228 lines
37 KiB
XML
228 lines
37 KiB
XML
<document id="AA4CA4B5CE6CBD7E4E3D042546776B39" ID-DOI="10.5281/zenodo.5393017" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393017" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630317797950" checkinUser="marcus" docAuthor="Duy-Jacquemin, Monique Nguyen" docDate="2002" docId="731987BCFFD9FFE8FF4725555E99D758" docLanguage="en" docName="Zoosystema.2002.24.2.451-470.pdf" docOrigin="Zoosystema 24 (2)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Lophoturus monserratensis Duy-Jacquemin 2002, n. sp." docType="treatment" docVersion="3" lastPageNumber="455" masterDocId="8F20FFC4FFDDFFEEFFB7275E5E46D61D" masterDocTitle="New species and distribution of the genera Lophoturus and Ancistroxenus (Myriapoda, Diplopoda, Penicillata) in the Caribbean and northern South America" masterLastPageNumber="470" masterPageNumber="451" pageNumber="455" updateTime="1699224897839" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="1D63964E81FDE1FBA92E15930E544714">
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<mods:title id="1C02F55790C68F813C9CAF97F15328C3">New species and distribution of the genera Lophoturus and Ancistroxenus (Myriapoda, Diplopoda, Penicillata) in the Caribbean and northern South America</mods:title>
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<mods:name id="829DE338AD7F4E37D0B12584B1CFEB88" type="personal">
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<mods:role id="5BE3B006B71D71AC89E1A256B5112510">
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<mods:roleTerm id="16C30E7160AB42507CEA09DE1948F0CE">Author</mods:roleTerm>
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<mods:namePart id="7DE47F9FB8E5127C0378CC25D935C59F">Duy-Jacquemin, Monique Nguyen</mods:namePart>
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<mods:affiliation id="FAA3261BAF2DC3A089BD32644DDE8616">Laboratoire de Zoologie (Arthropodes), Muséum national d’Histoire naturelle, 61 rue Buffon, F- 75231 Paris cedex 05 (France) monguyen @ mnhn. fr.</mods:affiliation>
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<mods:nameIdentifier id="3D0FA847202F4C7DC4A4B9ABABA34113" type="email">monguyen@mnhn.fr</mods:nameIdentifier>
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<mods:typeOfResource id="50B0BBD52CBD449AD286D44F94807F59">text</mods:typeOfResource>
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<mods:titleInfo id="474AF6F5E0762C8BA0AD7901B085CE71">
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<mods:title id="0C2E26A8B5A292B250521D2C8C25DE61">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="80A5DD90087C245F2E25DCCE36C39C54">
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<mods:date id="826C754C7FE92CD3708B250E2968C6F2">2002</mods:date>
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<mods:detail id="895B315446DE4D4BFCC44AD21A2D778F" type="volume">
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<mods:number id="4BC3AFCE37F242A0B2C1E76AC6CE2206">24</mods:number>
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<mods:detail id="4E9B7D3C10C0C679038DE2DFDAF44CEE" type="issue">
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<mods:number id="B18AA7E391E7A564E062BC5D12140835">2</mods:number>
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<mods:start id="BEFC2390102F0D4DF8A8CA7488D1A3EA">451</mods:start>
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<mods:end id="310FE7CEADC4CE851049EA14CC05B3ED">470</mods:end>
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<mods:classification id="B8E427CFD1A71252BCF026ADAD3E4F6C">journal article</mods:classification>
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<mods:identifier id="C24B35B8C6B739976BE746EA3E53A738" type="DOI">10.5281/zenodo.5393017</mods:identifier>
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<mods:identifier id="1871718891936D934E85340C450B8E9E" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="72019B74B4F9B1496ECAEB8EF1E9A0BA" type="Zenodo-Dep">5393017</mods:identifier>
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<treatment id="731987BCFFD9FFE8FF4725555E99D758" LSID="urn:lsid:plazi:treatment:731987BCFFD9FFE8FF4725555E99D758" httpUri="http://treatment.plazi.org/id/731987BCFFD9FFE8FF4725555E99D758" lastPageId="6" lastPageNumber="455" pageId="4" pageNumber="455">
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<subSubSection id="B3AA6521FFD9FFEAFF4725555F8FD458" pageId="4" pageNumber="455" type="nomenclature">
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<paragraph id="FB0F36AAFFD9FFEAFF4725555F8FD458" blockId="4.[240,595,521,581]" pageId="4" pageNumber="455">
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<taxonomicName id="3CB04D29FFD9FFEAFF4725555C56D43E" ID-CoL="3W4QG" authority="Duy-Jacquemin, 2002" authorityName="Duy-Jacquemin" authorityYear="2002" box="[240,528,521,549]" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="monserratensis" status="sp. nov.">
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<emphasis id="C9C4EAB8FFD9FFEAFF4725555C56D43E" bold="true" box="[240,528,521,549]" italics="true" pageId="4" pageNumber="455">Lophoturus monserratensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D2F757C3FFD9FFEAFDAF25545C15D439" box="[536,595,522,548]" pageId="4" pageNumber="455" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="638B2A2FFFD9FFEAFE3625755F84D458" box="[385,450,555,581]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B3AA6521FFD9FFEAFF20253D5F3ED4D5" pageId="4" pageNumber="455" type="materials_examined">
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<paragraph id="FB0F36AAFFD9FFEAFF20253D5F3ED4D5" blockId="4.[151,684,611,778]" pageId="4" pageNumber="455">
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<collectionCode id="9DA1AE6FFFD9FFEAFF20253D5F69D464" box="[151,303,611,634]" pageId="4" pageNumber="455">TYPE MATERIAL</collectionCode>
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. —
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<emphasis id="C9C4EAB8FFD9FFEAFED7253A5F8CD461" bold="true" box="[352,458,612,636]" pageId="4" pageNumber="455">
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<collectingCountry id="83A7763AFFD9FFEAFED7253A5F81D461" box="[352,455,612,636]" name="Colombia" pageId="4" pageNumber="455">Colombia</collectingCountry>
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.
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</emphasis>
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Near
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<collectingRegion id="3974F848FFD9FFEAFDBB253A5C13D461" box="[524,597,612,636]" country="Colombia" name="Bogota" pageId="4" pageNumber="455">Bogotá</collectingRegion>
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, Párámo de Monserrate,
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<emphasis id="C9C4EAB8FFD9FFEAFE8025235F01D489" box="[311,327,637,660]" italics="true" pageId="4" pageNumber="455">c.</emphasis>
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<quantity id="3C489B4FFFD9FFEAFEF825235FE7D489" box="[335,417,637,661]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="4" pageNumber="455" unit="m" value="2250.0">2250 m</quantity>
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,
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<taxonomicName id="3CB04D29FFD9FFEAFE1925235C33D488" box="[430,629,637,661]" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="455" phylum="Tracheophyta" rank="species" species="grandiflora">
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<emphasis id="C9C4EAB8FFD9FFEAFE1925235C33D488" box="[430,629,637,661]" italics="true" pageId="4" pageNumber="455">Espeletia grandiflora</emphasis>
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</taxonomicName>
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dead leaves,
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<date id="8F0E106AFFD9FFEAFF5725C95F24D4B2" box="[224,354,663,687]" pageId="4" pageNumber="455" value="1978-07-21">
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21.
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<collectionCode id="9DA1AE6FFFD9FFEAFF4925C95F61D4B2" box="[254,295,663,687]" pageId="4" pageNumber="455">VII</collectionCode>
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.1978
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</date>
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, H. Sturm leg., ad.
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<typeStatus id="240B8808FFD9FFEAFDE425C95CEAD4B2" box="[595,684,663,687]" pageId="4" pageNumber="455" type="holotype">holotype</typeStatus>
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(78/209 Kol.
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<collectionCode id="9DA1AE6FFFD9FFEAFEAA25EE5F2BD4D5" box="[285,365,688,712]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="4" pageNumber="455">ZMUC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="B3AA6521FFD9FFEAFF2025865F92D517" pageId="4" pageNumber="455" type="etymology">
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<paragraph id="FB0F36AAFFD9FFEAFF2025865F92D517" blockId="4.[151,684,611,778]" pageId="4" pageNumber="455">
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ETYMOLOGY. — The name refers to the locality where the
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<typeStatus id="240B8808FFD9FFEAFF0B25AF5EA0D514" box="[188,230,753,777]" pageId="4" pageNumber="455">type</typeStatus>
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specimen was collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="B3AA6521FFD9FFEAFF2024745D09D3F8" pageId="4" pageNumber="455" type="description">
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<paragraph id="FB0F36AAFFD9FFEAFF2024745F6ED55F" blockId="4.[151,684,810,933]" box="[151,296,810,836]" pageId="4" pageNumber="455">DESCRIPTION</paragraph>
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<paragraph id="FB0F36AAFFD9FFEAFF2024145F6FD57E" blockId="4.[151,684,810,933]" box="[151,297,842,867]" pageId="4" pageNumber="455">
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<emphasis id="C9C4EAB8FFD9FFEAFF2024145F6FD57E" box="[151,297,842,867]" italics="true" pageId="4" pageNumber="455">Measurements</emphasis>
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</paragraph>
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<paragraph id="FB0F36AAFFD9FFEAFF2024355C58D5B9" blockId="4.[151,684,810,933]" pageId="4" pageNumber="455">
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Body length (without caudal pencil)
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<quantity id="3C489B4FFFD9FFEAFD8B24355CEED599" box="[572,680,874,901]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="455" unit="mm" value="2.4">2.40 mm</quantity>
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. Tarsus II length of 13
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<superScript id="0CC59BE2FFD9FFEAFE3424D85FD2D584" attach="left" box="[387,404,902,921]" fontSize="8" pageId="4" pageNumber="455">th</superScript>
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leg 193 µm.
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</paragraph>
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<paragraph id="FB0F36AAFFD9FFEAFF2024955E96D5F8" blockId="4.[151,684,971,1669]" box="[151,208,971,997]" pageId="4" pageNumber="455">
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<emphasis id="C9C4EAB8FFD9FFEAFF2024955E96D5F8" box="[151,208,971,997]" italics="true" pageId="4" pageNumber="455">Head</emphasis>
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</paragraph>
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<paragraph id="FB0F36AAFFD9FFEAFF2024B55F7DD218" blockId="4.[151,684,971,1669]" box="[151,315,1003,1029]" pageId="4" pageNumber="455">Anophthalmic.</paragraph>
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<paragraph id="FB0F36AAFFD9FFEAFF2023545F5AD2B8" blockId="4.[151,684,971,1669]" pageId="4" pageNumber="455">
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Vertex with one pair of posterior tufts consisting of two rows: anterior row of 15 and 16 trichomes, posterior row with six trichomes (
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<figureCitation id="638B2A2FFFD9FFEAFDE723155CD8D278" box="[592,670,1099,1125]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2J</figureCitation>
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); the distance between each tuft is greater than their length.
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</paragraph>
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<paragraph id="FB0F36AAFFD9FFEAFF2023F55C6ED098" blockId="4.[151,684,971,1669]" pageId="4" pageNumber="455">
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The proportions of the antennal articles are represented in
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<figureCitation id="638B2A2FFFD9FFEAFEB523955F11D2F8" box="[258,343,1227,1253]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2A</figureCitation>
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. Antennal article VI with three dorsal basiconica sensilla (
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<figureCitation id="638B2A2FFFD9FFEAFE0F23B55C57D318" box="[440,529,1259,1285]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2C</figureCitation>
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): two are side by side, the anterior one
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<emphasis id="C9C4EAB8FFD9FFEAFE1922545FF0D33E" box="[430,438,1290,1315]" italics="true" pageId="4" pageNumber="455">i</emphasis>
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||
is slightly thicker and longer than the posterior
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<emphasis id="C9C4EAB8FFD9FFEAFE1922745FFCD35E" box="[430,442,1322,1347]" italics="true" pageId="4" pageNumber="455">p</emphasis>
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||
; the third sensillum
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<emphasis id="C9C4EAB8FFD9FFEAFD2822745CEDD35E" box="[671,683,1322,1347]" italics="true" pageId="4" pageNumber="455">a</emphasis>
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is more basal, anterior, thinner and half as long as the other two sensilla, its apex reaching their bases. One sensillum coeloconicum
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<emphasis id="C9C4EAB8FFD9FFEAFD9122D45C69D3BE" box="[550,559,1418,1443]" italics="true" pageId="4" pageNumber="455">c</emphasis>
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is near the posterior sensillum; setiform sensillum absent. Antennal article VII with two dorsal basiconica sensilla: one anterior thin
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<emphasis id="C9C4EAB8FFD9FFEAFE0622B45FFBD01E" box="[433,445,1514,1539]" italics="true" pageId="4" pageNumber="455">a</emphasis>
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||
, one posterior thick
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<emphasis id="C9C4EAB8FFD9FFEAFD2822B45CEAD01E" box="[671,684,1514,1539]" italics="true" pageId="4" pageNumber="455">p</emphasis>
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and one apical sensillum coeloconicum
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<emphasis id="C9C4EAB8FFD9FFEAFDD421545C2AD03E" box="[611,620,1546,1571]" italics="true" pageId="4" pageNumber="455">c</emphasis>
|
||
(
|
||
<figureCitation id="638B2A2FFFD9FFEAFDC821555EFFD058" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2B</figureCitation>
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). Left antenna has three sensilla coeloconica
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<emphasis id="C9C4EAB8FFD9FFEAFD1521745CEDD05E" box="[674,683,1578,1603]" italics="true" pageId="4" pageNumber="455">c</emphasis>
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on the article VII (
|
||
<figureCitation id="638B2A2FFFD9FFEAFEDD21155F84D078" box="[362,450,1611,1637]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2D</figureCitation>
|
||
) and none on article VI because it is regenerated (
|
||
<figureCitation id="638B2A2FFFD9FFEAFE7F21355C5DD098" box="[456,539,1643,1669]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D27F55A72D6F8" blockId="4.[714,1247,170,517]" pageId="4" pageNumber="455">
|
||
Anterior trichobothria with the dilated apex ending in a short spine-hair (
|
||
<figureCitation id="638B2A2FFFD9FFEAFC6C27955A6ED6F8" box="[987,1064,203,229]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D27B55D27D7D8" blockId="4.[714,1247,170,517]" pageId="4" pageNumber="455">
|
||
Whole area of labrum well ornamented with numerous flat papillae, more or less arranged in groups and covered by small granules (
|
||
<figureCitation id="638B2A2FFFD9FFEAFBC026755A94D758" box="[1143,1234,299,325]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2N</figureCitation>
|
||
). Two rows of cuticular setae on posterior edge of labrum, second row interrupted by spaces. Clypeo-labrum with 11 long setae along the posterior margin.
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D26955D9AD418" blockId="4.[714,1247,170,517]" pageId="4" pageNumber="455">
|
||
Palpi of gnathochilarium with 39 (left) and 45 (right) sensilla (
|
||
<figureCitation id="638B2A2FFFD9FFEAFCC526B55D89D418" box="[882,975,491,517]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D25755D48D458" blockId="4.[714,1247,555,901]" box="[714,782,555,581]" pageId="4" pageNumber="455">
|
||
<emphasis id="C9C4EAB8FFD9FFEAFD7D25755D48D458" box="[714,782,555,581]" italics="true" pageId="4" pageNumber="455">Trunk</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D25155D4CD598" blockId="4.[714,1247,555,901]" pageId="4" pageNumber="455">
|
||
The trichomes of each tergite are grouped into two, separate, oval clusters with an additional posterior row, subdivided in its middle (
|
||
<figureCitation id="638B2A2FFFD9FFEAFBCB25D55A94D4B8" box="[1148,1234,651,677]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2K</figureCitation>
|
||
), except on collum; distance between both clusters much greater than length of each. Collum with left tuft of 55 and right tuft of 56 trichomes; each lateral protuberance of collum with six trichomes; each oval cluster of tergites II to IV with 31 to 37 trichomes and posterior row with 33 to 37.
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D24F45CB3D5DE" blockId="4.[714,1247,938,1285]" box="[714,757,938,963]" pageId="4" pageNumber="455">
|
||
<emphasis id="C9C4EAB8FFD9FFEAFD7D24F45CB3D5DE" box="[714,757,938,963]" italics="true" pageId="4" pageNumber="455">Legs</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D24955D61D2F8" blockId="4.[714,1247,938,1285]" pageId="4" pageNumber="455">
|
||
Each leg article, except for femora, tibia and tarsi, bearing pubescent cylindrical setae (
|
||
<figureCitation id="638B2A2FFFD9FFEAFBD524B55AE1D218" box="[1122,1191,1003,1029]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2</figureCitation>
|
||
G-I); coxae II to XII bearing two to five setae; I and XIII with one seta; trochanters and prefemoras I to XIII with one seta. Spine of tarsus II at least 1.5 times longer than telotarsus (
|
||
<figureCitation id="638B2A2FFFD9FFEAFBD823355A97D298" box="[1135,1233,1131,1157]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2F</figureCitation>
|
||
). Telotarsus with two dorsal, subequal, anterior and posterior denticles. Claw with ventral denticle.
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D23B55A19D319" blockId="4.[714,1247,938,1285]" box="[714,1119,1258,1285]" pageId="4" pageNumber="455">All areas of penis with cuticular setae.</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D22755D4BD358" blockId="4.[714,1247,1323,1509]" box="[714,781,1323,1349]" pageId="4" pageNumber="455">
|
||
<taxonomicName id="3CB04D29FFD9FFEAFD7D22755D4BD358" authorityName="Pearse" authorityYear="1952" box="[714,781,1323,1349]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9C4EAB8FFD9FFEAFD7D22755D4BD358" box="[714,781,1323,1349]" italics="true" pageId="4" pageNumber="455">Telson</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D22155D09D3F8" blockId="4.[714,1247,1323,1509]" pageId="4" pageNumber="455">
|
||
Nine trichomes
|
||
<emphasis id="C9C4EAB8FFD9FFEAFCCE22145DC3D37E" box="[889,901,1354,1379]" italics="true" pageId="4" pageNumber="455">a</emphasis>
|
||
, six of which placed on internal side (
|
||
<emphasis id="C9C4EAB8FFD9FFEAFCB422345D4DD39E" box="[771,779,1386,1411]" italics="true" pageId="4" pageNumber="455">i</emphasis>
|
||
) and three on external side (
|
||
<emphasis id="C9C4EAB8FFD9FFEAFBF722345A0CD39E" box="[1088,1098,1386,1411]" italics="true" pageId="4" pageNumber="455">e</emphasis>
|
||
) of the group of trichomes
|
||
<emphasis id="C9C4EAB8FFD9FFEAFCE022D55DE3D3B8" box="[855,933,1419,1445]" italics="true" pageId="4" pageNumber="455">b-c1-c3</emphasis>
|
||
(
|
||
<figureCitation id="638B2A2FFFD9FFEAFC0322D55A40D3B8" box="[948,1030,1419,1445]" captionStart="FIG" captionStartId="5.[151,162,1545,1562]" captionTargetBox="[151,1248,194,1489]" captionTargetId="figure-20@5.[155,1245,193,1683]" captionTargetPageId="5" captionText="FIG. 2. — Lophoturus monserratensis n. sp., holotype; A, right antenna; B, C, detail of sensilla on articles VII and VI of antenna A; D, E, sensilla on left antennal VII and VI articles; F, distal part of tarsus II and telotarsus of left leg I; G, seta of prefemur left leg I; H, I, coxal setae of left legs III and I; J, insertions of trichomes of left posterior tuft on vertex and detail of anterior left trichobothrium; K, L, insertions of trichomes on left half tergite II and anterior part of telson; M, right palpus of gnathochilarium; N, labrum. Abbreviations: a, anterior sensillum basiconicum; a1-a6, insertions of trichomes on internal side i; a7-a9, on external side e of the group b-c1-c3; c, sensillum coeloconicum; i, p, intermediate and posterior sensilla basiconica. Scale bars: A, J-L, 50 µm; B-F, G-I, M, N, 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5393021" httpUri="https://zenodo.org/record/5393021/files/figure.png" pageId="4" pageNumber="455">Fig. 2L</figureCitation>
|
||
). Most trichomes of caudal penicil lost; those remaining with one to three hooks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3AA6521FFD9FFE8FD7D21545E99D758" lastPageId="6" lastPageNumber="457" pageId="4" pageNumber="455" type="discussion">
|
||
<paragraph id="FB0F36AAFFD9FFEAFD7D21545D68D03F" blockId="4.[714,1247,1546,1669]" box="[714,814,1546,1572]" pageId="4" pageNumber="455">REMARKS</paragraph>
|
||
<paragraph id="FB0F36AAFFD9FFE8FD7D21755E99D758" blockId="4.[714,1247,1546,1669]" lastBlockId="6.[151,684,171,325]" lastPageId="6" lastPageNumber="457" pageId="4" pageNumber="455">
|
||
The position of the antennal sensilla of article VI is the same as in
|
||
<taxonomicName id="3CB04D29FFD9FFEAFC1521145D1DD098" authority="Conde & Terver, 1964" authorityName="Conde & Terver" authorityYear="1964" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="anisorhabdus">
|
||
<emphasis id="C9C4EAB8FFD9FFEAFC1521145A27D078" box="[930,1121,1610,1637]" italics="true" pageId="4" pageNumber="455">L. anisorhabdus</emphasis>
|
||
<bibRefCitation id="9F214B5BFFD9FFEAFBDB21155D1DD098" author="CONDE B. & TERVER D." pageId="4" pageNumber="455" pagination="1 - 21" refId="ref10832" refString="CONDE B. & TERVER D. 1964. - Penicillates du Surinam et du Guatemala (Diplopodes, Lophoproctidae). Studies on the Fauna of Surinam and Other Guyanas 7 (22): 1 - 21." type="journal article" year="1964">Condé & Terver, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
. The labral ornamentations are spe- cific and differ from those known in other species of
|
||
<taxonomicName id="3CB04D29FFDBFFE8FF0127955F6CD6F8" authorityName="Brolemann" authorityYear="1931" box="[182,298,203,229]" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="6" pageNumber="457" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9C4EAB8FFDBFFE8FF0127955F6CD6F8" box="[182,298,203,229]" italics="true" pageId="6" pageNumber="457">Lophoturus</emphasis>
|
||
</taxonomicName>
|
||
, such as
|
||
<taxonomicName id="3CB04D29FFDBFFE8FE2627945CE1D6F8" authority="(Loomis, 1934)" baseAuthorityName="Loomis" baseAuthorityYear="1934" box="[401,679,202,229]" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="niveus">
|
||
<emphasis id="C9C4EAB8FFDBFFE8FE2627945FB7D6FE" box="[401,497,202,227]" italics="true" pageId="6" pageNumber="457">L. niveus</emphasis>
|
||
(
|
||
<bibRefCitation id="9F214B5BFFDBFFE8FDB627955CD9D6F8" author="LOOMIS H. F." box="[513,671,203,229]" pageId="6" pageNumber="457" pagination="357 - 363" refId="ref10992" refString="LOOMIS H. F. 1934. - Three new Cuban millipeds, with notes on two little-known species. Bulletin of the Museum of Comparative Zoology. 75 (9): 357 - 363." type="journal article" year="1934">Loomis, 1934</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3CB04D29FFDBFFE8FF2027B45FAED718" authority="(Loomis, 1936)" baseAuthorityName="Loomis" baseAuthorityYear="1936" box="[151,488,234,261]" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="aequatus">
|
||
<emphasis id="C9C4EAB8FFDBFFE8FF2027B45F5BD71E" box="[151,285,234,259]" italics="true" pageId="6" pageNumber="457">L. aequatus</emphasis>
|
||
(Loomis, 1936)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3CB04D29FFDBFFE8FE4027B45F30D738" authority="(Chamberlin, 1955)" baseAuthorityName="Chamberlin" baseAuthorityYear="1955" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="quebradanus">
|
||
<emphasis id="C9C4EAB8FFDBFFE8FE4027B45CEAD718" box="[503,684,234,261]" italics="true" pageId="6" pageNumber="457">L. quebradanus</emphasis>
|
||
(Chamberlin, 1955)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3CB04D29FFDBFFE8FE0626545E9AD758" authority="(Broleman, 1931)" baseAuthorityName="Broleman" baseAuthorityYear="1931" class="Diplopoda" family="Lophoproctidae" genus="Lophoturus" kingdom="Animalia" order="Polyxenida" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="obscurus">
|
||
<emphasis id="C9C4EAB8FFDBFFE8FE0626545C61D738" box="[433,551,266,293]" italics="true" pageId="6" pageNumber="457">L. obscurus</emphasis>
|
||
(Broleman, 1931)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |