treatments-xml/data/72/7B/0A/727B0AC3E6085C00A77F218EF6819BF7.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.128.1804" ID-PMC="PMC3175138" ID-Pensoft-Pub="1313-2970-128-15" ID-Pensoft-UUID="6157FFBE34536B410C79CB5C11170069" ID-PubMed="21998550" ID-Zenodo-Dep="577176" ID-ZooBank="837E761ADDC84351B1080F9B9CDC7ED1" ModsDocID="1313-2970-128-15" checkinTime="1451249919653" checkinUser="pensoft" docAuthor="Valim, Ye, Kavanaugh, David H., Shi, Hongliang &amp; Liang, Hongbin" docDate="2011" docId="727B0AC3E6085C00A77F218EF6819BF7" docLanguage="en" docName="ZooKeys 128: 15-52" docOrigin="ZooKeys 128" docPubDate="2011-09-09" docSource="http://dx.doi.org/10.3897/zookeys.128.1804" docTitle="agilis (Lithochlaenius) Chaudoir" docType="treatment" docVersion="3" id="6157FFBE34536B410C79CB5C11170069" lastPageNumber="27" masterDocId="6157FFBE34536B410C79CB5C11170069" masterDocTitle="A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species" masterLastPageNumber="52" masterPageNumber="15" pageNumber="26" updateTime="1668152074611" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Valim, Ye</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Key Laboratory for Plant Pests Management of Mountainous Region, Institute of Entomology, Guizhou, University, Guiyang 550025, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kavanaugh, David H.</mods:namePart>
<mods:affiliation>Department of Entomology, California Academy of Science, San Francisco, California 94118, U. S. A.</mods:affiliation>
<mods:nameIdentifier type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="pubDate">
<mods:number>2011-09-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>128</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>15</mods:start>
<mods:end>52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.128.1804</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.128.1804</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-128-15</mods:identifier>
<mods:identifier type="ZooBank">837E761ADDC84351B1080F9B9CDC7ED1</mods:identifier>
<mods:identifier type="Pensoft-UUID">6157FFBE34536B410C79CB5C11170069</mods:identifier>
<mods:identifier type="Zenodo-Dep">577176</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182236340" LSID="urn:lsid:plazi:treatment:727B0AC3E6085C00A77F218EF6819BF7" httpUri="http://treatment.plazi.org/id/727B0AC3E6085C00A77F218EF6819BF7" lastPageId="12" lastPageNumber="27" pageId="11" pageNumber="26">
<subSubSection pageId="11" pageNumber="26" type="nomenclature">
<paragraph pageId="11" pageNumber="26">
<taxonomicName LSID="727B0AC3-E608-5C00-A77F-218EF6819BF7" authority="agilis Chaudoir" authorityName="Chaudoir" genus="Agilis" lsidName="agilis (Lithochlaenius)" pageId="11" pageNumber="26" rank="genus">Chlaenius (Lithochlaenius) agilis Chaudoir</taxonomicName>
<figureCitation captionStart="Figures 22–27" captionStartId="F3" captionText="Figures 22 - 27. Habitus of Chlaenius (Lithochlaenius) spp 22 Chlaenius agilis Chaudoir, holotype, dorsal view 23 Chlaenius agilis Chaudoir, holotype, ventral view 24 Chlaenius formosensis Lorenz, male, in CCCC, dorsal view 25 Chlaenius formosensis Lorenz, ventral view 26 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, dorsal view 27 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17280" pageId="11" pageNumber="26">Figs 22-23</figureCitation>
<figureCitation captionStart="Figures 34–39" captionStartId="F5" captionText="Figures 34 - 39. Chlaenius (Lithochlaenius) spp 34 Chlaenius rambouseki Lutshnik, male, collected from Ussuri region, in IZCAS, dorsal view 35 Chlaenius rambouseki Lutshnik, male, collected from Ussuri region, in IZCAS, ventral view 36 Chlaenius anchomenoides Bates, cotype, male, in BMNH, dorsal view 37 Chlaenius anchomenoides Bates, cotype, male, in BMNH, ventral view 38 Chlaenius nuristanus Jedlicka, paratype, male, in MNHN, dorsal view 39 Chlaenius nuristanus Jedlicka, paratype, male, in MNHN, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17282" pageId="11" pageNumber="26">36</figureCitation>
<figureCitation captionStart="Figures 40–45" captionStartId="F6" captionText="Figures 40 - 45. Chlaenius (Lithochlaenius) spp 40 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, dorsal view 41 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, ventral view 42 Chlaenius agiloides Jedlicka, holotype, male, in NMPC, dorsal view 43 Chlaenius formosanus Jedlicka, holotype, male, in NMPC, dorsal view 44 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, dorsal view 45 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17283" pageId="11" pageNumber="26">41</figureCitation>
<figureCitation captionStart="Figures 46–51" captionStartId="F7" captionText="Figures 46 - 51. Labels of Chlaenius (Lithochlaenius) spp 46 holotype, Chlaenius agilis Chaudoir (see Figs 22 - 23) 47 Cotype, Chlaenius anchomenoides Bates (see Figs 40 - 41) 48 Paratype, Chlaenius nuristanus Jedlicka (see Figs 42 - 43) 49 Cotype, Chlaenius nuristanus Jedlicka (see Figs 44 - 45) 50 Holotype, Chlaenius agiloides Jedlicka (see Fig. 48) 51 Holotype, Chlaenius formosanus Jedlicka (see Fig 49)." httpUri="https://binary.pensoft.net/fig/17284" pageId="11" pageNumber="26">46-49</figureCitation>
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="11" pageNumber="26">55, 62-64</figureCitation>
<figureCitation captionStart="Figures 66–79" captionStartId="F9" captionText="Figures 66 - 79. Humeral region of elytra of Chlaenius (Lithochlaenius) spp 66 Chlaenius chuanqianensis sp. n., holotype 67 Chlaenius linwensini sp. n., holotype 68 Chlaenius propeagilis sp. n., holotype 69 Chlaenius agilis Chaudoir, holotype 70 Chlaenius formosensis Lorenz, CCCC 71 Chlaenius leishanensis Kirschenhofer, IZCAS 72 Chlaenius noguchii Bates, IZCAS 73 Chlaenius wrasei Kirschenhofer, IZCAS 74 Chlaenius agiloides Jedlicka, IZCAS 75 Chlaenius rambouseki Lutshnik from Ussuri region, ZCAS 76 Chlaenius anchomenoides Bates, paratype 77 Chlaenius nuristanus Jedlicka, paratype 78 Chlaenius nuristanus Jedlicka, paratype 79 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17286" pageId="11" pageNumber="26">69, 76-78</figureCitation>
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="11" pageNumber="26">83, 90-92</figureCitation>
<figureCitation captionStart="Figures 94–116" captionStartId="F11" captionText="Figures 94 - 116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species 94 Chlaenius chuanqianensis sp. n., male, holotype 95 Chlaenius chuanqianensis sp. n., female, paratype 96 Chlaenius linwensini sp. n., male, holotype 97 Chlaenius linwensini sp. n., female, paratype 98 Chlaenius propeagilis sp. n., male, holotype 99 Chlaenius propeagilis sp. n., female, paratype 100 Chlaenius agilis Chaudoir, male, holotype 101 Chlaenius formosensis Lorenz, male 102 Chlaenius formosensis Lorenz, female 103 Chlaenius leishanensis Kirschenhofer, male 104 Chlaenius leishanensis Kirschenhofer, female 105 Chlaenius noguchii Bates, male 106 Chlaenius noguchii Bates, female 107 Chlaenius wrasei Kirschenhofer, male 108 Chlaenius wrasei Kirschenhofer, female 109 Chlaenius agiloides Jedlicka, male 110 Chlaenius agiloides Jedlicka, female 111 Chlaenius rambouseki Lutshnik, from Ussuri region, male 112 Chlaenius rambouseki Lutshnik from Ussuri region, female 113 Chlaenius anchomenoides Bates, male, paratype 114 Chlaenius nuristanus Jedlicka, male, paratype 115 Chlaenius nuristanus Jedlicka, female, paratype 116 Chlaenius formosanus Jedlicka from Taiwan, male. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17288" pageId="11" pageNumber="26">100, 113-115</figureCitation>
<figureCitation captionStart="Figures 16–21" captionStartId="F2" captionText="Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17279" pageId="11" pageNumber="26">123-124</figureCitation>
<figureCitation captionStart="Figures 22–27" captionStartId="F3" captionText="Figures 22 - 27. Habitus of Chlaenius (Lithochlaenius) spp 22 Chlaenius agilis Chaudoir, holotype, dorsal view 23 Chlaenius agilis Chaudoir, holotype, ventral view 24 Chlaenius formosensis Lorenz, male, in CCCC, dorsal view 25 Chlaenius formosensis Lorenz, ventral view 26 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, dorsal view 27 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17280" pageId="11" pageNumber="26">138-143</figureCitation>
<figureCitation captionStart="Figures 146–151" captionStartId="F14" captionText="Figures 146 - 151. Photographs of habitats for Chlaenius (Lithochlaenius) spp. 146 Chlaenius chuanqianensis sp. n., Dabaitang, Xishui county, North Guizhou, China 147 Chlaenius anchomenoides Bates, Goorais valley, Pakistan (Provided by Dr. Muhammad Abbas in Pakistan Museum of Natural History) 148 Chlaenius propeagilis sp. n., Gaoligongshan, Yunnan, China 149 Chlaenius leishanensis Kirschenhofer, Xiaodanjiang, Leigongshan, Leishan county, Southeast Guizhou, China 150 Chlaenius chuanqianensis sp. n. and Chlaenius agiloides Jedlicka, Jinshajiang, Sichuan 151 Chlaenius rambouseki Lutshnik, Ussri river, Heilongjiang, China." httpUri="https://binary.pensoft.net/fig/17291" pageId="11" pageNumber="26">147</figureCitation>
<figureCitation captionStart="Figure 158" captionStartId="F16" captionText="Figure 158. Map showing the known geographical distributions of Chlaenius (Lithochlaenius) spp." httpUri="https://binary.pensoft.net/fig/17293" pageId="11" pageNumber="26">158</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="26" type="reference_group">
<paragraph pageId="11" pageNumber="26">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="latro">Chlaenius latro</taxonomicName>
<normalizedToken originalValue="LaFerté-Sénectère">LaFerte-Senectere</normalizedToken>
, 1851:250 (unavailable). Nomen nudum 
<emphasis bold="true" pageId="11" pageNumber="26">stat. n.</emphasis>
</paragraph>
<paragraph pageId="11" pageNumber="26">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="agilis">Chlaenius agilis</taxonomicName>
Chaudoir, 1856:246; 1876:193; Kryzhanovskij, 1976:12
</paragraph>
<paragraph pageId="11" pageNumber="26">
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="anchomenoides">Chlaenius anchomenoides</taxonomicName>
Bates, 1889:212; Kirschenhofer, 1997:116; Kirschenhofer, 2005:490. 
<emphasis bold="true" pageId="11" pageNumber="26">syn. n.</emphasis>
</paragraph>
<paragraph pageId="11" pageNumber="26">
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="nuristanus">Chlaenius nuristanus</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, 1956:194; Kirschenhofer, 2005:490. 
<emphasis bold="true" pageId="11" pageNumber="26">syn. n.</emphasis>
</paragraph>
<paragraph pageId="11" pageNumber="26">
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="nuristanus">Chlaenius nuristanus</taxonomicName>
a. 
<emphasis italics="true" pageId="11" pageNumber="26">rubridipes</emphasis>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, 1956:194 (not available, ICZN Articles 10A and 45.6.2).
</paragraph>
<paragraph pageId="11" pageNumber="26">
<taxonomicName class="Insecta" family="Carabidae" genus="Stenochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="anchomenoides">Stenochlaenius anchomenoides</taxonomicName>
Bates: Mandl, 1972:104
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="26" type="specimens examined">
<paragraph pageId="11" pageNumber="26">Specimens examined.</paragraph>
<paragraph pageId="11" pageNumber="26">
Total 15 specimens. 
<emphasis bold="true" pageId="11" pageNumber="26">India:</emphasis>
type: male (MNHN), &quot; 
<emphasis italics="true" pageId="11" pageNumber="26">agilis</emphasis>
Chaud, Ind. orient bor. C. Boys.&quot;/ &quot;TYPE 
<emphasis italics="true" pageId="11" pageNumber="26">agilis</emphasis>
&quot;/ &quot;Ex Musaeo Chaudoir&quot;/ &quot;agilis Chd&quot;; Cotype: female (MNHN), &quot; 
<emphasis italics="true" pageId="11" pageNumber="26">agilis</emphasis>
Chaud, Ind. orient bor. C. Boys.&quot;/ &quot;TYPE agilis&quot;/ &quot;Ex Musaeo Chaudoir&quot;/ &quot;agilis Chd&quot;; Cotype: male (MNHN), &quot; 
<emphasis italics="true" pageId="11" pageNumber="26">agilis</emphasis>
Chaud, Ind. orient bor. C. Boys.&quot;/ &quot; 
<emphasis italics="true" pageId="11" pageNumber="26">latro</emphasis>
&quot;; 2 males and 3 females (CASC), &quot;Bajaura, Kongra district (Indes Angl.)&quot;/ &quot;G. Babault, Juin 1914&quot;/ &quot;van Dyke collection&quot;/ &quot; 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="11" pageNumber="26">Chlaenius agilis</emphasis>
</taxonomicName>
, Chaud., H.E. Andrewes det.&quot;. 1 female (CASC), &quot;W. Almora Divn, Kumaon U.P., Apr. 1917, HGC.&quot;/ &quot;Van Dyke Collection&quot;. 
<emphasis bold="true" pageId="11" pageNumber="26">Pakistan:</emphasis>
Cotype: 2 males (BMNH), &quot;Goorais valley, 7000ft, V. 87&quot;/ &quot;H. E. Andrews Coll. B. M. 1945-97&quot;/ &quot;Ex coll. R. 
<normalizedToken originalValue="Oberthür”">Oberthuer&quot;</normalizedToken>
/ &quot; 
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="11" pageNumber="26">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates, cotype, H. E. Andrews det.&quot;/ &quot;Co-type&quot;. 
<emphasis bold="true" pageId="11" pageNumber="26">Afghanistan</emphasis>
: 1 female (MNHN), &quot;J. Klapperich, Bashgultal, 1100m, Nuristan, 22.4.53, Afghanistan&quot;/ &quot; 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="11" pageNumber="26">Chlaenius nuristanus</emphasis>
</taxonomicName>
sp. n, det. ING. JEDLICKA&quot;/ &quot;Type&quot;;
</paragraph>
<paragraph pageId="11" pageNumber="26">
Paratype: 2 males (MNHN), &quot;J. Klapperich, Asmar, 900m, Kunartal, 3.4.53, O-Afghanistan&quot;/ &quot; 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="11" pageNumber="26">Chlaenius nuristanus</emphasis>
</taxonomicName>
sp.n, det. ING. JEDLICKA&quot;/ &quot;PARATYPE&quot;; Type: 1 male (MNHN), &quot;J. Klapperich, Asmar, 900m, Kunartal, 3.4.53, O-Afghanistan&quot;/ &quot; 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="11" pageNumber="26">Chlaenius nuristanus</emphasis>
</taxonomicName>
a. 
<emphasis italics="true" pageId="11" pageNumber="26">rubridipes</emphasis>
n. det. ING. JEDLICKA&quot;/ &quot;TYPUS&quot;; Cotype: 1 male (MNHN), &quot;J. Klapperich, Asmar, 900m, Kunartal, 3.4.53, O-Afghanistan&quot;/ &quot; 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="11" pageNumber="26">Chlaenius nuristanus</emphasis>
</taxonomicName>
a. 
<emphasis italics="true" pageId="11" pageNumber="26">rubridipes</emphasis>
n. det. ING. JEDLICKA&quot;/ &quot;Cotype&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="26" type="diagnosis">
<paragraph pageId="11" pageNumber="26">Diagnosis.</paragraph>
<paragraph pageId="11" pageNumber="26">
Antennomere 1 elongate ovoid (
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="11" pageNumber="26">Figs 55, 62-64</figureCitation>
); basal two-thirds of intervals 1-5 glabrous medially, with one row of pubescence laterally near striae (
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="11" pageNumber="26">Figs 83, 90-92</figureCitation>
); aedeagus abruptly bent near base (
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="11" pageNumber="26">Figs 124</figureCitation>
, 
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="11" pageNumber="26">139, 141, 143</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="27" pageId="11" pageNumber="26" type="description">
<paragraph pageId="11" pageNumber="26">Description.</paragraph>
<paragraph pageId="11" pageNumber="26">Total length = 15.50-16.00 mm (mean = 15.50), width = 5.60-6.10 mm (mean = 5.73); HW = 2.75-2.95 mm (mean = 2.82), EYL = 1.00-1.15 mm (mean =1.00), ratio Ant3/Ant1 = 1.67-1.83 (mean = 1.71), PL/PW = 0.84-0.85 (mean = 0.85), EL/EW = 1.63-1.73 (mean = 1.71), EYL/ PL = 1.15-1.44 (mean = 1.34). Mensural data cited in the description were obtained from type and cotype specimens examined.</paragraph>
<paragraph pageId="12" pageNumber="27">
<pageBreakToken pageId="12" pageNumber="27" start="start">Head</pageBreakToken>
, pronotum and elytra black, with green or blue metallic luster; ventral surface black; mandibles and trochanters dark brown; palpomeres, femora, and tarsomeres brown to dark brown; antennae and tibiae yellow to brown (
<figureCitation captionStart="Figures 22–27" captionStartId="F3" captionText="Figures 22 - 27. Habitus of Chlaenius (Lithochlaenius) spp 22 Chlaenius agilis Chaudoir, holotype, dorsal view 23 Chlaenius agilis Chaudoir, holotype, ventral view 24 Chlaenius formosensis Lorenz, male, in CCCC, dorsal view 25 Chlaenius formosensis Lorenz, ventral view 26 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, dorsal view 27 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17280" pageId="12" pageNumber="27">Figs 22-23</figureCitation>
, 
<figureCitation captionStart="Figures 34–39" captionStartId="F5" captionText="Figures 34 - 39. Chlaenius (Lithochlaenius) spp 34 Chlaenius rambouseki Lutshnik, male, collected from Ussuri region, in IZCAS, dorsal view 35 Chlaenius rambouseki Lutshnik, male, collected from Ussuri region, in IZCAS, ventral view 36 Chlaenius anchomenoides Bates, cotype, male, in BMNH, dorsal view 37 Chlaenius anchomenoides Bates, cotype, male, in BMNH, ventral view 38 Chlaenius nuristanus Jedlicka, paratype, male, in MNHN, dorsal view 39 Chlaenius nuristanus Jedlicka, paratype, male, in MNHN, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17282" pageId="12" pageNumber="27">36</figureCitation>
- 
<figureCitation captionStart="Figures 40–45" captionStartId="F6" captionText="Figures 40 - 45. Chlaenius (Lithochlaenius) spp 40 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, dorsal view 41 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, ventral view 42 Chlaenius agiloides Jedlicka, holotype, male, in NMPC, dorsal view 43 Chlaenius formosanus Jedlicka, holotype, male, in NMPC, dorsal view 44 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, dorsal view 45 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17283" pageId="12" pageNumber="27">41</figureCitation>
).
</paragraph>
<paragraph pageId="12" pageNumber="27">
Head with vertex smooth or very sparsely punctate behind eyes; antennomere 1 elongate ovoid (
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="12" pageNumber="27">Figs 55, 62-64</figureCitation>
); labrum slightly emarginate at apex. Pronotum with disk smooth; basal foveae deep, sparsely punctate and pubescent (
<figureCitation captionStart="Figures 22–27" captionStartId="F3" captionText="Figures 22 - 27. Habitus of Chlaenius (Lithochlaenius) spp 22 Chlaenius agilis Chaudoir, holotype, dorsal view 23 Chlaenius agilis Chaudoir, holotype, ventral view 24 Chlaenius formosensis Lorenz, male, in CCCC, dorsal view 25 Chlaenius formosensis Lorenz, ventral view 26 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, dorsal view 27 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17280" pageId="12" pageNumber="27">Figs 22</figureCitation>
, 
<figureCitation captionStart="Figures 34–39" captionStartId="F5" captionText="Figures 34 - 39. Chlaenius (Lithochlaenius) spp 34 Chlaenius rambouseki Lutshnik, male, collected from Ussuri region, in IZCAS, dorsal view 35 Chlaenius rambouseki Lutshnik, male, collected from Ussuri region, in IZCAS, ventral view 36 Chlaenius anchomenoides Bates, cotype, male, in BMNH, dorsal view 37 Chlaenius anchomenoides Bates, cotype, male, in BMNH, ventral view 38 Chlaenius nuristanus Jedlicka, paratype, male, in MNHN, dorsal view 39 Chlaenius nuristanus Jedlicka, paratype, male, in MNHN, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17282" pageId="12" pageNumber="27">36, 38</figureCitation>
, 
<figureCitation captionStart="Figures 40–45" captionStartId="F6" captionText="Figures 40 - 45. Chlaenius (Lithochlaenius) spp 40 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, dorsal view 41 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, ventral view 42 Chlaenius agiloides Jedlicka, holotype, male, in NMPC, dorsal view 43 Chlaenius formosanus Jedlicka, holotype, male, in NMPC, dorsal view 44 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, dorsal view 45 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17283" pageId="12" pageNumber="27">40</figureCitation>
). Elytra with intervals convex, basal two-thirds of intervals 1-5 glabrous medially, with one row of pubescence laterally near striae (
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="12" pageNumber="27">Figs 83, 90-92</figureCitation>
), intervals 6-9 and apical one-third of intervals 1-5 densely pubescent; striae deep, punctate; humeral angle obtuse (
<figureCitation captionStart="Figures 66–79" captionStartId="F9" captionText="Figures 66 - 79. Humeral region of elytra of Chlaenius (Lithochlaenius) spp 66 Chlaenius chuanqianensis sp. n., holotype 67 Chlaenius linwensini sp. n., holotype 68 Chlaenius propeagilis sp. n., holotype 69 Chlaenius agilis Chaudoir, holotype 70 Chlaenius formosensis Lorenz, CCCC 71 Chlaenius leishanensis Kirschenhofer, IZCAS 72 Chlaenius noguchii Bates, IZCAS 73 Chlaenius wrasei Kirschenhofer, IZCAS 74 Chlaenius agiloides Jedlicka, IZCAS 75 Chlaenius rambouseki Lutshnik from Ussuri region, ZCAS 76 Chlaenius anchomenoides Bates, paratype 77 Chlaenius nuristanus Jedlicka, paratype 78 Chlaenius nuristanus Jedlicka, paratype 79 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17286" pageId="12" pageNumber="27">Figs 69, 76-78</figureCitation>
). Abdominal sterna IV-VI sparsely pubescent medially; sternum VII rugose-pubescent, narrowly rounded apically in males (
<figureCitation captionStart="Figures 94–116" captionStartId="F11" captionText="Figures 94 - 116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species 94 Chlaenius chuanqianensis sp. n., male, holotype 95 Chlaenius chuanqianensis sp. n., female, paratype 96 Chlaenius linwensini sp. n., male, holotype 97 Chlaenius linwensini sp. n., female, paratype 98 Chlaenius propeagilis sp. n., male, holotype 99 Chlaenius propeagilis sp. n., female, paratype 100 Chlaenius agilis Chaudoir, male, holotype 101 Chlaenius formosensis Lorenz, male 102 Chlaenius formosensis Lorenz, female 103 Chlaenius leishanensis Kirschenhofer, male 104 Chlaenius leishanensis Kirschenhofer, female 105 Chlaenius noguchii Bates, male 106 Chlaenius noguchii Bates, female 107 Chlaenius wrasei Kirschenhofer, male 108 Chlaenius wrasei Kirschenhofer, female 109 Chlaenius agiloides Jedlicka, male 110 Chlaenius agiloides Jedlicka, female 111 Chlaenius rambouseki Lutshnik, from Ussuri region, male 112 Chlaenius rambouseki Lutshnik from Ussuri region, female 113 Chlaenius anchomenoides Bates, male, paratype 114 Chlaenius nuristanus Jedlicka, male, paratype 115 Chlaenius nuristanus Jedlicka, female, paratype 116 Chlaenius formosanus Jedlicka from Taiwan, male. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17288" pageId="12" pageNumber="27">Figs 100, 113-114</figureCitation>
), broadly rounded apically in females (
<figureCitation captionStart="Figures 94–116" captionStartId="F11" captionText="Figures 94 - 116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species 94 Chlaenius chuanqianensis sp. n., male, holotype 95 Chlaenius chuanqianensis sp. n., female, paratype 96 Chlaenius linwensini sp. n., male, holotype 97 Chlaenius linwensini sp. n., female, paratype 98 Chlaenius propeagilis sp. n., male, holotype 99 Chlaenius propeagilis sp. n., female, paratype 100 Chlaenius agilis Chaudoir, male, holotype 101 Chlaenius formosensis Lorenz, male 102 Chlaenius formosensis Lorenz, female 103 Chlaenius leishanensis Kirschenhofer, male 104 Chlaenius leishanensis Kirschenhofer, female 105 Chlaenius noguchii Bates, male 106 Chlaenius noguchii Bates, female 107 Chlaenius wrasei Kirschenhofer, male 108 Chlaenius wrasei Kirschenhofer, female 109 Chlaenius agiloides Jedlicka, male 110 Chlaenius agiloides Jedlicka, female 111 Chlaenius rambouseki Lutshnik, from Ussuri region, male 112 Chlaenius rambouseki Lutshnik from Ussuri region, female 113 Chlaenius anchomenoides Bates, male, paratype 114 Chlaenius nuristanus Jedlicka, male, paratype 115 Chlaenius nuristanus Jedlicka, female, paratype 116 Chlaenius formosanus Jedlicka from Taiwan, male. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17288" pageId="12" pageNumber="27">Fig. 115</figureCitation>
). Aedeagus abruptly bent near base, apical lamella round (
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="12" pageNumber="27">Figs 123</figureCitation>
, 
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="12" pageNumber="27">138, 140, 142</figureCitation>
), thin and slight bent ventrally (
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="12" pageNumber="27">Figs 124</figureCitation>
, 
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="12" pageNumber="27">139, 141, 143</figureCitation>
).
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/17284" pageId="12" pageNumber="27" start="Figures 46–51" startId="F7">
<paragraph pageId="12" pageNumber="27">
<emphasis bold="true" pageId="12" pageNumber="27">Figures 46-51.</emphasis>
Labels of 
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius (Lithochlaenius)</emphasis>
</taxonomicName>
spp 
<emphasis bold="true" pageId="12" pageNumber="27">46</emphasis>
holotype, 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir (see Figs 22-23) 
<emphasis bold="true" pageId="12" pageNumber="27">47</emphasis>
Cotype, 
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates (see 
<figureCitation captionStart="Figures 40–45" captionStartId="F6" captionText="Figures 40 - 45. Chlaenius (Lithochlaenius) spp 40 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, dorsal view 41 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, ventral view 42 Chlaenius agiloides Jedlicka, holotype, male, in NMPC, dorsal view 43 Chlaenius formosanus Jedlicka, holotype, male, in NMPC, dorsal view 44 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, dorsal view 45 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17283" pageId="12" pageNumber="27">Figs 40-41</figureCitation>
) 
<emphasis bold="true" pageId="12" pageNumber="27">48</emphasis>
Paratype, 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
(see 
<figureCitation captionStart="Figures 40–45" captionStartId="F6" captionText="Figures 40 - 45. Chlaenius (Lithochlaenius) spp 40 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, dorsal view 41 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, ventral view 42 Chlaenius agiloides Jedlicka, holotype, male, in NMPC, dorsal view 43 Chlaenius formosanus Jedlicka, holotype, male, in NMPC, dorsal view 44 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, dorsal view 45 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17283" pageId="12" pageNumber="27">Figs 42-43</figureCitation>
) 
<emphasis bold="true" pageId="12" pageNumber="27">49</emphasis>
Cotype, 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
(see Figs 44-45) 
<emphasis bold="true" pageId="12" pageNumber="27">50</emphasis>
Holotype, 
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
(see 
<figureCitation captionStart="Figures 46–51" captionStartId="F7" captionText="Figures 46 - 51. Labels of Chlaenius (Lithochlaenius) spp 46 holotype, Chlaenius agilis Chaudoir (see Figs 22 - 23) 47 Cotype, Chlaenius anchomenoides Bates (see Figs 40 - 41) 48 Paratype, Chlaenius nuristanus Jedlicka (see Figs 42 - 43) 49 Cotype, Chlaenius nuristanus Jedlicka (see Figs 44 - 45) 50 Holotype, Chlaenius agiloides Jedlicka (see Fig. 48) 51 Holotype, Chlaenius formosanus Jedlicka (see Fig 49)." httpUri="https://binary.pensoft.net/fig/17284" pageId="12" pageNumber="27">Fig. 48</figureCitation>
) 
<emphasis bold="true" pageId="12" pageNumber="27">51</emphasis>
Holotype, 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
(see 
<figureCitation captionStart="Figures 46–51" captionStartId="F7" captionText="Figures 46 - 51. Labels of Chlaenius (Lithochlaenius) spp 46 holotype, Chlaenius agilis Chaudoir (see Figs 22 - 23) 47 Cotype, Chlaenius anchomenoides Bates (see Figs 40 - 41) 48 Paratype, Chlaenius nuristanus Jedlicka (see Figs 42 - 43) 49 Cotype, Chlaenius nuristanus Jedlicka (see Figs 44 - 45) 50 Holotype, Chlaenius agiloides Jedlicka (see Fig. 48) 51 Holotype, Chlaenius formosanus Jedlicka (see Fig 49)." httpUri="https://binary.pensoft.net/fig/17284" pageId="12" pageNumber="27">Fig 49</figureCitation>
).
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/17285" pageId="12" pageNumber="27" start="Figures 52–65" startId="F8">
<paragraph pageId="12" pageNumber="27">
<emphasis bold="true" pageId="12" pageNumber="27">Figures 52-65.</emphasis>
Antennomere 1 of 
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius (Lithochlaenius)</emphasis>
</taxonomicName>
species 
<emphasis bold="true" pageId="12" pageNumber="27">52</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="chuanqianensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius chuanqianensis</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">53</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="linwensini">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius linwensini</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">54</emphasis>
<taxonomicName authorityName="Liu &amp; Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="propeagilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius propeagilis</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">55</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir, holotype 
<emphasis bold="true" pageId="12" pageNumber="27">56</emphasis>
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosensis</emphasis>
</taxonomicName>
Lorenz, CCCC 
<emphasis bold="true" pageId="12" pageNumber="27">57</emphasis>
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="leishanensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius leishanensis</emphasis>
</taxonomicName>
Kirschenhofer, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">58</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="noguchii">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius noguchii</emphasis>
</taxonomicName>
Bates, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">59</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius wrasei</emphasis>
</taxonomicName>
Kirschenhofer, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">60</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">61</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Ussuri region, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">62</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">63</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">64</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
a. 
<emphasis italics="true" pageId="12" pageNumber="27">rubridipes</emphasis>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">65</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Taiwan,CCCC. Scale line = 1.0 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/17286" pageId="12" pageNumber="27" start="Figures 66–79" startId="F9">
<paragraph pageId="12" pageNumber="27">
<emphasis bold="true" pageId="12" pageNumber="27">Figures 66-79.</emphasis>
Humeral region of elytra of 
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius (Lithochlaenius)</emphasis>
</taxonomicName>
spp 
<emphasis bold="true" pageId="12" pageNumber="27">66</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="chuanqianensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius chuanqianensis</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">67</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="linwensini">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius linwensini</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">68</emphasis>
<taxonomicName authorityName="Liu &amp; Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="propeagilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius propeagilis</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">69</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir, holotype 
<emphasis bold="true" pageId="12" pageNumber="27">70</emphasis>
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosensis</emphasis>
</taxonomicName>
Lorenz, CCCC 
<emphasis bold="true" pageId="12" pageNumber="27">71</emphasis>
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="leishanensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius leishanensis</emphasis>
</taxonomicName>
Kirschenhofer, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">72</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="noguchii">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius noguchii</emphasis>
</taxonomicName>
Bates, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">73</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius wrasei</emphasis>
</taxonomicName>
Kirschenhofer, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">74</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">75</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Ussuri region, ZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">76</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">77</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">78</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">79</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/17287" pageId="12" pageNumber="27" start="Figures 80–93" startId="F10">
<paragraph pageId="12" pageNumber="27">
<emphasis bold="true" pageId="12" pageNumber="27">Figures 80-93.</emphasis>
Elytral intervals of 
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius (Lithochlaenius)</emphasis>
</taxonomicName>
spp. 
<emphasis bold="true" pageId="12" pageNumber="27">80</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="chuanqianensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius chuanqianensis</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">81</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="linwensini">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius linwensini</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">82</emphasis>
<taxonomicName authorityName="Liu &amp; Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="propeagilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius propeagilis</emphasis>
</taxonomicName>
sp. n., holotype 
<emphasis bold="true" pageId="12" pageNumber="27">83</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir, holotype 
<emphasis bold="true" pageId="12" pageNumber="27">84</emphasis>
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosensis</emphasis>
</taxonomicName>
Lorenz, CCCC 
<emphasis bold="true" pageId="12" pageNumber="27">85</emphasis>
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="leishanensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius leishanensis</emphasis>
</taxonomicName>
, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">86</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="noguchii">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius noguchii</emphasis>
</taxonomicName>
Bates, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">87</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius wrasei</emphasis>
</taxonomicName>
Kirschenhofer, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">88</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">89</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Ussuri region, IZCAS 
<emphasis bold="true" pageId="12" pageNumber="27">90</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">91</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">92</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">93</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Taiwan
<emphasis italics="true" pageId="12" pageNumber="27">,</emphasis>
CCCC. Scale line = 1.0 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/17288" pageId="12" pageNumber="27" start="Figures 94–116" startId="F11">
<paragraph pageId="12" pageNumber="27">
<emphasis bold="true" pageId="12" pageNumber="27">Figures 94-116.</emphasis>
Abdominal sternum VII (ventral aspect) of 
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius (Lithochlaenius)</emphasis>
</taxonomicName>
species 
<emphasis bold="true" pageId="12" pageNumber="27">94</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="chuanqianensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius chuanqianensis</emphasis>
</taxonomicName>
sp. n., male, holotype 
<emphasis bold="true" pageId="12" pageNumber="27">95</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="chuanqianensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius chuanqianensis</emphasis>
</taxonomicName>
sp. n., female, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">96</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="linwensini">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius linwensini</emphasis>
</taxonomicName>
sp. n., male, holotype 
<emphasis bold="true" pageId="12" pageNumber="27">97</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="linwensini">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius linwensini</emphasis>
</taxonomicName>
sp. n., female, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">98</emphasis>
<taxonomicName authorityName="Liu &amp; Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="propeagilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius propeagilis</emphasis>
</taxonomicName>
sp. n., male, holotype 
<emphasis bold="true" pageId="12" pageNumber="27">99</emphasis>
<taxonomicName authorityName="Liu &amp; Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="propeagilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius propeagilis</emphasis>
</taxonomicName>
sp. n., female, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">100</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir, male, holotype 
<emphasis bold="true" pageId="12" pageNumber="27">101</emphasis>
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosensis</emphasis>
</taxonomicName>
Lorenz, male 
<emphasis bold="true" pageId="12" pageNumber="27">102</emphasis>
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosensis</emphasis>
</taxonomicName>
Lorenz, female 
<emphasis bold="true" pageId="12" pageNumber="27">103</emphasis>
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="leishanensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius leishanensis</emphasis>
</taxonomicName>
Kirschenhofer, male 
<emphasis bold="true" pageId="12" pageNumber="27">104</emphasis>
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="leishanensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius leishanensis</emphasis>
</taxonomicName>
Kirschenhofer, female 
<emphasis bold="true" pageId="12" pageNumber="27">105</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="noguchii">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius noguchii</emphasis>
</taxonomicName>
Bates, male 
<emphasis bold="true" pageId="12" pageNumber="27">106</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="noguchii">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius noguchii</emphasis>
</taxonomicName>
Bates, female 
<emphasis bold="true" pageId="12" pageNumber="27">107</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius wrasei</emphasis>
</taxonomicName>
Kirschenhofer, male 
<emphasis bold="true" pageId="12" pageNumber="27">108</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius wrasei</emphasis>
</taxonomicName>
Kirschenhofer, female 
<emphasis bold="true" pageId="12" pageNumber="27">109</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, male 
<emphasis bold="true" pageId="12" pageNumber="27">110</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, female 
<emphasis bold="true" pageId="12" pageNumber="27">111</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik, from Ussuri region, male 
<emphasis bold="true" pageId="12" pageNumber="27">112</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Ussuri region, female 
<emphasis bold="true" pageId="12" pageNumber="27">113</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates, male, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">114</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, male, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">115</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, female, paratype 
<emphasis bold="true" pageId="12" pageNumber="27">116</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
from Taiwan, male. Scale line = 1.0 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/17289" pageId="12" pageNumber="27" start="Figures 117–130" startId="F12">
<paragraph pageId="12" pageNumber="27">
<emphasis bold="true" pageId="12" pageNumber="27">Figures 117-130.</emphasis>
Aedeagi of 
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius (Lithochlaenius)</emphasis>
</taxonomicName>
species 
<emphasis bold="true" pageId="12" pageNumber="27">117</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="chuanqianensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius chuanqianensis</emphasis>
</taxonomicName>
sp. n., holotype (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">118</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="chuanqianensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius chuanqianensis</emphasis>
</taxonomicName>
sp. n., holotype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">119</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="linwensini">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius linwensini</emphasis>
</taxonomicName>
sp. n., holotype (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">120</emphasis>
<taxonomicName authorityName="Liu &amp; Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="linwensini">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius linwensini</emphasis>
</taxonomicName>
sp. n., holotype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">121</emphasis>
<taxonomicName authorityName="Liu &amp; Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="propeagilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius propeagilis</emphasis>
</taxonomicName>
sp. n., holotype (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">122</emphasis>
<taxonomicName authorityName="Liu &amp; Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="propeagilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius propeagilis</emphasis>
</taxonomicName>
sp. n., holotype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">123</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir, holotype (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">124</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir, holotype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">125</emphasis>
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosensis</emphasis>
</taxonomicName>
Lorenz, holotype (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">126</emphasis>
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="formosensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius formosensis</emphasis>
</taxonomicName>
Lorenz, holotype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">127</emphasis>
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="leishanensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius leishanensis</emphasis>
</taxonomicName>
Kirschenhofer (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">128</emphasis>
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="leishanensis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius leishanensis</emphasis>
</taxonomicName>
Kirschenhofer (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">129</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="noguchii">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius noguchii</emphasis>
</taxonomicName>
Bates (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">130</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="noguchii">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius noguchii</emphasis>
</taxonomicName>
Bates (left lateral view). Scale lines: A = 1.0 mm (
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="12" pageNumber="27">Figs 117, 119, 121, 123, 125, 127, 129</figureCitation>
); B = 0.5 mm (
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="12" pageNumber="27">Figs 118, 120, 122, 124, 126, 128, 130</figureCitation>
).
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/17290" pageId="12" pageNumber="27" start="Figures 131–145" startId="F13">
<paragraph pageId="12" pageNumber="27">
<emphasis bold="true" pageId="12" pageNumber="27">Figures 131-145.</emphasis>
Aedeagi of 
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius (Lithochlaenius)</emphasis>
</taxonomicName>
species 
<emphasis bold="true" pageId="12" pageNumber="27">131</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius wrasei</emphasis>
</taxonomicName>
Kirschenhofer (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">132</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius wrasei</emphasis>
</taxonomicName>
Kirschenhofer (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">133</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
(dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">134</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, holotype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">135</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agiloides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agiloides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
(left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">136</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Ussuri region (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">137</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Ussuri region (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">138</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates, paratype (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">139</emphasis>
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates, paratype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">140</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype (dorsal view, black leg) 
<emphasis bold="true" pageId="12" pageNumber="27">141</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">142</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype (dorsal view, yellow leg) 
<emphasis bold="true" pageId="12" pageNumber="27">143</emphasis>
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, paratype (left lateral view) 
<emphasis bold="true" pageId="12" pageNumber="27">144</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Taiwan (dorsal view) 
<emphasis bold="true" pageId="12" pageNumber="27">145</emphasis>
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius rambouseki</emphasis>
</taxonomicName>
Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="12" pageNumber="27">Figs 131, 133, 136, 138, 140, 142, 144</figureCitation>
); B = 0.5 mm (
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="12" pageNumber="27">Figs 132, 134, 135, 137, 139, 141, 143, 145</figureCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="12" pageNumber="27" type="color variation">
<paragraph pageId="12" pageNumber="27">Color variation.</paragraph>
<paragraph pageId="12" pageNumber="27">
Color of antennae, palpomeres, femora, and tibiae varied from medium brown to dark brown or even black among different individuals (
<figureCitation captionStart="Figures 22–27" captionStartId="F3" captionText="Figures 22 - 27. Habitus of Chlaenius (Lithochlaenius) spp 22 Chlaenius agilis Chaudoir, holotype, dorsal view 23 Chlaenius agilis Chaudoir, holotype, ventral view 24 Chlaenius formosensis Lorenz, male, in CCCC, dorsal view 25 Chlaenius formosensis Lorenz, ventral view 26 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, dorsal view 27 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17280" pageId="12" pageNumber="27">Figs 22-23</figureCitation>
, 
<figureCitation captionStart="Figures 34–39" captionStartId="F5" captionText="Figures 34 - 39. Chlaenius (Lithochlaenius) spp 34 Chlaenius rambouseki Lutshnik, male, collected from Ussuri region, in IZCAS, dorsal view 35 Chlaenius rambouseki Lutshnik, male, collected from Ussuri region, in IZCAS, ventral view 36 Chlaenius anchomenoides Bates, cotype, male, in BMNH, dorsal view 37 Chlaenius anchomenoides Bates, cotype, male, in BMNH, ventral view 38 Chlaenius nuristanus Jedlicka, paratype, male, in MNHN, dorsal view 39 Chlaenius nuristanus Jedlicka, paratype, male, in MNHN, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17282" pageId="12" pageNumber="27">36</figureCitation>
- 
<figureCitation captionStart="Figures 40–45" captionStartId="F6" captionText="Figures 40 - 45. Chlaenius (Lithochlaenius) spp 40 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, dorsal view 41 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, ventral view 42 Chlaenius agiloides Jedlicka, holotype, male, in NMPC, dorsal view 43 Chlaenius formosanus Jedlicka, holotype, male, in NMPC, dorsal view 44 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, dorsal view 45 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17283" pageId="12" pageNumber="27">41</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="27" type="distribution">
<paragraph pageId="12" pageNumber="27">Geographical distribution.</paragraph>
<paragraph pageId="12" pageNumber="27">
<figureCitation captionStart="Figure 158" captionStartId="F16" captionText="Figure 158. Map showing the known geographical distributions of Chlaenius (Lithochlaenius) spp." httpUri="https://binary.pensoft.net/fig/17293" pageId="12" pageNumber="27">Fig. 158</figureCitation>
. Known only from eastern Afghanistan, Pakistan, and northern India.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="27" type="remarks">
<paragraph pageId="12" pageNumber="27">Remarks.</paragraph>
<paragraph pageId="12" pageNumber="27">
Mensural data cited in the description were obtained from type and cotype specimens examined. The name 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir was once treated as a junior synonym of 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="latro">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius latro</emphasis>
</taxonomicName>
<normalizedToken originalValue="LaFerté-Sénectère">LaFerte-Senectere</normalizedToken>
(
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen II. Carabidae II, W. Junk, Berlin" pageId="23" pageNumber="38" pagination="739 - 1022" refId="B9" refString="Csiki, E, 1931. Carabidae: Harpalinae V (Pars 115). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen II. Carabidae II, W. Junk, Berlin: 739 - 1022" title="Carabidae: Harpalinae V (Pars 115)." volumeTitle="Coleopterorum catalogus." year="1931">Csiki 1931</bibRefCitation>
, 
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="23" pageNumber="38" pagination="103 - 122" refId="B13" refString="Kirschenhofer, E, 1997. Contribution to the faunistics and taxonomy of Carabidae (Coleoptera) of Korea. Annales Historico-Naturales Musei Nationalis Hungarici 89: 103 - 122" title="Contribution to the faunistics and taxonomy of Carabidae (Coleoptera) of Korea." volume="89" year="1997">Kirschenhofer 1997</bibRefCitation>
). However, in his work, 
<bibRefCitation author="LaFerte-Senectere, FT de" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="23" pageNumber="38" pagination="209 - 294" refId="B18" refString="LaFerte-Senectere, FT de, 1851. Revision de la tribu des patellimanes de Dejean, coleopteres pentameres de la famille des carabiques. Annales de la Societe Entomologique de France 2 (9): 209 - 294" title="Revision de la tribu des patellimanes de Dejean, coleopteres pentameres de la famille des carabiques" volume="2" year="1851">
<normalizedToken originalValue="LaFerté-Sénectère">LaFerte-Senectere</normalizedToken>
(1851)
</bibRefCitation>
did not provide a specific description of 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="latro">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius latro</emphasis>
</taxonomicName>
. This means that 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="latro">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius latro</emphasis>
</taxonomicName>
<normalizedToken originalValue="LaFerté-Sénectère">LaFerte-Senectere</normalizedToken>
is a 
<emphasis italics="true" pageId="12" pageNumber="27">nomen nudum</emphasis>
, and therefore, unavailable.
</paragraph>
<paragraph pageId="12" pageNumber="27">
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="23" pageNumber="38" pagination="489 - 501" refId="B15" refString="Kirschenhofer, E, 2005. Weiter neue Arten der Gattung Chlaenius Bonelli 1810 aus China, Myanmar und Indien (Coleoptera, Carabidae). Linzer Biologische Beitraege 37: 489 - 501" title="Weiter neue Arten der Gattung Chlaenius Bonelli 1810 aus China, Myanmar und Indien (Coleoptera, Carabidae)." volume="37" year="2005">Kirschenhofer (2005)</bibRefCitation>
considered 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
to be a junior synonym of 
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
Bates. We have examined the types and/or cotypes of 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius agilis</emphasis>
</taxonomicName>
Chaudoir, 
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoide">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoide</emphasis>
</taxonomicName>
s, 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
and its aberration 
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="nuristanus">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius nuristanus</emphasis>
</taxonomicName>
a. 
<emphasis italics="true" pageId="12" pageNumber="27">rubridipes</emphasis>
, and found no significant differences between them, except for variation in the color of antennae and legs. The aedeagi are abruptly bent near the base in all dissected males (
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="12" pageNumber="27">Figs 124</figureCitation>
, 
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="12" pageNumber="27">139, 141, 143</figureCitation>
).
</paragraph>
<paragraph pageId="12" pageNumber="27">
<bibRefCitation author="Mandl, K" journalOrPublisher="Nachrichtenblatt der Bayerischen Entomologen" pageId="23" pageNumber="38" pagination="97 - 105" refId="B21" refString="Mandl, K, 1972. Beitrag zur Kenntnis des Genus Stenochlaenius Reitter. Die Arten der Gruppe coeruleus Steven. (Col. Carabidae). Nachrichtenblatt der Bayerischen Entomologen 21: 97 - 105" title="Beitrag zur Kenntnis des Genus Stenochlaenius Reitter. Die Arten der Gruppe coeruleus Steven. (Col. Carabidae)." volume="21" year="1972">Mandl (1972: 104)</bibRefCitation>
assigned 
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
to genus 
<taxonomicName authorityName="Reitter" authorityYear="1908" class="Insecta" family="Carabidae" genus="Stenochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="12" pageNumber="27">Stenochlaenius</emphasis>
</taxonomicName>
. Based on the pubescent elytral intervals and incomplete elytral basal margin in members of 
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="12" pageNumber="27" phylum="Arthropoda" rank="species" species="anchomenoides">
<emphasis italics="true" pageId="12" pageNumber="27">Chlaenius anchomenoides</emphasis>
</taxonomicName>
, we do not agree with this assignment.
</paragraph>
</subSubSection>
</treatment>
</document>