235218/files/figure.png" pageId="1" pageNumber="101">Figs. 12</figureCitation>
). Length of
<taxonomicName id="3CE64DF3FFB9FFEB837D0AE6FD6897C1" box="[605,740,1444,1469]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="1" pageNumber="101" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
20 (1820), width 24 (2225). Gnathosomal capsule rounded sub-trapezoid in dorsal aspect, slightly wider than long, dorsally with one pair of pointed and smooth cheliceral setae
<emphasis id="C992EA62FFB9FFEB81F80AAEFF7F9478" box="[216,243,1516,1540]" italics="true" pageId="1" pageNumber="101">ch</emphasis>
1 5 (46) and a weak longitudinal dorsal median apodeme in its anterior half (hardly discernible in some specimens), ventrally with one pair of pointed subcapitular setae
<emphasis id="C992EA62FFB9FFEB825E0956FC1B9457" box="[894,919,1556,1579]" italics="true" pageId="1" pageNumber="101">su</emphasis>
4 (44); palpi dorsally with a pair of smooth and pointed setae
<emphasis id="C992EA62FFB9FFEB80430975FE1E9433" box="[355,402,1591,1615]" italics="true" pageId="1" pageNumber="101">dGe</emphasis>
11 (911) and a pair of smooth and needle-like setae
<emphasis id="C992EA62FFB9FFEB82D60975FBAE9433" box="[1014,1058,1591,1615]" italics="true" pageId="1" pageNumber="101">dFe</emphasis>
6 (56),
<emphasis id="C992EA62FFB9FFEB85AB0975FB369433" box="[1163,1210,1591,1615]" italics="true" pageId="1" pageNumber="101">dGe</emphasis>
nearly twice longer than
<emphasis id="C992EA62FFB9FFEB81F20919FF72940F" box="[210,254,1627,1651]" italics="true" pageId="1" pageNumber="101">dFe</emphasis>
; tibiotarsus with single solenidion ventrally (barely visible in some specimens) and a terminal claw; chelicerae inconspicuous; pharyngeal system (
<figureCitation id="63DD2AF5FFB9FFEB8388093DFD6194EB" box="[680,749,1663,1688]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,880,903]" captionTargetBox="[302,1285,384,856]" captionTargetId="figure@2.[302,1285,384,857]" captionTargetPageId="2" captionText="FIGURES 1 3. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 1. Gnathosoma, dorsal view; 2. Gnathosoma, ventral view; 3. Pharyngeal system. Scale bar 10 µm." httpUri="https://zenodo.org/record/235218/files/figure.png" pageId="1" pageNumber="101">Fig. 3</figureCitation>
) including three striated pumps: pump 1 smallest and bowlike, pump 2 well developed and subcylindrical, pump 3 inverted cordate.
</paragraph>
<paragraph id="FB593670FFB9FFE881E70985FB3E933A" blockId="1.[151,1437,1258,2024]" lastBlockId="2.[151,1436,151,330]" lastPageId="2" lastPageNumber="102" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB81E70985FE1894A3" box="[199,404,1735,1759]" italics="true" pageId="1" pageNumber="101">Idiosomal dorsum</emphasis>
(
<figureCitation id="63DD2AF5FFB9FFEB80850985FE6094A3" box="[421,492,1735,1760]" captionStart="FIGURE 4" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[230,1356,481,1893]" captionTargetId="figure@3.[230,1356,481,1893]" captionTargetPageId="3" captionText="FIGURE 4. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): dorsum of the body. Scale bar 50 µm." httpUri="https://zenodo.org/record/235219/files/figure.png" pageId="1" pageNumber="101">Fig. 4</figureCitation>
). Anterior part of idiosoma overlapping posterior part of
<taxonomicName id="3CE64DF3FFB9FFEB855E0985FA89949C" box="[1150,1285,1735,1760]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="1" pageNumber="101" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
like a collar. All dorsal shields smooth, in a few specimens each tergite is posteriorly sparsely ornamented with barely visible small dimples; all dorsal setae slightly barbed and moderately blunt-ending, except smooth, attenuate and pointed setae
<date id="8F5810B0FFB9FFEB81F70876FF67952C" box="[215,235,1844,1872]" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB81F70876FF6F9537" box="[215,227,1844,1867]" italics="true" pageId="1" pageNumber="101">v</emphasis>
1
</date>
and smooth and needle-like setae
<date id="8F5810B0FFB9FFEB83500876FD09952C" box="[624,645,1844,1872]" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB83500876FDF09537" box="[624,636,1844,1867]" italics="true" pageId="1" pageNumber="101">v</emphasis>
2
</date>
; prodorsal shield (PrS) (
<figureCitation id="63DD2AF5FFB9FFEB82BD0871FC609537" box="[925,1004,1843,1868]" captionStart="FIGURE 4" captionStartId="3.[151,250,1915,1937]" captionTargetBox="[230,1356,481,1893]" captionTargetId="figure@3.[230,1356,481,1893]" captionTargetPageId="3" captionText="FIGURE 4. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): dorsum of the body. Scale bar 50 µm." httpUri="https://zenodo.org/record/235219/files/figure.png" pageId="1" pageNumber="101">Figs. 4</figureCitation>
and 15) broad flask-shaped, narrower and concave in its anterolateral margins, with one pair of thin-stemmed capitate trichobothria (
<emphasis id="C992EA62FFB9FFEB85E50818FB50950D" box="[1221,1244,1882,1905]" italics="true" pageId="1" pageNumber="101">sc</emphasis>
1) and three pairs of setae
<date id="8F5810B0FFB9FFEB81DB08C3FEBD95E5" box="[251,305,1921,1949]" pageId="1" pageNumber="101" value="1917-05-01">
<emphasis id="C992EA62FFB9FFEB81DB08C3FE8B95E4" box="[251,263,1921,1944]" italics="true" pageId="1" pageNumber="101">v</emphasis>
1 17
</date>
(1718),
<date id="8F5810B0FFB9FFEB808508C3FE6C95E5" box="[421,480,1920,1949]" pageId="1" pageNumber="101" value="1916-05-02">
<emphasis id="C992EA62FFB9FFEB808508C3FE3D95E4" box="[421,433,1921,1944]" italics="true" pageId="1" pageNumber="101">v</emphasis>
2 16
</date>
(1616) and
<emphasis id="C992EA62FFB9FFEB83A008C3FD1B95E4" box="[640,663,1921,1944]" italics="true" pageId="1" pageNumber="101">sc</emphasis>
2 39 (3739),
<date id="8F5810B0FFB9FFEB821608C3FCC695E1" box="[822,842,1921,1949]" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB821608C3FCCE95E4" box="[822,834,1921,1944]" italics="true" pageId="1" pageNumber="101">v</emphasis>
1
</date>
slightly longer
<date id="8F5810B0FFB9FFEB852008C3FB9895E1" box="[1024,1044,1921,1949]" pageId="1" pageNumber="101">
<emphasis id="C992EA62FFB9FFEB852008C3FB8095E4" box="[1024,1036,1921,1944]" italics="true" pageId="1" pageNumber="101">v</emphasis>
2
</date>
, both shorter than seta
<emphasis id="C992EA62FFB9FFEB840B08C3FACE95E4" box="[1323,1346,1921,1944]" italics="true" pageId="1" pageNumber="101">sc</emphasis>
2; ovoid stigmata located anterolaterally on prodorsum, associated with conspicuous atria; all cupuli (
<emphasis id="C992EA62FFB9FFEB858108EAFB3B95C3" box="[1185,1207,1960,1983]" italics="true" pageId="1" pageNumber="101">ia</emphasis>
,
<emphasis id="C992EA62FFB9FFEB85E408EAFB6C95C3" box="[1220,1248,1960,1983]" italics="true" pageId="1" pageNumber="101">im</emphasis>
, and
<emphasis id="C992EA62FFB9FFEB843D08E5FABF95C3" box="[1309,1331,1959,1983]" italics="true" pageId="1" pageNumber="101">ih</emphasis>
) evident, rounded; on tergite C
<emphasis id="C992EA62FFB9FFEB80BB088EFE2B959F" box="[411,423,1996,2019]" italics="true" pageId="1" pageNumber="101">c</emphasis>
2 39 (3639) slightly longer than
<emphasis id="C992EA62FFB9FFEB820C088EFCB4959F" box="[812,824,1996,2019]" italics="true" pageId="1" pageNumber="101">c</emphasis>
1 33 (2933); setae
<emphasis id="C992EA62FFB9FFEB853B0889FBA5959F" box="[1051,1065,1995,2019]" italics="true" pageId="1" pageNumber="101">d</emphasis>
37 (3236) on tergite D barely reaching the base of
<emphasis id="C992EA62FFBAFFE880A60FDAFE0292CC" box="[390,398,152,176]" italics="true" pageId="2" pageNumber="102">f</emphasis>
43 (3842) on tergite EF; tergite EF with barely visible small dimples, setae
<emphasis id="C992EA62FFBAFFE884290FDBFA9992CC" box="[1289,1301,153,176]" italics="true" pageId="2" pageNumber="102">e</emphasis>
19 (1721) barely exceeding the posterior margin of tergite EF, setae
<emphasis id="C992EA62FFBAFFE882380FFEFCAC92A8" box="[792,800,188,212]" italics="true" pageId="2" pageNumber="102">f</emphasis>
approximately 2.2 times longer than
<emphasis id="C992EA62FFBAFFE8859F0FFFFB4792A8" box="[1215,1227,189,212]" italics="true" pageId="2" pageNumber="102">e</emphasis>
; on tergite H setae
<emphasis id="C992EA62FFBAFFE881B70FA2FF299284" box="[151,165,224,248]" italics="true" pageId="2" pageNumber="102">h</emphasis>
1 43 (3843) more than 3.5 times longer than setae
<emphasis id="C992EA62FFBAFFE883FC0FA2FD669284" box="[732,746,224,248]" italics="true" pageId="2" pageNumber="102">h</emphasis>
2 12 (911). Distance between setae:
<date id="8F5810B0FFBAFFE885A20FA3FB569284" box="[1154,1242,224,252]" pageId="2" pageNumber="102" value="2011-05-01" valueMax="2011-05-01" valueMin="2011-05-01">
<emphasis id="C992EA62FFBAFFE885A20FA3FB029284" box="[1154,1166,225,248]" italics="true" pageId="2" pageNumber="102">v</emphasis>
1
<emphasis id="C992EA62FFBAFFE885840FA3FB3C9284" box="[1188,1200,225,248]" italics="true" pageId="2" pageNumber="102">v</emphasis>
1 11
</date>
(1114),
<date id="8F5810B0FFBAFFE884660FA3FA109284" box="[1350,1436,224,252]" pageId="2" pageNumber="102" value="1917-05-01" valueMax="1917-05-02" valueMin="1917-05-01">
<emphasis id="C992EA62FFBAFFE884660FA3FADE9284" box="[1350,1362,225,248]" italics="true" pageId="2" pageNumber="102">v</emphasis>
1
<emphasis id="C992EA62FFBAFFE884480FA3FAF89284" box="[1384,1396,225,248]" italics="true" pageId="2" pageNumber="102">v</emphasis>
2 17
</date>
(1517),
<date id="8F5810B0FFBAFFE881DD0E4AFED99363" box="[253,341,263,291]" pageId="2" pageNumber="102" value="1938-05-02" valueMax="1938-05-02" valueMin="1938-05-02">
<emphasis id="C992EA62FFBAFFE881DD0E4AFE859363" box="[253,265,264,287]" italics="true" pageId="2" pageNumber="102">v</emphasis>
2
<emphasis id="C992EA62FFBAFFE8803F0E4AFEA79363" box="[287,299,264,287]" italics="true" pageId="2" pageNumber="102">v</emphasis>
2 38
</date>
(3439),
<emphasis id="C992EA62FFBAFFE880E20E4AFE559363" box="[450,473,264,287]" italics="true" pageId="2" pageNumber="102">sc</emphasis>
2
<emphasis id="C992EA62FFBAFFE880CF0E4AFD8A9363" box="[495,518,264,287]" italics="true" pageId="2" pageNumber="102">sc</emphasis>
2 49 (4448),
<emphasis id="C992EA62FFBAFFE883BC0E4AFD249363" box="[668,680,264,287]" italics="true" pageId="2" pageNumber="102">c</emphasis>
1
<emphasis id="C992EA62FFBAFFE8839F0E4AFD479363" box="[703,715,264,287]" italics="true" pageId="2" pageNumber="102">c</emphasis>
1 38 (3439),
<emphasis id="C992EA62FFBAFFE8827F0E4AFCE79363" box="[863,875,264,287]" italics="true" pageId="2" pageNumber="102">c</emphasis>
1
<emphasis id="C992EA62FFBAFFE882A30E4AFC039363" box="[899,911,264,287]" italics="true" pageId="2" pageNumber="102">c</emphasis>
2 24 (2224),
<emphasis id="C992EA62FFBAFFE885050E4AFBBD9363" box="[1061,1073,264,287]" italics="true" pageId="2" pageNumber="102">c</emphasis>
2
<emphasis id="C992EA62FFBAFFE885690E4AFBD99363" box="[1097,1109,264,287]" italics="true" pageId="2" pageNumber="102">c</emphasis>
2 83 (7685),
<emphasis id="C992EA62FFBAFFE885C90E45FA9F9363" box="[1257,1299,263,287]" italics="true" pageId="2" pageNumber="102">dd</emphasis>
48 (4653),
<emphasis id="C992EA62FFBAFFE881B70E6CFF319339" box="[151,189,302,325]" italics="true" pageId="2" pageNumber="102">ee</emphasis>
60 (5462),
<emphasis id="C992EA62FFBAFFE8806E0E6CFEFD933A" box="[334,369,302,326]" italics="true" pageId="2" pageNumber="102">ef</emphasis>
7 (57),
<emphasis id="C992EA62FFBAFFE880F70E6CFE79933A" box="[471,501,302,326]" italics="true" pageId="2" pageNumber="102">ff</emphasis>
51 (4653),
<emphasis id="C992EA62FFBAFFE883A50E6FFD1F9339" box="[645,659,301,325]" italics="true" pageId="2" pageNumber="102">h</emphasis>
1
<emphasis id="C992EA62FFBAFFE883890E6FFD3B9339" box="[681,695,301,325]" italics="true" pageId="2" pageNumber="102">h</emphasis>
1 40 (3845),
<emphasis id="C992EA62FFBAFFE8826F0E6FFCD19339" box="[847,861,301,325]" italics="true" pageId="2" pageNumber="102">h</emphasis>
2
<emphasis id="C992EA62FFBAFFE882530E6FFC0D9339" box="[883,897,301,325]" italics="true" pageId="2" pageNumber="102">h</emphasis>
2 51 (4856),
<emphasis id="C992EA62FFBAFFE885390E6FFBAB9339" box="[1049,1063,301,325]" italics="true" pageId="2" pageNumber="102">h</emphasis>
1
<emphasis id="C992EA62FFBAFFE8851D0E6FFBC79339" box="[1085,1099,301,325]" italics="true" pageId="2" pageNumber="102">h</emphasis>
2 6 (57).
</paragraph>
<caption id="AF9966F8FFBAFFE881B70C32FC1C91D9" httpUri="https://zenodo.org/record/235218/files/figure.png" pageId="2" pageNumber="102" targetBox="[302,1285,384,856]" targetPageId="2">
<paragraph id="FB593670FFBAFFE881B70C32FC1C91D9" blockId="2.[151,1436,880,933]" pageId="2" pageNumber="102">
<emphasis id="C992EA62FFBAFFE881B70C32FEB291FA" bold="true" box="[151,318,880,903]" pageId="2" pageNumber="102">FIGURES 13.</emphasis>
<taxonomicName id="3CE64DF3FFBAFFE880690C33FC5291FB" authority="Katlav and Hajiqanbar" authorityName="Katlav and Hajiqanbar" box="[329,990,881,903]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="2" pageNumber="102" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFBAFFE880690C33FD6991FB" box="[329,741,881,903]" italics="true" pageId="2" pageNumber="102">Pseudopygmephorellus mazandaranicus</emphasis>
Katlav and Hajiqanbar
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFBAFFE882C90C33FBB191FA" box="[1001,1085,881,902]" pageId="2" pageNumber="102" rank="species">
<emphasis id="C992EA62FFBAFFE882C90C33FBB191FA" bold="true" box="[1001,1085,881,902]" pageId="2" pageNumber="102">sp. nov.</emphasis>
</taxonomicNameLabel>
(female): 1.
<taxonomicName id="3CE64DF3FFBAFFE885ED0C33FADC91FB" box="[1229,1360,881,903]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="101" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
, dorsal view; 2.
<taxonomicName id="3CE64DF3FFBAFFE881CF0CCDFEFD91D9" box="[239,369,911,933]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="101" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
, ventral view; 3. Pharyngeal system. Scale bar 10µm.
</paragraph>
</caption>
<paragraph id="FB593670FFBAFFE881E70C8FFF75979F" blockId="2.[151,1436,972,2017]" pageId="2" pageNumber="102">
<emphasis id="C992EA62FFBAFFE881E70C8FFE0B9199" box="[199,391,973,997]" italics="true" pageId="2" pageNumber="102">Idiosomal venter</emphasis>
(
<figureCitation id="63DD2AF5FFBAFFE880B80C8EFE539199" box="[408,479,972,997]" captionStart="FIGURE 5" captionStartId="4.[151,250,1480,1502]" captionTargetBox="[254,1326,193,1465]" captionTargetId="figure@4.[254,1327,193,1465]" captionTargetPageId="4" captionText="FIGURE 5. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): venter of the body. Scale bar 50 µm." httpUri="https://zenodo.org/record/235220/files/figure.png" pageId="2" pageNumber="102">Fig. 5</figureCitation>
). Ventral plates distinctly smooth; all propodosomal apodemes (ap1, ap2, appr and apsej) fully developed; ap1 reaching to presternal apodeme (appr), forming well scelerotized Y-shape juncture with appr; ap2 uniting with appr, with moderately thickened and arched lateral ends; sejugal apodeme (apsej) uniting to appr, medially forming a tiny protuberance; ap3 thickened and well sclerotized; ap4 partly developed, extending beyond seta 3
<emphasis id="C992EA62FFBAFFE880140B1CFECC9609" box="[308,320,1118,1141]" italics="true" pageId="2" pageNumber="102">c</emphasis>
; both ap4 and ap5 joined with poststernal apodeme (appo); ap5 reduced to a lateral fragment; all epimeral setae smooth and pointed except setae 1
<emphasis id="C992EA62FFBAFFE883EC0BC3FD5496E4" box="[716,728,1153,1176]" italics="true" pageId="2" pageNumber="102">c</emphasis>
and 1
<emphasis id="C992EA62FFBAFFE882010BC3FCA396E5" box="[801,815,1153,1177]" italics="true" pageId="2" pageNumber="102">b</emphasis>
which are moderately barbed and pointed; coxisternal plate I with three pairs of setae 1
<emphasis id="C992EA62FFBAFFE8832B0BE4FD9596C1" box="[523,537,1190,1213]" italics="true" pageId="2" pageNumber="102">a</emphasis>
11 (912), 1
<emphasis id="C992EA62FFBAFFE8838C0BE7FD3696C1" box="[684,698,1189,1213]" italics="true" pageId="2" pageNumber="102">b</emphasis>
14 (1316) and 1
<emphasis id="C992EA62FFBAFFE882A40BE4FC1C96C1" box="[900,912,1190,1213]" italics="true" pageId="2" pageNumber="102">c</emphasis>
12 (1113), setae 1
<emphasis id="C992EA62FFBAFFE885510BE7FBF396C1" box="[1137,1151,1189,1213]" italics="true" pageId="2" pageNumber="102">b</emphasis>
obviously longest on the plate, setae 1
<emphasis id="C992EA62FFBAFFE8800A0B8BFEBA969C" box="[298,310,1225,1248]" italics="true" pageId="2" pageNumber="102">c</emphasis>
situated adjacent to ap1; coxisternal plate II with two pairs of subequal setae 2
<emphasis id="C992EA62FFBAFFE885970B8BFB49969C" box="[1207,1221,1225,1248]" italics="true" pageId="2" pageNumber="102">a</emphasis>
13 (1113) and 2
<emphasis id="C992EA62FFBAFFE884AF0B8BFA17969C" box="[1423,1435,1225,1248]" italics="true" pageId="2" pageNumber="102">c</emphasis>
12 (1213); coxisternal plate III with three pairs of setae 3
<emphasis id="C992EA62FFBAFFE882760BACFCE89779" box="[854,868,1262,1285]" italics="true" pageId="2" pageNumber="102">a</emphasis>
14 (1213), 3
<emphasis id="C992EA62FFBAFFE885320BAFFBAC9779" box="[1042,1056,1261,1285]" italics="true" pageId="2" pageNumber="102">b</emphasis>
15 (1416) and 3
<emphasis id="C992EA62FFBAFFE885DC0BACFA849779" box="[1276,1288,1262,1285]" italics="true" pageId="2" pageNumber="102">c</emphasis>
13 (1213), coxisternal plate IV with three pairs of setae 4
<emphasis id="C992EA62FFBAFFE883860A53FD389754" box="[678,692,1297,1320]" italics="true" pageId="2" pageNumber="102">a</emphasis>
13 (1113), 4
<emphasis id="C992EA62FFBAFFE882760A53FCE89755" box="[854,868,1297,1321]" italics="true" pageId="2" pageNumber="102">b</emphasis>
15 (1314) and 4
<emphasis id="C992EA62FFBAFFE885110A53FBB19754" box="[1073,1085,1297,1320]" italics="true" pageId="2" pageNumber="102">c</emphasis>
18 (16
<emphasis id="C992EA62FFBAFFE885B00A52FB129755" bold="true" box="[1168,1182,1296,1321]" pageId="2" pageNumber="102"></emphasis>
19); 4
<emphasis id="C992EA62FFBAFFE885C10A53FB619754" box="[1249,1261,1297,1320]" italics="true" pageId="2" pageNumber="102">c</emphasis>
longest among epimeral setae; posterolateral margins of poststernal plate weakly convex and undulated; posterior margin of aggenital plate weakly concave medially; anterior genital sclerite (ags) small and mushroom-like, posterior genital sclerite (pgs) tiny and oval; pseudanal plate (PS) with three pairs of setae
<emphasis id="C992EA62FFBAFFE882F80A3CFC7D97E9" box="[984,1009,1406,1429]" italics="true" pageId="2" pageNumber="102">ps</emphasis>
1 12 (1213),
<emphasis id="C992EA62FFBAFFE885A20A3CFB1797E9" box="[1154,1179,1406,1429]" italics="true" pageId="2" pageNumber="102">ps</emphasis>
2 13 (1213) and
<emphasis id="C992EA62FFBAFFE8847B0A3CFAFF97E9" box="[1371,1395,1406,1429]" italics="true" pageId="2" pageNumber="102">ps</emphasis>
3 16 (1617), seta
<emphasis id="C992EA62FFBAFFE8801B0AE6FED897C7" box="[315,340,1444,1467]" italics="true" pageId="2" pageNumber="102">ps</emphasis>
3 weakly barbed and blunt ending, setae
<emphasis id="C992EA62FFBAFFE882180AE6FCDC97C7" box="[824,848,1444,1467]" italics="true" pageId="2" pageNumber="102">ps</emphasis>
1 and
<emphasis id="C992EA62FFBAFFE882B40AE6FC2097C7" box="[916,940,1444,1467]" italics="true" pageId="2" pageNumber="102">ps</emphasis>
2 somewhat delicate and needle-like, close together.
</paragraph>
<paragraph id="FB593670FFBAFFE881E70AADFC449551" blockId="2.[151,1436,972,2017]" pageId="2" pageNumber="102">
<emphasis id="C992EA62FFBAFFE881E70AADFF70947A" box="[199,252,1519,1542]" italics="true" pageId="2" pageNumber="102">Legs</emphasis>
(
<figureCitation id="63DD2AF5FFBAFFE8802B0AACFE09947B" box="[267,389,1518,1543]" captionStart-0="FIGURES 6 8" captionStart-1="FIGURES 9 10" captionStartId-0="5.[151,264,1948,1971]" captionStartId-1="6.[151,264,1687,1710]" captionTargetBox-0="[162,1425,517,1935]" captionTargetBox-1="[155,1424,237,1670]" captionTargetId-0="figure@5.[162,1426,517,1936]" captionTargetId-1="figure@6.[155,1424,193,1670]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 6 8. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 6 7. Legs I II respectively. Scale bar 20 µm; 8. Comparison of length of solenidia in two close species: a. P. mazandaranicus sp. nov, b. P. szekessyi (Mahunka, 1970). Scale bar 10 µm." captionText-1="FIGURES 9 10. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 9 10. Legs III IV respectively. Scale bar 20 µm." httpUri-0="https://zenodo.org/record/235221/files/figure.png" httpUri-1="https://zenodo.org/record/235222/files/figure.png" pageId="2" pageNumber="102">Figs. 610</figureCitation>
). Leg
<date id="8F5810B0FFBAFFE880EA0AADFE7A947B" box="[458,502,1519,1543]" pageId="2" pageNumber="102" value="1971-01">I 71</date>
(67
<emphasis id="C992EA62FFBAFFE883020AACFDBC947B" bold="true" box="[546,560,1518,1543]" pageId="2" pageNumber="102"></emphasis>
72) (
<figureCitation id="63DD2AF5FFBAFFE883440AACFD25947B" box="[612,681,1518,1543]" captionStart="FIGURES 6 8" captionStartId="5.[151,264,1948,1971]" captionTargetBox="[162,1425,517,1935]" captionTargetId="figure@5.[162,1426,517,1936]" captionTargetPageId="5" captionText="FIGURES 6 8. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 6 7. Legs I II respectively. Scale bar 20 µm; 8. Comparison of length of solenidia in two close species: a. P. mazandaranicus sp. nov, b. P. szekessyi (Mahunka, 1970). Scale bar 10 µm." httpUri="https://zenodo.org/record/235221/files/figure.png" pageId="2" pageNumber="102">Fig. 6</figureCitation>
): thicker and shorter than other legs with a massive sickle-like claw opposed to a thick counterpiece (forming clasper-like structure); setal formula (solenidia in parentheses): Tr1-Fe4- Ge3-TiTa17(
<emphasis id="C992EA62FFBAFFE880070975FEB79432" box="[295,315,1591,1614]" italics="true" pageId="2" pageNumber="102">ω</emphasis>
1,
<emphasis id="C992EA62FFBAFFE880740975FEE49432" box="[340,360,1591,1614]" italics="true" pageId="2" pageNumber="102">ω</emphasis>
2,
<emphasis id="C992EA62FFBAFFE880A10975FE1C9432" box="[385,400,1591,1614]" italics="true" pageId="2" pageNumber="102">φ</emphasis>
1,
<emphasis id="C992EA62FFBAFFE880890975FE349432" box="[425,440,1591,1614]" italics="true" pageId="2" pageNumber="102">φ</emphasis>
2). Tibiotarsus: all solenidia including
<emphasis id="C992EA62FFBAFFE882530975FC0B9432" box="[883,903,1591,1614]" italics="true" pageId="2" pageNumber="102">ω</emphasis>
1 6 (56),
<emphasis id="C992EA62FFBAFFE882DA0975FB829432" box="[1018,1038,1591,1614]" italics="true" pageId="2" pageNumber="102">ω</emphasis>
2 3 (33),
<emphasis id="C992EA62FFBAFFE885A10975FB1C9432" box="[1153,1168,1591,1614]" italics="true" pageId="2" pageNumber="102">φ</emphasis>
1 5 (55) and
<emphasis id="C992EA62FFBAFFE8840A0975FAB59432" box="[1322,1337,1591,1614]" italics="true" pageId="2" pageNumber="102">φ</emphasis>
2 3 (33) finger-shaped;
<emphasis id="C992EA62FFBAFFE8801E091CFEDE9409" box="[318,338,1630,1653]" italics="true" pageId="2" pageNumber="102">ω</emphasis>
2 and
<emphasis id="C992EA62FFBAFFE880AF091CFE129409" box="[399,414,1630,1653]" italics="true" pageId="2" pageNumber="102">φ</emphasis>
2 tiny and subequal;
<emphasis id="C992EA62FFBAFFE8835C091CFD079409" box="[636,651,1630,1653]" italics="true" pageId="2" pageNumber="102">φ</emphasis>
1 slightly shorter, but wider than
<emphasis id="C992EA62FFBAFFE882D2091CFB8A9409" box="[1010,1030,1630,1653]" italics="true" pageId="2" pageNumber="102">ω</emphasis>
1 (
<figureCitation id="63DD2AF5FFBAFFE8853A091FFBD7940A" box="[1050,1115,1629,1654]" captionStart="FIGURES 6 8" captionStartId="5.[151,264,1948,1971]" captionTargetBox="[162,1425,517,1935]" captionTargetId="figure@5.[162,1426,517,1936]" captionTargetPageId="5" captionText="FIGURES 6 8. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 6 7. Legs I II respectively. Scale bar 20 µm; 8. Comparison of length of solenidia in two close species: a. P. mazandaranicus sp. nov, b. P. szekessyi (Mahunka, 1970). Scale bar 10 µm." httpUri="https://zenodo.org/record/235221/files/figure.png" pageId="2" pageNumber="102">Fig. 8</figureCitation>
a); eupathidia
<emphasis id="C992EA62FFBAFFE885DA091FFA9F9409" box="[1274,1299,1629,1653]" italics="true" pageId="2" pageNumber="102">ft</emphasis>
,
<emphasis id="C992EA62FFBAFFE88401091FFAB39409" box="[1313,1343,1629,1653]" italics="true" pageId="2" pageNumber="102">ft”</emphasis>
,
<emphasis id="C992EA62FFBAFFE8846D091CFAE69409" box="[1357,1386,1630,1653]" italics="true" pageId="2" pageNumber="102">tc</emphasis>
,
<emphasis id="C992EA62FFBAFFE88458091CFA109409" box="[1400,1436,1630,1653]" italics="true" pageId="2" pageNumber="102">tc”</emphasis>
and
<emphasis id="C992EA62FFBAFFE881E909C6FF6C94E7" box="[201,224,1668,1691]" italics="true" pageId="2" pageNumber="102">p</emphasis>
moderately blunt-ending; setae
<emphasis id="C992EA62FFBAFFE8837309C6FDED94E0" box="[595,609,1668,1692]" italics="true" pageId="2" pageNumber="102">d</emphasis>
,
<emphasis id="C992EA62FFBAFFE8835209C6FD0894E0" box="[626,644,1668,1692]" italics="true" pageId="2" pageNumber="102">l</emphasis>
and
<emphasis id="C992EA62FFBAFFE8839F09C6FD5A94E0" box="[703,726,1668,1692]" italics="true" pageId="2" pageNumber="102">l”</emphasis>
barbed and pointed,
<emphasis id="C992EA62FFBAFFE882EC09C6FC5194E0" box="[972,989,1668,1692]" italics="true" pageId="2" pageNumber="102">l</emphasis>
28 (27
<emphasis id="C992EA62FFBAFFE8851209C6FBCC94E1" bold="true" box="[1074,1088,1668,1693]" pageId="2" pageNumber="102"></emphasis>
30) 1.5 times as long as
<emphasis id="C992EA62FFBAFFE8847F09C6FAFA94E0" box="[1375,1398,1668,1692]" italics="true" pageId="2" pageNumber="102">l”</emphasis>
18 (18
<emphasis id="C992EA62FFBAFFE8819C09EAFF4694BD" bold="true" box="[188,202,1704,1729]" pageId="2" pageNumber="102"></emphasis>
20); remaining tibiotarsal setae smooth and pointed;
<emphasis id="C992EA62FFBAFFE8821309EAFCC094BC" box="[819,844,1704,1728]" italics="true" pageId="2" pageNumber="102">ft</emphasis>
slightly longer than
<emphasis id="C992EA62FFBAFFE8856309EAFBEE94BC" box="[1091,1122,1704,1728]" italics="true" pageId="2" pageNumber="102">ft”</emphasis>
; setae
<emphasis id="C992EA62FFBAFFE8859709EAFB4294C3" box="[1207,1230,1704,1727]" italics="true" pageId="2" pageNumber="102">p</emphasis>
,
<emphasis id="C992EA62FFBAFFE885C109EAFA8994C3" box="[1249,1285,1704,1727]" italics="true" pageId="2" pageNumber="102">pv</emphasis>
,
<emphasis id="C992EA62FFBAFFE8843609EAFACC94C3" box="[1302,1344,1704,1727]" italics="true" pageId="2" pageNumber="102">pv”</emphasis>
and
<emphasis id="C992EA62FFBAFFE8845E09EAFA1794C3" box="[1406,1435,1704,1727]" italics="true" pageId="2" pageNumber="102">tc</emphasis>
subequal;
<emphasis id="C992EA62FFBAFFE88029098EFEA8949F" box="[265,292,1740,1763]" italics="true" pageId="2" pageNumber="102">v”</emphasis>
and
<emphasis id="C992EA62FFBAFFE8807C098EFEF79498" box="[348,379,1740,1764]" italics="true" pageId="2" pageNumber="102">pl</emphasis>
whipped and subequal;
<emphasis id="C992EA62FFBAFFE883AB098EFD2D949F" box="[651,673,1740,1763]" italics="true" pageId="2" pageNumber="102">v</emphasis>
shortest on tibiotarsus. Genu:
<emphasis id="C992EA62FFBAFFE882D7098EFB849498" box="[1015,1032,1740,1764]" italics="true" pageId="2" pageNumber="102">l</emphasis>
and
<emphasis id="C992EA62FFBAFFE8851F098EFBDA9498" box="[1087,1110,1740,1764]" italics="true" pageId="2" pageNumber="102">l”</emphasis>
moderately barbed,
<emphasis id="C992EA62FFBAFFE8841C098EFAC19498" box="[1340,1357,1740,1764]" italics="true" pageId="2" pageNumber="102">l</emphasis>
longer than
<emphasis id="C992EA62FFBAFFE881EF09B2FF6B9574" box="[207,231,1776,1800]" italics="true" pageId="2" pageNumber="102">l”</emphasis>
;
<emphasis id="C992EA62FFBAFFE881D709B2FE81957B" box="[247,269,1776,1799]" italics="true" pageId="2" pageNumber="102">v</emphasis>
smooth and pointed, shortest on genu. Femur: seta
<emphasis id="C992EA62FFBAFFE8827A09B2FCE49574" box="[858,872,1776,1800]" italics="true" pageId="2" pageNumber="102">d</emphasis>
hooked;
<emphasis id="C992EA62FFBAFFE882F209B2FC669574" box="[978,1002,1776,1800]" italics="true" pageId="2" pageNumber="102">l”</emphasis>
nearly twice longer than
<emphasis id="C992EA62FFBAFFE8842D09B2FA929574" box="[1293,1310,1776,1800]" italics="true" pageId="2" pageNumber="102">l</emphasis>
;
<emphasis id="C992EA62FFBAFFE8840E09B2FACC9574" box="[1326,1344,1776,1800]" italics="true" pageId="2" pageNumber="102">l</emphasis>
slightly curved and blunt-ending. Trochanter: seta
<emphasis id="C992EA62FFBAFFE883520856FD049557" box="[626,648,1812,1835]" italics="true" pageId="2" pageNumber="102">v</emphasis>
slightly barbed and pointed.
</paragraph>
<paragraph id="FB593670FFBAFFE981E7087BFB8C92A9" blockId="2.[151,1436,972,2017]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="103" pageId="2" pageNumber="102">
Leg
<date id="8F5810B0FFBAFFE881D9087BFEA2952C" box="[249,302,1848,1872]" pageId="2" pageNumber="102" value="1989-02">II 89</date>
(87
<emphasis id="C992EA62FFBAFFE8807A087AFEE4952D" bold="true" box="[346,360,1848,1873]" pageId="2" pageNumber="102"></emphasis>
92) (
<figureCitation id="63DD2AF5FFBAFFE880BB087AFE6C952C" box="[411,480,1848,1873]" captionStart="FIGURES 6 8" captionStartId="5.[151,264,1948,1971]" captionTargetBox="[162,1425,517,1935]" captionTargetId="figure@5.[162,1426,517,1936]" captionTargetPageId="5" captionText="FIGURES 6 8. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 6 7. Legs I II respectively. Scale bar 20 µm; 8. Comparison of length of solenidia in two close species: a. P. mazandaranicus sp. nov, b. P. szekessyi (Mahunka, 1970). Scale bar 10 µm." httpUri="https://zenodo.org/record/235221/files/figure.png" pageId="2" pageNumber="102">Fig. 7</figureCitation>
): setal formula: Tr1-Fe3-Ge2-Ti4(
<emphasis id="C992EA62FFBAFFE88241087BFCFC952C" box="[865,880,1849,1872]" italics="true" pageId="2" pageNumber="102">φ</emphasis>
)-Ta6(
<emphasis id="C992EA62FFBAFFE88296087BFC46952C" box="[950,970,1849,1872]" italics="true" pageId="2" pageNumber="102">ω</emphasis>
). Tarsus: dorso-distally with a sclerotized cone-like protrusion; solenidion
<emphasis id="C992EA62FFBAFFE8832A081FFD929508" box="[522,542,1885,1908]" italics="true" pageId="2" pageNumber="102">ω</emphasis>
6 (66), finger-shaped;
<emphasis id="C992EA62FFBAFFE88216081EFCD6950F" box="[822,858,1884,1907]" italics="true" pageId="2" pageNumber="102">pv</emphasis>
,
<emphasis id="C992EA62FFBAFFE8824A081EFC18950F" box="[874,916,1884,1907]" italics="true" pageId="2" pageNumber="102">pv”</emphasis>
,
<emphasis id="C992EA62FFBAFFE88285081EFC459508" box="[933,969,1884,1908]" italics="true" pageId="2" pageNumber="102">pl”</emphasis>
and
<emphasis id="C992EA62FFBAFFE88525081EFBAF950F" box="[1029,1059,1884,1907]" italics="true" pageId="2" pageNumber="102">tc</emphasis>
barbed and pointed;
<emphasis id="C992EA62FFBAFFE88438081EFAB0950F" box="[1304,1340,1884,1907]" italics="true" pageId="2" pageNumber="102">tc”</emphasis>
slender, smooth and pointed;
<emphasis id="C992EA62FFBAFFE880B108C2FE2495EB" box="[401,424,1920,1943]" italics="true" pageId="2" pageNumber="102">u</emphasis>
moderately blunt-ending. Tibia: solenidion
<emphasis id="C992EA62FFBAFFE8828D08C3FC3095E4" box="[941,956,1921,1944]" italics="true" pageId="2" pageNumber="102">φ</emphasis>
2 (22) very small and hard to discern, situated in a shallow depression;
<emphasis id="C992EA62FFBAFFE8832E08E6FDA895C7" box="[526,548,1956,1979]" italics="true" pageId="2" pageNumber="102">v</emphasis>
and
<emphasis id="C992EA62FFBAFFE8837D08E6FDE295C0" box="[605,622,1956,1980]" italics="true" pageId="2" pageNumber="102">l</emphasis>
barbed, pointed and subequal;
<emphasis id="C992EA62FFBAFFE882F008E6FC5295C0" box="[976,990,1956,1980]" italics="true" pageId="2" pageNumber="102">d</emphasis>
smooth and pointed;
<emphasis id="C992EA62FFBAFFE885F508E6FB7C95C7" box="[1237,1264,1956,1979]" italics="true" pageId="2" pageNumber="102">v”</emphasis>
attenuated and pointed, longest on tibia. Genu:
<emphasis id="C992EA62FFBAFFE88334088AFDA9959C" box="[532,549,1992,2016]" italics="true" pageId="2" pageNumber="102">l</emphasis>
barbed and pointed, reaching base of tarsal solenidion; seta
<emphasis id="C992EA62FFBAFFE885D3088AFA8595A3" box="[1267,1289,1992,2015]" italics="true" pageId="2" pageNumber="102">v</emphasis>
pointed and slightly barbed. Femur:
<emphasis id="C992EA62FFBBFFE9808A0FDAFE3492CC" box="[426,440,152,176]" italics="true" pageId="3" pageNumber="103">d</emphasis>
barbed and blunt-ending, seta
<emphasis id="C992EA62FFBBFFE9823A0FDAFCA092CC" box="[794,812,152,176]" italics="true" pageId="3" pageNumber="103">l</emphasis>
smooth, stiff and blunt-ending,
<emphasis id="C992EA62FFBBFFE985BC0FDAFB3492D3" box="[1180,1208,152,175]" italics="true" pageId="3" pageNumber="103">v”</emphasis>
slightly barbed and pointed, subequal to
<emphasis id="C992EA62FFBBFFE980A00FFEFE0292A8" box="[384,398,188,212]" italics="true" pageId="3" pageNumber="103">d</emphasis>
. Trochanter:
<emphasis id="C992EA62FFBBFFE983040FFEFDB692AF" box="[548,570,188,211]" italics="true" pageId="3" pageNumber="103">v</emphasis>
indistinctly slightly barbed and pointed.
</paragraph>
<paragraph id="FB593670FFBBFFE981E70FA2FD9793D1" blockId="3.[151,1436,151,429]" pageId="3" pageNumber="103">
Leg
<date id="8F5810B0FFBBFFE981D90FA2FEB49284" box="[249,312,224,248]" pageId="3" pageNumber="103" value="1988-03">III 88</date>
(86
<emphasis id="C992EA62FFBBFFE980440F9DFEFE9284" bold="true" box="[356,370,223,248]" pageId="3" pageNumber="103"></emphasis>
92) (
<figureCitation id="63DD2AF5FFBBFFE980860F9DFE679284" box="[422,491,223,248]" captionStart="FIGURES 9 10" captionStartId="6.[151,264,1687,1710]" captionTargetBox="[155,1424,237,1670]" captionTargetId="figure@6.[155,1424,193,1670]" captionTargetPageId="6" captionText="FIGURES 9 10. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 9 10. Legs III IV respectively. Scale bar 20 µm." httpUri="https://zenodo.org/record/235222/files/figure.png" pageId="3" pageNumber="103">Fig. 9</figureCitation>
): setal formula: Tr1-Fe2-Ge2-Ti4(
<emphasis id="C992EA62FFBBFFE9824D0FA3FCF09284" box="[877,892,225,248]" italics="true" pageId="3" pageNumber="103">φ</emphasis>
) -Ta6. Tarsus:
<emphasis id="C992EA62FFBBFFE9853D0FA2FBCC928B" box="[1053,1088,224,247]" italics="true" pageId="3" pageNumber="103">pv</emphasis>
and
<emphasis id="C992EA62FFBBFFE985570FA2FB2D928B" box="[1143,1185,224,247]" italics="true" pageId="3" pageNumber="103">pv”</emphasis>
smooth and subequal;
<emphasis id="C992EA62FFBBFFE981B70E46FF399367" box="[151,181,260,283]" italics="true" pageId="3" pageNumber="103">tc</emphasis>
moderately barbed, subequal to smooth
<emphasis id="C992EA62FFBBFFE9835B0E46FD129367" box="[635,670,260,283]" italics="true" pageId="3" pageNumber="103">tc”</emphasis>
;
<emphasis id="C992EA62FFBBFFE9838D0E46FD5D9367" box="[685,721,260,283]" italics="true" pageId="3" pageNumber="103">tc”</emphasis>
slightly longer than pointed
<emphasis id="C992EA62FFBBFFE985330E46FBB49360" box="[1043,1080,260,284]" italics="true" pageId="3" pageNumber="103">pl”</emphasis>
;
<emphasis id="C992EA62FFBBFFE985670E46FBD29367" box="[1095,1118,260,283]" italics="true" pageId="3" pageNumber="103">u</emphasis>
somewhat blunt-ending and shortest on tarsus. Tibia: solenidion
<emphasis id="C992EA62FFBBFFE983160E6BFDC9933C" box="[566,581,297,320]" italics="true" pageId="3" pageNumber="103">φ</emphasis>
2 (22) very small and hard to discern, situated in shallow depression; all setae pointed and moderately barbed; setae
<emphasis id="C992EA62FFBBFFE9835E0E0EFD039318" box="[638,655,332,356]" italics="true" pageId="3" pageNumber="103">l</emphasis>
,
<emphasis id="C992EA62FFBBFFE983BD0E0EFD3F931F" box="[669,691,332,355]" italics="true" pageId="3" pageNumber="103">v</emphasis>
and
<emphasis id="C992EA62FFBBFFE983CA0E0EFC89931F" box="[746,773,332,355]" italics="true" pageId="3" pageNumber="103">v”</emphasis>
subequal, slightly longer than
<emphasis id="C992EA62FFBBFFE9857E0E0EFBE09318" box="[1118,1132,332,356]" italics="true" pageId="3" pageNumber="103">d</emphasis>
. Genu: setae
<emphasis id="C992EA62FFBBFFE984240E0EFA999318" box="[1284,1301,332,356]" italics="true" pageId="3" pageNumber="103">l</emphasis>
and
<emphasis id="C992EA62FFBBFFE9846C0E0EFAEE931F" box="[1356,1378,332,355]" italics="true" pageId="3" pageNumber="103">v</emphasis>
little barbed, pointed and subequal. Femur:
<emphasis id="C992EA62FFBBFFE983630E32FDD593FB" box="[579,601,368,391]" italics="true" pageId="3" pageNumber="103">v</emphasis>
pointed and moderately barbed;
<emphasis id="C992EA62FFBBFFE982E70E32FC5993F4" box="[967,981,368,392]" italics="true" pageId="3" pageNumber="103">d</emphasis>
barbed and blunt-ending, longer than
<emphasis id="C992EA62FFBBFFE9845F0E32FA1993FB" box="[1407,1429,368,391]" italics="true" pageId="3" pageNumber="103">v</emphasis>
. Trochanter:
<emphasis id="C992EA62FFBBFFE9803F0ED6FEB993D7" box="[287,309,404,427]" italics="true" pageId="3" pageNumber="103">v</emphasis>
barbed and pointed.
</paragraph>
<caption id="AF9966F8FFBBFFE981B70839FF5395D3" httpUri="https://zenodo.org/record/235219/files/figure.png" pageId="3" pageNumber="103" targetBox="[230,1356,481,1893]" targetPageId="3">
<paragraph id="FB593670FFBBFFE981B70839FF5395D3" blockId="3.[151,1436,1915,1967]" pageId="3" pageNumber="103">
<emphasis id="C992EA62FFBBFFE981B70839FE9F95EC" bold="true" box="[151,275,1915,1937]" pageId="3" pageNumber="103">FIGURE 4.</emphasis>
<taxonomicName id="3CE64DF3FFBBFFE9803B0839FC2F95ED" authority="Katlav and Hajiqanbar" authorityName="Katlav and Hajiqanbar" box="[283,931,1915,1937]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="103" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFBBFFE9803B0839FD3995ED" box="[283,693,1915,1937]" italics="true" pageId="3" pageNumber="103">Pseudopygmephorellus mazandaranicus</emphasis>
Katlav and Hajiqanbar
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFBBFFE9828B0839FC7195EC" box="[939,1021,1915,1936]" pageId="3" pageNumber="103" rank="species">
<emphasis id="C992EA62FFBBFFE9828B0839FC7195EC" bold="true" box="[939,1021,1915,1936]" pageId="3" pageNumber="103">sp. nov.</emphasis>
</taxonomicNameLabel>
(female): dorsum of the body. Scale bar 50 µm.
</paragraph>
</caption>
<caption id="AF9966F8FFBCFFEE81B70A8AFF339780" httpUri="https://zenodo.org/record/235220/files/figure.png" pageId="4" pageNumber="104" targetBox="[254,1326,193,1465]" targetPageId="4">
<paragraph id="FB593670FFBCFFEE81B70A8AFF339780" blockId="4.[151,1435,1480,1532]" pageId="4" pageNumber="104">
<emphasis id="C992EA62FFBCFFEE81B70A8AFE9E97A1" bold="true" box="[151,274,1480,1502]" pageId="4" pageNumber="104">FIGURE 5.</emphasis>
<taxonomicName id="3CE64DF3FFBCFFEE80380A8AFC1097A2" authority="Katlav and Hajiqanbar" authorityName="Katlav and Hajiqanbar" box="[280,924,1480,1502]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="104" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFBCFFEE80380A8AFD3D97A2" box="[280,689,1480,1502]" italics="true" pageId="4" pageNumber="104">Pseudopygmephorellus mazandaranicus</emphasis>
Katlav and Hajiqanbar
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFBCFFEE82820A8AFC7E97A1" box="[930,1010,1480,1501]" pageId="4" pageNumber="104" rank="species">
<emphasis id="C992EA62FFBCFFEE82820A8AFC7E97A1" bold="true" box="[930,1010,1480,1501]" pageId="4" pageNumber="104">sp. nov.</emphasis>
</taxonomicNameLabel>
(female): venter of the body. Scale bar 50 µm.
</paragraph>
</caption>
<paragraph id="FB593670FFBCFFEE81E70967FEF9948D" blockId="4.[151,1437,1572,2031]" pageId="4" pageNumber="104">
Leg IV 112 (107
<emphasis id="C992EA62FFBCFFEE80A10966FE039441" bold="true" box="[385,399,1572,1597]" pageId="4" pageNumber="104"></emphasis>
115) (
<figureCitation id="63DD2AF5FFBCFFEE80F10966FDA99440" box="[465,549,1572,1597]" captionStart="FIGURES 9 10" captionStartId="6.[151,264,1687,1710]" captionTargetBox="[155,1424,237,1670]" captionTargetId="figure@6.[155,1424,193,1670]" captionTargetPageId="6" captionText="FIGURES 9 10. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 9 10. Legs III IV respectively. Scale bar 20 µm." httpUri="https://zenodo.org/record/235222/files/figure.png" pageId="4" pageNumber="104">Fig. 10</figureCitation>
): Longer than other legs; setal formula: Tr1-Fe2-Ge1-Ti4-Ta6. Tarsus: all setae moderately barbed and pointed except seta
<emphasis id="C992EA62FFBCFFEE83B4090AFD35941C" box="[660,697,1608,1632]" italics="true" pageId="4" pageNumber="104">pl”</emphasis>
which is considerably stiff and blunt-ending; setae
<emphasis id="C992EA62FFBCFFEE843C090AFAB59423" box="[1308,1337,1608,1631]" italics="true" pageId="4" pageNumber="104">tc</emphasis>
and
<emphasis id="C992EA62FFBCFFEE8458090AFA109423" box="[1400,1436,1608,1631]" italics="true" pageId="4" pageNumber="104">pv</emphasis>
subequal, slightly longer than subequal setae
<emphasis id="C992EA62FFBCFFEE83B8092EFD3094FF" box="[664,700,1644,1667]" italics="true" pageId="4" pageNumber="104">tc”</emphasis>
and
<emphasis id="C992EA62FFBCFFEE83D4092EFC9194FF" box="[756,797,1644,1667]" italics="true" pageId="4" pageNumber="104">pv”</emphasis>
. Tibia: solenidion
<emphasis id="C992EA62FFBCFFEE82D4092FFB8F94F8" box="[1012,1027,1645,1668]" italics="true" pageId="4" pageNumber="104">φ</emphasis>
not seen;
<emphasis id="C992EA62FFBCFFEE8558092EFB1894FF" box="[1144,1172,1644,1667]" italics="true" pageId="4" pageNumber="104">v”</emphasis>
barbed and with rather blunt apex; other tibial setae moderately barbed and pointed;
<emphasis id="C992EA62FFBCFFEE827109D2FCD394D4" box="[849,863,1680,1704]" italics="true" pageId="4" pageNumber="104">d</emphasis>
longest on the tibia. Genu:
<emphasis id="C992EA62FFBCFFEE85BD09D2FB3F94DB" box="[1181,1203,1680,1703]" italics="true" pageId="4" pageNumber="104">v</emphasis>
barbed and pointed. Femur:
<emphasis id="C992EA62FFBCFFEE81D109F6FE8A94B7" box="[241,262,1716,1739]" italics="true" pageId="4" pageNumber="104">v</emphasis>
moderately barbed and pointed,
<emphasis id="C992EA62FFBCFFEE83A409F6FD1E94B0" box="[644,658,1716,1740]" italics="true" pageId="4" pageNumber="104">d</emphasis>
barbed and indistinctly blunt-ending, longer than
<emphasis id="C992EA62FFBCFFEE85F509F6FB6794B7" box="[1237,1259,1716,1739]" italics="true" pageId="4" pageNumber="104">v</emphasis>
. Trochanter:
<emphasis id="C992EA62FFBCFFEE84A609F6FA1094B7" box="[1414,1436,1716,1739]" italics="true" pageId="4" pageNumber="104">v</emphasis>
barbed and pointed.
</paragraph>
<paragraph id="FB593670FFBCFFEE81E709BEFE6A9569" blockId="4.[151,1437,1572,2031]" box="[199,486,1788,1813]" pageId="4" pageNumber="104">Male and larva unknown.</paragraph>
<paragraph id="FB593670FFBCFFEF81E70862FD1193AC" blockId="4.[151,1437,1572,2031]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="105" pageId="4" pageNumber="104">
<emphasis id="C992EA62FFBCFFEE81E70862FE4A9545" bold="true" box="[199,454,1824,1849]" pageId="4" pageNumber="104">Differential diagnosis</emphasis>
. The new species is most similar to
<taxonomicName id="3CE64DF3FFBCFFEE82420863FB229544" authority="Mahunka, 1970" authorityName="Mahunka" authorityYear="1970" box="[866,1198,1824,1849]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="104" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBCFFEE82420863FCFC9544" box="[866,880,1825,1848]" italics="true" pageId="4" pageNumber="104">P</emphasis>
.
<emphasis id="C992EA62FFBCFFEE825E0862FC6F9544" box="[894,995,1824,1848]" italics="true" pageId="4" pageNumber="104">szekessyi</emphasis>
(
<bibRefCitation id="9F774B81FFBCFFEE82D20862FB2A9544" author="Mahunka" box="[1010,1190,1824,1849]" pageId="4" pageNumber="104" refString="Mahunka, S. (1970) Considerations on the systematics of the Tarsonemina and the description of new European taxa (Acari: Trombidiformes). Acta Zoologica Hungaricae, 16, 137 - 174." type="journal article" year="1970">Mahunka, 1970</bibRefCitation>
)
</taxonomicName>
but differs from it in some characters as follows: in
<taxonomicName id="3CE64DF3FFBCFFEE80CC0807FD4D9520" box="[492,705,1860,1884]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="104" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFBCFFEE80CC0807FE769520" box="[492,506,1861,1884]" italics="true" pageId="4" pageNumber="104">P</emphasis>
.
<emphasis id="C992EA62FFBCFFEE83280806FD4D9520" box="[520,705,1860,1884]" italics="true" pageId="4" pageNumber="104">mazandaranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFBCFFEE83E70806FCAC9521" box="[711,800,1860,1885]" pageId="4" pageNumber="104" rank="species">
<emphasis id="C992EA62FFBCFFEE83E70806FCAC9521" bold="true" box="[711,800,1860,1885]" pageId="4" pageNumber="104">sp. nov.</emphasis>
</taxonomicNameLabel>
, solenidia on TiTaI (
<emphasis id="C992EA62FFBCFFEE85280807FB909520" box="[1032,1052,1861,1884]" italics="true" pageId="4" pageNumber="104">ω</emphasis>
1,
<emphasis id="C992EA62FFBCFFEE85110807FBC99520" box="[1073,1093,1861,1884]" italics="true" pageId="4" pageNumber="104">ω</emphasis>
2,
<emphasis id="C992EA62FFBCFFEE857A0807FBE59520" box="[1114,1129,1861,1884]" italics="true" pageId="4" pageNumber="104">φ</emphasis>
1 and
<emphasis id="C992EA62FFBCFFEE85860807FB399520" box="[1190,1205,1861,1884]" italics="true" pageId="4" pageNumber="104">φ</emphasis>
2) and TaII (
<emphasis id="C992EA62FFBCFFEE841A0807FAC29520" box="[1338,1358,1861,1884]" italics="true" pageId="4" pageNumber="104">ω</emphasis>
) much shorter than those in
<taxonomicName id="3CE64DF3FFBCFFEE80A6082EFD8495FF" box="[390,520,1899,1923]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="104" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBCFFEE80A6082EFE1895FF" box="[390,404,1900,1923]" italics="true" pageId="4" pageNumber="104">P</emphasis>
.
<emphasis id="C992EA62FFBCFFEE80830829FD8495FF" box="[419,520,1899,1923]" italics="true" pageId="4" pageNumber="104">szekessyi</emphasis>
</taxonomicName>
(6
<emphasis id="C992EA62FFBCFFEE8310082EFDCB95FF" box="[560,583,1900,1923]" italics="true" pageId="4" pageNumber="104">vs</emphasis>
10, 3
<emphasis id="C992EA62FFBCFFEE83B0082EFD2B95FF" box="[656,679,1900,1923]" italics="true" pageId="4" pageNumber="104">vs</emphasis>
7, 5
<emphasis id="C992EA62FFBCFFEE83C3082EFD7695FF" box="[739,762,1900,1923]" italics="true" pageId="4" pageNumber="104">vs</emphasis>
8, 3
<emphasis id="C992EA62FFBCFFEE8215082EFCC095FF" box="[821,844,1900,1923]" italics="true" pageId="4" pageNumber="104">vs</emphasis>
6 and 6
<emphasis id="C992EA62FFBCFFEE8291082EFC4495FF" box="[945,968,1900,1923]" italics="true" pageId="4" pageNumber="104">vs</emphasis>
8, respectively) (
<figureCitation id="63DD2AF5FFBCFFEE85AF0829FB5F95F8" box="[1167,1235,1899,1924]" captionStart="FIGURES 6 8" captionStartId="5.[151,264,1948,1971]" captionTargetBox="[162,1425,517,1935]" captionTargetId="figure@5.[162,1426,517,1936]" captionTargetPageId="5" captionText="FIGURES 6 8. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov. (female): 6 7. Legs I II respectively. Scale bar 20 µm; 8. Comparison of length of solenidia in two close species: a. P. mazandaranicus sp. nov, b. P. szekessyi (Mahunka, 1970). Scale bar 10 µm." httpUri="https://zenodo.org/record/235221/files/figure.png" pageId="4" pageNumber="104">Fig. 8</figureCitation>
b); all ventral and dorsal idiosomal tergites somewhat smooth (all ventral and dorsal idiosomal tergites densely ornamented with tiny dimples in
<taxonomicName id="3CE64DF3FFBCFFEE803B08F6FE2C95B7" box="[283,416,1971,1995]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="104" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBCFFEE803B08F6FEA595B7" box="[283,297,1972,1995]" italics="true" pageId="4" pageNumber="104">P</emphasis>
.
<emphasis id="C992EA62FFBCFFEE801B08F1FE2C95B7" box="[315,416,1971,1995]" italics="true" pageId="4" pageNumber="104">szekessyi</emphasis>
</taxonomicName>
); posterior margin of tergite EF with barely visible small dimples (posterior margin of tergite EF with prominent densely-dimpled microsculpture in
<taxonomicName id="3CE64DF3FFBCFFEE8271089AFC479593" box="[849,971,2007,2031]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="104" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBCFFEE8271089AFCD39593" box="[849,863,2008,2031]" italics="true" pageId="4" pageNumber="104">P</emphasis>
.
<emphasis id="C992EA62FFBCFFEE82460895FC479593" box="[870,971,2007,2031]" italics="true" pageId="4" pageNumber="104">szekessyi</emphasis>
</taxonomicName>
); anterior genital sclerite (ags) small and mushroom-like, posterior genital sclerite (pgs) tiny and oval (ags small, much narrower anteriorly, forming arrowhead-like habitus, pgs larger, ellipsoidal, posteriorly with a tiny rounded protuberance in
<taxonomicName id="3CE64DF3FFBDFFEF85F60FFFFAD592A8" box="[1238,1369,188,212]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF85F60FFFFB6892A8" box="[1238,1252,189,212]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF85D40FFEFAD592A8" box="[1268,1369,188,212]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
</taxonomicName>
); ap5 considerably reduced (ap5 well developed in
<taxonomicName id="3CE64DF3FFBDFFEF83860FA3FCA69284" box="[678,810,224,248]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF83860FA3FD389284" box="[678,692,225,248]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF83E50FA2FCA69284" box="[709,810,224,248]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
</taxonomicName>
); seta TaII
<emphasis id="C992EA62FFBDFFEF82890FA3FC319284" box="[937,957,225,248]" italics="true" pageId="5" pageNumber="105">tc</emphasis>
' seta-like, with pointed apex (seta TaII
<emphasis id="C992EA62FFBDFFEF84A30FA3FA1B9284" box="[1411,1431,225,248]" italics="true" pageId="5" pageNumber="105">tc</emphasis>
' thickened, somewhat pilose, with blunt apex in
<taxonomicName id="3CE64DF3FFBDFFEF83960E47FCB49360" box="[694,824,260,284]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF83960E47FD489360" box="[694,708,261,284]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF83F30E46FCB49360" box="[723,824,260,284]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
</taxonomicName>
); seta FeII
<emphasis id="C992EA62FFBDFFEF82940E46FC499360" box="[948,965,260,284]" italics="true" pageId="5" pageNumber="105">l</emphasis>
rod-like and distinctly blunt-ending (seta FeII
<emphasis id="C992EA62FFBDFFEF81E50E6AFF5A933C" box="[197,214,296,320]" italics="true" pageId="5" pageNumber="105">l</emphasis>
seta-like and distinctly pointed in
<taxonomicName id="3CE64DF3FFBDFFEF83420E6BFD68933C" box="[610,740,296,320]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF83420E6BFDFC933C" box="[610,624,297,320]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF835F0E6AFD68933C" box="[639,740,296,320]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
</taxonomicName>
); setae TrI
<emphasis id="C992EA62FFBDFFEF82410E6BFCE1933C" box="[865,877,297,320]" italics="true" pageId="5" pageNumber="105">v</emphasis>
', TrII
<emphasis id="C992EA62FFBDFFEF828E0E6BFC36933C" box="[942,954,297,320]" italics="true" pageId="5" pageNumber="105">v</emphasis>
' and TrIV
<emphasis id="C992EA62FFBDFFEF85100E6BFBB0933C" box="[1072,1084,297,320]" italics="true" pageId="5" pageNumber="105">v</emphasis>
' distinctly pointed (setae TrI
<emphasis id="C992EA62FFBDFFEF84A30E6BFA03933C" box="[1411,1423,297,320]" italics="true" pageId="5" pageNumber="105">v</emphasis>
', TrII
<emphasis id="C992EA62FFBDFFEF81E30E0FFF439318" box="[195,207,333,356]" italics="true" pageId="5" pageNumber="105">v</emphasis>
' and TrIV
<emphasis id="C992EA62FFBDFFEF80630E0FFEC39318" box="[323,335,333,356]" italics="true" pageId="5" pageNumber="105">v</emphasis>
' somewhat blunt-ending in
<taxonomicName id="3CE64DF3FFBDFFEF83A90E0FFC879318" box="[649,779,332,356]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF83A90E0FFD1B9318" box="[649,663,333,356]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF83860E0EFC879318" box="[678,779,332,356]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
</taxonomicName>
); seta FeIV
<emphasis id="C992EA62FFBDFFEF82B10E0EFC2B931F" box="[913,935,332,355]" italics="true" pageId="5" pageNumber="105">v</emphasis>
seta-like and distinctly pointed (seta FeIV
<emphasis id="C992EA62FFBDFFEF84A60E0EFA10931F" box="[1414,1436,332,355]" italics="true" pageId="5" pageNumber="105">v</emphasis>
rod-like and distinctly blunt-ending in
<taxonomicName id="3CE64DF3FFBDFFEF836D0E33FD4393F4" box="[589,719,368,392]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF836D0E33FDD793F4" box="[589,603,369,392]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF834A0E32FD4393F4" box="[618,719,368,392]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
</taxonomicName>
); seta TaIV
<emphasis id="C992EA62FFBDFFEF82750E32FCFB93F4" box="[853,887,368,392]" italics="true" pageId="5" pageNumber="105">pl&quot;</emphasis>
almost 1/3 as long as seta TaIV
<emphasis id="C992EA62FFBDFFEF85C20E32FB7093FB" box="[1250,1276,368,391]" italics="true" pageId="5" pageNumber="105">tc'</emphasis>
(seta TaIV
<emphasis id="C992EA62FFBDFFEF845A0E32FA1093F4" box="[1402,1436,368,392]" italics="true" pageId="5" pageNumber="105">pl&quot;</emphasis>
approximately more than half as long as seta TaIV
<emphasis id="C992EA62FFBDFFEF83F50ED6FD7C93D7" box="[725,752,404,427]" italics="true" pageId="5" pageNumber="105">tc'</emphasis>
in
<taxonomicName id="3CE64DF3FFBDFFEF82360ED7FC1593D0" box="[790,921,404,428]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF82360ED7FCA893D0" box="[790,804,405,428]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF82140ED6FC1593D0" box="[820,921,404,428]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
</taxonomicName>
); seta TrI
<emphasis id="C992EA62FFBDFFEF85290ED6FB9093D7" box="[1033,1052,404,427]" italics="true" pageId="5" pageNumber="105">v'</emphasis>
almost twice as long as seta FeI
<emphasis id="C992EA62FFBDFFEF84AE0ED6FA1093D0" box="[1422,1436,404,428]" italics="true" pageId="5" pageNumber="105">l'</emphasis>
(setae TrI
<emphasis id="C992EA62FFBDFFEF80220EFAFE9993B3" box="[258,277,440,463]" italics="true" pageId="5" pageNumber="105">v'</emphasis>
and FeI
<emphasis id="C992EA62FFBDFFEF80500EFAFEF493AC" box="[368,376,440,464]" italics="true" pageId="5" pageNumber="105">l</emphasis>
' subequal in
<taxonomicName id="3CE64DF3FFBDFFEF832B0EFBFD0093AC" box="[523,652,440,464]" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF832B0EFBFD9593AC" box="[523,537,441,464]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF83070EFAFD0093AC" box="[551,652,440,464]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="AF9966F8FFBDFFEF81B708DEFE73958C" httpUri="https://zenodo.org/record/235221/files/figure.png" pageId="5" pageNumber="105" targetBox="[162,1425,517,1935]" targetPageId="5">
<paragraph id="FB593670FFBDFFEF81B708DEFE73958C" blockId="5.[151,1437,1948,2032]" pageId="5" pageNumber="105">
<emphasis id="C992EA62FFBDFFEF81B708DEFEB695CE" bold="true" box="[151,314,1948,1971]" pageId="5" pageNumber="105">FIGURES 68.</emphasis>
<taxonomicName id="3CE64DF3FFBDFFEF806008DFFC4895CF" authority="Katlav and Hajiqanbar" authorityName="Katlav and Hajiqanbar" box="[320,964,1949,1971]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFBDFFEF806008DFFD5595CF" box="[320,729,1949,1971]" italics="true" pageId="5" pageNumber="105">Pseudopygmephorellus mazandaranicus</emphasis>
Katlav and Hajiqanbar
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFBDFFEF82EC08DFFB9095CE" box="[972,1052,1949,1970]" pageId="5" pageNumber="105" rank="species">
<emphasis id="C992EA62FFBDFFEF82EC08DFFB9095CE" bold="true" box="[972,1052,1949,1970]" pageId="5" pageNumber="105">sp. nov.</emphasis>
</taxonomicNameLabel>
(female): 6
<emphasis id="C992EA62FFBDFFEF85B308DFFB1395CE" bold="true" box="[1171,1183,1949,1970]" pageId="5" pageNumber="105"></emphasis>
7. Legs I
<emphasis id="C992EA62FFBDFFEF85D908DFFA8995CE" bold="true" box="[1273,1285,1949,1970]" pageId="5" pageNumber="105"></emphasis>
II respectively. Scale bar 20 µm; 8. Comparison of length of solenidia in two close species: a.
<taxonomicName id="3CE64DF3FFBDFFEF82C508FFFB2595AE" box="[997,1193,1980,2002]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFBDFFEF82C508FFFB2595AE" box="[997,1193,1980,2002]" italics="true" pageId="5" pageNumber="105">P. mazandaranicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFBDFFEF859308FEFA8D95AD" box="[1203,1281,1980,2001]" pageId="5" pageNumber="105" rank="species">
<emphasis id="C992EA62FFBDFFEF859308FEFA8D95AD" bold="true" box="[1203,1281,1980,2001]" pageId="5" pageNumber="105">sp. nov</emphasis>
</taxonomicNameLabel>
, b.
<taxonomicName id="3CE64DF3FFBDFFEF840D08FFFEC5958C" authority="Mahunka, 1970" authorityName="Mahunka" authorityYear="1970" class="Arachnida" family="Pygmephoridae" genus="Pygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="105" phylum="Arthropoda" rank="species" species="szekessyi">
<emphasis id="C992EA62FFBDFFEF840D08FFFAB695AE" box="[1325,1338,1981,2002]" italics="true" pageId="5" pageNumber="105">P</emphasis>
.
<emphasis id="C992EA62FFBDFFEF846008FEFA1095AE" box="[1344,1436,1980,2002]" italics="true" pageId="5" pageNumber="105">szekessyi</emphasis>
(Mahunka, 1970)
</taxonomicName>
. Scale bar 10 µm.
</paragraph>
</caption>
<caption id="AF9966F8FFBEFFEC81B709D5FE4E94B0" httpUri="https://zenodo.org/record/235222/files/figure.png" pageId="6" pageNumber="106" targetBox="[155,1424,237,1670]" targetPageId="6">
<paragraph id="FB593670FFBEFFEC81B709D5FE4E94B0" blockId="6.[151,1436,1687,1740]" pageId="6" pageNumber="106">
<emphasis id="C992EA62FFBEFFEC81B709D5FEC294D1" bold="true" box="[151,334,1687,1710]" pageId="6" pageNumber="106">FIGURES 910.</emphasis>
<taxonomicName id="3CE64DF3FFBEFFEC807B09DAFC7794D2" authority="Katlav and Hajiqanbar" authorityName="Katlav and Hajiqanbar" box="[347,1019,1688,1710]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="106" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFBEFFEC807B09DAFD7694D2" box="[347,762,1688,1710]" italics="true" pageId="6" pageNumber="106">Pseudopygmephorellus mazandaranicus</emphasis>
Katlav and Hajiqanbar
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFBEFFEC852809DAFBD394D1" box="[1032,1119,1688,1709]" pageId="6" pageNumber="106" rank="species">
<emphasis id="C992EA62FFBEFFEC852809DAFBD394D1" bold="true" box="[1032,1119,1688,1709]" pageId="6" pageNumber="106">sp. nov.</emphasis>
</taxonomicNameLabel>
(female): 9
<emphasis id="C992EA62FFBEFFEC85C309DAFB7C94D1" bold="true" box="[1251,1264,1688,1709]" pageId="6" pageNumber="106"></emphasis>
10. Legs III
<emphasis id="C992EA62FFBEFFEC845409DAFA0D94D1" bold="true" box="[1396,1409,1688,1709]" pageId="6" pageNumber="106"></emphasis>
IV respectively. Scale bar 20µm.
</paragraph>
</caption>
<subSubSection id="B3FC65FBFFBEFFED81E709BBFB49954F" lastPageId="7" lastPageNumber="107" pageId="6" pageNumber="106" type="materials_examined">
<paragraph id="FB593670FFBEFFEC81E709BBFE3D95BA" blockId="6.[151,1437,1785,1990]" pageId="6" pageNumber="106">
<emphasis id="C992EA62FFBEFFEC81E709BBFEF6956E" bold="true" box="[199,378,1785,1810]" pageId="6" pageNumber="106">
<typeStatus id="245D88D2FFBEFFEC81E709BBFE8E956E" box="[199,258,1785,1810]" pageId="6" pageNumber="106">Type</typeStatus>
material.
</emphasis>
Female
<typeStatus id="245D88D2FFBEFFEC80C509BBFDCB956E" box="[485,583,1785,1810]" pageId="6" pageNumber="106" type="holotype">holotype</typeStatus>
AK-20130318-H and more than 180
<typeStatus id="245D88D2FFBEFFEC852E09B8FBF0956E" box="[1038,1148,1786,1810]" pageId="6" pageNumber="106" type="paratype">paratypes</typeStatus>
, as three colonies (each including more than
<specimenCount id="EDE0FDF9FFBEFFEC80B0085FFD99954A" box="[400,533,1821,1846]" pageId="6" pageNumber="106" type="female">60 females</specimenCount>
), attaching firmly on soft membranes between head and pronotum (cervical membrane) and between pronotum and elytra of three specimens of the beetle
<taxonomicName id="3CE64DF3FFBEFFEC85610803FB549525" box="[1089,1240,1857,1881]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="C992EA62FFBEFFEC85610803FB549525" box="[1089,1240,1857,1881]" italics="true" pageId="6" pageNumber="106">Onthophagus</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="3CE64DF3FFBEFFEC84380803FA159526" box="[1304,1433,1857,1882]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="106" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="3CE64DF3FFBEFFEC81B70827FEA19502" box="[151,301,1893,1918]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="106" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
) (
<figureCitation id="63DD2AF5FFBEFFEC80660827FE5D9502" box="[326,465,1893,1918]" captionStart="FIGURES 11 14" captionStartId="7.[151,264,1484,1507]" captionTargetBox="[150,1436,193,1462]" captionTargetId="figure@7.[150,1436,193,1464]" captionTargetPageId="7" captionText="FIGURES 11 14. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov and the beetle hosts: 11. Onthophagus sp. (Col., Scarabaeidae); 12. Colony of mite on cervical membrane and between pronotum and elytra of Onthophagus sp.; 13. Aphodius varians Duftschmid, 1805 (Col., Scarabaeidae); 14. Colony of mite on first abdominal sternite (near third coxa) of the beetle Aphodius varians." httpUri="https://zenodo.org/record/235223/files/figure.png" pageId="6" pageNumber="106">Figs. 1112</figureCitation>
). Host beetles were collected directly by forceps from cow-pads in Dohezar Forest, Tonekabon town, west of Mazandaran Province, northern
<collectingCountry id="83F176E0FFBEFFEC823C08C8FCC095DD" box="[796,844,1930,1953]" name="Iran" pageId="6" pageNumber="106">Iran</collectingCountry>
, (
<geoCoordinate id="9ED250B7FFBEFFEC827F08CBFC4B95DD" box="[863,967,1929,1954]" direction="north" orientation="latitude" pageId="6" pageNumber="106" precision="925" value="36.65">36°39 N</geoCoordinate>
,
<geoCoordinate id="9ED250B7FFBEFFEC82F108CBFBB995DD" box="[977,1077,1929,1954]" direction="east" orientation="longitude" pageId="6" pageNumber="106" precision="925" value="50.666668">50°40 E</geoCoordinate>
, altitude
<quantity id="3C1E9B95FFBEFFEC85BA08CBFB6C95DD" box="[1178,1248,1929,1954]" metricMagnitude="2" metricUnit="m" metricValue="5.779999999999999" pageId="6" pageNumber="106" unit="m" value="578.0">578 m</quantity>
a.s.l.),
<date id="8F5810B0FFBEFFEC841108C8FF5E95BA" pageId="6" pageNumber="106" value="2013-03-18">18 March 2013</date>
, leg. Alihan Katlav.
</paragraph>
<caption id="AF9966F8FFBFFFED81B70A8EFD0D9443" httpUri="https://zenodo.org/record/235223/files/figure.png" pageId="7" pageNumber="107" targetBox="[150,1436,193,1462]" targetPageId="7">
<paragraph id="FB593670FFBFFFED81B70A8EFD0D9443" blockId="7.[151,1436,1484,1599]" pageId="7" pageNumber="107">
<emphasis id="C992EA62FFBFFFED81B70A8EFED7979E" bold="true" box="[151,347,1484,1507]" pageId="7" pageNumber="107">FIGURES 1114.</emphasis>
<taxonomicName id="3CE64DF3FFBFFFED804A0A8FFB9E979F" authority="Katlav and Hajiqanbar" authorityName="Katlav and Hajiqanbar" box="[362,1042,1485,1507]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="107" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFBFFFED804A0A8FFC87979F" box="[362,779,1485,1507]" italics="true" pageId="7" pageNumber="107">Pseudopygmephorellus mazandaranicus</emphasis>
Katlav and Hajiqanbar
</taxonomicName>
<emphasis id="C992EA62FFBFFFED85010A8FFBFA979E" bold="true" box="[1057,1142,1485,1506]" pageId="7" pageNumber="107">
<taxonomicNameLabel id="D2A15719FFBFFFED85010A8FFBFA979E" box="[1057,1142,1485,1506]" pageId="7" pageNumber="107" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
and the beetle hosts: 11.
<taxonomicName id="3CE64DF3FFBFFFED81B70AAEFEAC947E" box="[151,288,1516,1538]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="C992EA62FFBFFFED81B70AAEFEAC947E" box="[151,288,1516,1538]" italics="true" pageId="7" pageNumber="107">Onthophagus</emphasis>
</taxonomicName>
sp. (Col.,
<taxonomicName id="3CE64DF3FFBFFFED80BA0AAEFDA8947E" box="[410,548,1516,1538]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
); 12. Colony of mite on cervical membrane and between pronotum and elytra of
<taxonomicName id="3CE64DF3FFBFFFED81B70948FEAC945C" box="[151,288,1546,1568]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="C992EA62FFBFFFED81B70948FEAC945C" box="[151,288,1546,1568]" italics="true" pageId="7" pageNumber="107">Onthophagus</emphasis>
</taxonomicName>
sp.; 13.
<taxonomicName id="3CE64DF3FFBFFFED80570948FD6B945C" authority="Duftschmid, 1805" authorityName="Duftschmid" authorityYear="1805" box="[375,743,1546,1568]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="C992EA62FFBFFFED80570948FDA5945C" box="[375,553,1546,1568]" italics="true" pageId="7" pageNumber="107">Aphodius varians</emphasis>
Duftschmid, 1805
</taxonomicName>
(Col.,
<taxonomicName id="3CE64DF3FFBFFFED820E0948FC34945C" box="[814,952,1546,1568]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
); 14. Colony of mite on first abdominal sternite (near third coxa) of the beetle
<taxonomicName id="3CE64DF3FFBFFFED80E9096BFDF79443" box="[457,635,1577,1599]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="C992EA62FFBFFFED80E9096BFDF79443" box="[457,635,1577,1599]" italics="true" pageId="7" pageNumber="107">Aphodius varians</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="FB593670FFBFFFED81E70924FB49954F" blockId="7.[151,1436,1638,2030]" pageId="7" pageNumber="107">
<emphasis id="C992EA62FFBFFFED81E70924FE369403" bold="true" box="[199,442,1638,1663]" pageId="7" pageNumber="107">Additional material.</emphasis>
Twelve females as two small colonies (one including
<specimenCount id="EDE0FDF9FFBFFFED853A0924FB0A9403" box="[1050,1158,1638,1663]" pageId="7" pageNumber="107" type="female">6 females</specimenCount>
and the other
<specimenCount id="EDE0FDF9FFBFFFED84070925FA199403" box="[1319,1429,1638,1663]" pageId="7" pageNumber="107" type="female">7 females</specimenCount>
) attaching firmly to the first abdominal sternite near third coxae of two specimens of the beetle
<taxonomicName id="3CE64DF3FFBFFFED85F509C8FEE494BB" authority="Duftschmid, 1805" authorityName="Duftschmid" authorityYear="1805" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="C992EA62FFBFFFED85F509C8FA1094DE" box="[1237,1436,1674,1698]" italics="true" pageId="7" pageNumber="107">Aphodius varians</emphasis>
Duftschmid, 1805
</taxonomicName>
(
<taxonomicName id="3CE64DF3FFBFFFED805E09ECFE7394BB" box="[382,511,1710,1735]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="3CE64DF3FFBFFFED833009ECFD2A94BB" box="[528,678,1710,1735]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
) (
<figureCitation id="63DD2AF5FFBFFFED83E409ECFCDD94BB" box="[708,849,1710,1735]" captionStart="FIGURES 11 14" captionStartId="7.[151,264,1484,1507]" captionTargetBox="[150,1436,193,1462]" captionTargetId="figure@7.[150,1436,193,1464]" captionTargetPageId="7" captionText="FIGURES 11 14. Pseudopygmephorellus mazandaranicus Katlav and Hajiqanbar sp. nov and the beetle hosts: 11. Onthophagus sp. (Col., Scarabaeidae); 12. Colony of mite on cervical membrane and between pronotum and elytra of Onthophagus sp.; 13. Aphodius varians Duftschmid, 1805 (Col., Scarabaeidae); 14. Colony of mite on first abdominal sternite (near third coxa) of the beetle Aphodius varians." httpUri="https://zenodo.org/record/235223/files/figure.png" pageId="7" pageNumber="107">Figs. 1314</figureCitation>
) collected directly by forceps from cow-pads in Kzemkela village, Abbas abad, west of Mazandaran Province, northern
<collectingCountry id="83F176E0FFBFFFED82F90991FB859496" box="[985,1033,1747,1770]" name="Iran" pageId="7" pageNumber="107">Iran</collectingCountry>
, (
<geoCoordinate id="9ED250B7FFBFFFED85000990FB009496" box="[1056,1164,1746,1771]" direction="north" orientation="latitude" pageId="7" pageNumber="107" precision="925" value="36.683334">36°41 N</geoCoordinate>
,
<geoCoordinate id="9ED250B7FFBFFFED85BA0990FA8D9496" box="[1178,1281,1746,1771]" direction="east" orientation="longitude" pageId="7" pageNumber="107" precision="925" value="51.083332">51°05 E</geoCoordinate>
, altitude
<quantity id="3C1E9B95FFBFFFED844E0991FA109496" box="[1390,1436,1747,1771]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="107" unit="m" value="9.0">9 m</quantity>
a.s.l.),
<date id="8F5810B0FFBFFFED81C809B5FE2F9573" box="[232,419,1782,1807]" pageId="7" pageNumber="107" value="2013-03-18">18 March 2013</date>
, leg. Alihan Katlav. Two extra females under elytra, on abdominal tergites of single specimen of the beetle
<taxonomicName id="3CE64DF3FFBFFFED80B90858FCCA954F" authority="Kugelann, 1792" authorityName="Kugelann" authorityYear="1792" box="[409,838,1818,1843]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="C992EA62FFBFFFED80B90858FDFB954E" box="[409,631,1818,1842]" italics="true" pageId="7" pageNumber="107">Aphodius depressus</emphasis>
(Kugelann, 1792)
</taxonomicName>
, same collection data as
<typeStatus id="245D88D2FFBFFFED857C0858FB4E954F" box="[1116,1218,1818,1843]" pageId="7" pageNumber="107" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="B3FC65FBFFBFFFED81E7087CFEB49506" pageId="7" pageNumber="107" type="etymology">
<paragraph id="FB593670FFBFFFED81E7087CFEB49506" blockId="7.[151,1436,1638,2030]" pageId="7" pageNumber="107">
<emphasis id="C992EA62FFBFFFED81E7087CFEC2952B" bold="true" box="[199,334,1854,1879]" pageId="7" pageNumber="107">Etymology.</emphasis>
The specific epithet refers to the locality of the origin (Mazandaran province), a large district in Northern
<collectingCountry id="83F176E0FFBFFFED80230821FEB89506" box="[259,308,1891,1914]" name="Iran" pageId="7" pageNumber="107">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B3FC65FBFFBFFFE281E708C4FC6F912E" lastPageId="8" lastPageNumber="108" pageId="7" pageNumber="107" type="discussion">
<paragraph id="FB593670FFBFFFE281E708C4FCDD9361" blockId="7.[151,1436,1638,2030]" lastBlockId="8.[151,1437,151,285]" lastPageId="8" lastPageNumber="108" pageId="7" pageNumber="107">
<emphasis id="C992EA62FFBFFFED81E708C4FEB795E3" bold="true" box="[199,315,1926,1951]" pageId="7" pageNumber="107">Remarks.</emphasis>
One of the 60 slide-mounted specimens, associated with
<taxonomicName id="3CE64DF3FFBFFFED82E508C5FBDE95E2" box="[965,1106,1926,1950]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="107" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="C992EA62FFBFFFED82E508C5FBDE95E2" box="[965,1106,1926,1950]" italics="true" pageId="7" pageNumber="107">A. depressus</emphasis>
</taxonomicName>
, had a setiform
<emphasis id="C992EA62FFBFFFED842B08C5FAAE95E2" box="[1291,1314,1927,1950]" italics="true" pageId="7" pageNumber="107">sc</emphasis>
1 (Fig. 16), however, the remaining characters were exactly the same. This anomaly could be a case of neoteny as larvae of other species have a setiform
<emphasis id="C992EA62FFBFFFED80C70890FE729595" box="[487,510,2002,2025]" italics="true" pageId="7" pageNumber="107">sc</emphasis>
1 (Evert E. Lindquist, personal communications) or may also be an implication for the phenomenon “atavism” in which an individual remarkably represent the tendency to revert to ancestral phenotypical features (
<bibRefCitation id="9F774B81FFB0FFE280BD0FFEFDC892A8" author="Le" box="[413,580,188,213]" pageId="8" pageNumber="108" refString="Le Page, M. (2007) The ancestor within. New Scientist, 193, 28 - 33." type="journal article" year="2007">Le Page 2007</bibRefCitation>
). These features reappear in some individuals after several generations of absence (
<bibRefCitation id="9F774B81FFB0FFE280240F9DFE4D9284" author="Liu" box="[260,449,223,248]" pageId="8" pageNumber="108" refString="Liu, Y. S., Zhou, X. M., Zhi, M. X., Li, X. J. &amp; Wang, Q. L. (2009) Darwin's contributions to genetics. Journal of Applied Genetics, 50 (3), 177 - 184. http: // dx. doi. org / 10.1007 / BF 03195671" type="journal article" year="2009">
Liu
<emphasis id="C992EA62FFB0FFE280190FA3FEE39284" box="[313,367,224,248]" italics="true" pageId="8" pageNumber="108">et al</emphasis>
. 2009
</bibRefCitation>
). Alternatively, this specimen could represent another species, but we refrain from considering this option with just one likely aberrant specimen.
</paragraph>
<paragraph id="FB593670FFB0FFE281B70C5EFC6F912E" blockId="8.[151,1436,796,853]" pageId="8" pageNumber="108">
<emphasis id="C992EA62FFB0FFE281B70C5EFED4914E" bold="true" box="[151,344,796,819]" pageId="8" pageNumber="108">FIGURES 1516.</emphasis>
<taxonomicName id="3CE64DF3FFB0FFE280420C5FFC74914F" authority="Katlav and Hajiqanbar" authorityName="Katlav and Hajiqanbar" box="[354,1016,797,819]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="108" phylum="Arthropoda" rank="species" species="mazandaranicus" status="sp. nov.">
<emphasis id="C992EA62FFB0FFE280420C5FFD73914F" box="[354,767,797,819]" italics="true" pageId="8" pageNumber="108">Pseudopygmephorellus mazandaranicus</emphasis>
Katlav and Hajiqanbar
</taxonomicName>
<taxonomicNameLabel id="D2A15719FFB0FFE285230C5FFBD4914E" box="[1027,1112,797,818]" pageId="8" pageNumber="108" rank="species">
<emphasis id="C992EA62FFB0FFE285230C5FFBD4914E" bold="true" box="[1027,1112,797,818]" pageId="8" pageNumber="108">sp. nov.</emphasis>
</taxonomicNameLabel>
(female): 15. Prodorsum with capitate and bothridial shape of
<emphasis id="C992EA62FFB0FFE280FB0C7FFE7C912E" box="[475,496,829,850]" italics="true" pageId="8" pageNumber="108">sc</emphasis>
<emphasis id="C992EA62FFB0FFE280D00C0BFE7B9129" bold="true" box="[496,503,841,853]" pageId="8" pageNumber="108">1</emphasis>
; 16. Prodorsum with setiform appearance of
<emphasis id="C992EA62FFB0FFE282E10C7FFC5A912E" box="[961,982,829,850]" italics="true" pageId="8" pageNumber="108">sc</emphasis>
<emphasis id="C992EA62FFB0FFE282F60C0BFC519129" bold="true" box="[982,989,841,853]" pageId="8" pageNumber="108">1</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>