259 lines
23 KiB
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259 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.875.37856" ID-GBIF-Dataset="b0d5590c-66a9-4126-b2b2-04eaf78f0b7a" ID-PMC="PMC6751220" ID-Pensoft-Pub="1313-2970-875-31" ID-Pensoft-UUID="258D336D48D15A3CA8C52670044D8899" ID-PubMed="31572044" ID-ZooBank="17D16DA3DEFC491AB4A12B49FACCFEA6" ModsDocID="1313-2970-875-31" checkinTime="1568292418084" checkinUser="pensoft" docAuthor="Chou, Ming-Hsun & Yeh, Wen-Bin" docDate="2019" docId="7297C7A08CE25121BDA33327F45FF95D" docLanguage="en" docName="ZooKeys 875: 31-62" docOrigin="ZooKeys 875" docSource="http://dx.doi.org/10.3897/zookeys.875.37856" docTitle="Cylindera (Cylindera) autumnalis Chou & Yeh, 2019, sp. nov." docType="treatment" docUuid="341884A9-BC65-4443-B269-A962E3472D0A" docUuidSource="ZooBank" docVersion="5" id="258D336D48D15A3CA8C52670044D8899" lastPageNumber="52" masterDocId="258D336D48D15A3CA8C52670044D8899" masterDocTitle="Delineation of two new, highly similar species of Taiwanese Cylindera tiger beetles (Coleoptera, Carabidae, Cicindelinae) based on morphological and molecular evidence" masterLastPageNumber="62" masterPageNumber="31" pageNumber="47" updateTime="1668167804062" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Delineation of two new, highly similar species of Taiwanese Cylindera tiger beetles (Coleoptera, Carabidae, Cicindelinae) based on morphological and molecular evidence</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chou, Ming-Hsun</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yeh, Wen-Bin</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>875</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>31</mods:start>
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<mods:end>62</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.875.37856</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.875.37856</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-875-31</mods:identifier>
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<mods:identifier type="ZooBank">17D16DA3DEFC491AB4A12B49FACCFEA6</mods:identifier>
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<mods:identifier type="Pensoft-UUID">258D336D48D15A3CA8C52670044D8899</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159412827" LSID="urn:lsid:zoobank.org:act:341884A9-BC65-4443-B269-A962E3472D0A" httpUri="http://treatment.plazi.org/id/7297C7A08CE25121BDA33327F45FF95D" lastPageId="22" lastPageNumber="52" pageId="16" pageNumber="47">
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<subSubSection pageId="16" pageNumber="47" type="nomenclature">
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<paragraph pageId="16" pageNumber="47">
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<taxonomicName LSID="7297C7A08CE25121BDA33327F45FF95D" class="Insecta" family="Carabidae" genus="Cylindera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cylindera (Cylindera) autumnalis" order="Coleoptera" pageId="16" pageNumber="47" phylum="Arthropoda" rank="species" species="autumnalis" subGenus="Cylindera">
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<pageBreakToken pageId="16" pageNumber="47" start="start">Cylindera</pageBreakToken>
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(Cylindera) autumnalis
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</taxonomicName>
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<taxonomicNameLabel pageId="16" pageNumber="47">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="18" lastPageNumber="49" pageId="17" pageNumber="48" type="type material">
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<paragraph pageId="17" pageNumber="48">
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<pageBreakToken pageId="17" pageNumber="48" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph lastPageId="18" lastPageNumber="49" pageId="17" pageNumber="48">
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<emphasis bold="true" italics="true" pageId="17" pageNumber="48">Holotype:</emphasis>
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male (
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<figureCitation captionStart="Figures 26–27" captionStartId="F14" captionText="Figures 26 - 27. Type specimens of Cylindera autumnalis sp. nov. 26 male holotype (AdeC 84 - 3) 27 female paratype (AdeC 78 - 2)." figureDoi="10.3897/zookeys.875.37856.figures26-27" httpUri="https://binary.pensoft.net/fig/339401" pageId="17" pageNumber="48" tableDoi="10.3897/zookeys.875.37856.figures26-27">Fig. 26</figureCitation>
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; specimen code: AdeC84-3; dry pinned, with aedeagus in glycerol in a separated microvial labeled
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<normalizedToken originalValue="“AdeC84-3”">"AdeC84-3"</normalizedToken>
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): Taiwan, Pintung, Shuangliu Forest Recreation Area, 03 Sep. 2018, Ming-Hsun Chou leg. Original label: "Locality:
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<normalizedToken originalValue="屏東雙流森林遊樂區">屏東雙流森林遊樂區</normalizedToken>
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/ Date: 2018.IX.03 / Collector:
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<normalizedToken originalValue="周明勳">周明勳</normalizedToken>
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/ Code: AdeC84-3"; " NCHU 0011-0736". Dry specimen and aedeagus of holotype deposited in NCHU.
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<emphasis bold="true" italics="true" pageId="17" pageNumber="48">Paratypes</emphasis>
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: 3 males (specimen code: AdeC48-4, AdeC48-5, and AdeC48-8, respectively; dry pinned, with aedeagus in glycerol in a separated microvial, respectively), 1 female (specimen code: AdeC48-1; dry pinned), and 1 female (specimen code: AdeC48-2; dry pinned, with genitalia preserved in glycerol in a separated microvial): Taiwan, Pintung, Shuangliu Forest Recreation Area, 10 Aug. 2017, Ming-Hsun Chou leg. 1 male (specimen code: AdeC78-1; dry pinned, with aedeagus in glycerol in a separated microvial) and 1 female (specimen code: AdeC78-2 (
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<figureCitation captionStart="Figures 26–27" captionStartId="F14" captionText="Figures 26 - 27. Type specimens of Cylindera autumnalis sp. nov. 26 male holotype (AdeC 84 - 3) 27 female paratype (AdeC 78 - 2)." figureDoi="10.3897/zookeys.875.37856.figures26-27" httpUri="https://binary.pensoft.net/fig/339401" pageId="17" pageNumber="48" tableDoi="10.3897/zookeys.875.37856.figures26-27">Fig. 27</figureCitation>
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); dry pinned): Taiwan, Pintung, Shuangliu Forest Recreation Area, 21 Jul. 2018, Ming-Hsun Chou leg. 1 male (specimen code: AdeC84-1; dry pinned, with aedeagus in glycerol in a separated microvial) and 1 female (specimen code: AdeC84-2; dry pinned, with geni
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<pageBreakToken pageId="18" pageNumber="49" start="start">talia</pageBreakToken>
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in glycerol in a separated microvial): same collecting information as for holotype. Original labels of paratypes see Table
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<tableCitation captionStartId="T2" httpUri="http://table.plazi.org/id/A3D62C522B559692263B921080AAB736" pageId="18" pageNumber="49" tableUuid="A3D62C522B559692263B921080AAB736">2</tableCitation>
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. All dry specimens and genitalia of paratypes deposited in NMNS.
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</paragraph>
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<caption doi="10.3897/zookeys.875.37856.figures26-27" httpUri="https://binary.pensoft.net/fig/339401" pageId="18" pageNumber="49" start="Figures 26–27" startId="F14">
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<paragraph pageId="18" pageNumber="49">
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<emphasis bold="true" pageId="18" pageNumber="49">Figures 26-27.</emphasis>
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Type specimens of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cylindera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cylindera autumnalis" order="Coleoptera" pageId="18" pageNumber="49" phylum="Arthropoda" rank="species" species="autumnalis">
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<emphasis italics="true" pageId="18" pageNumber="49">Cylindera autumnalis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="18" pageNumber="49">26</emphasis>
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male holotype (AdeC84-3)
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<emphasis bold="true" pageId="18" pageNumber="49">27</emphasis>
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female paratype (AdeC78-2).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="49" type="type locality">
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<paragraph pageId="18" pageNumber="49">Type locality.</paragraph>
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<paragraph pageId="18" pageNumber="49">Taiwan, Pintung, Shuangliu Forest Recreation Area.</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="49" type="diagnosis">
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<paragraph pageId="18" pageNumber="49">Diagnosis.</paragraph>
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<paragraph pageId="18" pageNumber="49">
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Elytra are metallic brownish and marked with obvious punctures. The apical lunula is thickened in both ends (subapical corner and apical end near suture).
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<taxonomicName class="Insecta" family="Carabidae" genus="Cylindera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cylindera autumnalis" order="Coleoptera" pageId="18" pageNumber="49" phylum="Arthropoda" rank="species" species="autumnalis">
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<emphasis italics="true" pageId="18" pageNumber="49">Cylindera autumnalis</emphasis>
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</taxonomicName>
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sp. nov. has a different body coloration and more obvious elytral maculation than
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<taxonomicName lsidName="C. pseudocylindriformis" pageId="18" pageNumber="49" rank="species" species="pseudocylindriformis">
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<emphasis italics="true" pageId="18" pageNumber="49">C. pseudocylindriformis</emphasis>
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</taxonomicName>
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(
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<figureCitation captionStart="Figure 28" captionStartId="F15" captionText="Figure 28. Dorsal habitus of Cylindera pseudocylindriformis (left - male; right - female)." figureDoi="10.3897/zookeys.875.37856.figure28" httpUri="https://binary.pensoft.net/fig/339402" pageId="18" pageNumber="49" tableDoi="10.3897/zookeys.875.37856.figure28">Fig. 28</figureCitation>
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). The former has few hairs on mesoepisterna in male and on metepisterna in both genders, but the
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<normalizedToken originalValue="latter’s">latter's</normalizedToken>
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mesoepisterna and metepisterna are glabrous in both genders. Body size of
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<taxonomicName lsidName="C. autumnalis" pageId="18" pageNumber="49" rank="species" species="autumnalis">
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<emphasis italics="true" pageId="18" pageNumber="49">C. autumnalis</emphasis>
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</taxonomicName>
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sp. nov. male was significantly larger than
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<taxonomicName lsidName="C. pseudocylindriformis" pageId="18" pageNumber="49" rank="species" species="pseudocylindriformis">
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<emphasis italics="true" pageId="18" pageNumber="49">C. pseudocylindriformis</emphasis>
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</taxonomicName>
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although is not statistically significant in female. Their labrum (
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<figureCitation captionStart="Figures 29–32" captionStartId="F16" captionText="Figures 29 - 32. Digital images (left) and line drawings (right) of labra of Cylindera autumnalis sp. nov. 29, 30 male (holotype, AdeC 84 - 3) 31, 32 female (paratype, AdeC 78 - 2)." figureDoi="10.3897/zookeys.875.37856.figures29-32" httpUri="https://binary.pensoft.net/fig/339403" pageId="18" pageNumber="49" tableDoi="10.3897/zookeys.875.37856.figures29-32">Figs 29-32</figureCitation>
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,
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<figureCitation captionStart="Figures 33–36" captionStartId="F17" captionText="Figures 33 - 36. Digital images (left) and line drawings (right) of labra of Cylindera pseudocylindriformis 33, 34 male 35, 36 female." figureDoi="10.3897/zookeys.875.37856.figures33-36" httpUri="https://binary.pensoft.net/fig/339404" pageId="18" pageNumber="49" tableDoi="10.3897/zookeys.875.37856.figures33-36">33-36</figureCitation>
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) and male genitalia (
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<figureCitation captionStart="Figures 12, 13" captionStartId="F9" captionText="Figures 12, 13. Digital image of aedeagus in left view of Pintung form (12) and Cylindera pseudocylindriformis (13). Abbreviations: ap arciform piece; cp central plate; f flagellum; p paramere; mt medial tooth; sr stiffening rib." figureDoi="10.3897/zookeys.875.37856.figures12-13" httpUri="https://binary.pensoft.net/fig/339396" pageId="18" pageNumber="49" tableDoi="10.3897/zookeys.875.37856.figures12-13">Figs 12</figureCitation>
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,
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<figureCitation captionStart="Figures 12, 13" captionStartId="F9" captionText="Figures 12, 13. Digital image of aedeagus in left view of Pintung form (12) and Cylindera pseudocylindriformis (13). Abbreviations: ap arciform piece; cp central plate; f flagellum; p paramere; mt medial tooth; sr stiffening rib." figureDoi="10.3897/zookeys.875.37856.figures12-13" httpUri="https://binary.pensoft.net/fig/339396" pageId="18" pageNumber="49" tableDoi="10.3897/zookeys.875.37856.figures12-13">13</figureCitation>
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) might be poorly distinctive.
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</paragraph>
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<caption doi="10.3897/zookeys.875.37856.figure28" httpUri="https://binary.pensoft.net/fig/339402" pageId="18" pageNumber="49" start="Figure 28" startId="F15">
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<paragraph pageId="18" pageNumber="49">
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<emphasis bold="true" pageId="18" pageNumber="49">Figure 28.</emphasis>
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Dorsal habitus of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cylindera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cylindera pseudocylindriformis" order="Coleoptera" pageId="18" pageNumber="49" phylum="Arthropoda" rank="species" species="pseudocylindriformis">
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<emphasis italics="true" pageId="18" pageNumber="49">Cylindera pseudocylindriformis</emphasis>
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</taxonomicName>
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(left - male; right - female).
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.875.37856.figures29-32" httpUri="https://binary.pensoft.net/fig/339403" pageId="18" pageNumber="49" start="Figures 29–32" startId="F16">
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<paragraph pageId="18" pageNumber="49">
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<emphasis bold="true" pageId="18" pageNumber="49">Figures 29-32.</emphasis>
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Digital images (left) and line drawings (right) of labra of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cylindera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cylindera autumnalis" order="Coleoptera" pageId="18" pageNumber="49" phylum="Arthropoda" rank="species" species="autumnalis">
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<emphasis italics="true" pageId="18" pageNumber="49">Cylindera autumnalis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="18" pageNumber="49">29, 30</emphasis>
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male (holotype, AdeC84-3)
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<emphasis bold="true" pageId="18" pageNumber="49">31, 32</emphasis>
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female (paratype, AdeC78-2).
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.875.37856.figures33-36" httpUri="https://binary.pensoft.net/fig/339404" pageId="18" pageNumber="49" start="Figures 33–36" startId="F17">
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<paragraph pageId="18" pageNumber="49">
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<emphasis bold="true" pageId="18" pageNumber="49">Figures 33-36.</emphasis>
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Digital images (left) and line drawings (right) of labra of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cylindera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cylindera pseudocylindriformis" order="Coleoptera" pageId="18" pageNumber="49" phylum="Arthropoda" rank="species" species="pseudocylindriformis">
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<emphasis italics="true" pageId="18" pageNumber="49">Cylindera pseudocylindriformis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="18" pageNumber="49">33, 34</emphasis>
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male
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<emphasis bold="true" pageId="18" pageNumber="49">35, 36</emphasis>
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female.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="21" lastPageNumber="52" pageId="19" pageNumber="50" type="description">
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<paragraph pageId="19" pageNumber="50">
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<pageBreakToken pageId="19" pageNumber="50" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph lastPageId="21" lastPageNumber="52" pageId="19" pageNumber="50">
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<emphasis bold="true" italics="true" pageId="19" pageNumber="50">Head</emphasis>
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metallic bronze with weak greenish luster; genae dark metallic green; canthus with one seta; rugae longitudinal along frons, canthi, vertex, and lateral neck, and becoming transverse on genae; clypeus patina and microsculptured.
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<emphasis bold="true" italics="true" pageId="19" pageNumber="50">Compound eyes</emphasis>
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large and protruding.
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<emphasis bold="true" italics="true" pageId="19" pageNumber="50">Antennae</emphasis>
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slender and filiform; scape with one apical seta; 1-4 antennomeres metallic dark brown; 5-11 ones darker.
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<emphasis bold="true" italics="true" pageId="19" pageNumber="50">Mandible</emphasis>
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yellowish pale with darker teeth, exceeding labrum when closed.
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<emphasis bold="true" italics="true" pageId="19" pageNumber="50">Maxillary palps</emphasis>
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yellowish; last palpomere metallic dark testaceous.
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<emphasis bold="true" italics="true" pageId="19" pageNumber="50">Labial palps</emphasis>
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yellowish; last palpomere metallic dark testaceous.
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<emphasis bold="true" italics="true" pageId="19" pageNumber="50">Labrum</emphasis>
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testaceous; anterior margin rounded and unidentate in female; anterior margin without noticeable tooth or even concaved in male; margin with three or four preapical and two lateral setae (
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<figureCitation captionStart="Figures 29–32" captionStartId="F16" captionText="Figures 29 - 32. Digital images (left) and line drawings (right) of labra of Cylindera autumnalis sp. nov. 29, 30 male (holotype, AdeC 84 - 3) 31, 32 female (paratype, AdeC 78 - 2)." figureDoi="10.3897/zookeys.875.37856.figures29-32" httpUri="https://binary.pensoft.net/fig/339403" pageId="19" pageNumber="50" tableDoi="10.3897/zookeys.875.37856.figures29-32">Figs 29-32</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="19" pageNumber="50">Pronotum</emphasis>
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cylindrical and metallic bronze with little greenish luster; dorsum transversely rugose; one transverse groove on each anterior and posterior dorsum portions, connected with one shal
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<pageBreakToken pageId="20" pageNumber="51" start="start">low</pageBreakToken>
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longitudinal groove.
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<emphasis bold="true" italics="true" pageId="20" pageNumber="51">Elytra</emphasis>
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bronze with metallic luster, slender, and marked with many obvious punctures; humeral spot present; posthumeral spot discoidal or irregular; one triangular spot on middle margin of elytron, connected with one clavate spot but disconnected in some individuals; apical lunula obvious, crescent; both subapical portion and apical end near suture of apical lunula thickened.
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<emphasis bold="true" italics="true" pageId="20" pageNumber="51">Legs</emphasis>
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slender and testaceous, except metallic dark green coxae; some white hairs on coxae and femurs; pro-, mesocoxae, pro- and mesotrochanters with one long seta; pro-tarsi sexually dimorphic, basal 1-3 tarsomeres with short brush-like ventral setae and little wider than four or five tarsomeres in male, all pro-tarsomeres equivalent in width roughly and without brush-like ventral setae in female.
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<emphasis bold="true" italics="true" pageId="20" pageNumber="51">Thoracic proepisternum</emphasis>
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dark metallic green, longitudinally rugose, and glabrous.
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<emphasis bold="true" italics="true" pageId="20" pageNumber="51">Prosternum</emphasis>
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dark metallic green, transversally rugose, glabrous.
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<emphasis bold="true" italics="true" pageId="20" pageNumber="51">Mesoepisternum</emphasis>
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dark metallic green, rugose and longitudinally depressed,
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<pageBreakToken pageId="21" pageNumber="52" start="start">with</pageBreakToken>
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two or three hairs in male but glabrous in female.
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<emphasis bold="true" italics="true" pageId="21" pageNumber="52">Mesosternum</emphasis>
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dark metallic green, transversally rugose, glabrous.
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<emphasis bold="true" italics="true" pageId="21" pageNumber="52">Metepisternum</emphasis>
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dark metallic green, coarsely sculptured, with few hairs.
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<emphasis bold="true" italics="true" pageId="21" pageNumber="52">Metasternum</emphasis>
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dark metallic green, microsculptured, and almost glabrous.
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<emphasis bold="true" italics="true" pageId="21" pageNumber="52">Abdomen sternum</emphasis>
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dark green with little metallic luster, almost hairless, except one pair of long hairs on 4-6 segments.
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<emphasis bold="true" italics="true" pageId="21" pageNumber="52">Aedeagus</emphasis>
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of holotype shown in
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<figureCitation captionStart="Figures 9, 10" captionStartId="F7" captionText="Figures 9, 10. Digital image of aedeagus in left view of Kosempo form (AdeC 66 - 1) (9) and Cylindera sauteri (10). Abbreviations: ap arciform piece; cp central plate; f flagellum; p paramere; mt medial tooth; sr stiffening rib." figureDoi="10.3897/zookeys.875.37856.figures9-10" httpUri="https://binary.pensoft.net/fig/339394" pageId="21" pageNumber="52" tableDoi="10.3897/zookeys.875.37856.figures9-10">Fig. 10</figureCitation>
|
||
. Description same as Results.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="21" pageNumber="52" type="etymology">
|
||
<paragraph pageId="21" pageNumber="52">Etymology.</paragraph>
|
||
<paragraph pageId="21" pageNumber="52">
|
||
During the collection period in 2017 and 2018, this species was collected mostly in August to early September, especially in September. Many individuals could be found in early September when other tiger beetle adults disappeared mostly in that habitat. Thus, the specific name "
|
||
<taxonomicName lsidName="autumnalis" pageId="21" pageNumber="52" rank="species" species="autumnalis">
|
||
<emphasis italics="true" pageId="21" pageNumber="52">autumnalis</emphasis>
|
||
</taxonomicName>
|
||
" means the autumnal tiger beetle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="21" pageNumber="52" type="distribution">
|
||
<paragraph pageId="21" pageNumber="52">Distribution.</paragraph>
|
||
<paragraph pageId="21" pageNumber="52">Only known from type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="22" lastPageNumber="53" pageId="21" pageNumber="52" type="ecology">
|
||
<paragraph pageId="21" pageNumber="52">Ecology.</paragraph>
|
||
<paragraph lastPageId="22" lastPageNumber="53" pageId="21" pageNumber="52">
|
||
According to field observation, adults live in forest trails in late summer to autumn (late July to September). They crawl on the open ground and fly away for a short distance when being bothered, sometimes hiding in the grass or litters. The other two tiger beetle species which could be also found in the same habitat are
|
||
<taxonomicName lsidName="C. cylindriformis" pageId="21" pageNumber="52" rank="species" species="cylindriformis">
|
||
<emphasis italics="true" pageId="21" pageNumber="52">C. cylindriformis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Therates" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Therates alboobliquatus" order="Coleoptera" pageId="21" pageNumber="52" phylum="Arthropoda" rank="subspecies" species="alboobliquatus" subspecies="alboobliquatus">
|
||
<emphasis italics="true" pageId="21" pageNumber="52">Therates alboobliquatus alboobliquatus</emphasis>
|
||
</taxonomicName>
|
||
Horn, 1909. However, adults
|
||
<pageBreakToken pageId="22" pageNumber="53" start="start">of</pageBreakToken>
|
||
these three tiger beetles seem to appear in different seasons.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cylindera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cylindera cylindriformis" order="Coleoptera" pageId="22" pageNumber="53" phylum="Arthropoda" rank="species" species="cylindriformis">
|
||
<emphasis italics="true" pageId="22" pageNumber="53">Cylindera cylindriformis</emphasis>
|
||
</taxonomicName>
|
||
adults appear in early to mid-summer, and
|
||
<taxonomicName lsidName="T. a." pageId="22" pageNumber="53" rank="subspecies" species="a." subspecies="alboobliquatus">
|
||
<emphasis italics="true" pageId="22" pageNumber="53">T. a. alboobliquatus</emphasis>
|
||
</taxonomicName>
|
||
was recorded mainly in mid-summer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |