301 lines
38 KiB
XML
301 lines
38 KiB
XML
<document ID-CLB-Dataset="51924" ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-GBIF-Dataset="c2e00414-9633-4890-ab72-cb2aa3e9d174" ID-PMC="PMC10208988" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-PubMed="37250203" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak & Sutcharit, Chirasak" docDate="2023" docId="7279C3DCA2715E4297DDD938D43BAF25" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Pterocyclos jousseaumei" docType="treatment" docVersion="5" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1732652259105" updateUser="ExternalLinkService">
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<mods:title id="5488A7C02E175424D73DB59872F59325">Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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<mods:namePart id="6436876158A1EA4FA3E7FDC13B780B0A">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="A20956DCCC640611B5FC80D6B4D4AFDE">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="6A05C84ADE55C54B10D8FD99D0BFAD3A">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="D3032EC800527A4124349C48A418E338">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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<mods:namePart id="9FB8061917E05743FE6F766C86A05302">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="5FED1F177EDF71D28391EC527B0033F8">Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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<mods:namePart id="75BA580975D0B3BE1A124D3DB26FA025">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="BB42D955C45D6E0CB8D5C135F323792C">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:namePart id="4B83916C97B6123BD72B926090CC17BE">Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation id="38662C0EB0278D58DE740D2CDE6D1296">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="F8D46C4E2F0A20CBE3C058148809394C">Panha, Somsak</mods:namePart>
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<mods:affiliation id="E2BB3DB6F49B69097E49448716F7803A">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="2A0D07FF66ECF61B7D2B75074AD72EC7" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:namePart id="362C45D03CCAC45BE4B5E0BBFD6A14F6">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="08F4353723F2476EBCB1FEC7C1FFC561">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="5A8F8758E8D0CCCF5E87944749839CB8" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:title id="812004C6C4FB37C5CB1C6740F86CCBF1">ZooKeys</mods:title>
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<mods:date id="6BEA909D47A9FEFAB5EAB8DB3CE7C471">2023</mods:date>
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<mods:number id="88A53F644026CD92F5E3CF99DDE14E7A">2023-01-24</mods:number>
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<mods:number id="20921916A32EB63E833A7FA9B1DF8027">1142</mods:number>
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<treatment id="7279C3DCA2715E4297DDD938D43BAF25" ID-GBIF-Taxon="209102223" LSID="urn:lsid:plazi:treatment:7279C3DCA2715E4297DDD938D43BAF25" httpUri="http://treatment.plazi.org/id/7279C3DCA2715E4297DDD938D43BAF25" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Cyclophoridae" scope_subClass="Caenogastropoda" scope_subFamily="Cyclophorinae">
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<subSubSection id="42558C6F6CC97E3E8E400307605DC213" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="ED35C8203C4A31B1896B5BB550BC1AC4" pageId="0" pageNumber="1">
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<taxonomicName id="5A376D402F0F5756EA8015ECC7A67B32" ID-CoL="B3CK6" LSID="7279C3DC-A271-5E42-97DD-D938D43BAF25" authority="(Morgan, 1885)" baseAuthorityName="Morgan" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos jousseaumei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">Pterocyclos jousseaumei (Morgan, 1885)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="180837A97BBB571DE7EFCB6DF40C1CFF" pageId="0" pageNumber="1" type="description">
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<paragraph id="0BA0E79EC424C8733E6B5E9E0993C2C3" pageId="0" pageNumber="1">
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<figureCitation id="EF81520B225D1999D006B8244FAEBD14" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">Fig. 56</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="882ACC026067A7EFBD3CF5D09542072E" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="0F49BF0738A6651BF6D71879936ACDB5" pageId="0" pageNumber="1">
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<taxonomicName id="853C59BD886E49FC3B64D4721AD237DF" authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma jousseaumei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">Rhiostoma jousseaumei</taxonomicName>
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||
Morgan, 1885: 400, pl. 8, fig. 2. Type locality: la haute
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<normalizedToken id="91F6C9A42B18333EC1D005AA838136A7" originalValue="vallée">vallee</normalizedToken>
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||
de Kinta (
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<normalizedToken id="B7729F381D5E341587F17C9743DBA3E2" originalValue="Pérak">Perak</normalizedToken>
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) [Kinta valley, Perak, Malaysia].
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||
<bibRefCitation id="4219FAB36D4D605D8669745B19BF42C3" DOI="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" author="Moellendorff, OF von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="330 - 348" refId="B96" refString="Moellendorff, OF von, 1891. On the land and freshwater shells of Perak. Proceedings of the Zoological Society of London 59 (3): 330 - 348, DOI: https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="59" year="1891">
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<normalizedToken id="01A3891D2D93C449F9AC9A684101A422" originalValue="Möllendorff">Moellendorff</normalizedToken>
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1891
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||
</bibRefCitation>
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||
: 340.
|
||
<bibRefCitation id="E1E5C2A2DFAFE7CFDBA77EA84B6D67A4" author="Kobelt, W" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="1" pagination="1 - 663" refId="B71" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich 16: 1 - 663" title="Cyclophoridae." volume="16" year="1902">Kobelt 1902</bibRefCitation>
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||
: 178. Kobelt 1911: 758, pl. 110, figs 11-13.
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||
<bibRefCitation id="72E87C15AC64E2457644D2FB0557BA73" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B80" refString="Laidlaw, FF, 1928. A list of land and fresh-water Mollusca of the Malay Peninsula with note. Journal of the Malaysian Branch of the Royal Asiatic Society 3: 25 - 37" title="A list of land and fresh-water Mollusca of the Malay Peninsula with note." volume="3" year="1928">Laidlaw 1928</bibRefCitation>
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||
: 31.
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||
<bibRefCitation id="50233C20A0EFA860A16142EFCE4782FF" author="Chang, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" refId="B27" refString="Chang, S-Y, 1995. A brief collection of land shells of Desaru anjung Penawar. The Papustyla 9: 4, 5." title="A brief collection of land shells of Desaru anjung Penawar. The Papustyla 9: 4, 5." year="1995">Chang 1995</bibRefCitation>
|
||
: 5 with text figure.
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||
<bibRefCitation id="8E1069D9F90895007F83598E1FF8B09C" DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B45" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
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||
: 24, fig. 8f.
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||
</paragraph>
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||
<paragraph id="78DE5A03CB8406C23B0EC32D087689F3" pageId="0" pageNumber="1">
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||
<taxonomicName id="07747BD089708EB429348426289991D9" authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhiostoma</taxonomicName>
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<taxonomicName id="92FDF124A8593508BE978EA2EFBA4D40" authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhiostoma</taxonomicName>
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||
macalpine-woodsi Laidlaw, 1939: 166, with text figure. Type locality: Sungei Siput, Perak.
|
||
<bibRefCitation id="D48D815A00A771DC998F7B6EB278D5A8" DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. 2019</bibRefCitation>
|
||
: 37, fig. 8g.
|
||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C458F2F8E2532968CF018B00495D64FD" pageId="0" pageNumber="1" type="type material">
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<paragraph id="1E8BE1345249D1FD42099762AF30B81A" pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph id="9EA7B4681CD7189F322171FD1BE937F3" pageId="0" pageNumber="1">
|
||
The type specimens
|
||
<taxonomicName id="E067390D0AACBFCF57232664E99B42A1" authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma jousseaumei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
|
||
<emphasis id="301378606964F2CB633DE7AA5C9B7C82" italics="true" pageId="0" pageNumber="1">Rhiostoma jousseaumei</emphasis>
|
||
</taxonomicName>
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||
Morgan, 1885 could not be located in the MNHN collection.
|
||
<emphasis id="006529EEF86761524DEE75C102B964A2" bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
|
||
(designation in
|
||
<bibRefCitation id="9BB0A51449C10CCC44214613CA20D762" DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. (2019)</bibRefCitation>
|
||
) NHMUK 1939.4.13.23 (Fig.
|
||
<figureCitation id="950113AA2CDBB3F148B46574B552A7E7" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56D</figureCitation>
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) of
|
||
<taxonomicName id="B70F8CAFE73F2CB86B4DDA18B06F1285" authorityName="Laidlaw" authorityYear="1939" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma macalpinewoodsi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="macalpinewoodsi">
|
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<emphasis id="C1672A42E1A5086C60380227CF88E2E5" italics="true" pageId="0" pageNumber="1">Rhiostoma macalpinewoodsi</emphasis>
|
||
</taxonomicName>
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||
Laidlaw, 1939, from Sungei Siput, Perak.
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="3286BBAFCB19BDA82BB78F54CAB39ED3" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="2A2F0D08BA4E3AA97FA3E7034CB37724" pageId="0" pageNumber="1">Other material examined.</paragraph>
|
||
<paragraph id="86ABB8082F8BC383B6D964A9485C4174" pageId="0" pageNumber="1">
|
||
<materialsCitation id="5859B3BDE1A8067580AA680FB0F38109" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Malaysia" location="Perak State" specimenCount="1" stateProvince="Perak State">
|
||
<emphasis id="5912670AF68A32958B3DA57DB7A57253" bold="true" pageId="0" pageNumber="1">
|
||
<collectingCountry id="EA4D8AC59CAAF0F1B03F7FD833429CDE" name="Malaysia">Malaysia</collectingCountry>
|
||
</emphasis>
|
||
: Specimen CUMZ 4752 (2 shells; Fig.
|
||
<figureCitation id="2DB5266F10986D95D235014247CC3BA9" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56B</figureCitation>
|
||
) from Ipoh,
|
||
<collectingRegion id="F668DD597E09BA6F608F1A1B7D242696" country="Malaysia" name="Perak">Perak State</collectingRegion>
|
||
</materialsCitation>
|
||
. CUMZ 7457 (15 shells; Fig.
|
||
<figureCitation id="C8D7ACCDCA7C527D0F03EECF4E2D59F0" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56C</figureCitation>
|
||
) from Tanjung Rambutan, Kinta,
|
||
<collectingRegion id="D248F69EFA06BAC568CBE1DD46557B2E" country="Malaysia" name="Perak">Perak State</collectingRegion>
|
||
. CUMZ 10213 (5 shells; Fig.
|
||
<figureCitation id="ABA97493272D22B50E5D9A0C720EF3AB" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56E, F</figureCitation>
|
||
) from Gunung Dato, Ipoh,
|
||
<collectingRegion id="35134D0311113F95A58C36BB97C99012" country="Malaysia" name="Perak">Perak State</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<caption id="2772058458A0252A43C61DE4B6384E67" doi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1" start="Figure 56" startId="F56">
|
||
<paragraph id="59FD4282A0AEA86849CB62638FD05BC1" pageId="0" pageNumber="1">
|
||
<emphasis id="A92535D6B261145487AA8A86C2C5AFBF" bold="true" pageId="0" pageNumber="1">Figure 56.</emphasis>
|
||
Shell of
|
||
<taxonomicName id="CF03658921A5692DB6D83497CEE2CB6F" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus jousseaumei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
|
||
<emphasis id="C71FFA7EF8C0EED07ADC4C5D0E3AEA62" italics="true" pageId="0" pageNumber="1">Opisthoporus jousseaumei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F20D37FA25FF6F805E0F9B5CC320C1F9" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
original figure (after
|
||
<bibRefCitation id="039BBC40A604192443A3E19F21CB19FB" DOI="https://doi.org/10.5962/bhl.part.14301" author="Morgan, J de" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="353 - 428" refId="B104" refString="Morgan, J de, 1885. Mollusques terrestres & fluviatiles du royaume de Perak et des pays voisins (presqueIle Malaise). Bulletin de la Societe Zoologique de France 10: 353 - 428, DOI: https://doi.org/10.5962/bhl.part.14301" title="Mollusques terrestres & fluviatiles du royaume de Perak et des pays voisins (presqueIle Malaise)." url="https://doi.org/10.5962/bhl.part.14301" volume="10" year="1885">Morgan 1885</bibRefCitation>
|
||
: pl. 8, fig. 2)
|
||
<emphasis id="8C4CAA9FB39024AC53C4BECB368CC18E" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
specimen CUMZ 4752 from Ipoh,
|
||
<collectingRegion id="1A06B9C0593C55758BED602356BDD5FF" country="Malaysia" name="Perak">Perak</collectingRegion>
|
||
,
|
||
<collectingCountry id="9D13A1580EDF745928C44E894F502287" name="Malaysia">Malaysia</collectingCountry>
|
||
<emphasis id="1F7BD70652C84BFD9BB6D6A0030AD1ED" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
specimen CUMZ 7457 from Kinta,
|
||
<collectingRegion id="71AFED3C9195FDF406A23E9A593536A3" country="Malaysia" name="Perak">Perak</collectingRegion>
|
||
,
|
||
<collectingCountry id="11FC611A7705829F63B085BEF4A4B904" name="Malaysia">Malaysia</collectingCountry>
|
||
<emphasis id="AB64C5FBD9BA2F40E4AFBFD10B14488B" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<typeStatus id="DA55301AD886CF63A1CA0C27D051764A">lectotype</typeStatus>
|
||
of "
|
||
<taxonomicName id="A32C5E263230534E270C03234FF8E10D" authorityName="Laidlaw" authorityYear="1939" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma macalpinewoodsi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="macalpinewoodsi">
|
||
<emphasis id="8A35512AA4F63C60457FC715F6542C6A" italics="true" pageId="0" pageNumber="1">Rhiostoma macalpinewoodsi</emphasis>
|
||
</taxonomicName>
|
||
Laidlaw, 1939" NHMUK 1939.4.13.23 from Sungei Siput,
|
||
<collectingRegion id="35D9F7AF11E2AF68B572C60609EA6476" country="Malaysia" name="Perak">Perak</collectingRegion>
|
||
<emphasis id="29F9D4C7F8CDD8F2BDB001DE4AD2CEED" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
||
specimens CUMZ 10213 from Gunung Dato, Ipoh,
|
||
<collectingRegion id="D06B6F9478B951EE1C99ABE8CF1CB736" country="Malaysia" name="Perak">Perak</collectingRegion>
|
||
,
|
||
<collectingCountry id="D2D3522A53826616FE7CED22BD6B445B" name="Malaysia">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="AF21ADB9EB6ED4D33A625644B0A6FBB4" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="81EE9380F68E60BFD5F765C7C2E40E2A" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="28CBC85C20533A79C0F03A58778B55DD" pageId="0" pageNumber="1">Periostracum thin, corneous, shell varying from uniform colour to dark brown zigzag pattern; periphery with a thin spiral band. Detached-whorl length similar to apertural width. Peristome circular and double or boundary hardly visible; lip thick or thin, and not expanded to slightly expanded. Breathing device with incomplete tube or tubular shape, curved posteriorly and its tip usually attached to preceding whorl. Operculum calcareous, multispiral, and slightly concave on both surfaces.</paragraph>
|
||
<paragraph id="D58DFA30A8927614BA52BA633ED53CAC" pageId="0" pageNumber="1">
|
||
An operculum accompanied the examined specimen and was also figured in
|
||
<bibRefCitation id="C47CD1F3C66EFC72F45A5E28C56050D6" DOI="https://doi.org/10.5962/bhl.part.14301" author="Morgan, J de" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="353 - 428" refId="B104" refString="Morgan, J de, 1885. Mollusques terrestres & fluviatiles du royaume de Perak et des pays voisins (presqueIle Malaise). Bulletin de la Societe Zoologique de France 10: 353 - 428, DOI: https://doi.org/10.5962/bhl.part.14301" title="Mollusques terrestres & fluviatiles du royaume de Perak et des pays voisins (presqueIle Malaise)." url="https://doi.org/10.5962/bhl.part.14301" volume="10" year="1885">Morgan (1885)</bibRefCitation>
|
||
: it was multispiral and plate-shaped (Fig.
|
||
<figureCitation id="648D8DE5687468456424CF3E94D23FF9" captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56A, C</figureCitation>
|
||
). With these observations, we suggest the re-assignment of this species to the
|
||
<taxonomicName id="2AD128B1E299F853D03915CCE4DD5492" authorityName="Benson" authorityYear="1832" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis id="30FAD6ACCE2F003E66BE49FB891E8249" italics="true" pageId="0" pageNumber="1">Pterocyclos</emphasis>
|
||
</taxonomicName>
|
||
. This species can be separated from
|
||
<taxonomicName id="5BD81F184E26887D98987F2F53C2D118" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos subalatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subalatus">
|
||
<emphasis id="7CE9D0D91DB598E62CF186F64BECD418" italics="true" pageId="0" pageNumber="1">Pterocyclos subalatus</emphasis>
|
||
</taxonomicName>
|
||
Sykes, 1903,
|
||
<taxonomicName id="92A0C3B7D1526F8452BDF1982DE19217" baseAuthorityName="Tomlin" baseAuthorityYear="1931" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos spelaeotes" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spelaeotes">
|
||
<emphasis id="EFFCA7E2E3980A6BA57D397431750380" italics="true" pageId="0" pageNumber="1">Pterocyclos spelaeotes</emphasis>
|
||
</taxonomicName>
|
||
(Tomlin, 1931), and
|
||
<taxonomicName id="4888B28172CE40EA353B68C325A1BF07" baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1949" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos umbraticus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="umbraticus">
|
||
<emphasis id="6CC5EA9ABF7A8733AFB664EEB23B6BC2" italics="true" pageId="0" pageNumber="1">Pterocyclos umbraticus</emphasis>
|
||
</taxonomicName>
|
||
(Benthem Jutting, 1949) from peninsular Malaysia in having the apertural lip thickened and usually not expanded (rarely slightly expanded), while the three species (Fig.
|
||
<figureCitation id="7B8707C804E01DE65BB7BC0283CDD154" captionStart="Figure 57" captionStartId="F57" captionText="Figure 57. A Pterocyclos subalatus Sykes, 1903, holotype UMZC 1032 from Gunong Inas, Perak, Malay Peninsula B Pterocyclos spelaeotes (Tomlin, 1931), lectotype NMW 1955.158.01107 from Batu Cave, Kula Lumpur, Selangor C Pterocyclos umbraticus (Benthem Jutting, 1949), holotype ZMA 135622 from Maxwell's Hill, Perak, Malaysia D Cyclotus boxalli syntype NHMUK 1894.5.23.1 from Kina Balu E Opisthoporus battambangensis holotype MNHN-IM- 2000 - 35506 from Battambang, Cambodia F Opisthoporus grohi holotype MNHN-IM- 2000 - 35507 from Dak Nong, Vietnam. Photograph: H Wood, NMW (B), M Caballer, A Lardeur and V Heros, MNHN (E, F)." figureDoi="10.3897/zookeys.1142.90097.figure57" httpUri="https://binary.pensoft.net/fig/799915" pageId="0" pageNumber="1">57A-C</figureCitation>
|
||
) tend to have a widely expanded lip (see also
|
||
<bibRefCitation id="B04539AC3E0BD47F4F8E9C1DDA5DFD2A" author="Sutcharit, C" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="330 - 338" refId="B159" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2014. The land snail genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with description of two new species. The Raffles Bulletin of Zoology 62: 330 - 338" title="The land snail genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with description of two new species." volume="62" year="2014">Sutcharit et al. 2014</bibRefCitation>
|
||
). Furthermore,
|
||
<taxonomicName id="7AB30B8DC512169421D0E574FE1F1F57" lsidName="P. subalatus" pageId="0" pageNumber="1" rank="species" species="subalatus">
|
||
<emphasis id="C66CB00812FB3ECA0DA73E714CB27798" italics="true" pageId="0" pageNumber="1">P. subalatus</emphasis>
|
||
</taxonomicName>
|
||
has no detached whorl but has a wide canal-shaped breathing device attached to the preceding whorl;
|
||
<taxonomicName id="0E70CF5B9517CD7A42411996928F0D1F" lsidName="P. umbraticus" pageId="0" pageNumber="1" rank="species" species="umbraticus">
|
||
<emphasis id="72F1225BB8565A856745AFBF0D99EAA5" italics="true" pageId="0" pageNumber="1">P. umbraticus</emphasis>
|
||
</taxonomicName>
|
||
has a larger shell (width 27 mm), without a detached whorl, and an undulated shell surface.
|
||
</paragraph>
|
||
<caption id="4CA5225E7552EB3B8F74B066ADDB7C6E" doi="10.3897/zookeys.1142.90097.figure57" httpUri="https://binary.pensoft.net/fig/799915" pageId="0" pageNumber="1" start="Figure 57" startId="F57">
|
||
<paragraph id="E8F9815524F94D945BA7594ABA086263" pageId="0" pageNumber="1">
|
||
<emphasis id="9DB1E2A8515F2EFE1951F89460FBC1F5" bold="true" pageId="0" pageNumber="1">Figure 57.</emphasis>
|
||
<emphasis id="258C0A5A82864CF70041A333944B2C09" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
<taxonomicName id="628D9810B49AA5CBD87BB7E47B5E998F" authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos subalatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subalatus">
|
||
<emphasis id="6252727616392259B765CFCF4C7CC21E" italics="true" pageId="0" pageNumber="1">Pterocyclos subalatus</emphasis>
|
||
</taxonomicName>
|
||
Sykes, 1903, holotype UMZC 1032 from Gunong Inas, Perak, Malay Peninsula
|
||
<emphasis id="0615D043294735DA20D9629863F58B35" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
<taxonomicName id="415A92FCE84FBB7D376C2FDF2E492DD5" baseAuthorityName="Tomlin" baseAuthorityYear="1931" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos spelaeotes" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spelaeotes">
|
||
<emphasis id="690AC79A48E81C56818D73B76A888C1A" italics="true" pageId="0" pageNumber="1">Pterocyclos spelaeotes</emphasis>
|
||
</taxonomicName>
|
||
(Tomlin, 1931), lectotype NMW 1955.158.01107 from Batu Cave, Kula Lumpur, Selangor
|
||
<emphasis id="CBA5F3B32333B3A6192E68A5ADCCEFFC" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
<taxonomicName id="365C0BF8205A7D418FCC70A7841F6390" baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1949" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos umbraticus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="umbraticus">
|
||
<emphasis id="4EFB19F0B2D34E27E4F5AE75FD490F55" italics="true" pageId="0" pageNumber="1">Pterocyclos umbraticus</emphasis>
|
||
</taxonomicName>
|
||
(Benthem Jutting, 1949), holotype ZMA 135622 from
|
||
<normalizedToken id="53187B228B6BF6E1BDB42A54AFBFDDDE" originalValue="Maxwell’s">Maxwell's</normalizedToken>
|
||
Hill, Perak, Malaysia
|
||
<emphasis id="4F9AEA43ABFC6EA2285B35AA0F17B8A2" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<taxonomicName id="BD54BF9880EDA5F51563E9AD996477DE" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Cyclotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclotus boxalli" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="boxalli">
|
||
<emphasis id="3E7336E309D74B2ADC28100F639D2BFF" italics="true" pageId="0" pageNumber="1">Cyclotus boxalli</emphasis>
|
||
</taxonomicName>
|
||
syntype NHMUK 1894.5.23.1 from Kina Balu
|
||
<emphasis id="EA95D064252C814BF5E651DCDD6AC82F" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
<taxonomicName id="16969C62305D830DFAA91BB2DBEC05D4" baseAuthorityName="Thach & Huber" baseAuthorityYear="2020" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus battambangensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="battambangensis">
|
||
<emphasis id="635B55724E1CF50AB2DC249FD0119506" italics="true" pageId="0" pageNumber="1">Opisthoporus battambangensis</emphasis>
|
||
</taxonomicName>
|
||
holotype MNHN-IM-2000-35506 from Battambang, Cambodia
|
||
<emphasis id="A46335B8ACCE6B38387C87A3D379A301" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
<taxonomicName id="5F92EB73D1F7731880DB1A8BAC281A2C" baseAuthorityName="Thach" baseAuthorityYear="2020" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus grohi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohi">
|
||
<emphasis id="9F7B47655551F8320718E4F737A5C171" italics="true" pageId="0" pageNumber="1">Opisthoporus grohi</emphasis>
|
||
</taxonomicName>
|
||
holotype MNHN-IM-2000-35507 from Dak Nong, Vietnam. Photograph: H Wood, NMW (B), M Caballer, A Lardeur and V Heros, MNHN (
|
||
<emphasis id="C1533A6CC5576AD53EA9BFC332236571" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CC4B9F1AD29473D68B87BD713446CE20" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="993B1CB0331E71C29CBC18931FEEFEF9" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" refId="B81" refString="Laidlaw, FF, 1939. A new Rhiostoma from Malay. Journal of Conchology 21: 116." title="A new Rhiostoma from Malay. Journal of Conchology 21: 116." year="1939">Laidlaw (1939)</bibRefCitation>
|
||
stated that
|
||
<taxonomicName id="42C51A4CEB72CE2AE168C09C31D50D96" lsidName="R. macalpinewoodsi" pageId="0" pageNumber="1" rank="species" species="macalpinewoodsi">
|
||
<emphasis id="9E6BF28815B899B7E1869EB12B180E6B" italics="true" pageId="0" pageNumber="1">R. macalpinewoodsi</emphasis>
|
||
</taxonomicName>
|
||
differed from
|
||
<taxonomicName id="8A2E9A5E545F8DEAA99B198074CC0541" lsidName="R. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
|
||
<emphasis id="A9B0C1BA289F0A1478E3AA379938C01B" italics="true" pageId="0" pageNumber="1">R. jousseaumei</emphasis>
|
||
</taxonomicName>
|
||
in having a relatively small shell (height 6 mm, width 12 mm), with slightly thickened but not expanded apertural lip, a breathing device channel-like to short tubular [= incomplete tube] shape. Later,
|
||
<bibRefCitation id="1D0098D8A488C6F78D13343F8C4841B4" DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B45" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. (2017)</bibRefCitation>
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examined several specimens and found an intermediate shell form; the type locality of both nominal species is situated in the Kinta Valley, Perak State, peninsular Malaysia. In addition, the specimens examined herein show a mix between smaller shell (with incomplete tube) and larger shell (with tubular) forms in a single collection locality, with the intermediates as reported in
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<bibRefCitation id="4CA1CC518A62D2BC28E14D5D97ADA876" DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B45" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. (2017)</bibRefCitation>
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. This distinctive shell size and breathing device could be attributed to intraspecific variation, and examination of more specimens and systematic study will help resolve this issue.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |