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<mods:title id="ACD13F956C12558DC17F56269C18D72E">Two new genera and two new species of Conopidae (Diptera) from China</mods:title>
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<mods:namePart id="3D6A2DBAAEAFAC7036F1F06EB9308DE4">Stuke, Jens-Hermann</mods:namePart>
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<subSubSection id="B38FBEBFFFA3A22DFF64F9E9FE6EF90F" box="[212,487,1642,1669]" pageId="1" pageNumber="302" type="nomenclature">
<paragraph id="FB2AED34FFA3A22DFF64F9E9FE6EF90F" blockId="1.[212,487,1642,1702]" box="[212,487,1642,1669]" pageId="1" pageNumber="302">
<heading id="A0625A58FFA3A22DFF64F9E9FE6EF90F" box="[212,487,1642,1669]" centered="true" fontSize="12" level="2" pageId="1" pageNumber="302" reason="2">
<taxonomicName id="3C9596B7FFA3A22DFF64F9E9FE09F90F" ID-CoL="89Q9S" authority="Stuke, 2019" authorityName="Stuke" authorityYear="2019" box="[212,384,1642,1669]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="C9E13126FFA3A22DFF64F9E9FE09F90F" box="[212,384,1642,1669]" italics="true" pageId="1" pageNumber="302">Gellergrimmellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D28C5DFFA3A22DFE36F9E9FE6EF90F" box="[390,487,1642,1669]" pageId="1" pageNumber="302" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B38FBEBFFFA3A22DFF64F911FEDDF972" pageId="1" pageNumber="302" type="description">
<paragraph id="FB2AED34FFA3A22DFF64F911FEA1F92C" blockId="1.[212,487,1642,1702]" box="[212,296,1682,1702]" pageId="1" pageNumber="302">
(
<figureCitation id="63AEF1B1FFA3A22DFF68F911FEAAF92C" box="[216,291,1682,1702]" captionStart-0="Figs 1–4" captionStart-1="Figs 5–8" captionStart-2="Figs 9–10" captionStart-3="Figs 11–13" captionStartId-0="2.[215,255,1718,1740]" captionStartId-1="3.[174,214,1777,1799]" captionStartId-2="4.[212,252,1713,1735]" captionStartId-3="5.[174,214,1065,1087]" captionTargetBox-0="[212,1459,255,1710]" captionTargetBox-1="[174,1421,255,1769]" captionTargetBox-2="[244,1442,299,1684]" captionTargetBox-3="[174,1421,255,1059]" captionTargetId-0="figure-251@2.[212,1459,255,1710]" captionTargetId-1="figure-220@3.[174,1421,255,1769]" captionTargetId-2="figure-256@4.[212,1459,252,1703]" captionTargetId-3="figure-594@5.[174,1421,255,1059]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," captionText-1="Figs 5–8: Gellergrimmellus fritzi spec. nov. – 5  abdomen, dorsal view ( holotype). – 6 wing, ventral view ( paratype). – 7 " captionText-2="Figs 9–10:  genitalia of Gellergrimmellus fritzi spec. nov. – 9  hypandrium and phallus, lateral view ( paratype); – 10  hypandrium and phallus, dorsal view ( paratype). bp - basiphallus; dbhy - dorsolateral bar of hypandrium; dp - distiphallus; ea - ejaculatory apodeme; edp - evagination of distiphallus; hya - hypandrial arm; hybr - hypandrial bridge; hyr - hypandrial bars; po - postgonite; poe - postgonite evagination; rs - ring sclerite." captionText-3="Figs 11–13:  postabdomen of Gellergrimmellus fritzi spec. nov. – 11  postabdomen, ventral view ( paratype). – 12  theca, posterior view ( paratype). – 13 spermathecae ( paratype). fsrd - fused spermathecae ducts; pS6 - posterior part of sternite 6; S6 - sternite 6; S7 - sternite 7; sr - spermatheca; srd - spermatheca duct; T6 - tergite 6; T7 - tergite 7." figureDoi-0="http://doi.org/10.5281/zenodo.10125033" figureDoi-1="http://doi.org/10.5281/zenodo.10125035" figureDoi-2="http://doi.org/10.5281/zenodo.10125037" figureDoi-3="http://doi.org/10.5281/zenodo.10125039" httpUri-0="https://zenodo.org/record/10125033/files/figure.png" httpUri-1="https://zenodo.org/record/10125035/files/figure.png" httpUri-2="https://zenodo.org/record/10125037/files/figure.png" httpUri-3="https://zenodo.org/record/10125039/files/figure.png" pageId="1" pageNumber="302">Figs 1−13</figureCitation>
)
</paragraph>
<paragraph id="FB2AED34FFA3A22DFF1EF93CFEDDF972" blockId="1.[174,778,1727,1784]" pageId="1" pageNumber="302">
<emphasis id="C9E13126FFA3A22DFF1EF93CFEB3F95D" bold="true" box="[174,314,1727,1751]" pageId="1" pageNumber="302">Type-species</emphasis>
: 
<taxonomicName id="3C9596B7FFA3A22DFEFCF943FDB7F952" authority="Stuke, 2019" authorityName="Stuke" authorityYear="2019" box="[332,574,1728,1752]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
<emphasis id="C9E13126FFA3A22DFEFCF943FDB7F952" box="[332,574,1728,1752]" italics="true" pageId="1" pageNumber="302">Gellergrimmellus fritzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D28C5DFFA3A22DFDFAF943FD33F95D" box="[586,698,1728,1751]" pageId="1" pageNumber="302" rank="species">spec. nov.</taxonomicNameLabel>
, designated herewith.
</paragraph>
</subSubSection>
<subSubSection id="B38FBEBFFFA3A22DFF1EF89CFBEAFDD2" pageId="1" pageNumber="302" type="diagnosis">
<paragraph id="FB2AED34FFA3A22DFF1EF89CFBEAFDD2" blockId="1.[174,779,1823,2104]" lastBlockId="1.[816,1422,320,600]" pageId="1" pageNumber="302">
<emphasis id="C9E13126FFA3A22DFF1EF89CFE92F8BD" bold="true" box="[174,283,1823,1847]" pageId="1" pageNumber="302">Diagnosis</emphasis>
: 
<taxonomicName id="3C9596B7FFA3A22DFE9DF8A3FE56F8B2" authorityName="Stuke" authorityYear="2019" box="[301,479,1824,1848]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA3A22DFE9DF8A3FE56F8B2" box="[301,479,1824,1848]" italics="true" pageId="1" pageNumber="302">Gellergrimmellus</emphasis>
</taxonomicName>
is easily identified by the combination of the following characters: three distinct ocelli (
<figureCitation id="63AEF1B1FFA3A22DFF4DF8E3FEB5F8F2" box="[253,316,1888,1912]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="1" pageNumber="302">Fig. 4</figureCitation>
), frons and vertex smooth, lacking any obvious grooves (
<figureCitation id="63AEF1B1FFA3A22DFEC5F803FE3AF812" box="[373,435,1920,1944]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="1" pageNumber="302">Fig. 4</figureCitation>
), proboscis shorter than head length (
<figureCitation id="63AEF1B1FFA3A22DFEBCF823FEC3F832" box="[268,330,1952,1976]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="1" pageNumber="302">Fig. 3</figureCitation>
), arista reduced to only one visible aristomere (
<figureCitation id="63AEF1B1FFA3A22DFE9DF843FEE5F852" box="[301,364,1984,2008]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="1" pageNumber="302">Fig. 2</figureCitation>
), wing completely covered with microtrichia, and tibiae dorsally with preapical setulae. At first glance 
<taxonomicName id="3C9596B7FFA3A22DFF47F783FE20F792" authorityName="Stuke" authorityYear="2019" box="[247,425,2048,2072]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA3A22DFF47F783FE20F792" box="[247,425,2048,2072]" italics="true" pageId="1" pageNumber="302">Gellergrimmellus</emphasis>
</taxonomicName>
resembles some of the Australian 
<taxonomicName id="3C9596B7FFA3A22DFF1EF7A3FEADF7B2" box="[174,292,2080,2104]" class="Insecta" family="Conopidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="subFamily" subFamily="Conopinae">Conopinae</taxonomicName>
which may be very small, often have ocelli and can have a reduced and inconspicuous arista (although always of at least with two distinct aristomeres). Only 
<taxonomicName id="3C9596B7FFA3A22DFC80FE03FBE4FE12" authority="SCHNEIDER, 2010" authorityName="SCHNEIDER" authorityYear="2010" box="[816,1133,384,408]" class="Insecta" family="Conopidae" genus="Tanyconops" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA3A22DFC80FE03FC23FE1D" box="[816,938,384,407]" italics="true" pageId="1" pageNumber="302">Tanyconops</emphasis>
<bibRefCitation id="9F0490C5FFA3A22DFC01FE03FBE4FE12" author="SCHNEIDER, M. A." box="[945,1133,384,408]" pageId="1" pageNumber="302" pagination="1 - 246" refId="ref9344" refString="SCHNEIDER, M. A. 2010: A taxonomic revision of Australian Conopidae (Insecta: Diptera). - Zootaxa 2581: 1 - 246. - https: // www. biotaxa. org / Zootaxa / art icle / view / zootaxa. 2581.1.1 / 38173." type="journal article" year="2010">SCHNEIDER, 2010</bibRefCitation>
</taxonomicName>
has a very short proboscis and lacks distinct grooves on the frons, but this has a very characteristic female postabdomen with an obviously prolonged and flattened tergite 7 (
<bibRefCitation id="9F0490C5FFA3A22DFAA2FE63FC11FD92" author="SCHNEIDER, M. A." pageId="1" pageNumber="302" pagination="1 - 246" refId="ref9344" refString="SCHNEIDER, M. A. 2010: A taxonomic revision of Australian Conopidae (Insecta: Diptera). - Zootaxa 2581: 1 - 246. - https: // www. biotaxa. org / Zootaxa / art icle / view / zootaxa. 2581.1.1 / 38173." type="journal article" year="2010">SCHNEIDER 2010: 241</bibRefCitation>
, Figs 321 &amp; 242, Fig. 325). 
<taxonomicName id="3C9596B7FFA3A22DFB6CFD83FADFFD9D" authorityName="SCHNEIDER" authorityYear="2010" box="[1244,1366,512,535]" class="Insecta" family="Conopidae" genus="Tanyconops" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA3A22DFB6CFD83FADFFD9D" box="[1244,1366,512,535]" italics="true" pageId="1" pageNumber="302">Tanyconops</emphasis>
</taxonomicName>
also lacks ocelli and has an obviously different wing venation (
<bibRefCitation id="9F0490C5FFA3A22DFC86FDC3FBAAFDD2" author="SCHNEIDER, M. A." box="[822,1059,576,600]" pageId="1" pageNumber="302" pagination="1 - 246" refId="ref9344" refString="SCHNEIDER, M. A. 2010: A taxonomic revision of Australian Conopidae (Insecta: Diptera). - Zootaxa 2581: 1 - 246. - https: // www. biotaxa. org / Zootaxa / art icle / view / zootaxa. 2581.1.1 / 38173." type="journal article" year="2010">SCHNEIDER 2010: 241</bibRefCitation>
: 320).
</paragraph>
</subSubSection>
<subSubSection id="B38FBEBFFFA3A22DFC80FD03FB99FCF2" pageId="1" pageNumber="302" type="etymology">
<paragraph id="FB2AED34FFA3A22DFC80FD03FB99FCF2" blockId="1.[816,1421,640,888]" pageId="1" pageNumber="302">
<emphasis id="C9E13126FFA3A22DFC80FD03FC2EFD12" bold="true" box="[816,935,640,664]" pageId="1" pageNumber="302">Etymology</emphasis>
: The genus 
<taxonomicName id="3C9596B7FFA3A22DFB9DFD03FB56FD12" authorityName="Stuke" authorityYear="2019" box="[1069,1247,640,664]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA3A22DFB9DFD03FB56FD12" box="[1069,1247,640,664]" italics="true" pageId="1" pageNumber="302">Gellergrimmellus</emphasis>
</taxonomicName>
is derived from the name „Geller-Grimm“. The Latinized name is in the diminuative, as signified by the ending „-ellus“, which refers to the small body size of the fly. This patronym is dedicated to Fritz Geller-Grimm (Frankfurt) to whom I am very grateful for his long-standing support of my dipterological work. The name 
<taxonomicName id="3C9596B7FFA3A22DFB34FCC3FABFFCD2" authorityName="Stuke" authorityYear="2019" box="[1156,1334,832,856]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA3A22DFB34FCC3FABFFCD2" box="[1156,1334,832,856]" italics="true" pageId="1" pageNumber="302">Gellergrimmellus</emphasis>
</taxonomicName>
is to be treated as masculine.
</paragraph>
</subSubSection>
<subSubSection id="B38FBEBFFFA3A229FC80FC1CFB11F932" lastPageId="5" lastPageNumber="306" pageId="1" pageNumber="302" type="description">
<paragraph id="FB2AED34FFA3A22DFC80FC1CFC70F7BD" blockId="1.[816,1422,927,2104]" pageId="1" pageNumber="302">
<emphasis id="C9E13126FFA3A22DFC80FC1CFC26FC3D" bold="true" box="[816,943,927,951]" pageId="1" pageNumber="302">Description</emphasis>
: 
<emphasis id="C9E13126FFA3A22DFC09FC23FC7BFC32" bold="true" box="[953,1010,928,952]" pageId="1" pageNumber="302">Head</emphasis>
: Arista with only one minute aristomere situated at tip of basal flagellomere (
<figureCitation id="63AEF1B1FFA3A22DFB0AFC43FB7EFC52" box="[1210,1271,960,984]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="1" pageNumber="302">Fig. 2</figureCitation>
). Scape about two times longer than maximum width, apically and ventrally with black setae. Pedicel about four times longer than maximum width, completely covered with black setae, lacking a blunt ridge at base, and slightly expanded towards the apex. Basal flagellomere long, about three times longer than high, and almost as long as pedicel. Basal flagellomere pointed, ventrally lacking membranous area, dorsally lacking setae. Lunule between base of antennae and ptilinal suture distinct, distinctly shorter than width of scape. Eye reddish brown, lacking ommatrichia. Facets all of about the same size. Posterior margin of eye convex, lacking an indentation. Ratio of gena height / eye height (measurements are taken from head in lateral view) ≈ 0.1. Ocellar tubercle distinct, with three ocelli (
<figureCitation id="63AEF1B1FFA3A22DFAA6FA03FADAFA12" box="[1302,1363,1408,1432]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="1" pageNumber="302">Fig. 4</figureCitation>
). No ocellar triangle. No frontofacial spot. Vertex only about 2/3 width of frons, separated from latter by a ridge, distinctly rounded anteriorly, bulging posteriorly, and covered with scattered black setulae (
<figureCitation id="63AEF1B1FFA3A22DFB94F983FBEBF992" box="[1060,1122,1536,1560]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="1" pageNumber="302">Fig. 4</figureCitation>
). Gena with a few long black setulae, parafacial with line of regularly arranged minute white and black setulae. Distinct facial grooves reaching mouth edge. Distinct facial carina reaching from base of antennae to a barely developed frontoclypeal tubercle. Ptilinal suture stretching a short way on either side beneath the antennal bases. The area surrounding the ptilinal suture is blackish. Mouth opening slightly tapering dorsally. Postcranium obviously invaginated. Postgena not widened and therefore not separated from occiput. Bottom portion of postcranium distinctly separated. No palp. Proboscis geniculated once only, at base. Labium shorter than headlength, hardly projecting out of mouth opening, distinctly thickened basally, anterior section not fused into a tube. Labrum not recognised. Labellum short, completely divided, broad, white and covered with long yellow setulae. Labellum with 8 indistinct pseudotracheae on each side. Head lacking setae.
</paragraph>
<paragraph id="FB2AED34FFA0A22EFD0BFF11FA3AFF22" blockId="2.[699,1459,146,169]" box="[699,1459,146,169]" pageId="2" pageNumber="303">
CONTRIBUTIONS TO ENTOMOLOGY: BEITRÄGE ZUR ENTOMOLOGIE — 
<emphasis id="C9E13126FFA0A22EFA81FF11FAC1FF23" bold="true" box="[1329,1352,146,169]" pageId="2" pageNumber="303">69</emphasis>
(2) 301–317
</paragraph>
<caption id="AFEABDBCFFA0A22EFF67F935FA3AF946" ID-DOI="http://doi.org/10.5281/zenodo.10125033" ID-Zenodo-Dep="10125033" box="[215,1459,1718,1740]" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="2" pageNumber="303" startId="2.[215,255,1718,1740]" targetBox="[212,1459,255,1710]" targetPageId="2">
<paragraph id="FB2AED34FFA0A22EFF67F935FA3AF946" blockId="2.[214,1459,1718,1770]" box="[215,1459,1718,1740]" pageId="2" pageNumber="303">
<emphasis id="C9E13126FFA0A22EFF67F935FEA3F941" bold="true" box="[215,298,1718,1740]" pageId="2" pageNumber="303">Figs 1–4</emphasis>
: 
<taxonomicName id="3C9596B7FFA0A22EFE86F935FD82F941" authority="Stuke, 2019" authorityName="Stuke" authorityYear="2019" box="[310,523,1718,1739]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
<emphasis id="C9E13126FFA0A22EFE86F935FD82F941" box="[310,523,1718,1739]" italics="true" pageId="2" pageNumber="303">Gellergrimmellus fritzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D28C5DFFA0A22EFDA2F934FDFBF946" box="[530,626,1719,1740]" pageId="2" pageNumber="303" rank="species">spec. nov.</taxonomicNameLabel>
– 
<emphasis id="C9E13126FFA0A22EFD3BF935FD1EF941" bold="true" box="[651,663,1718,1739]" pageId="2" pageNumber="303">1</emphasis>
habitus, lateral view ( holotype). – 
<emphasis id="C9E13126FFA0A22EFBB5F935FB98F941" bold="true" box="[1029,1041,1718,1739]" pageId="2" pageNumber="303">2</emphasis>
arista, lateral view ( paratype). – 
<emphasis id="C9E13126FFA0A22EFADDF935FAF0F941" bold="true" box="[1389,1401,1718,1739]" pageId="2" pageNumber="303">3</emphasis>
head,
</paragraph>
</caption>
<paragraph id="FB2AED34FFA0A22EFF66F957FA8CF960" blockId="2.[214,1459,1718,1770]" box="[214,1285,1747,1770]" pageId="2" pageNumber="303">
lateral view ( 
<typeStatus id="242E5396FFA0A22EFED9F957FE48F960" box="[361,449,1748,1770]" pageId="2" pageNumber="303" type="holotype">holotype</typeStatus>
). – 
<emphasis id="C9E13126FFA0A22EFE52F950FE67F962" bold="true" box="[482,494,1747,1768]" pageId="2" pageNumber="303">4</emphasis>
vertex and frons, dorsal view ( 
<typeStatus id="242E5396FFA0A22EFC9DF957FC0CF960" box="[813,901,1748,1770]" pageId="2" pageNumber="303" type="holotype">holotype</typeStatus>
). ar - aristomere; bf - basal flagellomere.
</paragraph>
<paragraph id="FB2AED34FFA0A22EFF64F8A3FA3AF7B2" blockId="2.[212,817,1824,2104]" lastBlockId="2.[854,1459,1824,2110]" pageId="2" pageNumber="303">
<emphasis id="C9E13126FFA0A22EFF64F8A3FEADF8B2" bold="true" box="[212,292,1824,1848]" pageId="2" pageNumber="303">Thorax</emphasis>
: Presternum distinct, about as wide as the compound basisterna. Presternum distinctly separated from basisternum. Basisternum broad, narrowed to a tip, lacking setulae. Proepisternum with 1 black seta ventrally, dorsally lacking setulae. Mediotergite convex, lacking setulae, and projecting distinctly over scutellum. Subscutellum not developed. Metakatepisternum, anepisternum and anepimeron lacking setulae. Subcostal-radial crossvein sc-r present (
<figureCitation id="63AEF1B1FFA0A22EFE65F7A3FD9DF7B2" box="[469,532,2080,2104]" captionStart="Figs 5–8" captionStartId="3.[174,214,1777,1799]" captionTargetBox="[174,1421,255,1769]" captionTargetId="figure-220@3.[174,1421,255,1769]" captionTargetPageId="3" captionText="Figs 5–8: Gellergrimmellus fritzi spec. nov. – 5  abdomen, dorsal view ( holotype). – 6 wing, ventral view ( paratype). – 7 " figureDoi="http://doi.org/10.5281/zenodo.10125035" httpUri="https://zenodo.org/record/10125035/files/figure.png" pageId="2" pageNumber="303">Fig. 6</figureCitation>
). Radial-medial crossvein r-m complete. Radius R 
<subScript id="6711EF71FFA0A22EFBE4F8B3FBD2F8B4" attach="left" box="[1108,1115,1840,1854]" fontSize="6" pageId="2" pageNumber="303">1</subScript>
and R 
<subScript id="6711EF71FFA0A22EFB12F8B3FB20F8B4" attach="left" box="[1186,1193,1840,1854]" fontSize="6" pageId="2" pageNumber="303">2</subScript>
terminate close together in costa, well beyond the end of the subcosta. Radius R 
<subScript id="6711EF71FFA0A22EFA2BF8D3FA3AF8D4" attach="left" box="[1435,1459,1872,1886]" fontSize="6" pageId="2" pageNumber="303">4+5</subScript>
with a shallow and even curve in the distal section which is directed to a point beyond the wing tip. Radial cell r 
<subScript id="6711EF71FFA0A22EFA2BF813FA3AF814" attach="left" box="[1435,1459,1936,1950]" fontSize="6" pageId="2" pageNumber="303">4+5</subScript>
pedunculate, with vein R 
<subScript id="6711EF71FFA0A22EFBD0F833FBF1F834" attach="left" box="[1120,1144,1968,1982]" fontSize="6" pageId="2" pageNumber="303">4+5</subScript>
+ M distinctly expressed and longer than radial-medial crossvein r-m. Cubital cell cup elongated (distinctly longer than vein A 
<subScript id="6711EF71FFA0A22EFA93F873FAA3F874" attach="left" box="[1315,1322,2032,2046]" fontSize="6" pageId="2" pageNumber="303">1</subScript>
+CuA 
<subScript id="6711EF71FFA0A22EFADCF873FAFAF874" attach="left" box="[1388,1395,2032,2046]" fontSize="6" pageId="2" pageNumber="303">2</subScript>
) and pointed distally (cubitus CuA 
<subScript id="6711EF71FFA0A22EFB26F793FB14F794" attach="left" box="[1174,1181,2064,2078]" fontSize="6" pageId="2" pageNumber="303">2</subScript>
and anal vein A 
<subScript id="6711EF71FFA0A22EFAE3F793FAD3F794" attach="left" box="[1363,1370,2064,2078]" fontSize="6" pageId="2" pageNumber="303">1</subScript>
meet at an acute angle). Cubital vein CuA 
<subScript id="6711EF71FFA0A22EFB74F7B3FB42F7B4" attach="left" box="[1220,1227,2096,2110]" fontSize="6" pageId="2" pageNumber="303">1</subScript>
and crossvein bm-cu
</paragraph>
<paragraph id="FB2AED34FFA1A22FFF1EFF10FB79F8AD" blockId="3.[174,915,147,169]" lastBlockId="3.[174,1421,1776,1831]" pageId="3" pageNumber="304">
STUKE, J.-H.: TWO neW geneRa and tWO neW SpecieS OF 
<taxonomicName id="3C9596B7FFA1A22FFD33FF10FCA6FF23" authority="(DipteRa)" authorityName="Latreille" authorityYear="1802" baseAuthorityName="DipteRa" box="[643,815,147,169]" class="Insecta" family="Conopidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="family">COnOpidae (DipteRa)</taxonomicName>
FROM 
<collectingCountry id="8382ADA4FFA1A22FFCD1FF10FC1AFF23" box="[865,915,147,169]" name="China" pageId="3" pageNumber="304">China</collectingCountry>
postabdomen, lateral view ( 
<typeStatus id="242E5396FFA1A22FFE7FF892FDAEF8AD" box="[463,551,1809,1831]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
). – 
<emphasis id="C9E13126FFA1A22FFDF9F892FDDCF8AC" bold="true" box="[585,597,1809,1830]" pageId="3" pageNumber="304">8</emphasis>
 epandrium, dorsal view ( 
<typeStatus id="242E5396FFA1A22FFCCEF892FC5FF8AC" box="[894,982,1809,1830]" pageId="3" pageNumber="304" type="paratype">paratype</typeStatus>
). ce - cercus; ep - epandrium.
</paragraph>
<caption id="AFEABDBCFFA1A22FFF1EF972FA04F88D" ID-DOI="http://doi.org/10.5281/zenodo.10125035" ID-Zenodo-Dep="10125035" box="[174,1421,1776,1799]" httpUri="https://zenodo.org/record/10125035/files/figure.png" pageId="3" pageNumber="304" startId="3.[174,214,1777,1799]" targetBox="[174,1421,255,1769]" targetPageId="3">
<paragraph id="FB2AED34FFA1A22FFF1EF972FA04F88D" blockId="3.[174,1421,1776,1831]" box="[174,1421,1776,1799]" pageId="3" pageNumber="304">
<emphasis id="C9E13126FFA1A22FFF1EF972FE89F88C" bold="true" box="[174,256,1777,1799]" pageId="3" pageNumber="304">Figs 5–8</emphasis>
: 
<taxonomicName id="3C9596B7FFA1A22FFEBDF972FE68F88C" authority="Stuke, 2019" authorityName="Stuke" authorityYear="2019" box="[269,481,1777,1798]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
<emphasis id="C9E13126FFA1A22FFEBDF972FE68F88C" box="[269,481,1777,1798]" italics="true" pageId="3" pageNumber="304">Gellergrimmellus fritzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D28C5DFFA1A22FFE57F972FDCFF88C" box="[487,582,1777,1798]" pageId="3" pageNumber="304" rank="species">spec. nov.</taxonomicNameLabel>
– 
<emphasis id="C9E13126FFA1A22FFDEEF972FDE3F88C" bold="true" box="[606,618,1777,1798]" pageId="3" pageNumber="304">5</emphasis>
 abdomen, dorsal view ( holotype). – 
<emphasis id="C9E13126FFA1A22FFBB1F972FB84F88C" bold="true" box="[1025,1037,1777,1798]" pageId="3" pageNumber="304">6</emphasis>
wing, ventral view ( paratype). – 
<emphasis id="C9E13126FFA1A22FFAD9F972FAFCF88C" bold="true" box="[1385,1397,1777,1798]" pageId="3" pageNumber="304">7</emphasis>

</paragraph>
</caption>
<paragraph id="FB2AED34FFA1A22FFF1EF8C3FA8DF792" blockId="3.[174,779,1856,2104]" lastBlockId="3.[816,1422,1856,2072]" pageId="3" pageNumber="304">distinctly separated. Alula minute, about 3 times broader than long, with white setulae on posterior margin. Vena spuria indistinctly evident only in cubital cell cup. Base and stem of haltere each with areas of sensillae. Knob of haltere with isolated white setulae. Posterior surfaces of fore and middle tibiae lacking obvious dusted fields distally. Areas with dense brown setulae ventrally at tip of fore tibia, and ventrally and posteriorly at tip of hind tibia. Middle femur posteriorly with regularly arranged, comb-like long setulae. Hind femur dorsally lacking outstanding long setulae. Tibiae dorsally with small preapical setulae. No setulae ventrally on tibiae. Femora ventrally lacking rows of short black setulae. Hind femur not obviously thickened in basal half. No lines of black, regularly arranged setulae ventrally on tarsi.</paragraph>
<caption id="AFEABDBCFFA6A228FF64F932FCA1F8A8" ID-DOI="http://doi.org/10.5281/zenodo.10125037" ID-Zenodo-Dep="10125037" httpUri="https://zenodo.org/record/10125037/files/figure.png" pageId="4" pageNumber="305" startId="4.[212,252,1713,1735]" targetBox="[244,1442,299,1684]" targetPageId="4">
<paragraph id="FB2AED34FFA6A228FF64F932FCA1F8A8" blockId="4.[212,1459,1712,1826]" pageId="4" pageNumber="305">
<emphasis id="C9E13126FFA6A228FF64F932FEBDF94C" bold="true" box="[212,308,1713,1735]" pageId="4" pageNumber="305">Figs 9–10</emphasis>
:  genitalia of 
<taxonomicName id="3C9596B7FFA6A228FE66F932FD25F94C" authority="Stuke, 2019" authorityName="Stuke" authorityYear="2019" box="[470,684,1713,1734]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
<emphasis id="C9E13126FFA6A228FE66F932FD25F94C" box="[470,684,1713,1734]" italics="true" pageId="4" pageNumber="305">Gellergrimmellus fritzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D28C5DFFA6A228FD05F931FC9FF94D" box="[693,790,1714,1735]" pageId="4" pageNumber="305" rank="species">spec. nov.</taxonomicNameLabel>
– 
<emphasis id="C9E13126FFA6A228FC82F932FCB7F94C" bold="true" box="[818,830,1713,1734]" pageId="4" pageNumber="305">9</emphasis>
 hypandrium and phallus, lateral view ( paratype); – 
<emphasis id="C9E13126FFA6A228FA31F932FA10F94C" bold="true" box="[1409,1433,1713,1734]" pageId="4" pageNumber="305">10</emphasis>
 hypandrium and phallus, dorsal view ( paratype). bp - basiphallus; dbhy - dorsolateral bar of hypandrium; dp - distiphallus; ea - ejaculatory apodeme; edp - evagination of distiphallus; hya - hypandrial arm; hybr - hypandrial bridge; hyr - hypandrial bars; po - postgonite; poe - postgonite evagination; rs - ring sclerite.
</paragraph>
</caption>
<paragraph id="FB2AED34FFA6A228FF64F8BCFEF0F812" blockId="4.[212,817,1855,2104]" pageId="4" pageNumber="305">
<emphasis id="C9E13126FFA6A228FF64F8BCFEC8F8DD" bold="true" box="[212,321,1855,1879]" pageId="4" pageNumber="305">Abdomen</emphasis>
: Tergites 1−3 fused but distinctly separated from each other (
<figureCitation id="63AEF1B1FFA6A228FE16F8E3FE6CF8F2" box="[422,485,1888,1912]" captionStart="Figs 5–8" captionStartId="3.[174,214,1777,1799]" captionTargetBox="[174,1421,255,1769]" captionTargetId="figure-220@3.[174,1421,255,1769]" captionTargetPageId="3" captionText="Figs 5–8: Gellergrimmellus fritzi spec. nov. – 5  abdomen, dorsal view ( holotype). – 6 wing, ventral view ( paratype). – 7 " figureDoi="http://doi.org/10.5281/zenodo.10125035" httpUri="https://zenodo.org/record/10125035/files/figure.png" pageId="4" pageNumber="305">Fig. 5</figureCitation>
). Lateral margins of tergites almost straight.
</paragraph>
<paragraph id="FB2AED34FFA6A228FF64F822FB6EF852" blockId="4.[212,817,1855,2104]" lastBlockId="4.[854,1460,1856,2104]" pageId="4" pageNumber="305"> abdomen with 5 tergites. Tergite 1 with obvious black setulae laterally on the bulbous projections. Tergite 2 distinctly elongated and parallel-sided, about 5 times as long as broad when viewed dorsally, and lacking obvious lateral tufts of setulae. Tergite 3 distinctly widened posteriorly, about 3.8 times as wide posteriorly than anteriorly. Sternites 1-5 present, sternites 1 and 2 not fused and distinctly separated. Tergite 5 and sternite 5 distinctly separated. Sternite 5 apically with a small field of thick setulae and several long black setulae.</paragraph>
<paragraph id="FB2AED34FFA6A229FCE6F862FDB2F812" blockId="4.[854,1460,1856,2104]" lastBlockId="5.[174,779,1184,2104]" lastPageId="5" lastPageNumber="306" pageId="4" pageNumber="305">
 postabdomen with protandrium obviously broader than epandrium and therefore projecting over it. Sternite 8 distinctly delimited from protandrium. Ventrally the lateral edges of the protandrium are fused by a narrow sclerotised strip, which is not widened medially. Cercus distinct, completely sclerotised and covered with setulae (
<figureCitation id="63AEF1B1FFA7A229FEF8FA83FE0CFA92" box="[328,389,1280,1304]" captionStart="Figs 5–8" captionStartId="3.[174,214,1777,1799]" captionTargetBox="[174,1421,255,1769]" captionTargetId="figure-220@3.[174,1421,255,1769]" captionTargetPageId="3" captionText="Figs 5–8: Gellergrimmellus fritzi spec. nov. – 5  abdomen, dorsal view ( holotype). – 6 wing, ventral view ( paratype). – 7 " figureDoi="http://doi.org/10.5281/zenodo.10125035" httpUri="https://zenodo.org/record/10125035/files/figure.png" pageId="5" pageNumber="306">Fig. 8</figureCitation>
). No hypoproct evident. Posterior and anterior surstyli absent. No obvious strong black setae nor long black setulae which would indicate the base of a surstylus. Subepandrial plate not sclerotised nor covered with microtrichia and therefore not evident. Hypandrium slightly sclerotised or hyaline with the exception of two unique dorsolateral bars (
<figureCitation id="63AEF1B1FFA7A229FDCCFA43FD35FA52" box="[636,700,1472,1496]" captionStart="Figs 9–10" captionStartId="4.[212,252,1713,1735]" captionTargetBox="[244,1442,299,1684]" captionTargetId="figure-256@4.[212,1459,252,1703]" captionTargetPageId="4" captionText="Figs 9–10:  genitalia of Gellergrimmellus fritzi spec. nov. – 9  hypandrium and phallus, lateral view ( paratype); – 10  hypandrium and phallus, dorsal view ( paratype). bp - basiphallus; dbhy - dorsolateral bar of hypandrium; dp - distiphallus; ea - ejaculatory apodeme; edp - evagination of distiphallus; hya - hypandrial arm; hybr - hypandrial bridge; hyr - hypandrial bars; po - postgonite; poe - postgonite evagination; rs - ring sclerite." figureDoi="http://doi.org/10.5281/zenodo.10125037" httpUri="https://zenodo.org/record/10125037/files/figure.png" pageId="5" pageNumber="306">Fig. 9</figureCitation>
: dbhy). These bars are curved upwards apically, bear a broad tooth basally and are connected by a small, slightly sclerotised hypandrial bridge (
<figureCitation id="63AEF1B1FFA7A229FE65F9A3FD9DF9B2" box="[469,532,1568,1592]" captionStart="Figs 9–10" captionStartId="4.[212,252,1713,1735]" captionTargetBox="[244,1442,299,1684]" captionTargetId="figure-256@4.[212,1459,252,1703]" captionTargetPageId="4" captionText="Figs 9–10:  genitalia of Gellergrimmellus fritzi spec. nov. – 9  hypandrium and phallus, lateral view ( paratype); – 10  hypandrium and phallus, dorsal view ( paratype). bp - basiphallus; dbhy - dorsolateral bar of hypandrium; dp - distiphallus; ea - ejaculatory apodeme; edp - evagination of distiphallus; hya - hypandrial arm; hybr - hypandrial bridge; hyr - hypandrial bars; po - postgonite; poe - postgonite evagination; rs - ring sclerite." figureDoi="http://doi.org/10.5281/zenodo.10125037" httpUri="https://zenodo.org/record/10125037/files/figure.png" pageId="5" pageNumber="306">Fig. 9</figureCitation>
: hybr). No hypandrial lobe evident. Hypandrial bars fused distally to a hypandrial arm (
<figureCitation id="63AEF1B1FFA7A229FE82F9E3FEF4F9F2" box="[306,381,1632,1656]" captionStart="Figs 9–10" captionStartId="4.[212,252,1713,1735]" captionTargetBox="[244,1442,299,1684]" captionTargetId="figure-256@4.[212,1459,252,1703]" captionTargetPageId="4" captionText="Figs 9–10:  genitalia of Gellergrimmellus fritzi spec. nov. – 9  hypandrium and phallus, lateral view ( paratype); – 10  hypandrium and phallus, dorsal view ( paratype). bp - basiphallus; dbhy - dorsolateral bar of hypandrium; dp - distiphallus; ea - ejaculatory apodeme; edp - evagination of distiphallus; hya - hypandrial arm; hybr - hypandrial bridge; hyr - hypandrial bars; po - postgonite; poe - postgonite evagination; rs - ring sclerite." figureDoi="http://doi.org/10.5281/zenodo.10125037" httpUri="https://zenodo.org/record/10125037/files/figure.png" pageId="5" pageNumber="306">Fig. 10</figureCitation>
: hya, hyr). Hypandrial membrane reduced, lacking microtrichia. Phallus sheath not fused dorsally; lacking any evagination or setulae. Postgonite small. Postgonite evagination not sclerotised, not projecting above distiphallus and lacking obvious microtrichia (
<figureCitation id="63AEF1B1FFA7A229FF05F883FF7BF892" box="[181,242,1792,1816]" captionStart="Figs 9–10" captionStartId="4.[212,252,1713,1735]" captionTargetBox="[244,1442,299,1684]" captionTargetId="figure-256@4.[212,1459,252,1703]" captionTargetPageId="4" captionText="Figs 9–10:  genitalia of Gellergrimmellus fritzi spec. nov. – 9  hypandrium and phallus, lateral view ( paratype); – 10  hypandrium and phallus, dorsal view ( paratype). bp - basiphallus; dbhy - dorsolateral bar of hypandrium; dp - distiphallus; ea - ejaculatory apodeme; edp - evagination of distiphallus; hya - hypandrial arm; hybr - hypandrial bridge; hyr - hypandrial bars; po - postgonite; poe - postgonite evagination; rs - ring sclerite." figureDoi="http://doi.org/10.5281/zenodo.10125037" httpUri="https://zenodo.org/record/10125037/files/figure.png" pageId="5" pageNumber="306">Fig. 9</figureCitation>
: poe). No plate on inner side of postgonite evagination. Ring sclerite developed (
<figureCitation id="63AEF1B1FFA7A229FE44F8A3FDB6F8B2" box="[500,575,1824,1848]" captionStart="Figs 9–10" captionStartId="4.[212,252,1713,1735]" captionTargetBox="[244,1442,299,1684]" captionTargetId="figure-256@4.[212,1459,252,1703]" captionTargetPageId="4" captionText="Figs 9–10:  genitalia of Gellergrimmellus fritzi spec. nov. – 9  hypandrium and phallus, lateral view ( paratype); – 10  hypandrium and phallus, dorsal view ( paratype). bp - basiphallus; dbhy - dorsolateral bar of hypandrium; dp - distiphallus; ea - ejaculatory apodeme; edp - evagination of distiphallus; hya - hypandrial arm; hybr - hypandrial bridge; hyr - hypandrial bars; po - postgonite; poe - postgonite evagination; rs - ring sclerite." figureDoi="http://doi.org/10.5281/zenodo.10125037" httpUri="https://zenodo.org/record/10125037/files/figure.png" pageId="5" pageNumber="306">Fig. 10</figureCitation>
: rs). No epiphallus recognised. Phallus apodeme longer than hypandrium arm. Ejaculatory apodeme elongate, lacking distinct attachment to sperm sac (
<figureCitation id="63AEF1B1FFA7A229FE0FF803FD82F812" box="[447,523,1920,1944]" captionStart="Figs 9–10" captionStartId="4.[212,252,1713,1735]" captionTargetBox="[244,1442,299,1684]" captionTargetId="figure-256@4.[212,1459,252,1703]" captionTargetPageId="4" captionText="Figs 9–10:  genitalia of Gellergrimmellus fritzi spec. nov. – 9  hypandrium and phallus, lateral view ( paratype); – 10  hypandrium and phallus, dorsal view ( paratype). bp - basiphallus; dbhy - dorsolateral bar of hypandrium; dp - distiphallus; ea - ejaculatory apodeme; edp - evagination of distiphallus; hya - hypandrial arm; hybr - hypandrial bridge; hyr - hypandrial bars; po - postgonite; poe - postgonite evagination; rs - ring sclerite." figureDoi="http://doi.org/10.5281/zenodo.10125037" httpUri="https://zenodo.org/record/10125037/files/figure.png" pageId="5" pageNumber="306">Fig. 10</figureCitation>
: ea).
</paragraph>
<caption id="AFEABDBCFFA7A229FF1EFBAAFBEBFBF0" ID-DOI="http://doi.org/10.5281/zenodo.10125039" ID-Zenodo-Dep="10125039" httpUri="https://zenodo.org/record/10125039/files/figure.png" pageId="5" pageNumber="306" startId="5.[174,214,1065,1087]" targetBox="[174,1421,255,1059]" targetPageId="5">
<paragraph id="FB2AED34FFA7A229FF1EFBAAFBEBFBF0" blockId="5.[174,1422,1065,1146]" pageId="5" pageNumber="306">
<emphasis id="C9E13126FFA7A229FF1EFBAAFE90FBB4" bold="true" box="[174,281,1065,1087]" pageId="5" pageNumber="306">Figs 11–13</emphasis>
:  postabdomen of 
<taxonomicName id="3C9596B7FFA7A229FE54FBAAFD33FBB4" authority="Stuke, 2019" authorityName="Stuke" authorityYear="2019" box="[484,698,1065,1086]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="fritzi" status="sp. nov.">
<emphasis id="C9E13126FFA7A229FE54FBAAFD33FBB4" box="[484,698,1065,1086]" italics="true" pageId="5" pageNumber="306">Gellergrimmellus fritzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D28C5DFFA7A229FD71FBA9FCA9FBB5" box="[705,800,1066,1087]" pageId="5" pageNumber="306" rank="species">spec. nov.</taxonomicNameLabel>
– 
<emphasis id="C9E13126FFA7A229FC8AFBAAFCDBFBB4" bold="true" box="[826,850,1065,1086]" pageId="5" pageNumber="306">11</emphasis>
 postabdomen, ventral view ( paratype). – 
<emphasis id="C9E13126FFA7A229FAADFBAAFABCFBB4" bold="true" box="[1309,1333,1065,1086]" pageId="5" pageNumber="306">12</emphasis>
 theca, posterior view ( paratype). – 
<emphasis id="C9E13126FFA7A229FE6CFBC4FE7DFBD6" bold="true" box="[476,500,1095,1116]" pageId="5" pageNumber="306">13</emphasis>
spermathecae ( paratype). fsrd - fused spermathecae ducts; pS6 - posterior part of sternite 6; S6 - sternite 6; S7 - sternite 7; sr - spermatheca; srd - spermatheca duct; T6 - tergite 6; T7 - tergite 7.
</paragraph>
</caption>
<paragraph id="FB2AED34FFA7A229FF1EF822FB11F932" blockId="5.[174,779,1184,2104]" lastBlockId="5.[816,1423,1184,1720]" pageId="5" pageNumber="306">
 abdomen with sternites 1−2 fused. Sternites 3−4 not protruding ventrally, posterior parts of sternites inconspicuous. Tergites 3 and 4 lacking any protuberances. Tergite 5 and sternite 5 fused laterally to form a syntergosternite with obvious theca below (
<figureCitation id="63AEF1B1FFA7A229FDF8F7A3FD1DF7B2" box="[584,660,2080,2104]" captionStart="Figs 11–13" captionStartId="5.[174,214,1065,1087]" captionTargetBox="[174,1421,255,1059]" captionTargetId="figure-594@5.[174,1421,255,1059]" captionTargetPageId="5" captionText="Figs 11–13:  postabdomen of Gellergrimmellus fritzi spec. nov. – 11  postabdomen, ventral view ( paratype). – 12  theca, posterior view ( paratype). – 13 spermathecae ( paratype). fsrd - fused spermathecae ducts; pS6 - posterior part of sternite 6; S6 - sternite 6; S7 - sternite 7; sr - spermatheca; srd - spermatheca duct; T6 - tergite 6; T7 - tergite 7." figureDoi="http://doi.org/10.5281/zenodo.10125039" httpUri="https://zenodo.org/record/10125039/files/figure.png" pageId="5" pageNumber="306">Fig. 12</figureCitation>
). Posterior part of sternite 6 conspicuous sclerotised, slightly bent dorsally, and distinctly divided medially (
<figureCitation id="63AEF1B1FFA7A229FAB1FB43FAC4FB52" box="[1281,1357,1216,1240]" captionStart="Figs 11–13" captionStartId="5.[174,214,1065,1087]" captionTargetBox="[174,1421,255,1059]" captionTargetId="figure-594@5.[174,1421,255,1059]" captionTargetPageId="5" captionText="Figs 11–13:  postabdomen of Gellergrimmellus fritzi spec. nov. – 11  postabdomen, ventral view ( paratype). – 12  theca, posterior view ( paratype). – 13 spermathecae ( paratype). fsrd - fused spermathecae ducts; pS6 - posterior part of sternite 6; S6 - sternite 6; S7 - sternite 7; sr - spermatheca; srd - spermatheca duct; T6 - tergite 6; T7 - tergite 7." figureDoi="http://doi.org/10.5281/zenodo.10125039" httpUri="https://zenodo.org/record/10125039/files/figure.png" pageId="5" pageNumber="306">Fig. 11</figureCitation>
: pS6). Tergite 7 distinctly bent ventrally, without a longitudinal gap, and with a hardly protruding blunt tooth in middle of posterior margin. Sternite 8 not fused with syntergite 8+9 and therefore not connecting on its sides. Tooth on syntergite 8+9 distinct, its base elongated anteriorly. Sternite 9 slightly bulging posteriorly, lacking strong long black setae but with several long setulae. Paired cerci distinct. Sack-like ventral protrusion of vagina with distinct annular sclerotisation. Sack-like ventral protrusion is hardly larger than annular sclerotisation and no evagination or additional sclerotisations are recognised. 2 pairs of round spermathecae, spermathecal ducts fused some distance before reaching spermathecae. Spermathecal ducts sclerotised a long distance before reaching spermathecae, this sclerotised part winding (
<figureCitation id="63AEF1B1FFA7A229FB8FF923FB02F932" box="[1087,1163,1696,1720]" captionStart="Figs 11–13" captionStartId="5.[174,214,1065,1087]" captionTargetBox="[174,1421,255,1059]" captionTargetId="figure-594@5.[174,1421,255,1059]" captionTargetPageId="5" captionText="Figs 11–13:  postabdomen of Gellergrimmellus fritzi spec. nov. – 11  postabdomen, ventral view ( paratype). – 12  theca, posterior view ( paratype). – 13 spermathecae ( paratype). fsrd - fused spermathecae ducts; pS6 - posterior part of sternite 6; S6 - sternite 6; S7 - sternite 7; sr - spermatheca; srd - spermatheca duct; T6 - tergite 6; T7 - tergite 7." figureDoi="http://doi.org/10.5281/zenodo.10125039" httpUri="https://zenodo.org/record/10125039/files/figure.png" pageId="5" pageNumber="306">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B38FBEBFFFA7A229FC80F95CFAB1F852" pageId="5" pageNumber="306" type="distribution">
<paragraph id="FB2AED34FFA7A229FC80F95CFAB1F852" blockId="5.[816,1422,1759,2008]" pageId="5" pageNumber="306">
<emphasis id="C9E13126FFA7A229FC80F95CFC3FF97D" bold="true" box="[816,950,1759,1783]" pageId="5" pageNumber="306">Distribution</emphasis>
: The genus 
<taxonomicName id="3C9596B7FFA7A229FB8DF963FB62F972" authorityName="Stuke" authorityYear="2019" box="[1085,1259,1760,1784]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA7A229FB8DF963FB62F972" box="[1085,1259,1760,1784]" italics="true" pageId="5" pageNumber="306">Gellergrimmellus</emphasis>
</taxonomicName>
is only known from the 
<emphasis id="C9E13126FFA7A229FC12F883FBA5F89D" box="[930,1068,1792,1816]" italics="true" pageId="5" pageNumber="306">locus typicus</emphasis>
of 
<taxonomicName id="3C9596B7FFA7A229FBEDF883FAC3F892" authorityName="Stuke" authorityYear="2019" box="[1117,1354,1792,1816]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="fritzi">
<emphasis id="C9E13126FFA7A229FBEDF883FAC3F892" box="[1117,1354,1792,1816]" italics="true" pageId="5" pageNumber="306">Gellergrimmellus fritzi</emphasis>
</taxonomicName>
. This location is placed in the Min Mountains (Minshan). The biogeographical classification of this location is not clear. Sensu 
<bibRefCitation id="9F0490C5FFA7A229FCC3F8E3FB0EF8F2" author="HEISER, M. &amp; SCHMITT, M." box="[883,1159,1888,1912]" pageId="5" pageNumber="306" pagination="2047 - 2058" refId="ref8965" refString="HEISER, M. &amp; SCHMITT, M. 2013: Tracking the boundary between the Palaearctic and the Oriental region: new insights from dragonflies and damselflies (Odonata). - Journal of Biogeography 40: 2047 - 2058. - https: // onlinelibrary. wiley. com / doi / epdf / 10.1111 / jbi. 12133." type="journal article" year="2013">HEISER &amp; SCHMITT (2013)</bibRefCitation>
the locus typicus belongs to the transition zone of the Oriental and Palaearctic regions, whilst sensu 
<bibRefCitation id="9F0490C5FFA7A229FBA2F823FB5AF832" author="CHEN, L. &amp; SONG, Y. &amp; XU, S." box="[1042,1235,1952,1976]" pageId="5" pageNumber="306" pagination="833 - 841" refId="ref8631" refString="CHEN, L.; SONG, Y. &amp; XU, S. 2008: The boundary of palaearctic and oriental realms in western China. - Progress in Natural Science 18: 833 - 841. - http: // w w w. ns fc. gov. c n / c s c / 2 0 3 4 5 / 2 4 3 7 1 / p d f / 2 0 0 8 / The % 20 boundary % 20 of % 20 palaearctic % 20 and % 20 oriental % 20 realms % 20 in % 20 western % 20 China. pdf." type="journal article" year="2008">CHEN et al. (2008)</bibRefCitation>
or 
<bibRefCitation id="9F0490C5FFA7A229FB48F823FCFCF852" author="KREFT, H. &amp; JETZ, W." pageId="5" pageNumber="306" pagination="2029 - 2053" refId="ref9027" refString="KREFT, H. &amp; JETZ, W. 2010: A framework for delineating biogeographical regions based on species distributions. - Journal of Biogeography 37: 2029 - 2053. - https: // onlinelibrary. wiley. com / doi / epdf / 10.1111 / j. 1365 - 2699.2010.02375. x." type="journal article" year="2010">KREFT &amp; JETZ (2010)</bibRefCitation>
the 
<emphasis id="C9E13126FFA7A229FC11F843FBABF85D" box="[929,1058,1984,2008]" italics="true" pageId="5" pageNumber="306">locus typicus</emphasis>
lies in the Oriental region.
</paragraph>
</subSubSection>
<subSubSection id="B38FBEBFFFA7A22AFC80F87CFE4BFC92" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306" type="biology_ecology">
<paragraph id="FB2AED34FFA7A229FC80F87CFCFCF7B2" blockId="5.[816,1421,2047,2104]" pageId="5" pageNumber="306">
<emphasis id="C9E13126FFA7A229FC80F87CFC0CF79D" bold="true" box="[816,901,2047,2071]" pageId="5" pageNumber="306">Biology</emphasis>
: Nothing is known about the biology of this genus.
</paragraph>
<paragraph id="FB2AED34FFA4A22AFF64FF7CFE4BFC92" blockId="6.[212,817,255,792]" pageId="6" pageNumber="307">
<emphasis id="C9E13126FFA4A22AFF64FF7CFD5AFE9D" bold="true" box="[212,723,255,280]" pageId="6" pageNumber="307">
Phylogenetic placement of 
<taxonomicName id="3C9596B7FFA4A22AFDAAFF7CFD5AFE9D" authorityName="Stuke" authorityYear="2019" box="[538,723,255,279]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA4A22AFDAAFF7CFD5AFE9D" bold="true" box="[538,723,255,279]" italics="true" pageId="6" pageNumber="307">Gellergrimmellus</emphasis>
</taxonomicName>
</emphasis>
: 
<taxonomicName id="3C9596B7FFA4A22AFD5AFE83FEC1FEB2" authorityName="Stuke" authorityYear="2019" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA4A22AFD5AFE83FEC1FEB2" italics="true" pageId="6" pageNumber="307">Gellergrimmellus</emphasis>
</taxonomicName>
belongs in the subfamily 
<taxonomicName id="3C9596B7FFA4A22AFDDAFEA3FD69FEB2" box="[618,736,288,312]" class="Insecta" family="Conopidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Conopinae">Conopinae</taxonomicName>
due to the following characters: Stylate arista placed apically on basal flagellomere (
<figureCitation id="63AEF1B1FFA4A22AFE77FEE3FD8EFEF2" box="[455,519,352,376]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="6" pageNumber="307">Fig. 2</figureCitation>
); no ocellar triangle; lunule distinct (
<figureCitation id="63AEF1B1FFA4A22AFE83FE03FEFAFE12" box="[307,371,384,408]" captionStart="Figs 1–4" captionStartId="2.[215,255,1718,1740]" captionTargetBox="[212,1459,255,1710]" captionTargetId="figure-251@2.[212,1459,255,1710]" captionTargetPageId="2" captionText="Figs 1–4: Gellergrimmellus fritzi spec. nov. – 1 habitus, lateral view ( holotype). – 2 arista, lateral view ( paratype). – 3 head," figureDoi="http://doi.org/10.5281/zenodo.10125033" httpUri="https://zenodo.org/record/10125033/files/figure.png" pageId="6" pageNumber="307">Fig. 4</figureCitation>
); facial grooves reaching to mouth opening, divided by a central carina which widens ventrally; mouth opening tapering dorsally; postgena not widened and therefore not separate from the narrow occiput; chaetotaxy reduced, no obvious setae on the head; radial cell r 
<subScript id="6711EF71FFA4A22AFEE5FDB3FEE4FDB4" attach="left" box="[341,365,560,574]" fontSize="6" pageId="6" pageNumber="307">4+5</subScript>
petiolate and R 
<subScript id="6711EF71FFA4A22AFD92FDB3FDB3FDB4" attach="left" box="[546,570,560,574]" fontSize="6" pageId="6" pageNumber="307">4+5</subScript>
+ M distinct (
<figureCitation id="63AEF1B1FFA4A22AFD55FDA3FCAAFDB2" box="[741,803,544,568]" captionStart="Figs 5–8" captionStartId="3.[174,214,1777,1799]" captionTargetBox="[174,1421,255,1769]" captionTargetId="figure-220@3.[174,1421,255,1769]" captionTargetPageId="3" captionText="Figs 5–8: Gellergrimmellus fritzi spec. nov. – 5  abdomen, dorsal view ( holotype). – 6 wing, ventral view ( paratype). – 7 " figureDoi="http://doi.org/10.5281/zenodo.10125035" httpUri="https://zenodo.org/record/10125035/files/figure.png" pageId="6" pageNumber="307">Fig. 6</figureCitation>
); subcosta-radial crossvein sc-r well developed (
<figureCitation id="63AEF1B1FFA4A22AFD55FDC3FCAAFDD2" box="[741,803,576,600]" captionStart="Figs 5–8" captionStartId="3.[174,214,1777,1799]" captionTargetBox="[174,1421,255,1769]" captionTargetId="figure-220@3.[174,1421,255,1769]" captionTargetPageId="3" captionText="Figs 5–8: Gellergrimmellus fritzi spec. nov. – 5  abdomen, dorsal view ( holotype). – 6 wing, ventral view ( paratype). – 7 " figureDoi="http://doi.org/10.5281/zenodo.10125035" httpUri="https://zenodo.org/record/10125035/files/figure.png" pageId="6" pageNumber="307">Fig. 6</figureCitation>
);  protandrium obviously broader than epandrium and therefore projecting over it. As discussed in detail under 
<taxonomicName id="3C9596B7FFA4A22AFF64FD23FED2FD3D" authorityName=", Taxonomie" authorityYear="2019" box="[212,347,672,695]" class="Insecta" family="Conopidae" genus="Hauserimyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA4A22AFF64FD23FED2FD3D" box="[212,347,672,695]" italics="true" pageId="6" pageNumber="307">Hauserimyia</emphasis>
</taxonomicName>
a more detailed phylogenetic classification is not possible at the time of writing. Within the 
<taxonomicName id="3C9596B7FFA4A22AFD0BFD43FCB8FD52" box="[699,817,704,728]" class="Insecta" family="Conopidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="subFamily" subFamily="Conopinae">Conopinae</taxonomicName>
<taxonomicName id="3C9596B7FFA4A22AFF64FD63FE0FFD72" authorityName="Stuke" authorityYear="2019" box="[212,390,736,760]" class="Insecta" family="Conopidae" genus="Gellergrimmellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="C9E13126FFA4A22AFF64FD63FE0FFD72" box="[212,390,736,760]" italics="true" pageId="6" pageNumber="307">Gellergrimmellus</emphasis>
</taxonomicName>
is an isolated genus with no related or similar genera known.
</paragraph>
</subSubSection>
</treatment>
</document>