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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62593" ID-GBIF-Dataset="2b58bee7-f25a-42c9-8651-3060ada2112e" ID-PMC="PMC8222207" ID-Pensoft-Pub="1313-2970-1044-563" ID-Pensoft-UUID="8787843B77CB51DEB6FCF0B8F3A09AC3" ID-PubMed="34183886" ID-ZooBank="71DCAFC2B2BB475B91662BCDC83C7F07" ModsDocID="1313-2970-1044-563" checkinTime="1623875370007" checkinUser="pensoft" docAuthor="Neri, Paolo &amp; Toledano, Luca" docDate="2021" docId="72A96E572D165EF9862C6B110F5D5552" docLanguage="en" docName="ZooKeys 1044: 563-587" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62593" docTitle="Bembidion (Plataphus) pseudolucillum Netolitzky 1938, comb. nov." docType="treatment" docVersion="5" id="8787843B77CB51DEB6FCF0B8F3A09AC3" lastPageNumber="563" masterDocId="8787843B77CB51DEB6FCF0B8F3A09AC3" masterDocTitle="Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 ' Catalogue of Palaearctic Coleoptera' (Coleoptera, Carabidae, Bembidiina)" masterLastPageNumber="587" masterPageNumber="563" pageNumber="563" updateTime="1732487846666" updateUser="ExternalLinkService">
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<mods:title id="357502EDF9F9DEA66BDDBD9385F53E6E">Geographic and taxonomic notes, addenda and corrigenda on the subtribe Bembidiina Stephens, 1827 of the 2017 ' Catalogue of Palaearctic Coleoptera' (Coleoptera, Carabidae, Bembidiina)</mods:title>
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<mods:namePart id="6E7638860976C87C04070EA6DDA77E0B">Neri, Paolo</mods:namePart>
<mods:affiliation id="C2B1EA19EC2DF99D9AE89D826A89B2CC">Via Guido Rossa 21 &quot; San Lorenzo &quot;, 47121 Forli (FC), Italy</mods:affiliation>
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<mods:namePart id="4E853DD13D36C356B5158941428D77D6">Toledano, Luca</mods:namePart>
<mods:affiliation id="6A964AB204DF4EB584ACC6A77419782F">Museo Civico di Storia Naturale di Verona, Lungadige Porta Vittoria 9, 37129 Verona, Italy</mods:affiliation>
<mods:nameIdentifier id="5BD749F079A177278F2EF6245C48F85F" type="email">lucatole2@libero.it</mods:nameIdentifier>
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<treatment id="72A96E572D165EF9862C6B110F5D5552" ID-GBIF-Taxon="182753947" LSID="urn:lsid:plazi:treatment:72A96E572D165EF9862C6B110F5D5552" httpUri="http://treatment.plazi.org/id/72A96E572D165EF9862C6B110F5D5552" lastPageNumber="563" pageId="0" pageNumber="563" scope_family="Carabidae" scope_order="Coleoptera">
<subSubSection id="80E415C8A8BC7F45DECDB857D6398872" pageId="0" pageNumber="563" type="nomenclature">
<paragraph id="345781505921D54438D9E112087C9893" pageId="0" pageNumber="563">
<taxonomicName id="F0F5FE397ED1F9BA4EB3765344AEE019" ID-CoL="8MLYV" LSID="72A96E57-2D16-5EF9-862C-6B110F5D5552" authority="Netolitzky, 1938" authorityName="Netolitzky" authorityYear="1938" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="1938" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Plataphus) pseudolucillum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="pseudolucillum" status="comb. nov." subGenus="Plataphus">Bembidion (Plataphus) pseudolucillum Netolitzky, 1938</taxonomicName>
<taxonomicNameLabel id="86FAA8CC25ACF6F26DDCE642BAAE36DA" pageId="0" pageNumber="563">comb. nov.</taxonomicNameLabel>
<figureCitation id="B07ECCDDB24620326A47F8E17B32D3AC" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 5.60 mm 2 B. (Plataphus) pseudolucillum Netolitzky, 1938 from Kibune, Kyoto (NHMW), 4.20 mm 3 B. (Peryphus) psuchrum Andrewes, 1935 Inde, Himachal Pradesh, Mahari, 3500 m (CTVR), 3.50 mm 4 labels of the holotype of Bembidion sapporense Jedlicka (NMPC). Photographs: Luca Toledano." figureDoi="10.3897/zookeys.1044.62593.figures1-4" httpUri="https://binary.pensoft.net/fig/555046" pageId="0" pageNumber="563">Figures 2</figureCitation>
<figureCitation id="ABF8A060ADF64870E080AEE470F239EC" captionStart="Figures 59" captionStartId="F2" captionText="Figures 5 - 9. Median lobe of the aedeagi of 5 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 1.55 mm 6 B. (P.) dostali Kirschenhofer, 1984, paratype (NHMW), 1.53 mm 7 B. (P.) lulinense Habu, 1973 (drawing from Habu 1973), 1.41 mm 8 Bembidion (Peryphus) psuchrum Andrewes, 1935, holotype (NHMUK), 0.91 mm 9 Bembidion (Plataphus) pseudolucillum Netolitzky, paratype (NHMW), 0.80 mm." figureDoi="10.3897/zookeys.1044.62593.figures5-9" httpUri="https://binary.pensoft.net/fig/555047" pageId="0" pageNumber="563">, 9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5A6132EE3855E8097499A3389EEB5C81" pageId="0" pageNumber="563" type="reference_group">
<paragraph id="C7987BD7DA19D244502EBCA90FB457A3" pageId="0" pageNumber="563">
<taxonomicName id="EA570C6A3F6FF470AE66A38B703799A1" authorityName="Neri &amp; Toledano" authorityYear="2021" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="1938" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) pseudolucillum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="pseudolucillum" status="comb. nov." subGenus="Peryphus">Bembidion (Peryphus) pseudolucillum</taxonomicName>
Netolitzky, 1938
</paragraph>
</subSubSection>
<subSubSection id="F0AEB766D3F6DB702BD1ED3A8A7C6B66" pageId="0" pageNumber="563" type="materials_examined">
<paragraph id="27EA6CF89B39A3249367436CEC7847B5" pageId="0" pageNumber="563">Material examined.</paragraph>
<paragraph id="2713BB98FB548FD45EFEAF7965A2AAAA" pageId="0" pageNumber="563">
<materialsCitation id="48693D4F310C1AF76EAD89584BD0025F" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectingMethod="pseudolucillum Net" collectorName="J. E. A. Lewis, H. E. Andrewes" country="Japan" location="Japan" specimenCount="1" specimenCount-female="1">
<specimenCount id="AD52E598ACCE462D77B39DE26F70F7DF" type="female">1 ♀</specimenCount>
, &quot;
<collectingCountry id="8CEA990875938FCC7CDB29AF7C807803" name="Japan">Japan</collectingCountry>
/ Settsu / Katsuoji / 16.VI.24 /
<collectorName id="13BDBCAB95C2C14D570AD78A5ACCD040">J.E.A. Lewis</collectorName>
[printed] // 199 [handwritten] // Type [red, printed] //
<collectingMethod id="D27379EB0AA51737D3B45C7933CBD856">
<emphasis id="146CB34726A80AC7BE866B0B84AF7BB2" italics="true" pageId="0" pageNumber="563">pseudolucillum</emphasis>
Net
</collectingMethod>
. / Type Netolitzky 37 //
<emphasis id="DBAD07A2AAC7FDF2AE3FAF3E5F0A3B4A" italics="true" pageId="0" pageNumber="563">Peryphus</emphasis>
//
<collectorName id="74EF534FBE0513D52CB40EB81B9B6E23">H.E. Andrewes</collectorName>
coll. / B.M. 1945-97 [printed]&quot; (NHMUK)
</materialsCitation>
.
<materialsCitation id="436B36DD63B95538E0FD5BDE14712051" collectingDate="1937-09-08" collectorName="H. E. Andrewes" country="Japan" determinerName="Netolitzky" specimenCount="3" specimenCount-female="3">
<specimenCount id="29BC765FC1CC58BD0AB240B70A307503" type="female">3 ♀♀</specimenCount>
, on the same pin, &quot;Oku Nikko / Nabagawa /
<collectingDate id="CD079B59C18B2C31510EB20D2E3BA169" value="1937-09-08">8.9.1937</collectingDate>
/ Jano, Jap. [handwritten] //
<emphasis id="B32FBF9CE1BBA5A0D1CD028239BFFE5F" italics="true" pageId="0" pageNumber="563">pseudolucillum</emphasis>
/ det.
<determinerName id="9FE89AC8DFBB4EACC13FF1D5B211B028">Netolitzky</determinerName>
[handwritten] //
<collectorName id="C93CA17569AE8234499C77B105884C9E">H.E. Andrewes</collectorName>
coll. / B.M. 1945-97 [printed]&quot; (NHMUK)
</materialsCitation>
.
<materialsCitation id="3C3DD8B635D00B5E516DF824D57410C5" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectingMethod="identisch Net, pseudolucillum Net" collectorName="J. E. A. Lewis" country="Japan" location="Japan" specimenCount="1" specimenCount-female="1">
<specimenCount id="BFA2492C784C1552FAFA868D4101267A" type="female">1 ♀</specimenCount>
, &quot;
<collectingCountry id="1793858F4075052A7D56C29C1BDBC3FB" name="Japan">Japan</collectingCountry>
/ Settsu / Katsuoji /
<collectorName id="D8925D0B330B4CA05BD2C784A42A031B">J.E.A. Lewis</collectorName>
[handwritten] // [one deleted label] //
<emphasis id="683D492F7EE9CD2ADA1F762ACEEADAA2" italics="true" pageId="0" pageNumber="563">Peryphus</emphasis>
sp. / striis mediocr- / punctat [illegible] / dt Netolitzky [handwritten] // mit Type / Brit. Museum /
<collectingMethod id="E320622E0446066C8BDFA74C949570A4">identisch Net</collectingMethod>
. [handwritten] // Coll. / Netolitzky [printed] //
<collectingMethod id="8F80519E1F22E4B6E51A45783F0AD259">
<emphasis id="DC83DFCEC66AF23E3E62962F99994BC4" italics="true" pageId="0" pageNumber="563">pseudolucillum</emphasis>
Net
</collectingMethod>
/ Cotypus Netolitzky 37 [handwritten] // CO / TYPUS [red, printed] // Coll. / Netolitzky [printed]&quot; (NHMW)
</materialsCitation>
.
<materialsCitation id="25DD91FBC21AF81D1C0CB14BF5FACB9A" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Netolitzky" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9DF09F7DED9D81FA9EC0ACD8CD02E4A3" type="male">1 ♂</specimenCount>
, &quot;Oku Nikko / Nabagawa / 8.9.37 Jano [handwritten] //
<emphasis id="9CF375C545DC0D4A324EE2E21E5150F0" italics="true" pageId="0" pageNumber="563">pseudolucillum</emphasis>
/ dt. Netolitzky [handwritten, contoured in red] // coll. / Netolitzky [printed]&quot; (NHMW); from the handwritten label contoured in red we deduce that this last specimen was considered by
<location id="4C1F15094BACE011D54561CAD54F3616" LSID="urn:lsid:plazi:treatment:72A96E572D165EF9862C6B110F5D5552:4C1F15094BACE011D54561CAD54F3616" country="Japan" name="Netolitzky">Netolitzky</location>
as a
<typeStatus id="626BCF45D49C3D8ED281BE4F7A81973E">paratype</typeStatus>
</materialsCitation>
.
<materialsCitation id="BEB0AC26A0704A41AE46504958D22C22" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Netolitzky" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="1881821B51152488C6FD69900120E022" type="male">1 ♂</specimenCount>
,
<specimenCount id="F9410AF4936652807A64D685E0D009E2" type="female">2 ♀♀</specimenCount>
, on the same pin, &quot;Oku Nikko / Nabagawa / 8.9.37 Jano [handwritten] // coll. / Netolitzky [printed] //
<emphasis id="808779D5F6192902EE752976FAA7E10A" italics="true" pageId="0" pageNumber="563">pseudolucillum</emphasis>
/ dt.
<location id="F9533FD39B693FDAABCC679C244257AA" LSID="urn:lsid:plazi:treatment:72A96E572D165EF9862C6B110F5D5552:F9533FD39B693FDAABCC679C244257AA" country="Japan" name="Netolitzky">Netolitzky</location>
[handwritten]&quot; (NHMW); we mounted the
</materialsCitation>
<specimenCount id="6655483326FF2C2D44AF1E8F521B6FA3"></specimenCount>
on a separate pin, the labels have been photocopied in order to mount the specimens on separate pins, and the
<specimenCount id="DB518CA09A2555A67BB8C9639CA0C54D"></specimenCount>
, dissected, bears the slide with the aedeagus mounted in euparal on the same pin.
<materialsCitation id="C49AC835B397316CE852F597517ACF60" collectingDate="1950-04-12" collectorName="Leg. S. Ueno" country="Japan" determinerName="Det. S. Ueno" location="Kibune" specimenCount="1" specimenCount-male="1" stateProvince="Kyoto" typeStatus="paratype">
<specimenCount id="B7AF14516DF37AF150443E38A890D23A" type="male">1 ♂</specimenCount>
, &quot;
<location id="FDCE121CEB4C00198AC7D227B8BCCE75" LSID="urn:lsid:plazi:treatment:72A96E572D165EF9862C6B110F5D5552:FDCE121CEB4C00198AC7D227B8BCCE75" country="Japan" name="Kibune" stateProvince="Kyoto">Kibune</location>
,
<collectingRegion id="286B77CB817791D4FB6ECEC369E780F1" country="Japan" name="Kyoto">Kyoto</collectingRegion>
/
<collectingDate id="129AEED3AB5BE99375B4D73E4E7A39CA" value="1950-04-12">12-IV-1950</collectingDate>
/
<collectorName id="AF84687D43D76C06FF987F4650F437BF">
Leg. S.
<normalizedToken id="C2679FC6AF66F78BBFFE04F7ABC64896" originalValue="Uéno">Ueno</normalizedToken>
</collectorName>
[handwritten] // Acqu.-Nr. / 1951-23 [printed] // Coll. / Meyer //
<emphasis id="CBB8E3A98544862B9180F5B8A2A5DB7A" italics="true" pageId="0" pageNumber="563">pseudolucillum</emphasis>
/
<bibRefCitation id="6BA2EB01291CC40D1B68A45016E5CA93" DOI="https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" author="Netolitzky, F" journalOrPublisher="Proceedings of the Royal Entomological Society of London - Series B: Taxonomy" pageId="0" pageNumber="563" pagination="37 - 39" refId="B44" refString="Netolitzky, F, 1938. Beitraege zur Kenntnis der Bembidion -Arten des fernen ostens (Japan, Korea, Ostsibirien) (Coleoptera). I. Mitteilung. Proceedings of the Royal Entomological Society of London - Series B: Taxonomy 7: 37 - 39, DOI: https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" title="Beitraege zur Kenntnis der Bembidion - Arten des fernen ostens (Japan, Korea, Ostsibirien) (Coleoptera). I. Mitteilung." url="https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" volume="7" year="1938">Netolitzky 1938</bibRefCitation>
[handwritten] /
<determinerName id="5D4E858FBBD34B1E1E2E88492B6F1F3D">
Det. S.
<normalizedToken id="58197B9C6B384CD979C3FE50AFE37D90" originalValue="Uéno">Ueno</normalizedToken>
</determinerName>
[printed]&quot; (NHMW) (Fig.
<figureCitation id="7A17AEA2C18F6941462752618A9EA3DF" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 5.60 mm 2 B. (Plataphus) pseudolucillum Netolitzky, 1938 from Kibune, Kyoto (NHMW), 4.20 mm 3 B. (Peryphus) psuchrum Andrewes, 1935 Inde, Himachal Pradesh, Mahari, 3500 m (CTVR), 3.50 mm 4 labels of the holotype of Bembidion sapporense Jedlicka (NMPC). Photographs: Luca Toledano." figureDoi="10.3897/zookeys.1044.62593.figures1-4" httpUri="https://binary.pensoft.net/fig/555046" pageId="0" pageNumber="563">2</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3D1F4A2FBBBAB373E01F722375DE6AC4" collectingDate="1950-04-12" collectorName="Leg. S. Ueno" country="Japan" determinerName="Paul Meyer" location="Kibune" specimenCount="2" specimenCount-female="2" stateProvince="Kyoto" typeStatus="paratype">
<specimenCount id="6A80C2F630D7AA6A7950BF644DF617EA" type="female">2 ♀♀</specimenCount>
, &quot;
<location id="6E4C31517D52041A4F9533E96586DAB5" LSID="urn:lsid:plazi:treatment:72A96E572D165EF9862C6B110F5D5552:6E4C31517D52041A4F9533E96586DAB5" country="Japan" name="Kibune" stateProvince="Kyoto">Kibune</location>
,
<collectingRegion id="AA3DA0D2974C6739CAC1A401A875C66B" country="Japan" name="Kyoto">Kyoto</collectingRegion>
/
<collectingDate id="A2D727B68048EDBFC8D6BC7093CAF0E9" value="1950-04-12">12-IV-1950</collectingDate>
/
<collectorName id="5AA6EE53927FD0290C0FB54B31BD6F3E">
Leg. S.
<normalizedToken id="94C5CC6C684A8E96B71C775C1A4E94DB" originalValue="Uéno">Ueno</normalizedToken>
</collectorName>
[handwritten] // Acqu.-Nr. / 1951-23 [printed] // Coll. / Meyer //
<emphasis id="6227D97D43F933B64964046FE50531CF" italics="true" pageId="0" pageNumber="563">pseudolucillum</emphasis>
[handwritten] / det.
<determinerName id="3B307EAE9315D05B30E3D3454B5FC331">Paul Meyer</determinerName>
[printed]&quot; (NHMW)
</materialsCitation>
.
<materialsCitation id="EE9F2BAF18A30F8596E15922E0476602" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="G. Lewis, Coll. Toledano Verona" country="Japan" location="Japan" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="3D6FBA184649A4749E6469B8C11FAF19" type="male">1 ♂</specimenCount>
, &quot;
<collectingCountry id="2BB0546AC9A7EA25FDBB82E03115CDAC" name="Japan">Japan</collectingCountry>
<collectorName id="54F48ACA7DAE08861B6AA02AC3CF0A97">G. Lewis</collectorName>
//
<collectingCountry id="810EF54D271AD8AA9B9BBC5C206D9DC7" name="Japan">Japan</collectingCountry>
. Lewis // Sharp Coll. 1905-313 //
<emphasis id="AAD705845F7ECA25E91AE3C745C9F9AA" italics="true" pageId="0" pageNumber="563">Bembidion</emphasis>
cfr.
<emphasis id="A416352B0B77C45F894B5EF5388C3FBB" italics="true" pageId="0" pageNumber="563">pseudolucillum</emphasis>
det. L.
<bibRefCitation id="F0283D9638EC8308631C4EE60F8D4307" author="Toledano, L" editor="Toledano, L" journalOrPublisher="Sezione Scienze della Vita" pageId="0" pageNumber="563" pagination="5 - 46" publicationUrl="http://www.museostorianaturaleverona.it/media/_Musei/_StoriaNaturale/_Allegati/Biblioteca/Altre%20pubblicazioni/Pubblicazioni%20collaboratori/Toledano/2008_Systematic_notes_Palaearctic_Bembidion_Latrelle.pdf" refId="B54" refString="Toledano, L, 2008. Systematic notes on the Palaearctic Bembidion Latreille, 1802 (Coleoptera, Carabidae) with particular reference to the fauna of China. In: Toledano, L, Ed., Contributions to the knowledge of the eastern Palaearctic Bembidiina (Insecta, Coleoptera, Carabidae). Memorie del Museo Civico di Storia Naturale di Verona 2. Serie. Sezione Scienze della Vita 18: 5 - 46, http://www.museostorianaturaleverona.it/media/_Musei/_StoriaNaturale/_Allegati/Biblioteca/Altre%20pubblicazioni/Pubblicazioni%20collaboratori/Toledano/2008_Systematic_notes_Palaearctic_Bembidion_Latrelle.pdf" title="Systematic notes on the Palaearctic Bembidion Latreille, 1802 (Coleoptera, Carabidae) with particular reference to the fauna of China." url="http://www.museostorianaturaleverona.it/media/_Musei/_StoriaNaturale/_Allegati/Biblioteca/Altre%20pubblicazioni/Pubblicazioni%20collaboratori/Toledano/2008_Systematic_notes_Palaearctic_Bembidion_Latrelle.pdf" volume="18" volumeTitle="Contributions to the knowledge of the eastern Palaearctic Bembidiina (Insecta, Coleoptera, Carabidae). Memorie del Museo Civico di Storia Naturale di Verona 2. Serie." year="2008">Toledano 2008</bibRefCitation>
// CTVR
<collectorName id="80BDB05DD066A54AB934707131CC1ED5">Coll. Toledano Verona</collectorName>
-
<collectingCountry id="CB38126F6A3B86CB17FC3B5C81840E65" name="Italy">Italy</collectingCountry>
&quot; (CTVR)
</materialsCitation>
.
<materialsCitation id="B3B7A6632AFC37CB050B561BF4B23D80" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Coll. Toledano" country="Japan" determinerName="L. Toledano" location="Verona" specimenCount="1" specimenCount-male="1" stateProvince="Aomori Pref." typeStatus="paratype">
<specimenCount id="B374092BA292313ADFA98B7D45D71F83" type="male">1 ♂</specimenCount>
, &quot;Kujukawa /
<collectingRegion id="F4484E080F293F0401026F4001C2B716" country="Japan" name="Aomori">Aomori Pref.</collectingRegion>
/ Honehu
<collectingCountry id="3C9D7CA0291B4C62C65F61F25158FF05" name="Japan">Japan</collectingCountry>
/ 10.9.54 [handwritten] // Bembidion / pseudolucillum /
<bibRefCitation id="2A0764D5306DFACF6215FFC4F1FFE91D" DOI="https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" author="Netolitzky, F" journalOrPublisher="Proceedings of the Royal Entomological Society of London - Series B: Taxonomy" pageId="0" pageNumber="563" pagination="37 - 39" refId="B44" refString="Netolitzky, F, 1938. Beitraege zur Kenntnis der Bembidion -Arten des fernen ostens (Japan, Korea, Ostsibirien) (Coleoptera). I. Mitteilung. Proceedings of the Royal Entomological Society of London - Series B: Taxonomy 7: 37 - 39, DOI: https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" title="Beitraege zur Kenntnis der Bembidion - Arten des fernen ostens (Japan, Korea, Ostsibirien) (Coleoptera). I. Mitteilung." url="https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" volume="7" year="1938">Netolitzky 1938</bibRefCitation>
/ det.
<determinerName id="3521EE680B189EB783792B8D0624F49A">L. Toledano</determinerName>
2011 [printed] // CTVR /
<collectorName id="E03153847D11A257010166C5FC9EB128">Coll. Toledano</collectorName>
/
<location id="F50A17E3CB7DD7E65E026602976A7B05" LSID="urn:lsid:plazi:treatment:72A96E572D165EF9862C6B110F5D5552:F50A17E3CB7DD7E65E026602976A7B05" country="Japan" name="Verona" stateProvince="Aomori Pref.">Verona</location>
-
<collectingCountry id="9EBA49FB2A72105ED64C40767FBEB2BB" name="Italy">Italy</collectingCountry>
[printed]&quot; (CTVR)
</materialsCitation>
.
<materialsCitation id="F0DFA07DCDAE6DA1E5BED859F023BF98" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Japan" specimenCount="7" specimenCount-female="7" typeStatus="paratype">
<specimenCount id="03DF0CFB67D0A94368C465220EC01871" type="female">7 ♀♀</specimenCount>
, &quot;Numata, Yumma Pref., Honshu
<collectingCountry id="C8C53F8AEECA9E9216EF2F1EA5CB3E1A" name="Japan">Japan</collectingCountry>
&quot; (CTVR, PN)
</materialsCitation>
.
<materialsCitation id="0E9C44069627194543808BA5E96161C4" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Shizuoka pref." specimenCount="1" specimenCount-female="1" stateProvince="Shizuoka pref." typeStatus="paratype">
<specimenCount id="6F298E99DD275C7840E15CE06157A03C" type="female">1 ♀</specimenCount>
, &quot;Amagi-Pass, Kawazu-cho,
<collectingRegion id="DA79E421917FBCCB2C5DDB8A32FCE392" country="Japan" name="Shizuoka">Shizuoka pref.</collectingRegion>
&quot; (CTVR)
</materialsCitation>
.
<materialsCitation id="9FF6AF1740B079732AA9586B8A63F441" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Yamanashi-pref." specimenCount="1" specimenCount-female="1" stateProvince="Shizuoka" typeStatus="paratype">
<specimenCount id="7E7063F7791F2CCE37CE19FDEDA4346B" type="female">1 ♀</specimenCount>
, &quot;Sanjo-valley, Tabayama vill.,
<location id="0BEE4B93252209FDA759590E1BF913BA" LSID="urn:lsid:plazi:treatment:72A96E572D165EF9862C6B110F5D5552:0BEE4B93252209FDA759590E1BF913BA" country="Japan" name="Yamanashi-pref." stateProvince="Shizuoka">Yamanashi-pref.</location>
&quot; (CTVR)
</materialsCitation>
.
<materialsCitation id="A6A6DD3AEB37F3D249327730B7F8E1AD" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Honshu" country="Japan" elevation="700" specimenCount="1" specimenCount-female="1" stateProvince="Shizuoka" typeStatus="paratype">
<specimenCount id="BEDFB75F275C98471377A45CB0DCE6B7" type="female">1 ♀</specimenCount>
, &quot;
<collectorName id="A20F5BE64BEB2AA2A7931BA50F04DAE3">J. Honshu</collectorName>
, Shiga-ken, Mikunidakeyama, E slope,
<quantity id="96F29967AC61D500DFE3268A71434509" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">
<elevation id="609B6EB383AF95DA0AF550455F80C789" metricMagnitude="2" metricUnit="m" metricValue="7.0" unit="m" value="700.0">700 m</elevation>
</quantity>
&quot; (CTVR)
</materialsCitation>
.
<materialsCitation id="54D8DBAEF9154253D5BAE8A6D9448F6A" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Japan" location="Mt. Hiba" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Hiroshima pref." typeStatus="paratype">
<specimenCount id="B08EE3FF21E979356B7BC261AE5E0276" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="4E958748AEB6F6E6F29B545006C7ED3E" type="male">3 ♂♂</specimenCount>
, &quot;
<collectingCountry id="2C9B0F82096FDA5266B631EC2EDA51A9" name="Japan">Japan</collectingCountry>
,
<location id="2D413C26106D1456284D0AB21B6A4ABB" LSID="urn:lsid:plazi:treatment:72A96E572D165EF9862C6B110F5D5552:2D413C26106D1456284D0AB21B6A4ABB" country="Japan" name="Mt. Hiba" stateProvince="Hiroshima pref.">Mt. Hiba</location>
, Saljo,
<collectingRegion id="859FE03704DEC092F0E620ABAFF42BD1" country="Japan" name="Hiroshima">Hiroshima pref.</collectingRegion>
&quot; (CTVR, PN)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B787C8CE3DE96A24A24521C586B4D9BD" pageId="0" pageNumber="563" type="notes">
<paragraph id="2C335CAB2C4FEBF7028B3359C80F7970" pageId="0" pageNumber="563">Notes.</paragraph>
<paragraph id="80326BDCBB03FD1BC051EB807A7A494E" pageId="0" pageNumber="563">
<bibRefCitation id="C4A59B1FA789552F9C66883FF2C3F182" DOI="https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" author="Netolitzky, F" journalOrPublisher="Proceedings of the Royal Entomological Society of London - Series B: Taxonomy" pageId="0" pageNumber="563" pagination="37 - 39" refId="B44" refString="Netolitzky, F, 1938. Beitraege zur Kenntnis der Bembidion -Arten des fernen ostens (Japan, Korea, Ostsibirien) (Coleoptera). I. Mitteilung. Proceedings of the Royal Entomological Society of London - Series B: Taxonomy 7: 37 - 39, DOI: https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" title="Beitraege zur Kenntnis der Bembidion - Arten des fernen ostens (Japan, Korea, Ostsibirien) (Coleoptera). I. Mitteilung." url="https://doi.org/10.1111/j.1365-3113.1938.tb01241.x" volume="7" year="1938">Netolitzky (1938)</bibRefCitation>
describes
<taxonomicName id="ED5151091DE7E7509AD4AA64561DA356" authorityName="Neri &amp; Toledano" authorityYear="2021" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="1938" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Peryphus) pseudolucillum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="pseudolucillum" subGenus="Peryphus">Bembidion (Peryphus) pseudolucillum</taxonomicName>
from Settsu (Japan), assigning it to the group of
<taxonomicName id="3D47CB1F2494A2184473460B7AA18F19" lsidName="B. nitidulum" pageId="0" pageNumber="563" rank="species" species="nitidulum">
<emphasis id="2B5F29FC08B7DE63C53EC412E105793E" italics="true" pageId="0" pageNumber="563">B. nitidulum</emphasis>
</taxonomicName>
Marsham, 1802, s. l. (=
<taxonomicName id="1D435219FFBDD92D6D9202C848329E16" lsidName="B. deletum" pageId="0" pageNumber="563" rank="species" species="deletum">
<emphasis id="F7771B92234B700BD9BBF8768BADBE6A" italics="true" pageId="0" pageNumber="563">B. deletum</emphasis>
</taxonomicName>
Serville, 1821); again
<bibRefCitation id="EA04F2B20F152657406850EA4D81A808" author="Netolitzky, F" journalOrPublisher="Proceedings of the Royal Entomological Society of London - Series B: Taxonomy" pageId="0" pageNumber="563" refId="B45" refString="Netolitzky, F, 1943a. Bestimmungs-Tabellen europaeischer Kaefer (9. Stück). II. Fam. Carabidae. Subfam. Bembidiinae. 66. Gattung: Bembidion Latr. Bestimmungstabelle der Bembidion-Arten des palaearktischen Gebietes. (Mit Hinweisen auf holarktische, aethiopische und orientalische Arten). Koleopterologische Rundschau 28(4/6): 69/41-124/96." title="Bestimmungs-Tabellen europaeischer Kaefer (9. Stu ̈ ck). II. Fam. Carabidae. Subfam. Bembidiinae. 66. Gattung: Bembidion Latr. Bestimmungstabelle der Bembidion-Arten des palaearktischen Gebietes. (Mit Hinweisen auf holarktische, aethiopische und orientalische Arten). Koleopterologische Rundschau 28 (4 / 6): 69 / 41 - 124 / 96." year="1943 a">Netolitzky (1943a</bibRefCitation>
,
<bibRefCitation id="867435A781F93CF7A2BC1493A80AFD48" author="Netolitzky, F" journalOrPublisher="Proceedings of the Royal Entomological Society of London - Series B: Taxonomy" pageId="0" pageNumber="563" refId="B46" refString="Netolitzky, F, 1943b. Bestimmungs-Tabellen europaeischer Kaefer (9. Stueck). II. Fam. Carabidae, Subfam. Bembidiinae. 66. Gattung: Bembidion Latr. Bestimmungstabelle der Bembidion-Arten des palaearktischen Gebietes. (Mit Hinweisen auf holarktische, aethiopische und orientalische Arten). Koleopterologische Rundschau 29: 1/97-70/166." title="Bestimmungs-Tabellen europaeischer Kaefer (9. Stueck). II. Fam. Carabidae, Subfam. Bembidiinae. 66. Gattung: Bembidion Latr. Bestimmungstabelle der Bembidion-Arten des palaearktischen Gebietes. (Mit Hinweisen auf holarktische, aethiopische und orientalische Arten). Koleopterologische Rundschau 29: 1 / 97 - 70 / 166." year="1943 b">1943b</bibRefCitation>
) found it difficult to find a correct position for the species in his Bestimmungs-Tabellen: he mentions similarities both with species of two subgenera,
<taxonomicName id="52A7A388E1181DD72163ECC836AA83A0" genus="Bembidionetolitzya" lsidName="Bembidionetolitzya" pageId="0" pageNumber="563" rank="genus">
<emphasis id="94B54C7EF65716E8C0BB2943B781E83A" italics="true" pageId="0" pageNumber="563">Bembidionetolitzya</emphasis>
</taxonomicName>
Strand, 1929 and
<taxonomicName id="5B2800685EF57560B3861D27B33B6BC8" authorityName="Dejean" authorityYear="1821" class="Insecta" family="Carabidae" genus="Peryphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Peryphus" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="genus">
<emphasis id="CC19150E139C9A39893247FCE956823D" italics="true" pageId="0" pageNumber="563">Peryphus</emphasis>
</taxonomicName>
Dejean, 1821, but preferred the latter.
<bibRefCitation id="28620638CA96037B20065FB8A306DFA3" author="Jedlicka, A" journalOrPublisher="Entomologisk Tidskrift" pageId="0" pageNumber="563" pagination="202 - 208" refId="B18" refString="Jedlicka, A, 1965. Neue Carabiden aus den Sammlungen des Riksmuseums in Stockholm (Coleoptera - Carabidae). Entomologisk Tidskrift 86: 202 - 208" title="Neue Carabiden aus den Sammlungen des Riksmuseums in Stockholm (Coleoptera - Carabidae)." volume="86" year="1965">
<normalizedToken id="0E725178297A4EC5F7C339164122D397" originalValue="Jedlička">Jedlicka</normalizedToken>
(1965)
</bibRefCitation>
, in his monograph on the
<taxonomicName id="A7FFCB4042808F4DCC4EC8A056F8F9DC" authorityName="Stephens" authorityYear="1827" lsidName="" pageId="0" pageNumber="563" rank="tribe" tribe="Bembidiini">Bembidiini</taxonomicName>
of Eastern Asia, includes the species in the keys for subgenus
<taxonomicName id="0AE188A2B24661B5E74FCF93B7F1564A" class="Insecta" family="Carabidae" genus="Plataphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Plataphus" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="genus">Plataphus</taxonomicName>
Motschulsky, 1864 and that of the subgenus
<taxonomicName id="BFC477F8FE114BD074E561E3E2BFF8AB" class="Insecta" family="Carabidae" genus="Plataphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Plataphus (Peryphus)" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subGenus" subGenus="Peryphus">Plataphus Peryphus</taxonomicName>
. The species is listed in &quot;incertae sedis&quot; in the last catalogues (e.g.,
<bibRefCitation id="BDC94AE04F42D2932AE82A45A1A5D658" author="Marggi, W" journalOrPublisher="Apollo Books, Stenstrup" pageId="0" pageNumber="563" refId="B28" refString="Marggi, W, Huber, C, Mueller-Motzfeld, G, Hartmann, M, 2003. Carabidae: Bembidiini: Bembidiina: 242-273. In: Loebl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera (Vol. 1). Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup" title="Carabidae: Bembidiini: Bembidiina: 242 - 273. In: Loebl I, Smetana A (Eds) Catalogue of Palaearctic Coleoptera (Vol. 1). Archostemata - Myxophaga - Adephaga." year="2003">Marggi et al. 2003</bibRefCitation>
;
<bibRefCitation id="F3191DD644666D4899A9A4A95193981B" author="Lorenz, W" journalOrPublisher="Brill, Leiden-Boston" pageId="0" pageNumber="563" refId="B26" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (2nd edn.). Tutzing, 530 pp." title="Systematic List of Extant Ground Beetles of the World (2 nd edn.). Tutzing, 530 pp." year="2005">Lorenz 2005</bibRefCitation>
;
<bibRefCitation id="CCD1A185D41FB0E5EEA62FEE4331B02A" author="Marggi, W" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="563" refId="B29" refString="Marggi, W, Toledano, L, Neri, P, 2017. Carabidae: Bembidiini: Bembidiina: 294-342. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera (Vol. I). Archostemata - Myxophaga - Adephaga. Brill, Leiden" title="Carabidae: Bembidiini: Bembidiina: 294 - 342. In: Loebl I, Loebl D (Eds) Catalogue of Palaearctic Coleoptera (Vol. I). Archostemata - Myxophaga - Adephaga." year="2017">Marggi et al. 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="608C18A3FFA7DCD10909E6E67E6F4838" pageId="0" pageNumber="563" type="conclusions">
<paragraph id="A1A2CF632439003265C2B7E7587AA63E" pageId="0" pageNumber="563">Conclusions.</paragraph>
<paragraph id="F3045575C04C78D2E88A720F68ADA0D3" pageId="0" pageNumber="563">
The examination of the holotype and of the aedeagus (Fig.
<figureCitation id="D4EB7DD773AA4518AAB3ED50698CF799" captionStart="Figures 59" captionStartId="F2" captionText="Figures 5 - 9. Median lobe of the aedeagi of 5 Bembidion (Peryphanes) sanatum Bates, 1883, holotype (NHMUK), 1.55 mm 6 B. (P.) dostali Kirschenhofer, 1984, paratype (NHMW), 1.53 mm 7 B. (P.) lulinense Habu, 1973 (drawing from Habu 1973), 1.41 mm 8 Bembidion (Peryphus) psuchrum Andrewes, 1935, holotype (NHMUK), 0.91 mm 9 Bembidion (Plataphus) pseudolucillum Netolitzky, paratype (NHMW), 0.80 mm." figureDoi="10.3897/zookeys.1044.62593.figures5-9" httpUri="https://binary.pensoft.net/fig/555047" pageId="0" pageNumber="563">9</figureCitation>
) shows that the species belongs to subgenus
<taxonomicName id="88C3F9C0EDC52EF5904FDE445C32CA55" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Plataphus)" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="subGenus" subGenus="Plataphus">Bembidion Plataphus</taxonomicName>
Motschulsky, 1864. The doubts regarding the systematic position of the species are due to its pronotal convexity, more evident than in other species of the subgenus. We added to all the non-type specimens the following label:
<taxonomicName id="5F2699ACEEE376408BB2D7DAA8C33A58" authorityName="Neri &amp; Toledano" authorityYear="2021" baseAuthorityName="Neri &amp; Toledano" baseAuthorityYear="1938" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bembidion (Plataphus) pseudolucillum" order="Coleoptera" pageId="0" pageNumber="563" phylum="Arthropoda" rank="species" species="pseudolucillum" subGenus="Plataphus">Bembidion (Plataphus) pseudolucillum</taxonomicName>
Net. - det. Neri and Toledano 2020. The species length varies from 3.6 to 4.7 mm, the aedeagus from 0.75 to 0.84 mm.
</paragraph>
</subSubSection>
</treatment>
</document>