236 lines
25 KiB
XML
236 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.134.1707" ID-GBIF-Dataset="aa324fa8-f6d9-4bd1-aa15-3db7a315ed7c" ID-PMC="PMC3229209" ID-Pensoft-Pub="1313-2970-134-15" ID-PubMed="22140340" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-134-15" ModsDocOrigin="ZooKeys 134" ModsDocTitle="Two new Typhloreicheia species from Sardinia and their biogeographical significance (Coleoptera, Carabidae, Scaritinae)" checkinTime="1451249810322" checkinUser="pensoft" docAuthor="Casale, Achille & Marcia, Paolo" docDate="2011" docId="7312F73EAEBFB7880B54982EB17FF66E" docLanguage="en" docName="ZooKeys 134: 15-31" docOrigin="ZooKeys 134" docSource="http://dx.doi.org/10.3897/zookeys.134.1707" docTitle="Typhloreicheia ilianae Casale & Marcia, 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="21" masterDocId="CB1CA16F0E76FF96FF95FE5B7C1E9860" masterDocTitle="Two new Typhloreicheia species from Sardinia and their biogeographical significance (Coleoptera, Carabidae, Scaritinae)" masterLastPageNumber="31" masterPageNumber="15" pageNumber="18" updateTime="1668152335975" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new Typhloreicheia species from Sardinia and their biogeographical significance (Coleoptera, Carabidae, Scaritinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Casale, Achille</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marcia, Paolo</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>134</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>31</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.134.1707</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.134.1707</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-134-15</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152031308" LSID="urn:lsid:zoobank.org:act:B5A1C382-9E73-4F50-B15A-611522B1F9A7" httpUri="http://treatment.plazi.org/id/7312F73EAEBFB7880B54982EB17FF66E" lastPageId="7" lastPageNumber="21" pageId="3" pageNumber="18">
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<subSubSection pageId="3" pageNumber="18" type="nomenclature">
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<paragraph pageId="3" pageNumber="18">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:B5A1C382-9E73-4F50-B15A-611522B1F9A7" class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="18">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="18" type="type locality">
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<paragraph pageId="3" pageNumber="18">Type locality.</paragraph>
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<paragraph pageId="3" pageNumber="18">
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Italy, Central-Eastern Sardinia: Oliena (Nuoro province), Corrasi Mt.: Nurra 'e Pradu cave (speleological inventory number: 3083 Sa/NU), 1220 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="40.2575">40°15'27.0"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="9.429251">009°25'45.3"E</geoCoordinate>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="18" type="type material">
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<paragraph pageId="3" pageNumber="18">Type material.</paragraph>
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<paragraph pageId="3" pageNumber="18">Holotype male with the following data: I - Sardegna, Oliena (NU), M. Corrasi 26.I.2008 P. Marcia", "Gr. Nurra 'e Pradu" (CCa).</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="18" type="etymology">
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<paragraph pageId="3" pageNumber="18">Etymology.</paragraph>
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<paragraph pageId="3" pageNumber="18">The name derives from the Oliena village at the base of the Corrasi mountain in which this new species, and many other hypogean organisms were discovered. An ancient legend says that the inhabitants (Ilienses in Latin) of Troy (Ilios in ancient Greek), after the fall of the town to the Greeks, reached Sardinia and founded a village named Iliana, which subsequently became Oliena.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="18" type="diagnosis">
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<paragraph pageId="3" pageNumber="18">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="18">
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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is the largest in size among the Sardinian congeners known so far, with its TL: 3.64 and L: 4.05 (male holotype). It is evidently related to
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia henroti" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="henroti">Typhloreicheia henroti</taxonomicName>
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Jeannel, 1957 (see features of the median lobes of aedeagus in respective species: Figures 5-6), but can be distinguished from the latter mainly by its larger size (TL: 3.10 - 3.65, L: 3.40 - 3.86 in
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia henroti" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="henroti">Typhloreicheia henroti</taxonomicName>
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); by wider and more robust head, with supra-antennal plates more prominent laterally, by more convex genae markedly constricted to the neck, by antennomeres 3-11 more thickened, and by anterior angles of both clypeus and labrum more acutely prominent; and by the different shape of the median lobe of aedeagus (Figures 5-6).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="20" pageId="3" pageNumber="18" type="description">
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<paragraph pageId="3" pageNumber="18">Description.</paragraph>
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<paragraph pageId="3" pageNumber="18">
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A large sized
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">Typhloreicheia</taxonomicName>
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species (TL: 3.64; L: 4.05 mm, in male holotype).
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</paragraph>
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<paragraph pageId="4" pageNumber="19">
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<pageBreakToken pageId="4" pageNumber="19" start="start">Body</pageBreakToken>
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elongate but robust, convex (Figure 2). Colour testaceous reddish, antennae and mouthparts slightly paler.
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</paragraph>
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<paragraph pageId="4" pageNumber="19">Integument shiny, highly polished; microsculpure with fine, hardly visible microlines in form of isodiametric mesh pattern on head and elytra, almost vanished on pronotum.</paragraph>
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<paragraph pageId="4" pageNumber="19">Head robust, with ocular part of genae markedly convex, constricted toward the neck. Eyes absent. Anterior angles of clypeus acutely prominent. Supra-antennal plates prominent laterally, with outer margin beaded, separated from genae by deep and broad furrow; frontal furrows very deep, with shallow wrinkles in the posterior tract; vertex with an evident, convex tubercle in the middle; antennae elongate (AL: 1.58 mm in male holotype) but robust, thickened; antennomeres 6-10 slightly longer than wide.</paragraph>
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<paragraph pageId="4" pageNumber="19">Pronotum markedly convex, relatively wide (PL/PW: 0.95), with its maximum width at the basal third; sides moderately rounded, slightly narrowed in front, markedly constricted to the basal peduncle; anterior angles acutely prominent; median furrow very shallow; lateral furrows very narrow and superficial.</paragraph>
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<paragraph lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
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Elytraelongate-ovate (EL/EW: 1.62), distinctly wider than pronotum (EW/PW: 1.24), convex, with their maximum width in the middle; humeri broadly rounded;
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<pageBreakToken pageId="5" pageNumber="20" start="start">lateral</pageBreakToken>
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furrows wide, flattened, not narrowed at apex; lateral margin reflexed, elytra with lateral margins with numerous (24) prominent marginal teeth; striae deep, deeply punctuate, all evident, gradually disappearing et apex; elytral intervals convex, intervals 2-7 each bearing a series of short, erected setae; umbilicate series of 16-19 punctures along stria 8.
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</paragraph>
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<paragraph pageId="5" pageNumber="20">Male genitalia as in Figure 5. Median lobe of aedeagus markedly curved, with long apical lamina, which is widened, hatched-like distally. Endophallus with a reduced, inconspicuous apical copulatory piece and an elongate packet of serrate scales in the middle. Parameres each with two apical setae.</paragraph>
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<paragraph pageId="5" pageNumber="20">Female genitalia: unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="20" type="distribution and habitat">
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<paragraph pageId="5" pageNumber="20">Distribution and habitat.</paragraph>
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<paragraph pageId="5" pageNumber="20">
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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sp. n. is known only from the type locality. The holotype was sampled in wet soil under big stones in a small pit (- 3.7 m), which represents the entrance of a large hypogean system, reaching the depth of 101 m. The associated subterranean fauna includes some of the most specialised troglomorphic endemic Sardinian elements, such as
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<taxonomicName class="Insecta" family="Carabidae" genus="Sardaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sardaphaenops supramontanus subsp. supramontanus" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="supramontanus" subSpecies="supramontanus">Sardaphaenops supramontanus supramontanus</taxonomicName>
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Cerruti & Henrot, 1956 (Coleoptera, Carabidae, Trechini) and
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<taxonomicName class="Insecta" family="Leiodidae" genus="Patriziella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patriziella sardoa" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="sardoa">Patriziella sardoa</taxonomicName>
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Jeannel, 1956 (Coleoptera, Cholevidae, Leptodirini), both described from a nearby cave Nurra 'e Sas Palumbas. The Nurra 'e Pradu cave is also one of the few localities of Sardinia from which the large sized, trogloxenic centipede
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<taxonomicName class="Chilopoda" family="Cryptopidae" genus="Plutonium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plutonium zwierleini" order="Scolopendromorpha" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="zwierleini">Plutonium zwierleini</taxonomicName>
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Cavanna, 1881 is reported (
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<bibRefCitation pageId="5" pageNumber="20">Zapparoli 2009</bibRefCitation>
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). We sampled in this cave the following taxa, all endemic to central-eastern Sardinia and representing new faunal records: the orthopteran
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<taxonomicName class="Insecta" family="Gryllidae" genus="Acroneuroptila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acroneuroptila" order="Orthoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus">Acroneuroptila</taxonomicName>
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cf. sardoa Baccetti, 1960, the sphodrine carabid beetle
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<taxonomicName class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laemostenus pippiai" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="pippiai">Laemostenus pippiai</taxonomicName>
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(G. Fiori, 1961) (A. Casale det.), the cholevid beetle
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<taxonomicName class="Insecta" family="Leiodidae" genus="Ovobathysciola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ovobathysciola majori" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="majori">Ovobathysciola majori</taxonomicName>
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(Reitter, 1885) (A. Casale det.), and the terrestrial snail
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<taxonomicName class="Gastropoda" family="Helicidae" genus="Tacheocampylaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tacheocampylaea carotii" order="Stylommatophora" pageId="5" pageNumber="20" phylum="Mollusca" rank="species" species="carotii">Tacheocampylaea carotii</taxonomicName>
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(Paulucci, 1882) (S. Birindelli det.). Subsequent trapping in this cave using pitfall traps, did not produce additional individuals of
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="22" pageId="5" pageNumber="20" type="relationships">
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<paragraph pageId="5" pageNumber="20">Relationships.</paragraph>
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<paragraph pageId="5" pageNumber="20">
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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is the largest among the Sardinian species known so far, exceeding the large size of
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia kraussei" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="kraussei">Typhloreicheia kraussei</taxonomicName>
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(Reitter, 1914), a typical endogean, not hypogean species reaching 3.69 mm in length (
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<bibRefCitation author="Leo, P" journalOrPublisher="Bollettino della Societa entomologica italiana" pageId="11" pageNumber="26" pagination="167 - 203" title="Materiali per lo studio delle Typhloreicheia della Sardegna con descrizione di nove specie nuove (Coleoptera Carabidae)." volume="137" year="2005">Leo et al. 2005</bibRefCitation>
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). Nevertheless, any relationship of
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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sp. n. with
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia kraussei" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="kraussei">Typhloreicheia kraussei</taxonomicName>
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(Reitter, 1914) and its adelphotaxon
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia manto" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="manto">Typhloreicheia manto</taxonomicName>
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(Holdhaus, 1924), both typically endogean, not hypogean species, are highly unlikely. Both latter species have elytral setiferous pores present in intervals 2, 3, 5 and 7 only, and the lateral margins serrate only in the basal half. Furthermore, in these taxa the features of the median lobe of aedeagus are markedly dissimilar to those of
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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sp. n., and the apical copulatory piece in endophallus is lacking in
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia kraussei" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="kraussei">Typhloreicheia kraussei</taxonomicName>
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(
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<bibRefCitation author="Leo, P" journalOrPublisher="Bollettino della Societa entomologica italiana" pageId="11" pageNumber="26" pagination="167 - 203" title="Materiali per lo studio delle Typhloreicheia della Sardegna con descrizione di nove specie nuove (Coleoptera Carabidae)." volume="137" year="2005">Leo et al. 2005</bibRefCitation>
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).
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</paragraph>
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<paragraph lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="20">
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The new taxon appears related to
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia henroti" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="henroti">Typhloreicheia henroti</taxonomicName>
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Jeannel, 1957, known from the Gurennoro cave (= Pisanu cave, 215 Sa/NU, near Dorgali). Both species are similar in having large-sized body and elytral lateral margins bearing numerous (21-24) teeth extending all the way from the humeral angle to elytral apex. Also both species share the peculiar shape of the median lobe of aedeagus, which is unique among the Sardinian species (Figures 5-6). These species form a pair of adelphotaxa very isolated from the rest of species known so far on the island, here indicated as henroti species group, excluding
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<pageBreakToken pageId="6" pageNumber="21" start="start">the</pageBreakToken>
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other specialised hypogean species of Sardinia known so far (
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia elegans" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="elegans">Typhloreicheia elegans</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia onnisi" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="onnisi">Typhloreicheia onnisi</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia monacha" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="monacha">Typhloreicheia monacha</taxonomicName>
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sp. n.) treated above aselegans species group in the narrow sense.
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</paragraph>
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<caption pageId="6" pageNumber="21">
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<paragraph pageId="6" pageNumber="21">
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Figures 3-6.
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus">Typhloreicheia</taxonomicName>
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spp., male genitalia, right lateral aspect: 3
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia monacha" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="monacha">Typhloreicheia monacha</taxonomicName>
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sp. n., median lobe f aedeagus and parameres 4
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia onnisi" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="onnisi">Typhloreicheia onnisi</taxonomicName>
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Casale & Magrini, median lobe of aedeagus 5
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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sp. n., median lobe of aedeagus and parameres 6
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia henroti" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="henroti">Typhloreicheia henroti</taxonomicName>
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Jeannel, median lobe of aedeagus.
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</paragraph>
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</caption>
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<paragraph pageId="6" pageNumber="21">
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The following operative and provisional key is provided to distinguish the cave dweller (deep hypogean, or troglophilic)
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus">Typhloreicheia</taxonomicName>
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species known so far in Central Eastern Sardinia:
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</paragraph>
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<paragraph lastPageId="7" lastPageNumber="22" pageId="6" pageNumber="21">
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<table lastPageId="7" lastPageNumber="22" pageId="6" pageNumber="21">
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<tr pageId="6" pageNumber="21">
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<td colspan="1" pageId="6" pageNumber="21" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia henroti" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="henroti">Typhloreicheia henroti</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="21">
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<td colspan="1" pageId="6" pageNumber="21" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia elegans" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="elegans">Typhloreicheia elegans</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="21">
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<td colspan="1" pageId="6" pageNumber="21" rowspan="1">Figure 5</td>
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<td colspan="1" pageId="6" pageNumber="21" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia ilianae" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="ilianae">Typhloreicheia ilianae</taxonomicName>
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</td>
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</tr>
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<tr pageId="6" pageNumber="21">
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<td colspan="1" pageId="6" pageNumber="21" rowspan="1">Figure 6</td>
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<td colspan="1" pageId="6" pageNumber="21" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia henroti" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="henroti">Typhloreicheia henroti</taxonomicName>
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</td>
|
|
</tr>
|
|
<tr pageId="6" pageNumber="21">
|
|
<td colspan="1" pageId="6" pageNumber="21" rowspan="1">
|
|
<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia elegans" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="elegans">Typhloreicheia elegans</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr pageId="6" pageNumber="21">
|
|
<td colspan="1" pageId="6" pageNumber="21" rowspan="1">Figure 3</td>
|
|
<td colspan="1" pageId="6" pageNumber="21" rowspan="1">
|
|
<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia monacha" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="monacha">Typhloreicheia monacha</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr pageId="6" pageNumber="21">
|
|
<td colspan="1" pageId="6" pageNumber="21" rowspan="1">Figure 4</td>
|
|
<td colspan="1" pageId="6" pageNumber="21" rowspan="1">
|
|
<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia onnisi" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="onnisi">Typhloreicheia onnisi</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr pageId="7" pageNumber="22">
|
|
<td colspan="1" pageId="7" pageNumber="22" rowspan="1">
|
|
<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia jana" order="Coleoptera" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="jana">
|
|
<pageBreakToken pageId="7" pageNumber="22" start="start">Typhloreicheia</pageBreakToken>
|
|
jana
|
|
</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
<tr pageId="7" pageNumber="22">
|
|
<td colspan="1" pageId="7" pageNumber="22" rowspan="1">
|
|
<taxonomicName class="Insecta" family="Carabidae" genus="Typhloreicheia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typhloreicheia pandora" order="Coleoptera" pageId="7" pageNumber="22" phylum="Arthropoda" rank="species" species="pandora">Typhloreicheia pandora</taxonomicName>
|
|
</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
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</treatment>
|
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</document> |