271 lines
32 KiB
XML
271 lines
32 KiB
XML
<document id="9881644F9A227E663F996F4D2AC4A2D9" ID-DOI="10.5281/zenodo.189228" ID-GBIF-Dataset="29e7002a-ee86-4b7b-804f-2e95f678e6a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="189228" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461100418165" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho" docDate="2009" docId="727C676A6A55FFEDFF23F88EFC87FABC" docLanguage="en" docName="zt02172p044.pdf" docOrigin="Zootaxa 2172" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptinopterus assimilis Grossi, new species" docType="treatment" docVersion="9" lastPageNumber="42" masterDocId="8E451F126A5DFFE7FFB4FA54FFC0FF86" masterDocTitle="Description of two new species of Leptinopterus Hope (Coleoptera: Lucanidae: Lucaninae) with notes on the taxonomy and natural history of the genus" masterLastPageNumber="44" masterPageNumber="32" pageNumber="40" updateTime="1698591109854" updateUser="plazi">
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<mods:mods id="0E0CD64E58C36D8A59AD42A2DF918A06" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="147721C24B0AA1562BCFE9997B4C177F">
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<mods:title id="F26ADF7D5CF420CFC15E932DA5604B4E">Description of two new species of Leptinopterus Hope (Coleoptera: Lucanidae: Lucaninae) with notes on the taxonomy and natural history of the genus</mods:title>
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</mods:titleInfo>
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<mods:name id="BF0393B6637E58A0B35FD66C48DBAEFA" type="personal">
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<mods:role id="FD2852D440DD6123871857225F227D9A">
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<mods:roleTerm id="2EDB92AF56A9E8737C2D2C089367EAA2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1B69FA0C13E90FF809CF1059328B1C06">Grossi, Paschoal Coelho</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="369ED21B7255492BF1B520ABF430AC73">text</mods:typeOfResource>
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<mods:titleInfo id="C7EB0FF7016C39787E641393D48D1BF2">
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<mods:title id="EEB821089C56B7DF9BA929D4CE2F3D36">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0F70BC5A528A6503192E08303613E3EA">
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<mods:date id="B61001138328FFAF03929447963A1185">2009</mods:date>
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<mods:detail id="EEEBD9E6CFADC8A19C558470090581DB" type="volume">
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<mods:number id="42BF56AB4694761374D24936F3BC523A">2172</mods:number>
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</mods:detail>
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<mods:extent id="3A20181793ECBD029A5FE3334E28A038" unit="page">
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<mods:start id="2BB13DB6825AC22B239FB59AAF585F4A">32</mods:start>
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<mods:end id="5BC1687F2C4D50E6A64B6BE580D9B466">44</mods:end>
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<mods:classification id="DF438EB3BA4C8A61FC01977CCB10FF54">journal article</mods:classification>
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<mods:identifier id="474B8FDD70ADDBD843EAC0390685597A" type="DOI">10.5281/zenodo.189228</mods:identifier>
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<mods:identifier id="D574B21F1C196ECC3E2A412B3F3ABA26" type="GBIF-Dataset">29e7002a-ee86-4b7b-804f-2e95f678e6a3</mods:identifier>
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<mods:identifier id="DCDA741FA1B2F26AF5807107A06D20C5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="FE98694C94B291872A8A440A30561D82" type="Zenodo-Dep">189228</mods:identifier>
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</mods:mods>
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<treatment id="727C676A6A55FFEDFF23F88EFC87FABC" ID-DOI="http://doi.org/10.5281/zenodo.6224806" ID-GBIF-Taxon="119621722" ID-Zenodo-Dep="6224806" LSID="urn:lsid:plazi:treatment:727C676A6A55FFEDFF23F88EFC87FABC" httpUri="http://treatment.plazi.org/id/727C676A6A55FFEDFF23F88EFC87FABC" lastPageId="10" lastPageNumber="42" pageId="8" pageNumber="40">
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<subSubSection id="B2CF85F76A55FFEFFF23F88EFE78FC91" pageId="8" pageNumber="40" type="nomenclature">
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||
<paragraph id="FA6AD67C6A55FFEFFF23F88EFD5DFD72" blockId="8.[151,669,730,791]" box="[151,669,730,756]" pageId="8" pageNumber="40">
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||
<heading id="A12261106A55FFEFFF23F88EFD5DFD72" bold="true" box="[151,669,730,756]" fontSize="11" level="1" pageId="8" pageNumber="40" reason="1">
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||
<emphasis id="C8A10A6E6A55FFEFFF23F88EFD5DFD72" bold="true" box="[151,669,730,756]" pageId="8" pageNumber="40">
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<taxonomicName id="3DD5ADFF6A55FFEFFF23F88EFDC6FD72" ID-CoL="3TJS6" authority="Grossi" authorityName="Grossi" box="[151,518,730,756]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="40" phylum="Arthropoda" rank="species" species="assimilis" status="sp. nov.">
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<emphasis id="C8A10A6E6A55FFEFFF23F88EFE6EFD72" bold="true" box="[151,430,730,756]" italics="true" pageId="8" pageNumber="40">Leptinopterus assimilis</emphasis>
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Grossi
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</taxonomicName>
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<taxonomicNameLabel id="D392B7156A55FFEFFDB9F88EFD5DFD72" box="[525,669,730,756]" pageId="8" pageNumber="40" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="FA6AD67C6A55FFEFFF23F8A9FE78FC91" blockId="8.[151,669,730,791]" box="[151,440,765,791]" pageId="8" pageNumber="40">
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||
(
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<figureCitation id="62EECAF96A55FFEFFF2BF8A9FED0FC91" box="[159,272,765,791]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,272,1976,2000]" captionTargetBox="[341,1243,204,1940]" captionTargetId="figure@2.[331,1255,195,1953]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Dorsal habitus of Leptinopterus species: 1 and 3) L. asketus, male and female; 2 and 4) L. assimilis, male and female." httpUri="https://zenodo.org/record/189229/files/figure.png" pageId="8" pageNumber="40">Figs. 2, 4</figureCitation>
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,
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<figureCitation id="62EECAF96A55FFEFFEA8F8A9FEECFC91" box="[284,300,765,791]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,271,787,811]" captionTargetBox="[463,1100,226,749]" captionTargetId="figure@3.[445,1143,194,763]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Left mandible of male majors of four species of Leptinopterus: 5) L. assimilis; 6) L. femoratus; 7) L. tibialis; 8) L. nitidus." httpUri="https://zenodo.org/record/189230/files/figure.png" pageId="8" pageNumber="40">5</figureCitation>
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,
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<figureCitation id="62EECAF96A55FFEFFE8DF8A9FE98FC91" box="[313,344,765,791]" captionStart="FIGURES 15 – 20" captionStartId="7.[151,269,1879,1903]" captionTargetBox="[239,1363,210,1831]" captionTargetId="figure@7.[210,1382,197,1841]" captionTargetPageId="7" captionText="FIGURES 15 – 20. Ventral and lateral views of the aedeagus of the new species and ventral view of median lobe: 15 and 16) Aedeagus of L. asketus; 18 and 19) Aedeagus of L. assimilis. (Figs. 15 and 18, setae and apex of internal sac omitted). Scale bar = 1 mm. 17) Ventral view of median lobe of L. asketus; 20) Ventral view of median lobe of L. assimilis. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/189232/files/figure.png" pageId="8" pageNumber="40">18</figureCitation>
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, 19, 20)
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</paragraph>
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</subSubSection>
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<subSubSection id="B2CF85F76A55FFEDFF23F91EFE27FD70" lastPageId="10" lastPageNumber="42" pageId="8" pageNumber="40" type="materials_examined">
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<paragraph id="FA6AD67C6A55FFEFFF23F91EFEC9FBA0" blockId="8.[151,1437,842,2028]" pageId="8" pageNumber="40">
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<emphasis id="C8A10A6E6A55FFEFFF23F91EFCCCFCE2" bold="true" box="[151,780,842,868]" pageId="8" pageNumber="40">
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<typeStatus id="256E68DE6A55FFEFFF23F91EFF15FCE2" box="[151,213,842,868]" pageId="8" pageNumber="40">Type</typeStatus>
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material.
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<typeStatus id="256E68DE6A55FFEFFEECF91EFE0CFCE2" box="[344,460,842,868]" pageId="8" pageNumber="40" type="holotype">Holotype</typeStatus>
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male labeled (dissected):
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</emphasis>
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<collectingCountry id="82C296EC6A55FFEFFCA1F91EFC9CFCE2" box="[789,860,842,868]" name="Brazil" pageId="8" pageNumber="40">Brasil</collectingCountry>
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Paraná, Piraquara, Mananciais da Serra,
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<date id="8E6BF0BC6A55FFEFFAFFF91EFF15FC0D" pageId="8" pageNumber="40" value="2005-11-23">23-XI- 2005</date>
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,
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<quantity id="3D2D7B996A55FFEFFF56F925FEFFFC0D" box="[226,319,881,907]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="40" unit="m" value="1000.0">1000 m</quantity>
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, P. L. Pastori col.
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<emphasis id="C8A10A6E6A55FFEFFDADF924FC8CFC0C" bold="true" box="[537,844,880,906]" pageId="8" pageNumber="40">Allotype female labeled:</emphasis>
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Data as
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<typeStatus id="256E68DE6A55FFEFFC02F925FBDEFC0D" box="[950,1054,881,907]" pageId="8" pageNumber="40" type="holotype">holotype</typeStatus>
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except for
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<date id="8E6BF0BC6A55FFEFFB12F925FA92FC0D" box="[1190,1362,881,907]" pageId="8" pageNumber="40" value="2005-10-24" valueMax="2005-10-29" valueMin="2005-10-24">24-29-X-2005</date>
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, L. C. Rocha Filho col., Arm. Emergência.
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<emphasis id="C8A10A6E6A55FFEFFDF9F9C3FCB3FC37" bold="true" box="[589,883,919,945]" pageId="8" pageNumber="40">
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Two
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<typeStatus id="256E68DE6A55FFEFFD3FF9C3FCC7FC37" box="[651,775,919,945]" pageId="8" pageNumber="40" type="paratype">paratypes</typeStatus>
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labeled:
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</emphasis>
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Same data as
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<typeStatus id="256E68DE6A55FFEFFBAAF9CCFB45FC34" box="[1054,1157,920,946]" pageId="8" pageNumber="40" type="holotype">holotype</typeStatus>
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and allotype except
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<date id="8E6BF0BC6A55FFEFFAC8F9CCFF15FC5E" pageId="8" pageNumber="40" value="2005-10">X- 2005</date>
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, P. C. Grossi col. Both
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<typeStatus id="256E68DE6A55FFEFFE50F9EAFD94FC5E" box="[484,596,958,984]" pageId="8" pageNumber="40" type="paratype">paratypes</typeStatus>
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are fragments of males; one
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<typeStatus id="256E68DE6A55FFEFFC13F9EAFBCCFC5E" box="[935,1036,958,984]" pageId="8" pageNumber="40" type="paratype">paratype</typeStatus>
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consists of a head and the other is an almost complete body, lacking antennal funicle, tarsi, and abdomen (with the remaining parts all glued together).
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</paragraph>
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<paragraph id="FA6AD67C6A55FFEFFF72FE66FB3DFBF5" blockId="8.[151,1437,842,2028]" pageId="8" pageNumber="40">
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<typeStatus id="256E68DE6A55FFEFFF72FE66FEF3FBCA" box="[198,307,1074,1100]" pageId="8" pageNumber="40" type="holotype">Holotype</typeStatus>
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and allotype ex
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<collectionCode id="9CC44EB96A55FFEFFDB1FE66FD90FBCA" box="[517,592,1074,1100]" pageId="8" pageNumber="40">EPGC</collectionCode>
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deposited at Coleção de Entomologia Pe. J. S. Moure, Universidade Federal do Paraná, Curitiba, Paraná,
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<collectingCountry id="82C296EC6A55FFEFFDF2FE0DFD4EFBF5" box="[582,654,1113,1139]" name="Brazil" pageId="8" pageNumber="40">Brazil</collectingCountry>
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(
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<collectionCode id="9CC44EB96A55FFEFFD29FE0DFD30FBF5" box="[669,752,1113,1139]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="8" pageNumber="40">DZUP</collectionCode>
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).
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<typeStatus id="256E68DE6A55FFEFFCB0FE0DFCB6FBF5" box="[772,886,1113,1139]" pageId="8" pageNumber="40" type="paratype">Paratypes</typeStatus>
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at
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<collectionCode id="9CC44EB96A55FFEFFC2DFE0DFC27FBF5" box="[921,999,1113,1139]" pageId="8" pageNumber="40">EPGC</collectionCode>
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, Nova Friburgo,
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<collectingCountry id="82C296EC6A55FFEFFB1AFE0DFB39FBF5" box="[1198,1273,1113,1139]" name="Brazil" pageId="8" pageNumber="40">Brazil</collectingCountry>
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.
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</paragraph>
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<paragraph id="FA6AD67C6A55FFEEFF72FE2BFF2BFECA" blockId="8.[151,1437,842,2028]" lastBlockId="9.[151,1436,152,332]" lastPageId="9" lastPageNumber="41" pageId="8" pageNumber="40">
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<emphasis id="C8A10A6E6A55FFEFFF72FE2BFE18FB1F" bold="true" box="[198,472,1151,1177]" pageId="8" pageNumber="40">
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||
<typeStatus id="256E68DE6A55FFEFFF72FE2BFEF8FB1F" box="[198,312,1151,1177]" pageId="8" pageNumber="40" type="holotype">Holotype</typeStatus>
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description.
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</emphasis>
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Body somewhat elongate, dorsally slightly convex.
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<emphasis id="C8A10A6E6A55FFEFFBD1FE2BFB05FB1F" bold="true" box="[1125,1221,1151,1177]" italics="true" pageId="8" pageNumber="40">Length:</emphasis>
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27.0 mm.
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<emphasis id="C8A10A6E6A55FFEFFAE0FE2BFEF1FB46" bold="true" pageId="8" pageNumber="40">
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<emphasis id="C8A10A6E6A55FFEFFAE0FE2BFA5CFB1F" bold="true" box="[1364,1436,1151,1177]" italics="true" pageId="8" pageNumber="40">Width</emphasis>
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(pronotum):
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</emphasis>
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8.0 mm
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<emphasis id="C8A10A6E6A55FFEFFE2EFEF2FE28FB46" bold="true" box="[410,488,1190,1216]" italics="true" pageId="8" pageNumber="40">Color:</emphasis>
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Almost totally brown; mandibles, head and pronotum darker; margins, legs, and venter dark brown.
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<emphasis id="C8A10A6E6A55FFEFFEC8FE99FE2DFB61" bold="true" box="[380,493,1229,1255]" italics="true" pageId="8" pageNumber="40">Ve st it ur e:</emphasis>
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Dorsal surface of pronotum and elytra setose; setae light yellow, simple, denser on lateral edges.
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<emphasis id="C8A10A6E6A55FFEFFED1FEA0FE71FA88" bold="true" box="[357,433,1268,1294]" italics="true" pageId="8" pageNumber="40">Head:</emphasis>
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Shape transverse, 1.4 times wider than long; anterior margin depressed, somewhat elongate and concave. Surface microgranulate, disc weakly concave; laterally slightly convex. Frons widely excavated; excavation as wide as third of head. Around excavation, posteriorly finely punctate; punctures fine. Anterior sides with a black spot on each side. Anterior angles round, somewhat projected above mandibles. Ocular canthi angled; angles straight, laterally slightly rounded, protruding into eyes until first third. Gena surface granulate, sparsely punctate; punctures fine, becoming large anteriorly. Antennae with scattered setae on each antennomere. Club with apical antennomere rounded, glabrous basally, from middle to apex tomentose; two proximal club antennomeres less tomentose, transversely elongated, not round. Mandibles elongate, two times longer than head, apparently symmetric, flattened; basal tooth bifid, the left one widely, above it a short conical tooth; apical fourth with an elongated clearly bifid tooth, above it with an acutely backward-turned tooth; apex acute, incurved. Labrum transverse, roundly expanded at middle and with transverse row of setae. Mentum transverse, trapezoidal; surface granulate, anteriorly punctate, setose; punctures moderate, coarse, setae exceeding anterior margin.
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<emphasis id="C8A10A6E6A55FFEFFCC3FC90FC3CF958" bold="true" box="[887,1020,1732,1758]" italics="true" pageId="8" pageNumber="40">Pronotum:</emphasis>
|
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Form transverse, 1.84 times wider than long; disc almost smooth, finely punctate; punctures fine; near sides surface granulate and setose; punctures moderate, large. Anterior margin weakly projected, almost round; border incomplete. Anterior angles round; lateral angles obtuse; posterior angles acute and somewhat backwards pointed. Each side with an elongate black spot near median angles.
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<taxonomicName id="3DD5ADFF6A55FFEFFD10FD0AFCBDF8FE" authority="Shape" authorityName="Shape" box="[676,893,1886,1912]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="8" pageNumber="40" phylum="Arthropoda" rank="genus">
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||
<emphasis id="C8A10A6E6A55FFEFFD10FD0AFCEBF8FE" bold="true" box="[676,811,1886,1912]" italics="true" pageId="8" pageNumber="40">Scutellum:</emphasis>
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Shape
|
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</taxonomicName>
|
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round. Surface totally covered by long, light yellow setae.
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<emphasis id="C8A10A6E6A55FFEFFEF5FDD1FE5BF819" bold="true" box="[321,411,1925,1951]" italics="true" pageId="8" pageNumber="40">Elytra:</emphasis>
|
||
Shape elongate, approximately 1.3 times longer than head and pronotum together, dorsally convex, setose and finely punctate; punctures fine becoming bigger anteriorly, setae denser at borders, longer than puncture diameter. Elytral suture dark brown.
|
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<emphasis id="C8A10A6E6A55FFEFFC00FD86FC38F86A" bold="true" box="[948,1016,2002,2028]" italics="true" pageId="8" pageNumber="40">Legs:</emphasis>
|
||
Protibiae with three external teeth, proximally serrate until base. Mesotibiae and metatibiae disarmed with external apex acutely pointed, more so on mesotibiae. Arolia with two simple bristles.
|
||
<emphasis id="C8A10A6E6A54FFEEFD67FAEAFC8EFF5E" bold="true" box="[723,846,190,216]" italics="true" pageId="9" pageNumber="41">Genitalia:</emphasis>
|
||
Form simple. Genital capsule similar to those of other
|
||
<taxonomicName id="3DD5ADFF6A54FFEEFF6FFAB1FE95FF79" box="[219,341,229,255]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Lucaninae">Lucaninae</taxonomicName>
|
||
species. Aedeagus symmetric (
|
||
<figureCitation id="62EECAF96A54FFEEFD7DFAB1FC98FF79" box="[713,856,229,255]" captionStart="FIGURES 15 – 20" captionStartId="7.[151,269,1879,1903]" captionTargetBox="[239,1363,210,1831]" captionTargetId="figure@7.[210,1382,197,1841]" captionTargetPageId="7" captionText="FIGURES 15 – 20. Ventral and lateral views of the aedeagus of the new species and ventral view of median lobe: 15 and 16) Aedeagus of L. asketus; 18 and 19) Aedeagus of L. assimilis. (Figs. 15 and 18, setae and apex of internal sac omitted). Scale bar = 1 mm. 17) Ventral view of median lobe of L. asketus; 20) Ventral view of median lobe of L. assimilis. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/189232/files/figure.png" pageId="9" pageNumber="41">Figs. 18–19</figureCitation>
|
||
); everted internal sac as long as aedeagus; paired papillae flat. Median lobe (
|
||
<figureCitation id="62EECAF96A54FFEEFE68FB58FDF4FEA0" box="[476,564,268,294]" captionStart="FIGURES 15 – 20" captionStartId="7.[151,269,1879,1903]" captionTargetBox="[239,1363,210,1831]" captionTargetId="figure@7.[210,1382,197,1841]" captionTargetPageId="7" captionText="FIGURES 15 – 20. Ventral and lateral views of the aedeagus of the new species and ventral view of median lobe: 15 and 16) Aedeagus of L. asketus; 18 and 19) Aedeagus of L. assimilis. (Figs. 15 and 18, setae and apex of internal sac omitted). Scale bar = 1 mm. 17) Ventral view of median lobe of L. asketus; 20) Ventral view of median lobe of L. assimilis. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/189232/files/figure.png" pageId="9" pageNumber="41">Fig. 20</figureCitation>
|
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) with a bifurcate projection at middle; laterally with four folds, the third deeper.
|
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</paragraph>
|
||
<caption id="AEAA86F46A54FFEEFF23FDF8FEA9F865" httpUri="https://zenodo.org/record/189233/files/figure.png" pageId="9" pageNumber="41" targetBox="[226,1357,374,1937]" targetPageId="9">
|
||
<paragraph id="FA6AD67C6A54FFEEFF23FDF8FEA9F865" blockId="9.[151,1436,1964,2020]" pageId="9" pageNumber="41">
|
||
<emphasis id="C8A10A6E6A54FFEEFF23FDF8FE9EF842" bold="true" box="[151,350,1964,1988]" pageId="9" pageNumber="41">FIGURES 21–24.</emphasis>
|
||
<taxonomicName id="3DD5ADFF6A54FFEEFED0FDF9FE38F842" box="[356,504,1965,1988]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C8A10A6E6A54FFEEFED0FDF9FE38F842" box="[356,504,1965,1988]" italics="true" pageId="9" pageNumber="41">Leptinopterus</emphasis>
|
||
</taxonomicName>
|
||
species feeding at sap flow on
|
||
<taxonomicName id="3DD5ADFF6A54FFEEFCFCFDF8FC7DF845" box="[840,957,1964,1987]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="41" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||
species: 21)
|
||
<taxonomicName id="3DD5ADFF6A54FFEEFBFDFDF9FB6FF842" box="[1097,1199,1965,1988]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="41" phylum="Arthropoda" rank="species" species="tibialis">
|
||
<emphasis id="C8A10A6E6A54FFEEFBFDFDF9FB6FF842" box="[1097,1199,1965,1988]" italics="true" pageId="9" pageNumber="41">L. tibialis</emphasis>
|
||
</taxonomicName>
|
||
; 22)
|
||
<taxonomicName id="3DD5ADFF6A54FFEEFB5CFDF9FA8DF842" box="[1256,1357,1965,1988]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="41" phylum="Arthropoda" rank="species" species="nitidus">
|
||
<emphasis id="C8A10A6E6A54FFEEFB5CFDF9FA8DF842" box="[1256,1357,1965,1988]" italics="true" pageId="9" pageNumber="41">L. nitidus</emphasis>
|
||
</taxonomicName>
|
||
; 23)
|
||
<taxonomicName id="3DD5ADFF6A54FFEEFA32FDF9FF28F862" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="41" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="C8A10A6E6A54FFEEFA32FDF9FF28F862" italics="true" pageId="9" pageNumber="41">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
; 24)
|
||
<taxonomicName id="3DD5ADFF6A54FFEEFE94FD99FEA1F862" box="[288,353,1997,2020]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="41" phylum="Arthropoda" rank="species" species="fryi">
|
||
<emphasis id="C8A10A6E6A54FFEEFE94FD99FEA1F862" box="[288,353,1997,2020]" italics="true" pageId="9" pageNumber="41">L. fryi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA6AD67C6A57FFEDFF72FAC3FE27FD70" blockId="10.[151,1437,151,1338]" pageId="10" pageNumber="42">
|
||
<emphasis id="C8A10A6E6A57FFEDFF72FAC3FDE8FF37" bold="true" box="[198,552,151,177]" pageId="10" pageNumber="42">Allotype female description.</emphasis>
|
||
As male in general aspects as vestiture and coloration.
|
||
<emphasis id="C8A10A6E6A57FFEDFB76FAC3FAE1FF37" bold="true" box="[1218,1313,151,177]" italics="true" pageId="10" pageNumber="42">Length:</emphasis>
|
||
13.0 mm.
|
||
<emphasis id="C8A10A6E6A57FFEDFF23FAEAFF27FF5E" bold="true" box="[151,231,190,216]" italics="true" pageId="10" pageNumber="42">Width:</emphasis>
|
||
7.3 mm.
|
||
<emphasis id="C8A10A6E6A57FFEDFEE3FAEAFE65FF5E" bold="true" box="[343,421,190,216]" italics="true" pageId="10" pageNumber="42">Color:</emphasis>
|
||
Somewhat dark.
|
||
<emphasis id="C8A10A6E6A57FFEDFDC5FAEAFD24FF5E" bold="true" box="[625,740,190,216]" italics="true" pageId="10" pageNumber="42">Vestiture:</emphasis>
|
||
Setae yellowish white.
|
||
<emphasis id="C8A10A6E6A57FFEDFC4EFAEAFB85FF5E" bold="true" box="[1018,1093,190,216]" italics="true" pageId="10" pageNumber="42">Head:</emphasis>
|
||
Shape less transverse, nearly quadrate, 0.41 times wider than long. Surface totally punctate, anteriorly by deep coarse punctures with a strong ridged appearance among punctures; punctures setose. Near lateral and posterior margins with smaller punctures, moderate, large. Anterior and lateral margins black. Vertex slightly concave. Supra-antennal brow darkened, distinct and round. Anterior angles less projected and not above mandibles. Canthi weakly expanded, rounder and with straight external sides, protruding into eyes until first fifth. Mandibles smaller than head, wide, with apex truncate; internally with one bifid tooth on each; apical tooth acute and internally directed. Dorsal furrow shallow, moderately punctate; punctures moderate. Longitudinal carinae visible, internal carina incomplete just before apex; external carina complete, ending apically. Labrum elongate, somewhat triangular with coarse to coalescent setose punctures. Mentum transverse and totally coarsely punctate.
|
||
<emphasis id="C8A10A6E6A57FFEDFEB8F815FE53FDDD" bold="true" box="[268,403,577,603]" italics="true" pageId="10" pageNumber="42">Pronotum:</emphasis>
|
||
Shape more rounded and more convex. Lateral borders crenulate. Surface with larger punctures laterally. Anterior angles acute; median angles nearly round.
|
||
<emphasis id="C8A10A6E6A57FFEDFC6BF833FBF3FD07" bold="true" box="[991,1075,615,641]" italics="true" pageId="10" pageNumber="42">Elytra:</emphasis>
|
||
Shape somewhat shorter, 1.53 times longer than both head and pronotum together.
|
||
<emphasis id="C8A10A6E6A57FFEDFD42F8DAFCF9FD2E" bold="true" box="[758,825,654,680]" italics="true" pageId="10" pageNumber="42">Legs:</emphasis>
|
||
Protibiae wider and strongly externally dentate with 5 teeth. Mesotibiae with 1 large external tooth, above it another smaller tooth. Metatibiae with very small, median tooth, almost absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2CF85F76A57FFEDFF72F956FA59FCC5" pageId="10" pageNumber="42" type="etymology">
|
||
<paragraph id="FA6AD67C6A57FFEDFF72F956FA59FCC5" blockId="10.[151,1437,151,1338]" pageId="10" pageNumber="42">
|
||
<emphasis id="C8A10A6E6A57FFEDFF72F956FE93FC9A" bold="true" box="[198,339,770,796]" pageId="10" pageNumber="42">Etymology.</emphasis>
|
||
The specific epithet is derived from the Greek
|
||
<emphasis id="C8A10A6E6A57FFEDFC25F956FC60FC9D" box="[913,928,770,795]" italics="true" pageId="10" pageNumber="42">a</emphasis>
|
||
- meaning ‘not’ and from the Latin
|
||
<emphasis id="C8A10A6E6A57FFEDFAFBF956FA5BFC9D" box="[1359,1435,770,795]" italics="true" pageId="10" pageNumber="42">similis</emphasis>
|
||
meaning ‘like’ in reference to the male mandibles, which are different from all the other species of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2CF85F76A57FFEDFF72F91BFC87FABC" pageId="10" pageNumber="42" type="discussion">
|
||
<paragraph id="FA6AD67C6A57FFEDFF72F91BFC87FABC" blockId="10.[151,1437,151,1338]" pageId="10" pageNumber="42">
|
||
<emphasis id="C8A10A6E6A57FFEDFF72F91BFE80FCEF" bold="true" box="[198,320,847,873]" pageId="10" pageNumber="42">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="256E68DE6A57FFEDFEC9F904FE23FCEC" box="[381,483,848,874]" pageId="10" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
male was found dead on the ground and is lacking some appendages such as the right antennal funicle, 4–5 tarsomeres of the right middle leg, and 2–5 tarsomeres of right fore leg. The only known female was collected accidentally with an emergence trap used to collect bees on the ground. The other specimens are two male fragments collected at the foot of a
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFCE1F990FC0CFC5B" box="[853,972,964,989]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="42" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="C8A10A6E6A57FFEDFCE1F990FC0CFC5B" box="[853,972,964,989]" italics="true" pageId="10" pageNumber="42">Baccharis</emphasis>
|
||
</taxonomicName>
|
||
sp. shrub, indicating that this species is probably attracted by the sap of this species of
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFD78F9BEFC92FB82" box="[716,850,1002,1028]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="10" pageNumber="42" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
|
||
.
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFCEAF9BEFBB3FB85" box="[862,1139,1002,1027]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="assimilis">
|
||
<emphasis id="C8A10A6E6A57FFEDFCEAF9BEFBB3FB85" box="[862,1139,1002,1027]" italics="true" pageId="10" pageNumber="42">Leptinopterus assimilis</emphasis>
|
||
</taxonomicName>
|
||
has a very characteristic mandibular shape in males not found in any other species of the genus with a tooth between the bifurcate subapical tooth and the apex. Only very large males of
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFC9FFE6CFC5CFBD7" box="[811,924,1080,1105]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="nitidus">
|
||
<emphasis id="C8A10A6E6A57FFEDFC9FFE6CFC5CFBD7" box="[811,924,1080,1105]" italics="true" pageId="10" pageNumber="42">L. nitidus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="62EECAF96A57FFEDFC18FE6CFC33FBD4" box="[940,1011,1080,1106]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,271,787,811]" captionTargetBox="[463,1100,226,749]" captionTargetId="figure@3.[445,1143,194,763]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Left mandible of male majors of four species of Leptinopterus: 5) L. assimilis; 6) L. femoratus; 7) L. tibialis; 8) L. nitidus." httpUri="https://zenodo.org/record/189230/files/figure.png" pageId="10" pageNumber="42">Fig. 8</figureCitation>
|
||
) approach this condition but with a much smaller tooth. The condition of a bifurcate tooth near the mandibular apex is also found in
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFA9CFE0BFEF6FB19" authority="Eschscholtz" authorityName="Eschscholtz" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="tibialis">
|
||
<emphasis id="C8A10A6E6A57FFEDFA9CFE0BFA5CFBFE" box="[1320,1436,1119,1144]" italics="true" pageId="10" pageNumber="42">L. tibialis</emphasis>
|
||
(Eschscholtz)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="62EECAF96A57FFEDFEF3FED1FE50FB19" box="[327,400,1157,1183]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,271,787,811]" captionTargetBox="[463,1100,226,749]" captionTargetId="figure@3.[445,1143,194,763]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Left mandible of male majors of four species of Leptinopterus: 5) L. assimilis; 6) L. femoratus; 7) L. tibialis; 8) L. nitidus." httpUri="https://zenodo.org/record/189230/files/figure.png" pageId="10" pageNumber="42">Fig. 7</figureCitation>
|
||
),
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFE13FED1FCD5FB19" authority="Burmeister" authorityName="Burmeister" box="[423,789,1157,1183]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="erythrocnemus">
|
||
<emphasis id="C8A10A6E6A57FFEDFE13FED1FDB4FB18" box="[423,628,1157,1182]" italics="true" pageId="10" pageNumber="42">L. erythrocnemus</emphasis>
|
||
(Burmeister)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFC95FED1FBE5FB19" authority="Olivier" authorityName="Olivier" box="[801,1061,1157,1183]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="femoratus">
|
||
<emphasis id="C8A10A6E6A57FFEDFC95FED1FC76FB18" box="[801,950,1157,1182]" italics="true" pageId="10" pageNumber="42">L. femoratus</emphasis>
|
||
(Olivier)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="62EECAF96A57FFEDFB82FED1FBBFFB19" box="[1078,1151,1157,1183]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,271,787,811]" captionTargetBox="[463,1100,226,749]" captionTargetId="figure@3.[445,1143,194,763]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Left mandible of male majors of four species of Leptinopterus: 5) L. assimilis; 6) L. femoratus; 7) L. tibialis; 8) L. nitidus." httpUri="https://zenodo.org/record/189230/files/figure.png" pageId="10" pageNumber="42">Fig. 6</figureCitation>
|
||
),
|
||
<emphasis id="C8A10A6E6A57FFEDFB21FED1FB6DFB18" box="[1173,1197,1157,1182]" italics="true" pageId="10" pageNumber="42">L.</emphasis>
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFB01FED1FA58FB19" authority="Luderwaldt" authorityName="Luderwaldt" box="[1205,1432,1157,1183]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="nitidus">nitidus Lüderwaldt</taxonomicName>
|
||
, and in
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFF5CFEF8FE5AFB40" authority="Parry" authorityName="Parry" box="[232,410,1196,1222]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="C8A10A6E6A57FFEDFF5CFEF8FE91FB43" box="[232,337,1196,1221]" italics="true" pageId="10" pageNumber="42">L. affinis</emphasis>
|
||
Parry
|
||
</taxonomicName>
|
||
, but in the three first species the mandibles are asymmetrical. The new species is most similar to
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFEB9FE87FEB6FB6A" box="[269,374,1235,1260]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="C8A10A6E6A57FFEDFEB9FE87FEB6FB6A" box="[269,374,1235,1260]" italics="true" pageId="10" pageNumber="42">L. affinis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFE19FE87FDDCFB6A" box="[429,540,1235,1260]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="nitidus">
|
||
<emphasis id="C8A10A6E6A57FFEDFE19FE87FDDCFB6A" box="[429,540,1235,1260]" italics="true" pageId="10" pageNumber="42">L. nitidus</emphasis>
|
||
</taxonomicName>
|
||
, with which it shares the symmetry of mandibles, however
|
||
<taxonomicName id="3DD5ADFF6A57FFEDFB60FE87FA99FB6A" box="[1236,1369,1235,1260]" class="Insecta" family="Lucanidae" genus="Leptinopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="42" phylum="Arthropoda" rank="species" species="assimilis">
|
||
<emphasis id="C8A10A6E6A57FFEDFB60FE87FA99FB6A" box="[1236,1369,1235,1260]" italics="true" pageId="10" pageNumber="42">L. assimilis</emphasis>
|
||
</taxonomicName>
|
||
has a setose body in both sexes while the other two species are almost glabrous. They are sympatric and have been collected in the same orographic region in southern
|
||
<collectingCountry id="82C296EC6A57FFEDFD43FF74FC83FABC" box="[759,835,1312,1338]" name="Brazil" pageId="10" pageNumber="42">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |