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<document id="D19362D9FE33EB4ECA63E3D6570F41AC" ID-DOI="10.5281/zenodo.189228" ID-GBIF-Dataset="29e7002a-ee86-4b7b-804f-2e95f678e6a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="189228" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461100418165" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho" docDate="2009" docId="727C676A6A5EFFE4FF23FF1FFB0EF877" docLanguage="en" docName="zt02172p044.pdf" docOrigin="Zootaxa 2172" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptinopterus affinis Parry 1872" docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="8E451F126A5DFFE7FFB4FA54FFC0FF86" masterDocTitle="Description of two new species of Leptinopterus Hope (Coleoptera: Lucanidae: Lucaninae) with notes on the taxonomy and natural history of the genus" masterLastPageNumber="44" masterPageNumber="32" pageNumber="35" updateTime="1698591109854" updateUser="plazi">
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<mods:title id="B675E5DF8F2D5B8244507D97A5BC8ED2">Description of two new species of Leptinopterus Hope (Coleoptera: Lucanidae: Lucaninae) with notes on the taxonomy and natural history of the genus</mods:title>
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<mods:date id="3CB499A5D88334922957272DB4FC50FE">2009</mods:date>
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<subSubSection id="B2CF85F76A5EFFE4FF23FF1FFDECFAE3" box="[151,556,1355,1381]" pageId="3" pageNumber="35" type="nomenclature">
<paragraph id="FA6AD67C6A5EFFE4FF23FF1FFDECFAE3" blockId="3.[151,556,1355,1381]" box="[151,556,1355,1381]" pageId="3" pageNumber="35">
<heading id="A12261106A5EFFE4FF23FF1FFDECFAE3" bold="true" box="[151,556,1355,1381]" fontSize="11" level="1" pageId="3" pageNumber="35" reason="1">
<taxonomicName id="3DD5ADFF6A5EFFE4FF23FF1FFDECFAE3" ID-CoL="3TJS4" authority="Parry, 1872" authorityName="Parry" authorityYear="1872" box="[151,556,1355,1381]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="C8A10A6E6A5EFFE4FF23FF1FFDECFAE3" bold="true" box="[151,556,1355,1381]" pageId="3" pageNumber="35">
<emphasis id="C8A10A6E6A5EFFE4FF23FF1FFE54FAE3" bold="true" box="[151,404,1355,1381]" italics="true" pageId="3" pageNumber="35">Leptinopterus affinis</emphasis>
<bibRefCitation id="9E44AB8D6A5EFFE4FE2EFF1FFDECFAE3" author="Parry" box="[410,556,1355,1381]" pageId="3" pageNumber="44" refString="Parry, F. J. S. (1872) Descriptions of new species of lucanoid Coleoptera: with remarks on the genus Cantharolethrus, and supplementary lists. Transactions of the Royal Entomological Society of London, series 1, volume 4, 73 - 84." type="journal article" year="1872">Parry, 1872</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="B2CF85F76A5EFFE4FF23FFC1FCC0F98D" pageId="3" pageNumber="35" type="reference_group">
<paragraph id="FA6AD67C6A5EFFE4FF23FFC1FAC5FA6D" blockId="3.[151,1436,1428,1612]" pageId="3" pageNumber="35">
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<taxonomicName id="3DD5ADFF6A5EFFE4FF23FFC1FDE3FA2D" ID-CoL="3TJS4" authority="Parry, 1872: 80" authorityName="Parry" authorityPageNumber="80" authorityYear="1872" box="[151,547,1428,1452]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="C8A10A6E6A5EFFE4FF23FFC1FEB5FA2A" box="[151,373,1429,1452]" italics="true" pageId="3" pageNumber="35">Leptinopterus affinis</emphasis>
<treatmentCitation id="7B74F06D6A5EFFE4FEC8FFC0FDE3FA2D" author="Parry" box="[380,547,1428,1451]" page="80" pageId="3" pageNumber="35" year="1872">
<bibRefCitation id="9E44AB8D6A5EFFE4FEC8FFC0FE3BFA2D" author="Parry" box="[380,507,1428,1451]" pageId="3" pageNumber="44" refString="Parry, F. J. S. (1872) Descriptions of new species of lucanoid Coleoptera: with remarks on the genus Cantharolethrus, and supplementary lists. Transactions of the Royal Entomological Society of London, series 1, volume 4, 73 - 84." type="journal article" year="1872">Parry, 1872</bibRefCitation>
: 80
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</taxonomicName>
, plate 1 (description and illustration);
<treatmentCitation id="7B74F06D6A5EFFE4FC0AFFC0FAD6FA2D" author="Heyne" box="[958,1302,1428,1451]" page="54" pageId="3" pageNumber="35" year="1908">
<bibRefCitation id="9E44AB8D6A5EFFE4FC0AFFC0FB2DFA2D" author="Heyne" box="[958,1261,1428,1451]" pageId="3" pageNumber="43" refString="Heyne, A. &amp; Taschenberg, O. (1908) Die exotischen Kafer in Wort und Bild. G. Reusche, Leipzig, 262 + L pp." type="journal volume" year="1908">Heyne &amp; Taschenberg 1908</bibRefCitation>
: 54
</treatmentCitation>
;
<treatmentCitation id="7B74F06D6A5EFFE4FA95FFC0FEF8FA4D" author="Luderwaldt" page="475" pageId="3" pageNumber="35" year="1935">
<bibRefCitation id="9E44AB8D6A5EFFE4FA95FFC0FEC0FA4D" author="Luderwaldt" pageId="3" pageNumber="44" refString="Luderwaldt, H. (1935) Monographia dos Lucanideos brasileiros. Revista do Museu Paulista, 19, 448 - 574." type="journal article" year="1935">Lüderwaldt 1935</bibRefCitation>
: 475
</treatmentCitation>
, plate 2;
<treatmentCitation id="7B74F06D6A5EFFE4FE14FFE0FD61FA4D" author="Blackwelder" box="[416,673,1460,1483]" page="196" pageId="3" pageNumber="35" year="1944">
<bibRefCitation id="9E44AB8D6A5EFFE4FE14FFE0FDA9FA4D" author="Blackwelder" box="[416,617,1460,1483]" pageId="3" pageNumber="43" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 265." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
: 196
</treatmentCitation>
;
<treatmentCitation id="7B74F06D6A5EFFE4FD1BFFE0FC14FA4D" author="Didier" box="[687,980,1460,1483]" page="128" pageId="3" pageNumber="35" year="1953">
<bibRefCitation id="9E44AB8D6A5EFFE4FD1BFFE0FC5FFA4D" author="Didier" box="[687,927,1460,1483]" pageId="3" pageNumber="43" refString="Didier, R. &amp; Seguy, E. (1953) Catalogue illustre des Lucanides du Globe. Texte. Encyclopedie Entomologique, Serie A 27, 1 - 223." type="journal article" year="1953">Didier &amp; Séguy 1953</bibRefCitation>
: 128
</treatmentCitation>
(462);
<treatmentCitation id="7B74F06D6A5EFFE4FB9CFFE0FB22FA4D" author="Benesh" box="[1064,1250,1460,1483]" page="51" pageId="3" pageNumber="35" year="1960">
<bibRefCitation id="9E44AB8D6A5EFFE4FB9CFFE0FB78FA4D" author="Benesh" box="[1064,1208,1460,1483]" pageId="3" pageNumber="43" refString="Benesh, B. (1960) Lucanidea. Coleopterorum Catalogus Supplementa (second edition), 8, 1 - 178." type="journal article" year="1960">Benesh 1960</bibRefCitation>
: 51
</treatmentCitation>
;
<treatmentCitation id="7B74F06D6A5EFFE4FB45FFE0FA58FA4D" author="Maes" box="[1265,1432,1460,1483]" page="62" pageId="3" pageNumber="35" year="1992">
<bibRefCitation id="9E44AB8D6A5EFFE4FB45FFE0FAADFA4D" author="Maes" box="[1265,1389,1460,1483]" pageId="3" pageNumber="44" refString="Maes, J. M. (1992) Lista de los Lucanidae (Coleoptera) del mundo. Revista Nicaraguense de Entomologia, 22, 1 - 121." type="journal article" year="1992">Maes 1992</bibRefCitation>
: 62
</treatmentCitation>
;
<treatmentCitation id="7B74F06D6A5EFFE4FF73FF80FDD1FA6D" author="Mizunuma" box="[199,529,1492,1515]" page="278" pageId="3" pageNumber="35" year="1994">
<bibRefCitation id="9E44AB8D6A5EFFE4FF73FF80FE1CFA6D" author="Mizunuma" box="[199,476,1492,1515]" pageId="3" pageNumber="44" refString="Mizunuma, T. &amp; Nagai, S. (1994) The Lucanid Beetles of the World. Mushi-sha Iconographic Series of Insects, volume 1. Mushi-sha, Tokyo, 338 pp." type="book" year="1994">Mizunuma &amp; Nagai 1994</bibRefCitation>
: 278
</treatmentCitation>
, (116);
<treatmentCitation id="7B74F06D6A5EFFE4FDD6FF80FCD9FA6D" author="Krajcik" box="[610,793,1492,1515]" page="27" pageId="3" pageNumber="35" year="2001">
<bibRefCitation id="9E44AB8D6A5EFFE4FDD6FF80FD32FA6D" author="Krajcik" box="[610,754,1492,1515]" pageId="3" pageNumber="43" refString="Krajcik, M. (2001) Lucanidae of the World. Catalogue-Part I. Checklist of the Stag Beetles of the World. Milan Krajcik, Most, Czech Republic, 108 pp." type="book" year="2001">Krajcik 2001</bibRefCitation>
: 27
</treatmentCitation>
;
<treatmentCitation id="7B74F06D6A5EFFE4FC91FF80FC1DFA6D" author="Krajcik" box="[805,989,1492,1515]" page="93" pageId="3" pageNumber="35" year="2003">
<bibRefCitation id="9E44AB8D6A5EFFE4FC91FF80FC75FA6D" author="Krajcik" box="[805,949,1492,1515]" pageId="3" pageNumber="43" refString="Krajcik, M. (2003) Lucanidae of the World. Catalog-Part II. Encyclopedia of the Lucanidae (Coleoptera: Lucanidae). Milan Krajcik, Most, Czech Republic, 197 pp." type="book" year="2003">Krajcik 2003</bibRefCitation>
: 93
</treatmentCitation>
;
<treatmentCitation id="7B74F06D6A5EFFE4FC5CFF80FB43FA6D" author="Baba" box="[1000,1155,1492,1515]" page="20" pageId="3" pageNumber="35" year="2003">
<bibRefCitation id="9E44AB8D6A5EFFE4FC5CFF80FB9EFA6D" author="Baba" box="[1000,1118,1492,1515]" pageId="3" pageNumber="43" refString="Baba, M. (2003) Notes on the genus Leptinopterus (Coleoptera, Lucanidae) from South America. Gekkan-Mushi, 390, 19 - 30." type="journal article" year="2003">Baba 2003</bibRefCitation>
: 20
</treatmentCitation>
(
<figureCitation id="62EECAF96A5EFFE4FB25FF80FB39FA6D" box="[1169,1273,1492,1515]" captionStart="FIGURES 1 4" captionStartId="2.[151,272,1976,2000]" captionTargetBox="[341,1243,204,1940]" captionTargetId="figure@2.[331,1255,195,1953]" captionTargetPageId="2" captionText="FIGURES 1 4. Dorsal habitus of Leptinopterus species: 1 and 3) L. asketus, male and female; 2 and 4) L. assimilis, male and female." httpUri="https://zenodo.org/record/189229/files/figure.png" pageId="3" pageNumber="35">Figs. 14</figureCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA6AD67C6A5EFFE4FF23FFA0FCC0F98D" blockId="3.[151,1436,1428,1612]" box="[151,768,1524,1548]" pageId="3" pageNumber="35">
<emphasis id="C8A10A6E6A5EFFE4FF23FFA0FE85F98A" bold="true" box="[151,325,1524,1548]" pageId="3" pageNumber="35">nomen nudum:</emphasis>
<taxonomicName id="3DD5ADFF6A5EFFE4FEF8FFA1FD3CF98D" authority="Parry 1864: 85" authorityName="Parry" authorityPageNumber="85" authorityYear="1864" box="[332,764,1524,1548]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="C8A10A6E6A5EFFE4FEF8FFA1FD93F98A" box="[332,595,1525,1548]" italics="true" pageId="3" pageNumber="35">Leptinopterus pulchellus</emphasis>
<bibRefCitation id="9E44AB8D6A5EFFE4FDEFFFA0FD14F98D" author="Parry" box="[603,724,1524,1547]" pageId="3" pageNumber="44" refString="Parry, F. J. S. (1864) A catalogue of lucanoid Coleoptera; with illustrations and descriptions of various new and interesting species. Transactions of the Entomological Society of London, series 3, volume 2, 1 - 113." type="journal article" year="1864">Parry 1864</bibRefCitation>
: 85
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B2CF85F76A5EFFE4FF23FC40FC65F9CD" pageId="3" pageNumber="35" type="description">
<paragraph id="FA6AD67C6A5EFFE4FF23FC40FC7DF9AD" blockId="3.[151,1436,1428,1612]" box="[151,957,1556,1580]" pageId="3" pageNumber="35">
<emphasis id="C8A10A6E6A5EFFE4FF23FC40FE85F9AA" bold="true" box="[151,325,1556,1580]" pageId="3" pageNumber="35">nomen nudum:</emphasis>
<taxonomicName id="3DD5ADFF6A5EFFE4FEF8FC41FD50F9AD" authority="Klug" authorityName="Klug" box="[332,656,1556,1580]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="pulchellus">
<emphasis id="C8A10A6E6A5EFFE4FEF8FC41FD93F9AA" box="[332,595,1557,1580]" italics="true" pageId="3" pageNumber="35">Leptinopterus pulchellus</emphasis>
Klug
</taxonomicName>
(ined.)
<bibRefCitation id="9E44AB8D6A5EFFE4FD50FC40FC43F9AD" author="Nonfried" box="[740,899,1556,1579]" pageId="3" pageNumber="44" refString="Nonfried, A. F. (1891) Verzeichniss der Lucaniden, beschrieben von 1875 bis Ende des Jahres 1889. Deutsche Entomologische Zeitschrift, 1891, 277 - 281." type="journal article" year="1891">Nonfried 1891</bibRefCitation>
: 281.
</paragraph>
<paragraph id="FA6AD67C6A5EFFE4FF23FC60FC65F9CD" blockId="3.[151,1436,1428,1612]" box="[151,933,1588,1612]" pageId="3" pageNumber="35">
<emphasis id="C8A10A6E6A5EFFE4FF23FC60FE85F9CA" bold="true" box="[151,325,1588,1612]" pageId="3" pageNumber="35">nomen nudum:</emphasis>
<emphasis id="C8A10A6E6A5EFFE4FEF8FC61FD6AF9CA" box="[332,682,1589,1612]" italics="true" pageId="3" pageNumber="35">
<taxonomicName id="3DD5ADFF6A5EFFE4FEF8FC61FDEFF9CA" box="[332,559,1589,1612]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidus">Leptinopterus nitidus</taxonomicName>
ab. lepidus
</emphasis>
<bibRefCitation id="9E44AB8D6A5EFFE4FD05FC60FCABF9CD" author="Luderwaldt" box="[689,875,1588,1611]" pageId="3" pageNumber="43" refString="Luderwaldt, H. (1930) Novos Lucanideos brasileiros (Col.). Boletim do Museu Nacional, Rio de Janeiro, 6 (2), 123 - 127." type="journal article" year="1930">Lüderwaldt 1930</bibRefCitation>
: 124.
</paragraph>
</subSubSection>
<subSubSection id="B2CF85F76A5EFFE4FF23FC28FB0EF877" pageId="3" pageNumber="35" type="reference_group">
<paragraph id="FA6AD67C6A5EFFE4FF23FC28FB57F84D" blockId="3.[151,1437,1659,2033]" pageId="3" pageNumber="35">
<taxonomicName id="3DD5ADFF6A5EFFE4FF23FC28FE0FF913" ID-CoL="3TJS4" authority="Parry" authorityName="Parry" box="[151,463,1659,1685]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="C8A10A6E6A5EFFE4FF23FC28FE49F913" box="[151,393,1660,1685]" italics="true" pageId="3" pageNumber="35">Leptinopterus affinis</emphasis>
Parry
</taxonomicName>
and
<taxonomicName id="3DD5ADFF6A5EFFE4FDB3FC28FCC3F913" ID-CoL="3TJSS" authority="Luderwaldt" authorityName="Luderwaldt" box="[519,771,1659,1685]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="C8A10A6E6A5EFFE4FDB3FC28FDB6F913" box="[519,630,1660,1685]" italics="true" pageId="3" pageNumber="35">L. nitidus</emphasis>
Lüderwaldt
</taxonomicName>
are very similar except for the femoral coloration, which is the main character that can be used to distinguish these two species. The femora in
<taxonomicName id="3DD5ADFF6A5EFFE4FB15FCF6FACCF93D" ID-CoL="3TJS4" box="[1185,1292,1698,1723]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="C8A10A6E6A5EFFE4FB15FCF6FACCF93D" box="[1185,1292,1698,1723]" italics="true" pageId="3" pageNumber="35">L. affinis</emphasis>
</taxonomicName>
are entirely dark yellow, while in specimens of
<taxonomicName id="3DD5ADFF6A5EFFE4FD8EFC9DFD6AF964" ID-CoL="3TJSS" box="[570,682,1737,1762]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="C8A10A6E6A5EFFE4FD8EFC9DFD6AF964" box="[570,682,1737,1762]" italics="true" pageId="3" pageNumber="35">L. nitidus</emphasis>
</taxonomicName>
they are totally black. After examining the
<typeStatus id="256E68DE6A5EFFE4FB1BFC9DFB2DF965" box="[1199,1261,1737,1763]" pageId="3" pageNumber="35">types</typeStatus>
of Lüderwaldt at MZSP, I concluded that “
<emphasis id="C8A10A6E6A5EFFE4FE6FFCA4FD11F88F" box="[475,721,1776,1801]" italics="true" pageId="3" pageNumber="35">
<taxonomicName id="3DD5ADFF6A5EFFE4FE6FFCA4FD89F88F" ID-CoL="3TJSS" box="[475,585,1776,1801]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidus">L. nitidus</taxonomicName>
ab. lepidus
</emphasis>
<bibRefCitation id="9E44AB8D6A5EFFE4FD6CFCBBFC6AF88F" author="Luderwaldt" box="[728,938,1775,1801]" pageId="3" pageNumber="43" refString="Luderwaldt, H. (1930) Novos Lucanideos brasileiros (Col.). Boletim do Museu Nacional, Rio de Janeiro, 6 (2), 123 - 127." type="journal article" year="1930">Lüderwaldt, 1930</bibRefCitation>
” is actually
<taxonomicName id="3DD5ADFF6A5EFFE4FB8EFCA4FB62F88F" ID-CoL="3TJS4" box="[1082,1186,1776,1801]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="C8A10A6E6A5EFFE4FB8EFCA4FB62F88F" box="[1082,1186,1776,1801]" italics="true" pageId="3" pageNumber="35">L. affinis</emphasis>
</taxonomicName>
, and not
<taxonomicName id="3DD5ADFF6A5EFFE4FABAFCA4FABCF88F" ID-CoL="3TJSS" box="[1294,1404,1776,1801]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="C8A10A6E6A5EFFE4FABAFCA4FABCF88F" box="[1294,1404,1776,1801]" italics="true" pageId="3" pageNumber="35">L. nitidus</emphasis>
</taxonomicName>
as previously concluded by
<bibRefCitation id="9E44AB8D6A5EFFE4FE67FD42FD49F8B6" author="Benesh" box="[467,649,1814,1840]" pageId="3" pageNumber="43" refString="Benesh, B. (1960) Lucanidea. Coleopterorum Catalogus Supplementa (second edition), 8, 1 - 178." type="journal article" year="1960">Benesh (1960)</bibRefCitation>
and
<bibRefCitation id="9E44AB8D6A5EFFE4FD73FD42FC43F8B6" author="Krajcik" box="[711,899,1814,1840]" pageId="3" pageNumber="43" refString="Krajcik, M. (2001) Lucanidae of the World. Catalogue-Part I. Checklist of the Stag Beetles of the World. Milan Krajcik, Most, Czech Republic, 108 pp." type="book" year="2001">Krajcik (2001)</bibRefCitation>
. In addition, some specimens identified by
<bibRefCitation id="9E44AB8D6A5EFFE4FF23FD69FEBBF8D1" author="Luderwaldt" box="[151,379,1853,1879]" pageId="3" pageNumber="44" refString="Luderwaldt, H. (1935) Monographia dos Lucanideos brasileiros. Revista do Museu Paulista, 19, 448 - 574." type="journal article" year="1935">Lüderwaldt (1935)</bibRefCitation>
as
<emphasis id="C8A10A6E6A5EFFE4FE11FD69FDDDF8D0" box="[421,541,1853,1878]" italics="true" pageId="3" pageNumber="35">L. v-niger</emphasis>
(Westwood) from Santa Catarina State were also
<taxonomicName id="3DD5ADFF6A5EFFE4FBCBFD69FB2CF8D0" ID-CoL="3TJS4" box="[1151,1260,1853,1878]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="C8A10A6E6A5EFFE4FBCBFD69FB2CF8D0" box="[1151,1260,1853,1878]" italics="true" pageId="3" pageNumber="35">L. affinis</emphasis>
</taxonomicName>
and
<taxonomicName id="3DD5ADFF6A5EFFE4FA9DFD69FA5DF8D0" ID-CoL="3TJSS" box="[1321,1437,1853,1878]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="C8A10A6E6A5EFFE4FA9DFD69FA5DF8D0" box="[1321,1437,1853,1878]" italics="true" pageId="3" pageNumber="35">L. nitidus</emphasis>
</taxonomicName>
specimens.
<emphasis id="C8A10A6E6A5EFFE4FE96FD30FDE5F8FB" box="[290,549,1892,1917]" italics="true" pageId="3" pageNumber="35">
<taxonomicName id="3DD5ADFF6A5EFFE4FE96FD30FE07F8FB" ID-CoL="62X9P" box="[290,455,1892,1917]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="genus">Leptinopterus</taxonomicName>
v-niger
</emphasis>
has its distribution restricted to eastern
<collectingCountry id="82C296EC6A5EFFE4FBB2FD37FB92F8FB" box="[1030,1106,1891,1917]" name="Brazil" pageId="3" pageNumber="35">Brazil</collectingCountry>
, not extending as far south, and is also not morphologically similar to the other species cited here. The elytral pattern is variable within these species and is not a good character for the identification of
<taxonomicName id="3DD5ADFF6A5EFFE4FC24FDE5FBF3F84C" ID-CoL="62X9P" box="[912,1075,1969,1994]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="C8A10A6E6A5EFFE4FC24FDE5FBF3F84C" box="[912,1075,1969,1994]" italics="true" pageId="3" pageNumber="35">Leptinopterus</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="FA6AD67C6A5EFFE4FF72FD8CFB0EF877" blockId="3.[151,1437,1659,2033]" box="[198,1230,2007,2033]" pageId="3" pageNumber="35">
<taxonomicName id="3DD5ADFF6A5EFFE4FF72FD8CFE79F877" ID-CoL="3TJS4" box="[198,441,2008,2033]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="C8A10A6E6A5EFFE4FF72FD8CFE79F877" box="[198,441,2008,2033]" italics="true" pageId="3" pageNumber="35">Leptinopterus affinis</emphasis>
</taxonomicName>
and
<taxonomicName id="3DD5ADFF6A5EFFE4FE47FD8CFDA2F877" ID-CoL="3TJSS" box="[499,610,2008,2033]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="C8A10A6E6A5EFFE4FE47FD8CFDA2F877" box="[499,610,2008,2033]" italics="true" pageId="3" pageNumber="35">L. nitidus</emphasis>
</taxonomicName>
are sympatric, found in the southern states of
<collectingCountry id="82C296EC6A5EFFE4FBCBFD83FB0AF877" box="[1151,1226,2007,2033]" name="Brazil" pageId="3" pageNumber="35">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>