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<mods:title id="B9E0E4BB5701B00AABF922243A9CCCB6">Description of two new species of Leptinopterus Hope (Coleoptera: Lucanidae: Lucaninae) with notes on the taxonomy and natural history of the genus</mods:title>
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<heading id="A12261106A5CFFE6FF23FE45FD4DFBAC" bold="true" box="[151,653,1040,1067]" fontSize="11" level="1" pageId="1" pageNumber="33" reason="1">
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<emphasis id="C8A10A6E6A5CFFE6FF23FE45FE58FBAD" bold="true" box="[151,408,1041,1067]" italics="true" pageId="1" pageNumber="33">Leptinopterus tibialis</emphasis>
(
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(
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)
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<emphasis id="C8A10A6E6A5CFFE6FF23FED1FE89FB1A" box="[151,329,1157,1180]" italics="true" pageId="1" pageNumber="33">Lucanus tibialis</emphasis>
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<bibRefCitation id="9E44AB8D6A5CFFE6FEE2FED1FDE3FB1A" author="Eschscholtz" box="[342,547,1157,1180]" pageId="1" pageNumber="43" refString="Eschscholtz, J. F. (1822) Entomographien, Erste Lieferung. O. Reimer, Berlin. 128 pp." type="book" year="1822">Eschscholtz, 1822</bibRefCitation>
: 5
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,
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, male (illustration);
<treatmentCitation id="7B74F06D6A5CFFE6FC3FFED1FBB3FB1A" author="Eschscholtz" box="[907,1139,1157,1180]" page="4" pageId="1" pageNumber="33" year="1835">
<bibRefCitation id="9E44AB8D6A5CFFE6FC3FFED1FB91FB1A" author="Eschscholtz" box="[907,1105,1157,1180]" pageId="1" pageNumber="43" refString="Eschscholtz, J. F. (1835) Descriptions d'insects nouveaux. Oeuvres Entomologique, 1, 3 - 9." type="journal article" year="1835">Eschscholtz 1835</bibRefCitation>
: 4
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;
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<bibRefCitation id="9E44AB8D6A5CFFE6FB31FED1FAFBFB1A" author="Westwood" box="[1157,1339,1157,1180]" pageId="1" pageNumber="44" refString="Westwood, J. O. (1845) A Catalogue of the Lucanoid Coleoptera in the Collection of the Rev. F. W. Hope, M. A., F. R. S., &amp; c., together with Descriptions of the New Species Therein Contained. J. C. Bridgewater, London, 31 pp." type="book" year="1845">Westwood 1845</bibRefCitation>
: 5
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(51);
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<bibRefCitation id="9E44AB8D6A5CFFE6FF73FEF1FEBEFB3A" author="Burmeister" box="[199,382,1189,1212]" pageId="1" pageNumber="43" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie, volume 5. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1847">Burmeister 1847</bibRefCitation>
: 379
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;
<treatmentCitation id="7B74F06D6A5CFFE6FE75FEF1FD42FB3A" author="Harold" box="[449,642,1189,1212]" page="954" pageId="1" pageNumber="33" year="1868">
<bibRefCitation id="9E44AB8D6A5CFFE6FE75FEF1FD8BFB3A" author="Harold" box="[449,587,1189,1212]" pageId="1" pageNumber="43" refString="Harold, E. von (1868) Lucanidae, pp. 941 - 978. In: M. Gemminger and E. Harold (eds.), Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus, volume 3. E. H. Gummi, Munich." type="journal volume" year="1868">Harold 1868</bibRefCitation>
: 954
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;
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<bibRefCitation id="9E44AB8D6A5CFFE6FD3AFEF1FC7EFB3A" author="Heyne" box="[654,958,1189,1212]" pageId="1" pageNumber="43" refString="Heyne, A. &amp; Taschenberg, O. (1908) Die exotischen Kafer in Wort und Bild. G. Reusche, Leipzig, 262 + L pp." type="journal volume" year="1908">Heyne &amp; Taschenberg 1908</bibRefCitation>
: 54
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, plate 8;
<treatmentCitation id="7B74F06D6A5CFFE6FBFFFEF1FAD5FB3A" author="Boileau" box="[1099,1301,1189,1212]" page="237" pageId="1" pageNumber="33" year="1913">
<bibRefCitation id="9E44AB8D6A5CFFE6FBFFFEF1FB1FFB3A" author="Boileau" box="[1099,1247,1189,1212]" pageId="1" pageNumber="43" refString="Boileau, M. H. (1913) Note sur Lucanides conserves dans les collections de l'Universites d'Oxford et du British Museum. Transactions of the Royal Entomological Society of London, 2, 213 - 272" type="journal article" year="1913">Boileau 1913</bibRefCitation>
: 237
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;
<treatmentCitation id="7B74F06D6A5CFFE6FA95FEF1FEF7FB5A" author="Luderwaldt" page="124" pageId="1" pageNumber="33" year="1930">
<bibRefCitation id="9E44AB8D6A5CFFE6FA95FEF1FEC0FB5A" author="Luderwaldt" pageId="1" pageNumber="43" refString="Luderwaldt, H. (1930) Novos Lucanideos brasileiros (Col.). Boletim do Museu Nacional, Rio de Janeiro, 6 (2), 123 - 127." type="journal article" year="1930">Lüderwaldt 1930</bibRefCitation>
: 124
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125;
<treatmentCitation id="7B74F06D6A5CFFE6FEC9FE91FDB2FB5A" author="Luderwaldt" box="[381,626,1221,1244]" page="492" pageId="1" pageNumber="33" year="1935">
<bibRefCitation id="9E44AB8D6A5CFFE6FEC9FE91FDFBFB5A" author="Luderwaldt" box="[381,571,1221,1244]" pageId="1" pageNumber="44" refString="Luderwaldt, H. (1935) Monographia dos Lucanideos brasileiros. Revista do Museu Paulista, 19, 448 - 574." type="journal article" year="1935">Lüderwaldt 1935</bibRefCitation>
: 492
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494, plate 3;
<treatmentCitation id="7B74F06D6A5CFFE6FCA1FE91FBD8FB5A" author="Blackwelder" box="[789,1048,1221,1244]" page="196" pageId="1" pageNumber="33" year="1944">
<bibRefCitation id="9E44AB8D6A5CFFE6FCA1FE91FC1FFB5A" author="Blackwelder" box="[789,991,1221,1244]" pageId="1" pageNumber="43" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin, 185, 189 - 265." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
: 196
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;
<treatmentCitation id="7B74F06D6A5CFFE6FB93FE91FA90FB5A" author="Didier" box="[1063,1360,1221,1244]" page="131" pageId="1" pageNumber="33" year="1953">
<bibRefCitation id="9E44AB8D6A5CFFE6FB93FE91FADAFB5A" author="Didier" box="[1063,1306,1221,1244]" pageId="1" pageNumber="43" refString="Didier, R. &amp; Seguy, E. (1953) Catalogue illustre des Lucanides du Globe. Texte. Encyclopedie Entomologique, Serie A 27, 1 - 223." type="journal article" year="1953">Didier &amp; Séguy 1953</bibRefCitation>
: 131
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(486);
<treatmentCitation id="7B74F06D6A5CFFE6FF73FEB1FE4DFB7A" author="Martinez" box="[199,397,1253,1276]" page="43" pageId="1" pageNumber="33" year="1953">
<bibRefCitation id="9E44AB8D6A5CFFE6FF73FEB1FEA6FB7A" author="Martinez" box="[199,358,1253,1276]" pageId="1" pageNumber="44" refString="Martinez, A. (1953) Insectos nuevos o poco conocidos X. Algunas notas sobre Lucanidos con descripcion de una especie nueva y el alotipo de Sclerostomus tucumanus Nagel (Col. Scarabaeoid. Lucanidae). Revista de la Sociedad Entomologica Argentina, 16, 42 - 48." type="journal article" year="1953">Martinez 1953</bibRefCitation>
: 43
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;
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<bibRefCitation id="9E44AB8D6A5CFFE6FE2CFEB1FDE6FB7A" author="Benesh" box="[408,550,1253,1276]" pageId="1" pageNumber="43" refString="Benesh, B. (1960) Lucanidea. Coleopterorum Catalogus Supplementa (second edition), 8, 1 - 178." type="journal article" year="1960">Benesh 1960</bibRefCitation>
: 53
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54;
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<bibRefCitation id="9E44AB8D6A5CFFE6FD34FEB1FCFAFB7A" author="Klausnitzer" box="[640,826,1253,1276]" pageId="1" pageNumber="43" refString="Klausnitzer, B. (1982) Hirschkafer oder Schroter. Lucanidae. A. Ziemsen Verlag, Wittenberg Lutherstadt, 83 pp." type="book" year="1982">Klausnitzer 1982</bibRefCitation>
: 43
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;
<treatmentCitation id="7B74F06D6A5CFFE6FCDFFEB1FBCCFB7A" author="Maes" box="[875,1036,1253,1276]" page="64" pageId="1" pageNumber="33" year="1992">
<bibRefCitation id="9E44AB8D6A5CFFE6FCDFFEB1FC25FB7A" author="Maes" box="[875,997,1253,1276]" pageId="1" pageNumber="44" refString="Maes, J. M. (1992) Lista de los Lucanidae (Coleoptera) del mundo. Revista Nicaraguense de Entomologia, 22, 1 - 121." type="journal article" year="1992">Maes 1992</bibRefCitation>
: 64
</treatmentCitation>
;
<treatmentCitation id="7B74F06D6A5CFFE6FBA2FEB1FA9DFB7A" author="Mizunuma" box="[1046,1373,1253,1276]" page="278" pageId="1" pageNumber="33" year="1994">
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: 278
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, plate 116;
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: 28
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;
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: 94
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;
<treatmentCitation id="7B74F06D6A5CFFE6FD35FF51FCDDFA9A" author="Baba" box="[641,797,1285,1308]" page="28" pageId="1" pageNumber="33" year="2003">
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: 28
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(Figs. 3845).
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</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF23FF70FD2FFABA" blockId="1.[151,1436,1157,1564]" box="[151,751,1316,1340]" pageId="1" pageNumber="33">
<emphasis id="C8A10A6E6A5CFFE6FF23FF70FEC6FABA" bold="true" box="[151,262,1316,1340]" pageId="1" pageNumber="33">synonym:</emphasis>
<taxonomicName id="3DD5ADFF6A5CFFE6FEB8FF71FD2FFABA" ID-CoL="6PFPR" authority="Thomson 1862: 397" authorityName="Thomson" authorityPageNumber="397" authorityYear="1862" box="[268,751,1317,1340]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="femoratus">
<emphasis id="C8A10A6E6A5CFFE6FEB8FF71FDD1FABA" box="[268,529,1317,1340]" italics="true" pageId="1" pageNumber="33">Leptinopterus femoratus</emphasis>
<bibRefCitation id="9E44AB8D6A5CFFE6FDADFF71FD7DFABA" author="Thomson" box="[537,701,1317,1340]" pageId="1" pageNumber="44" refString="Thomson, J. (1862) Catalogue des Lucanides. Annales de la Societe Entomologique de France, (4 th series), 2, 392 - 436." type="journal article" year="1862">Thomson 1862</bibRefCitation>
: 397
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="FA6AD67C6A5CFFE6FF23FF10FC6AFADA" blockId="1.[151,1436,1157,1564]" box="[151,938,1348,1372]" pageId="1" pageNumber="33">
<emphasis id="C8A10A6E6A5CFFE6FF23FF10FE85FADA" bold="true" box="[151,325,1348,1372]" pageId="1" pageNumber="33">nomen nudum:</emphasis>
<taxonomicName id="3DD5ADFF6A5CFFE6FEF8FF11FD55FADA" authority="Hope" authorityName="Hope" box="[332,661,1349,1372]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="ochraceus">
<emphasis id="C8A10A6E6A5CFFE6FEF8FF11FD94FADA" box="[332,596,1349,1372]" italics="true" pageId="1" pageNumber="33">Leptinopterus ochraceus</emphasis>
Hope
</taxonomicName>
(ined.),
<bibRefCitation id="9E44AB8D6A5CFFE6FD5AFF11FCB8FADA" author="Harold" box="[750,888,1349,1372]" pageId="1" pageNumber="43" refString="Harold, E. von (1868) Lucanidae, pp. 941 - 978. In: M. Gemminger and E. Harold (eds.), Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus, volume 3. E. H. Gummi, Munich." type="journal volume" year="1868">Harold 1868</bibRefCitation>
: 954
</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF23FF30FC06FAFA" blockId="1.[151,1436,1157,1564]" box="[151,966,1380,1404]" pageId="1" pageNumber="33">
<emphasis id="C8A10A6E6A5CFFE6FF23FF30FE85FAFA" bold="true" box="[151,325,1380,1404]" pageId="1" pageNumber="33">nomen nudum:</emphasis>
<taxonomicName id="3DD5ADFF6A5CFFE6FEF8FF31FD67FAFA" authority="Hope" authorityName="Hope" box="[332,679,1381,1404]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="ochropterus">
<emphasis id="C8A10A6E6A5CFFE6FEF8FF31FDA6FAFA" box="[332,614,1381,1404]" italics="true" pageId="1" pageNumber="33">Leptinopterus ochropterus</emphasis>
Hope
</taxonomicName>
(ined.),
<bibRefCitation id="9E44AB8D6A5CFFE6FCB5FF31FC53FAFA" author="Boileau" box="[769,915,1381,1404]" pageId="1" pageNumber="43" refString="Boileau, M. H. (1913) Note sur Lucanides conserves dans les collections de l'Universites d'Oxford et du British Museum. Transactions of the Royal Entomological Society of London, 2, 213 - 272" type="journal article" year="1913">Boileau 1913</bibRefCitation>
: 237
</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF23FFD0FC52FA1A" blockId="1.[151,1436,1157,1564]" box="[151,914,1412,1436]" pageId="1" pageNumber="33">
<emphasis id="C8A10A6E6A5CFFE6FF23FFD0FE85FA1A" bold="true" box="[151,325,1412,1436]" pageId="1" pageNumber="33">nomen nudum:</emphasis>
<emphasis id="C8A10A6E6A5CFFE6FEF8FFD1FD5EFA1A" box="[332,670,1413,1436]" italics="true" pageId="1" pageNumber="33">
<taxonomicName id="3DD5ADFF6A5CFFE6FEF8FFD1FDF0FA1A" box="[332,560,1413,1436]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="tibialis">Leptinopterus tibialis</taxonomicName>
ab. fulvus
</emphasis>
<bibRefCitation id="9E44AB8D6A5CFFE6FD11FFD1FC9FFA1A" author="Luderwaldt" box="[677,863,1413,1436]" pageId="1" pageNumber="43" refString="Luderwaldt, H. (1930) Novos Lucanideos brasileiros (Col.). Boletim do Museu Nacional, Rio de Janeiro, 6 (2), 123 - 127." type="journal article" year="1930">Lüderwaldt 1930</bibRefCitation>
: 125
</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF23FFF0FC75FA3A" blockId="1.[151,1436,1157,1564]" box="[151,949,1444,1468]" pageId="1" pageNumber="33">
<emphasis id="C8A10A6E6A5CFFE6FF23FFF0FE85FA3A" bold="true" box="[151,325,1444,1468]" pageId="1" pageNumber="33">nomen nudum:</emphasis>
<emphasis id="C8A10A6E6A5CFFE6FEF8FFF1FD08FA3A" box="[332,712,1445,1468]" italics="true" pageId="1" pageNumber="33">
<taxonomicName id="3DD5ADFF6A5CFFE6FEF8FFF1FDF0FA3A" box="[332,560,1445,1468]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="tibialis">Leptinopterus tibialis</taxonomicName>
ab. castaneus
</emphasis>
<bibRefCitation id="9E44AB8D6A5CFFE6FD7BFFF1FC49FA3A" author="Luderwaldt" box="[719,905,1445,1468]" pageId="1" pageNumber="43" refString="Luderwaldt, H. (1930) Novos Lucanideos brasileiros (Col.). Boletim do Museu Nacional, Rio de Janeiro, 6 (2), 123 - 127." type="journal article" year="1930">Lüderwaldt 1930</bibRefCitation>
:125
</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF23FF90FC52FA5A" blockId="1.[151,1436,1157,1564]" box="[151,914,1476,1500]" pageId="1" pageNumber="33">
<emphasis id="C8A10A6E6A5CFFE6FF23FF90FE85FA5A" bold="true" box="[151,325,1476,1500]" pageId="1" pageNumber="33">nomen nudum:</emphasis>
<taxonomicName id="3DD5ADFF6A5CFFE6FEF8FF91FC52FA5A" authority="Luderwaldt 1935: 493" authorityName="Luderwaldt" authorityPageNumber="493" authorityYear="1935" box="[332,914,1477,1500]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="variety" species="tibialis" variety="bucki">
<emphasis id="C8A10A6E6A5CFFE6FEF8FF91FD5FFA5A" box="[332,671,1477,1500]" italics="true" pageId="1" pageNumber="33">Leptinopterus tibialis var. bucki</emphasis>
<bibRefCitation id="9E44AB8D6A5CFFE6FD11FF91FC9FFA5A" author="Luderwaldt" box="[677,863,1477,1500]" pageId="1" pageNumber="44" refString="Luderwaldt, H. (1935) Monographia dos Lucanideos brasileiros. Revista do Museu Paulista, 19, 448 - 574." type="journal article" year="1935">Lüderwaldt 1935</bibRefCitation>
: 493
</taxonomicName>
</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF23FFB0FCE9FA7A" blockId="1.[151,1436,1157,1564]" box="[151,809,1508,1532]" pageId="1" pageNumber="33">
<emphasis id="C8A10A6E6A5CFFE6FF23FFB0FEC6FA7A" bold="true" box="[151,262,1508,1532]" pageId="1" pageNumber="33">synonym:</emphasis>
<taxonomicName id="3DD5ADFF6A5CFFE6FEB8FFB1FCE9FA7A" authority="Luderwaldt, 1930: 124" authorityName="Luderwaldt" authorityPageNumber="124" authorityYear="1930" box="[268,809,1509,1532]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="C8A10A6E6A5CFFE6FEB8FFB1FDEEFA7A" box="[268,558,1509,1532]" italics="true" pageId="1" pageNumber="33">Leptinopterus rotundicollis</emphasis>
<bibRefCitation id="9E44AB8D6A5CFFE6FD81FFB1FD37FA7A" author="Luderwaldt" box="[565,759,1509,1532]" pageId="1" pageNumber="43" refString="Luderwaldt, H. (1930) Novos Lucanideos brasileiros (Col.). Boletim do Museu Nacional, Rio de Janeiro, 6 (2), 123 - 127." type="journal article" year="1930">Lüderwaldt, 1930</bibRefCitation>
: 124
</taxonomicName>
</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF23FC50FC7AF99A" blockId="1.[151,1436,1157,1564]" box="[151,954,1540,1564]" pageId="1" pageNumber="33">
<emphasis id="C8A10A6E6A5CFFE6FF23FC50FEC6F99A" bold="true" box="[151,262,1540,1564]" pageId="1" pageNumber="33">synonym:</emphasis>
<taxonomicName id="3DD5ADFF6A5CFFE6FEB8FC51FDEEF99A" box="[268,558,1541,1564]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="C8A10A6E6A5CFFE6FEB8FC51FDEEF99A" box="[268,558,1541,1564]" italics="true" pageId="1" pageNumber="33">Leptinopterus luederwaldti</emphasis>
</taxonomicName>
de Moraes
<emphasis id="C8A10A6E6A5CFFE6FD1FFC51FD00F99A" box="[683,704,1541,1564]" italics="true" pageId="1" pageNumber="33">in</emphasis>
<bibRefCitation id="9E44AB8D6A5CFFE6FD72FC51FC48F99A" author="Luderwaldt" box="[710,904,1541,1564]" pageId="1" pageNumber="44" refString="Luderwaldt, H. (1935) Monographia dos Lucanideos brasileiros. Revista do Museu Paulista, 19, 448 - 574." type="journal article" year="1935">Lüderwaldt, 1935</bibRefCitation>
: 506
</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF23FC18FD09F8A1" blockId="1.[151,1436,1612,1986]" pageId="1" pageNumber="33">
<taxonomicName id="3DD5ADFF6A5CFFE6FF23FC18FE5DF9E3" box="[151,413,1612,1637]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="C8A10A6E6A5CFFE6FF23FC18FE5DF9E3" box="[151,413,1612,1637]" italics="true" pageId="1" pageNumber="33">Leptinopterus tibialis</emphasis>
</taxonomicName>
is probably the most common species in the genus, found especially from southern Brazilian States to southern São Paulo State. Based on the examination of the
<typeStatus id="256E68DE6A5CFFE6FB8CFC26FBABF90A" box="[1080,1131,1650,1676]" pageId="1" pageNumber="33">type</typeStatus>
specimens, two available names,
<taxonomicName id="3DD5ADFF6A5CFFE6FF41FCCDFD86F935" authority="Luderwaldt" authorityName="Luderwaldt" box="[245,582,1689,1715]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="C8A10A6E6A5CFFE6FF41FCCDFE71F934" box="[245,433,1689,1714]" italics="true" pageId="1" pageNumber="33">L. rotundicollis</emphasis>
Lüderwaldt
</taxonomicName>
and
<taxonomicName id="3DD5ADFF6A5CFFE6FD30FCCDFC80F934" box="[644,832,1689,1714]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="C8A10A6E6A5CFFE6FD30FCCDFC80F934" box="[644,832,1689,1714]" italics="true" pageId="1" pageNumber="33">L. luederwaldti</emphasis>
</taxonomicName>
de Moraes are here synonymized with
<taxonomicName id="3DD5ADFF6A5CFFE6FA91FCCDFEF6F95C" authority="Eschscholtz" authorityName="Eschscholtz" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="C8A10A6E6A5CFFE6FA91FCCDFA5CF934" box="[1317,1436,1689,1714]" italics="true" pageId="1" pageNumber="33">L. tibialis</emphasis>
(Eschscholtz)
</taxonomicName>
<emphasis id="C8A10A6E6A5CFFE6FE8AFCEBFDC3F95F" bold="true" box="[318,515,1727,1753]" italics="true" pageId="1" pageNumber="33">new synonymies</emphasis>
. The two species here synonymized were both described from Santa Catarina State, where hundreds of
<taxonomicName id="3DD5ADFF6A5CFFE6FE0BFCB3FDEEF886" box="[447,558,1767,1792]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="C8A10A6E6A5CFFE6FE0BFCB3FDEEF886" box="[447,558,1767,1792]" italics="true" pageId="1" pageNumber="33">L. tibialis</emphasis>
</taxonomicName>
have been either collected or examined from the main Brazilian
<taxonomicName id="3DD5ADFF6A5CFFE6FA96FCB2FA5CF886" box="[1314,1436,1766,1792]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="family">Lucanidae</taxonomicName>
collections (MZSP, MNRJ, IOC, DZUP, MCN).
</paragraph>
<paragraph id="FA6AD67C6A5CFFE6FF72FD60FEE0F844" blockId="1.[151,1436,1612,1986]" pageId="1" pageNumber="33">
The
<typeStatus id="256E68DE6A5CFFE6FF49FD60FEA3F8C8" box="[253,355,1844,1870]" pageId="1" pageNumber="33" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3DD5ADFF6A5CFFE6FE39FD60FD83F8CB" box="[397,579,1844,1869]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="33" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="C8A10A6E6A5CFFE6FE39FD60FD83F8CB" box="[397,579,1844,1869]" italics="true" pageId="1" pageNumber="33">L. rotundicollis</emphasis>
</taxonomicName>
is a very small female (less than
<quantity id="3D2D7B996A5CFFE6FC63FD60FBEBF8C8" box="[983,1067,1844,1870]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="33" unit="mm" value="13.0">13 mm</quantity>
in length), and was apparently found dead. The specimen lacks the left mandible, right antennal flagellum, right protibia, three last anterior left tarsomeres, four last median left and right tarsomeres, left metatarsus, last right metatarsomere, and some tibial spurs.
</paragraph>
<caption id="AEAA86F46A5FFFE5FF23FDECFE8FF874" httpUri="https://zenodo.org/record/189229/files/figure.png" pageId="2" pageNumber="34" targetBox="[341,1243,204,1940]" targetPageId="2">
<paragraph id="FA6AD67C6A5FFFE5FF23FDECFE8FF874" blockId="2.[151,1436,1976,2034]" pageId="2" pageNumber="34">
<emphasis id="C8A10A6E6A5FFFE5FF23FDECFE86F856" bold="true" box="[151,326,1976,2000]" pageId="2" pageNumber="34">FIGURES 14.</emphasis>
Dorsal habitus of
<taxonomicName id="3DD5ADFF6A5FFFE5FDA0FDECFD6CF849" box="[532,684,1976,1999]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="C8A10A6E6A5FFFE5FDA0FDECFD6CF849" box="[532,684,1976,1999]" italics="true" pageId="2" pageNumber="34">Leptinopterus</emphasis>
</taxonomicName>
species: 1 and 3)
<taxonomicName id="3DD5ADFF6A5FFFE5FCC0FDECFC21F849" box="[884,993,1976,1999]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="asketus">
<emphasis id="C8A10A6E6A5FFFE5FCC0FDECFC21F849" box="[884,993,1976,1999]" italics="true" pageId="2" pageNumber="34">L. asketus</emphasis>
</taxonomicName>
, male and female; 2 and 4)
<taxonomicName id="3DD5ADFF6A5FFFE5FAACFDECFA55F849" box="[1304,1429,1976,1999]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="34" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="C8A10A6E6A5FFFE5FAACFDECFA55F849" box="[1304,1429,1976,1999]" italics="true" pageId="2" pageNumber="34">L. assimilis</emphasis>
</taxonomicName>
, male and female.
</paragraph>
</caption>
<caption id="AEAA86F46A5EFFE4FF23F947FEB7FCCB" httpUri="https://zenodo.org/record/189230/files/figure.png" pageId="3" pageNumber="35" targetBox="[463,1100,226,749]" targetPageId="3">
<paragraph id="FA6AD67C6A5EFFE4FF23F947FEB7FCCB" blockId="3.[151,1437,787,845]" pageId="3" pageNumber="35">
<emphasis id="C8A10A6E6A5EFFE4FF23F947FE86FCAD" bold="true" box="[151,326,787,811]" pageId="3" pageNumber="35">FIGURES 58.</emphasis>
Left mandible of male majors of four species of
<taxonomicName id="3DD5ADFF6A5EFFE4FCD5F947FC38FCAC" box="[865,1016,787,810]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="C8A10A6E6A5EFFE4FCD5F947FC38FCAC" box="[865,1016,787,810]" italics="true" pageId="3" pageNumber="35">Leptinopterus</emphasis>
</taxonomicName>
: 5)
<taxonomicName id="3DD5ADFF6A5EFFE4FB91F947FB62FCAC" box="[1061,1186,787,810]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="C8A10A6E6A5EFFE4FB91F947FB62FCAC" box="[1061,1186,787,810]" italics="true" pageId="3" pageNumber="35">L. assimilis</emphasis>
</taxonomicName>
; 6)
<taxonomicName id="3DD5ADFF6A5EFFE4FB7BF947FA98FCAC" box="[1231,1368,787,810]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="femoratus">
<emphasis id="C8A10A6E6A5EFFE4FB7BF947FA98FCAC" box="[1231,1368,787,810]" italics="true" pageId="3" pageNumber="35">L. femoratus</emphasis>
</taxonomicName>
; 7)
<taxonomicName id="3DD5ADFF6A5EFFE4FA32F947FF21FCCB" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="C8A10A6E6A5EFFE4FA32F947FF21FCCB" italics="true" pageId="3" pageNumber="35">L. tibialis</emphasis>
</taxonomicName>
; 8)
<taxonomicName id="3DD5ADFF6A5EFFE4FEB8F962FEB0FCCB" box="[268,368,822,845]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="C8A10A6E6A5EFFE4FEB8F962FEB0FCCB" box="[268,368,822,845]" italics="true" pageId="3" pageNumber="35">L. nitidus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="FA6AD67C6A5EFFE4FF72F92FFB66FC65" blockId="3.[151,1437,891,1265]" pageId="3" pageNumber="35">
The species name of
<taxonomicName id="3DD5ADFF6A5EFFE4FE75F92FFDB6FC12" box="[449,630,891,916]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="C8A10A6E6A5EFFE4FE75F92FFDB6FC12" box="[449,630,891,916]" italics="true" pageId="3" pageNumber="35">L. rotundicollis</emphasis>
</taxonomicName>
was given with reference to the rounded sides of pronotum, lacking the lateral angles. This condition is commonly found in small female specimens of
<taxonomicName id="3DD5ADFF6A5EFFE4FB30F9F6FAEAFC3D" box="[1156,1322,930,955]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="C8A10A6E6A5EFFE4FB30F9F6FAEAFC3D" box="[1156,1322,930,955]" italics="true" pageId="3" pageNumber="35">Leptinopterus</emphasis>
</taxonomicName>
, and was also observed in
<taxonomicName id="3DD5ADFF6A5EFFE4FEE8F99DFE0DFC64" box="[348,461,969,994]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="C8A10A6E6A5EFFE4FEE8F99DFE0DFC64" box="[348,461,969,994]" italics="true" pageId="3" pageNumber="35">L. tibialis</emphasis>
</taxonomicName>
. Larger females always have well pronounced median angles.
</paragraph>
<paragraph id="FA6AD67C6A5EFFE4FF72F9BBFE37FB77" blockId="3.[151,1437,891,1265]" pageId="3" pageNumber="35">
Since its description, no additional specimens of
<taxonomicName id="3DD5ADFF6A5EFFE4FCA4F9A4FC05FB8F" box="[784,965,1008,1033]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="C8A10A6E6A5EFFE4FCA4F9A4FC05FB8F" box="[784,965,1008,1033]" italics="true" pageId="3" pageNumber="35">L. luederwaldti</emphasis>
</taxonomicName>
have been collected, which is certainly due to the fact that the single specimen of
<taxonomicName id="3DD5ADFF6A5EFFE4FD1DFE42FCA7FBA9" box="[681,871,1046,1071]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="C8A10A6E6A5EFFE4FD1DFE42FCA7FBA9" box="[681,871,1046,1071]" italics="true" pageId="3" pageNumber="35">L. luederwaldti</emphasis>
</taxonomicName>
is a damaged female of
<taxonomicName id="3DD5ADFF6A5EFFE4FB2CFE42FAD0FBA9" box="[1176,1296,1046,1071]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="C8A10A6E6A5EFFE4FB2CFE42FAD0FBA9" box="[1176,1296,1046,1071]" italics="true" pageId="3" pageNumber="35">L. tibialis</emphasis>
</taxonomicName>
. There is a handwritten label below the
<typeStatus id="256E68DE6A5EFFE4FE46FE69FD9CFBD1" box="[498,604,1085,1111]" pageId="3" pageNumber="35" type="holotype">holotype</typeStatus>
indicating “
<emphasis id="C8A10A6E6A5EFFE4FD47FE69FCAFFBD0" box="[755,879,1085,1110]" italics="true" pageId="3" pageNumber="35">Leptinopt.</emphasis>
female symbol / sp. nov. / Lueder. det. 1930” which probably induced de Moraes to describe the specimen in the Lüderwaldt monograph after his death. The specimen is very oily, dark colored, and somewhat shiny, but clearly morphologically a specimen of
<emphasis id="C8A10A6E6A5EFFE4FA30FEDEFF36FB4C" italics="true" pageId="3" pageNumber="35">
<taxonomicName id="3DD5ADFF6A5EFFE4FA30FEDEFF32FB4C" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="species" species="tibialis">L. tibialis</taxonomicName>
.
</emphasis>
Fatty specimens are commonly found in
<taxonomicName id="3DD5ADFF6A5EFFE4FD4EFEE5FC65FB4C" box="[762,933,1201,1226]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="C8A10A6E6A5EFFE4FD4EFEE5FC65FB4C" box="[762,933,1201,1226]" italics="true" pageId="3" pageNumber="35">Leptinopterus</emphasis>
</taxonomicName>
probably because of their usual habit of feeding on sap flows or fruits.
</paragraph>
</subSubSection>
</treatment>
</document>