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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e83035" ID-Pensoft-Pub="1864-8312-80-439" ID-Pensoft-UUID="3F4E21B524465471831D09738C2304A6" ID-ZooBank="62FFB94CEBF441639F22881435EFC37C" ModsDocID="1864-8312-80-439" checkinTime="1663021027595" checkinUser="pensoft" docAuthor="Santos, Daubian, Santos, Rodrigo dos Reis &amp; Ribeiro, Guilherme Cunha" docDate="2022" docId="72C07D55E408597BB1B4FA94365ECF99" docLanguage="en" docName="ArthropodSystPhyl 80: 439-494" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-09-12" docSource="http://dx.doi.org/10.3897/asp.80.e83035" docTitle="Amphineurus (Rhamphoneurus) triangularis Santos &amp; Santos &amp; Ribeiro 2022, sp. nov." docType="treatment" docUuid="F3B376D0-C8CE-4145-AD33-A1BA5DDF5AAB" docUuidSource="ZooBank" docVersion="1" id="3F4E21B524465471831D09738C2304A6" lastPageNumber="439" masterDocId="3F4E21B524465471831D09738C2304A6" masterDocTitle="Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="494" masterPageNumber="439" pageNumber="439" updateTime="1663021027595" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Daubian</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1220-1267</mods:nameIdentifier>
<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">daubians@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>439</mods:start>
<mods:end>494</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e83035</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e83035</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-439</mods:identifier>
<mods:identifier type="ZooBank">62FFB94CEBF441639F22881435EFC37C</mods:identifier>
<mods:identifier type="Pensoft-UUID">3F4E21B524465471831D09738C2304A6</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:F3B376D0-C8CE-4145-AD33-A1BA5DDF5AAB" httpUri="http://treatment.plazi.org/id/72C07D55E408597BB1B4FA94365ECF99" lastPageNumber="439" pageId="0" pageNumber="439">
<subSubSection pageId="0" pageNumber="439" type="nomenclature">
<paragraph pageId="0" pageNumber="439">
3.3.26.
<taxonomicName LSID="http://zoobank.org/F3B376D0-C8CE-4145-AD33-A1BA5DDF5AAB" authority="Santos &amp; Santos &amp; Ribeiro, 2022" authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) triangularis" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov." subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) triangularis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="439">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="materials_examined">
<paragraph pageId="0" pageNumber="439">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="439">
<emphasis bold="true" pageId="0" pageNumber="439">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<specimenCount></specimenCount>
, Chile,
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is., Chepu [
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2S</geoCoordinate>
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1W</geoCoordinate>
],
<date value="1952-02-08">8-II-1952</date>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM)*.
<emphasis bold="true" pageId="0" pageNumber="439">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
<specimenCount type="female">1 ♀</specimenCount>
, Chile,
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is., Dalcahue [
<geoCoordinate degrees="42" direction="south" minutes="22" orientation="latitude" precision="925" value="-42.366665">42°22S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="39" orientation="longitude" precision="925" value="-73.65">73°39W</geoCoordinate>
], 1/
<date value="1968-04-04">4-IV-1968</date>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM);
<specimenCount type="male">1 ♂</specimenCount>
, Chile,
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is., Chepu [
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2S</geoCoordinate>
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1W</geoCoordinate>
], 10/
<date value="1952-02-14">14-II-1952</date>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM)*. -
<emphasis bold="true" pageId="0" pageNumber="439">Additional material</emphasis>
:
<materialsCitation collectingDate="1955-03-10" collectorName="Pena" country="CHILE" elevation="60" latitude="-40.533333" location="Pucotrihue" longLatPrecision="1162" longitude="-73.7" specimenCount="2" specimenCount-male="2">
<emphasis bold="true" pageId="0" pageNumber="439">
<collectingCountry name="Chile">CHILE</collectingCountry>
.
</emphasis>
<specimenCount type="male">2 ♂</specimenCount>
and 1 [sex unknown],
<location LSID="urn:lsid:plazi:treatment:72C07D55E408597BB1B4FA94365ECF99:1CAA84D5686B6ED2129541D8B135F03D" country="CHILE" latitude="-40.533333" longLatPrecision="1162" longitude="-73.7" name="Pucotrihue">Pucotrihue</location>
[
<geoCoordinate degrees="40" direction="south" minutes="32" orientation="latitude" precision="925" value="-40.533333">40°32S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="42" orientation="longitude" precision="925" value="-73.7">73°42W</geoCoordinate>
],
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</elevation>
</quantity>
,
<collectingDate value="1955-03-10">10-III-1955</collectingDate>
,
<normalizedToken originalValue="Peña">
<collectorName>Pena</collectorName>
</normalizedToken>
(USNM)
</materialsCitation>
;
<specimenCount type="male">1 ♂</specimenCount>
,
<specimenCount type="female">1 ♀</specimenCount>
and 1 [sex unknown], Llanquihue, Carelmapu [
<geoCoordinate degrees="41" direction="south" minutes="44" orientation="latitude" precision="925" value="-41.733334">41°44S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="40" orientation="longitude" precision="925" value="-73.666664">73°40W</geoCoordinate>
], Canal Chacao,
<date value="1957-03-18">18-III-1957</date>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM); 1 [sex unknown],
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
Is., Chepu [
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2S</geoCoordinate>
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1W</geoCoordinate>
],
<date value="1952-02-08">8-II-1952</date>
,
<normalizedToken originalValue="Peña">Pena</normalizedToken>
(USNM).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="etymology">
<paragraph pageId="0" pageNumber="439">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="439">
The word
<taxonomicName authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus triangularis" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="triangularis">
<emphasis italics="true" pageId="0" pageNumber="439">Amphineurus triangularis</emphasis>
</taxonomicName>
is Latin for
<normalizedToken originalValue="“triangular”">&quot;triangular&quot;</normalizedToken>
. This name references the triangular shape of the spur of sheath of aedeagus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="diagnosis">
<paragraph pageId="0" pageNumber="439">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="439">This species is distinguished by only a dark spot near R2 and M1+2 as long as the basal deflection of M1. Furthermore, this species is characterized by male terminalia with sheath of aedeagus with a triangular spur, tip of both mesal lobes bent horizontally, straightened lateral branch of clasper of gonostylus and cercus uniformly wide and abruptly angled dorsad.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="description">
<paragraph pageId="0" pageNumber="439">Description.</paragraph>
<paragraph pageId="0" pageNumber="439">
Wing length 5.76 mm, width 1.74 mm. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Coloration</emphasis>
: General coloration brownish yellow. Rostrum and palpus dark brown. First segment of antenna yellowish brown, remainding segments brown. Head brown. Eye black. Thorax brownish yellow, pleura pale. Scutum with one black stripe. Halter dark yellow with knob yellowish brown. Coxae yellow and legs brown. Abdomen yellowish brown. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Head</emphasis>
(dorsal view Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Amphineurus (Rhamphoneurus) triangularis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439">31B</figureCitation>
, ventral view Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Amphineurus (Rhamphoneurus) triangularis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439">31A</figureCitation>
): Rostrum bifurcated with long appendices; first palpal segment shorter than terminal segment; scape thinner than pedicel; terminal flagellomere longer than penultimate. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Thorax</emphasis>
(lateral view Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Amphineurus (Rhamphoneurus) triangularis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439">31E</figureCitation>
, dorsal view Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Amphineurus (Rhamphoneurus) triangularis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439">31F</figureCitation>
): Division line of laterotergite almost vertical, anatergite as tall as katatergite. Halter with dilated knob. Wing (Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Amphineurus (Rhamphoneurus) triangularis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439">31D</figureCitation>
) almost clear of markings, except marking on R2; veins near fork of bM (Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Amphineurus (Rhamphoneurus) triangularis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439">31C</figureCitation>
) faded; M1+2 as long as basal deflection of M1; R2+3+4 longer than R2+3; basal deflection of R5 shorter than R2+3. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Female terminalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Amphineurus (Rhamphoneurus) triangularis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439">31H</figureCitation>
): Female tergite IX shorter, wider than tergite X; cercus bent at right angle. Hypogynial valve slender and longer than cercus. -
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Male terminalia</emphasis>
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Amphineurus (Rhamphoneurus) triangularis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439">31G</figureCitation>
): Male tergite IX slightly darkened along posterior margin, with wide V-shaped median notch. Ventral branch of gonocoxite slightly sinuous, with concentration of setae on distal tip. Dorsal branch of gonocoxite shorter than half-length of ventral branch, slightly inclined medially. Lobe of gonostylus with rounded lobule; lobule longer than wide, longer than half-length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved. Mesal lobes of gonocoxite asymmetrical; left mesal lobe curved; right mesal lobe short with sideways projection. Sheath of aedeagus straight with triangular projection, darkened near tip.
</paragraph>
<caption doi="10.3897/asp.80.e83035.figure31" httpUri="https://binary.pensoft.net/fig/740584" pageId="0" pageNumber="439" start="Figure 31" startId="F31">
<paragraph pageId="0" pageNumber="439">
<emphasis bold="true" pageId="0" pageNumber="439">Figure 31.</emphasis>
<taxonomicName authorityName="Santos &amp; Santos &amp; Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) triangularis" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="triangularis" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) triangularis</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
head (ventral view);
<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
head (dorsal view);
<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
detail of wing;
<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
wing;
<emphasis bold="true" pageId="0" pageNumber="439">E</emphasis>
thorax (lateral view);
<emphasis bold="true" pageId="0" pageNumber="439">F</emphasis>
thorax (dorsal view);
<emphasis bold="true" pageId="0" pageNumber="439">G</emphasis>
male tergite IX (dorsal view) and remainder of male terminalia (lateral view);
<emphasis bold="true" pageId="0" pageNumber="439">H</emphasis>
female terminalia (dorsal view). - Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
, sheath of aedeagus;
<emphasis bold="true" pageId="0" pageNumber="439">anatg</emphasis>
, anatergite;
<emphasis bold="true" pageId="0" pageNumber="439">anepm</emphasis>
, anepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">anepst</emphasis>
, anepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">aprn</emphasis>
, anteropronotum;
<emphasis bold="true" pageId="0" pageNumber="439">cer</emphasis>
, cercus;
<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
, clasper of gonostylus;
<emphasis bold="true" pageId="0" pageNumber="439">cx</emphasis>
, coxa;
<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
, gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">hlt</emphasis>
, halter;
<emphasis bold="true" pageId="0" pageNumber="439">hyp val</emphasis>
, hypogynial valve;
<emphasis bold="true" pageId="0" pageNumber="439">k</emphasis>
, knob of halter;
<emphasis bold="true" pageId="0" pageNumber="439">kepm</emphasis>
, katepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">kepst</emphasis>
, katepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">ktg</emphasis>
, katatergite;
<emphasis bold="true" pageId="0" pageNumber="439">l ms</emphasis>
, left mesal lobe of gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
; lobe of gonostylus;
<emphasis bold="true" pageId="0" pageNumber="439">mr</emphasis>
, meron;
<emphasis bold="true" pageId="0" pageNumber="439">mtanepst</emphasis>
, metanepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">mtepm</emphasis>
, metepimeron;
<emphasis bold="true" pageId="0" pageNumber="439">mtg</emphasis>
, mediotergite;
<emphasis bold="true" pageId="0" pageNumber="439">mtkepst</emphasis>
, metakatepisternum;
<emphasis bold="true" pageId="0" pageNumber="439">mtn</emphasis>
, metanotum;
<emphasis bold="true" pageId="0" pageNumber="439">p</emphasis>
, posterior basalare;
<emphasis bold="true" pageId="0" pageNumber="439">patg</emphasis>
, paratergite;
<emphasis bold="true" pageId="0" pageNumber="439">pprn</emphasis>
, postpronotum;
<emphasis bold="true" pageId="0" pageNumber="439">presct</emphasis>
, prescutum;
<emphasis bold="true" pageId="0" pageNumber="439">r ms</emphasis>
, right mesal lobe of gonocoxite;
<emphasis bold="true" pageId="0" pageNumber="439">sct</emphasis>
, scutum;
<emphasis bold="true" pageId="0" pageNumber="439">sctl</emphasis>
, scutellum;
<emphasis bold="true" pageId="0" pageNumber="439">st</emphasis>
, sternite;
<emphasis bold="true" pageId="0" pageNumber="439">tg</emphasis>
, tergite;
<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
, male tergite IX.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="439" type="remarks">
<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="439">
Some specimens of this new species were previously identified by C.P. Alexander as
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) insanus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="insanus" subGenus="R.">A. (R.) insanus</taxonomicName>
Alexander and some as
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) glabristylatus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="glabristylatus" subGenus="R.">A. (R.) glabristylatus</taxonomicName>
Alexander. This species resembles
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) podenasi" order="Diptera" pageId="0" pageNumber="439" rank="species" species="podenasi" subGenus="R.">A. (R.) podenasi</taxonomicName>
sp. nov. but differs mainly in the length of M1+2 and R2+3+4, shape of cercus and ventral branch of gonocoxite.
</paragraph>
</subSubSection>
</treatment>
</document>