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<mods:title id="BE5988037C221B75BEDE8BB7E6A83A72">Messinian rodents from Moncucco Torinese, NW Italy: palaeobiodiversity and biochronology</mods:title>
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<treatment id="72738785FFBDFFD5BBCEAB76FC2015B6" ID-DOI="http://doi.org/10.5281/zenodo.4836535" ID-GBIF-Taxon="182404777" ID-Zenodo-Dep="4836535" LSID="urn:lsid:plazi:treatment:72738785FFBDFFD5BBCEAB76FC2015B6" httpUri="http://treatment.plazi.org/id/72738785FFBDFFD5BBCEAB76FC2015B6" lastPageId="29" lastPageNumber="450" pageId="28" pageNumber="449">
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<paragraph id="FA653693FFBDFFD4BBCEAB76FE161247" blockId="28.[135,625,279,338]" pageId="28" pageNumber="449">
<taxonomicName id="3DDA4D10FFBDFFD4BBCEAB76FDDF1227" authority="Hugueney &amp; Mein, 1965" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[135,625,279,306]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="449" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBDFFD4BBCEAB76FEFA1224" bold="true" box="[135,340,279,306]" italics="true" pageId="28" pageNumber="449">Muscardinus vireti</emphasis>
<bibRefCitation id="9E4B4B62FFBDFFD4BA12AB79FDDF1227" author="HUGUENEY M. &amp; MEIN P." box="[347,625,279,306]" pageId="28" pageNumber="449" pagination="109 - 123" refId="ref32228" refString="HUGUENEY M. &amp; MEIN P. 1965. - Lagomorphes et rongeurs du Neogene de Lissieu (Rhone). Travaux du Laboratoire de Geologie de la Faculte des Sciences de Lyon Nouvelle Serie 12: 109 - 123." type="journal article" year="1965">Hugueney &amp; Mein, 1965</bibRefCitation>
</taxonomicName>
(
<figureCitation id="62E12A16FFBDFFD4BA01AB56FE291247" box="[328,391,311,338]" captionStart="FIG" captionStartId="26.[113,124,913,931]" captionTargetBox="[122,1206,227,849]" captionTargetId="figure-331@26.[661,1211,213,777]" captionTargetPageId="26" captionText="FIG. 6. — Isolated teeth of Eliomys Wagner, 1840 and Muscardinus Kaup, 1829 from Moncucco Torinese:A-E, Eliomys aff.intermedius Friant, 1953; A, MGPT-PU12128348, P4 dex.; B, MGPT-PU128270 M1 sin.; C, MGPT-PU127495, M2 sin.; D, MGPT-PU128269, m1 dex.; E, MGPT-PU127127368, m2 dex.; F-P, Muscardinus vireti Hugueney &amp; Mein, 1965; F, MGPT-PU127604, P4 dex.; G, MGPT- PU12392, M1 dex.; H, MGPT-PU127606, M2 sin.; I, MGPT-PU128178, M2 dex.; J, MGPT-PU127607, M3 dex.; K, MGPT-PU128182, M3 dex.; L, MGPT-PU127608, p4 sin.; M, MGPT-PU127609, m1 dex.; N, MGPT-PU127610, m2 sin.; O, MGPT-PU127611, m3 sin.; P, MGPT-PU127513, m3 dex. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4538575" httpUri="https://zenodo.org/record/4538575/files/figure.png" pageId="28" pageNumber="449">Fig. 6</figureCitation>
F-P)
</paragraph>
</subSubSection>
<subSubSection id="B2C06518FFBDFFD4BB38AB11FD291109" pageId="28" pageNumber="449" type="reference_group">
<paragraph id="FA653693FFBDFFD4BB38AB11FF3F12B7" blockId="28.[113,647,368,420]" pageId="28" pageNumber="449">
<treatmentCitationGroup id="DACA11BDFFBDFFD4BB38AB11FF3F12B7" pageId="28" pageNumber="449">
<treatmentCitation id="7B7B1082FFBDFFD4BB38AB11FD94129C" author="HUGUENEY M. &amp; MEIN P." box="[113,570,368,393]" page="118" pageId="28" pageNumber="449" year="1965">
<taxonomicName id="3DDA4D10FFBDFFD4BB38AB11FD94129C" authority="Hugueney &amp; Mein, 1965: 118" authorityName="Hugueney &amp; Mein" authorityPageNumber="118" authorityYear="1965" box="[113,570,368,393]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="449" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBDFFD4BB38AB11FEB6129D" box="[113,280,368,392]" italics="true" pageId="28" pageNumber="449">Muscardinus vireti</emphasis>
<bibRefCitation id="9E4B4B62FFBDFFD4BA55AB11FD94129C" author="HUGUENEY M. &amp; MEIN P." box="[284,570,368,393]" pageId="28" pageNumber="449" pagination="109 - 123" refId="ref32228" refString="HUGUENEY M. &amp; MEIN P. 1965. - Lagomorphes et rongeurs du Neogene de Lissieu (Rhone). Travaux du Laboratoire de Geologie de la Faculte des Sciences de Lyon Nouvelle Serie 12: 109 - 123." type="journal article" year="1965">Hugueney &amp; Mein, 1965: 118</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 64- 78.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38ABDBFE4312F9" blockId="28.[113,646,442,492]" pageId="28" pageNumber="449">
<treatmentCitationGroup id="DACA11BDFFBDFFD4BB38ABDBFE4312F9" pageId="28" pageNumber="449">
<taxonomicName id="3DDA4D10FFBDFFD4BB38ABDBFEE812C7" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[113,326,442,467]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="449" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBDFFD4BB38ABDBFF4612C7" box="[113,232,442,466]" italics="true" pageId="28" pageNumber="449">Muscardinus</emphasis>
aff.
<emphasis id="C8AEEA81FFBDFFD4BA5CABDAFEE812C7" box="[277,326,443,466]" italics="true" pageId="28" pageNumber="449">vireti</emphasis>
</taxonomicName>
<treatmentCitation id="7B7B1082FFBDFFD4BA29ABDBFD8D12C7" author="ENGESSER B." box="[352,547,442,466]" page="776" pageId="28" pageNumber="449" year="1983">
<bibRefCitation id="9E4B4B62FFBDFFD4BA29ABDBFD8D12C7" author="ENGESSER B." box="[352,547,442,466]" pageId="28" pageNumber="449" pagination="763 - 780" refId="ref31261" refString="ENGESSER B. 1983. - Die jungtertiaren Kleinsauger des Gebietes der Maremma (Toskana, Italien). 1. Teil: Gliridae (Rodentia, Mammalia). Eclogae Geologicae Helvetiae 76: 763 - 780." type="journal article" year="1983">Engesser 1983: 776</bibRefCitation>
</treatmentCitation>
, fig. 6. — Colombero
<emphasis id="C8AEEA81FFBDFFD4BBAEABB4FEB812F9" box="[231,278,468,492]" italics="true" pageId="28" pageNumber="449">et al.</emphasis>
2013: 122, fig. 5E, F.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38A865FD291109" blockId="28.[113,647,516,541]" box="[113,647,516,541]" pageId="28" pageNumber="449">
<treatmentCitationGroup id="DACA11BDFFBDFFD4BB38A865FD291109" box="[113,647,516,541]" pageId="28" pageNumber="449">
<taxonomicName id="3DDA4D10FFBDFFD4BB38A865FE811109" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[113,303,516,541]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="449" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBDFFD4BB38A865FF4C1109" box="[113,226,516,540]" italics="true" pageId="28" pageNumber="449">Muscardinus</emphasis>
cf.
<emphasis id="C8AEEA81FFBDFFD4BA48A864FE811109" box="[257,303,517,540]" italics="true" pageId="28" pageNumber="449">vireti</emphasis>
</taxonomicName>
<emphasis id="C8AEEA81FFBDFFD4BA7BA864FE901109" box="[306,318,517,540]" italics="true" pageId="28" pageNumber="449"></emphasis>
<treatmentCitation id="7B7B1082FFBDFFD4BA0AA865FD8E1109" author="ANGELONE C. &amp; COLOMBERO S. &amp; ESU D. &amp; GIUNTELLI P. &amp; MARCOLINI F. &amp; PAVIA M. &amp; TRENKWALDER S. &amp; VAN DEN HOEK OSTENDE L. W. &amp; ZUNINO M. &amp; PAVIA G." box="[323,544,516,540]" page="99" pageId="28" pageNumber="449" year="2011">
<bibRefCitation id="9E4B4B62FFBDFFD4BA0AA865FD8E1109" author="ANGELONE C. &amp; COLOMBERO S. &amp; ESU D. &amp; GIUNTELLI P. &amp; MARCOLINI F. &amp; PAVIA M. &amp; TRENKWALDER S. &amp; VAN DEN HOEK OSTENDE L. W. &amp; ZUNINO M. &amp; PAVIA G." box="[323,544,516,540]" pageId="28" pageNumber="449" pagination="89 - 104" refId="ref29204" refString="ANGELONE C., COLOMBERO S., ESU D., GIUNTELLI P., MARCOLINI F., PAVIA M., TRENKWALDER S., VAN DEN HOEK OSTENDE L. W., ZUNINO M. &amp; PAVIA G. 2011. - Moncucco Torinese, a new post-evaporitic Messinian fossiliferous site from Piedmont (NW Italy). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 259: 89 - 104." type="journal article" year="2011">
Angelone
<emphasis id="C8AEEA81FFBDFFD4BAD7A864FE671109" box="[414,457,516,540]" italics="true" pageId="28" pageNumber="449">et al.</emphasis>
2011: 99
</bibRefCitation>
</treatmentCitation>
, fig. 6 (16).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B2C06518FFBDFFD4BB38A855FEBE1183" pageId="28" pageNumber="449" type="materials_examined">
<paragraph id="FA653693FFBDFFD4BB38A855FE69115E" blockId="28.[113,455,564,588]" box="[113,455,564,588]" pageId="28" pageNumber="449">
<collectionCode id="9CCBAE56FFBDFFD4BB38A855FEAA115F" box="[113,260,564,587]" pageId="28" pageNumber="449">
<typeStatus id="25618831FFBDFFD4BB38A855FF0C115C" box="[113,162,564,587]" pageId="28" pageNumber="449">TYPE</typeStatus>
LOCALITY
</collectionCode>
. — Lissieu,
<collectingCountry id="82CD7603FFBDFFD4BA36A855FE6D115E" box="[383,451,564,587]" name="France" pageId="28" pageNumber="449">France</collectingCountry>
.
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38A804FEBE1183" blockId="28.[111,647,611,662]" pageId="28" pageNumber="449">OCCURRENCE IN THE STUDIED LAYERS. — MCC3, MCC4, MCC5, MCC7.</paragraph>
</subSubSection>
<subSubSection id="B2C06518FFBDFFD4BB38A8CFFC0D17F2" pageId="28" pageNumber="449" type="description">
<paragraph id="FA653693FFBDFFD4BB38A8CFFD991000" blockId="28.[113,648,685,789]" pageId="28" pageNumber="449">REFERRED MATERIAL. — A single maxillary fragment bearing M1 and M2; two isolated P4; 18 isolated M1; 15 isolated M2; three isolated M3; a single isolated p4; 22 isolated m1; 25 isolated m2; 9 isolated m3.</paragraph>
<paragraph id="FA653693FFBDFFD4BB38A94CFE271051" blockId="28.[113,393,812,836]" box="[113,393,812,836]" pageId="28" pageNumber="449">
MEASUREMENTS. —
<tableCitation id="B7580328FFBDFFD4BA73A94DFE271051" box="[314,393,812,836]" captionStart="TABLE" captionStartId="27.[113,122,219,237]" captionTargetPageId="27" captionText="TABLE 9. — Dental measurements (in mm) of Muscardinus vireti Hugueney &amp; Mein, 1965 from Moncucco Torinese. Abbreviations: see Table 1." pageId="28" pageNumber="449">Table 9.</tableCitation>
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38A913FEAC109F" blockId="28.[111,646,882,1003]" box="[113,258,882,908]" pageId="28" pageNumber="449">DESCRIPTION</paragraph>
<paragraph id="FA653693FFBDFFD4BB38A9F0FF2110BE" blockId="28.[111,646,882,1003]" box="[113,143,913,939]" pageId="28" pageNumber="449">
<emphasis id="C8AEEA81FFBDFFD4BB38A9F0FF2110BE" box="[113,143,913,939]" italics="true" pageId="28" pageNumber="449">P4</emphasis>
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38A9D1FE5410FF" blockId="28.[111,646,882,1003]" pageId="28" pageNumber="449">Ovoid in outline; three main ridges; first and second ridges connected on the lingual side.</paragraph>
<paragraph id="FA653693FFBDFFD4BB38AE71FF36173F" blockId="28.[111,649,1040,1319]" box="[113,152,1040,1066]" pageId="28" pageNumber="449">
<emphasis id="C8AEEA81FFBDFFD4BB38AE71FF36173F" box="[113,152,1040,1066]" italics="true" pageId="28" pageNumber="449">M1</emphasis>
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38AE4EFE511632" blockId="28.[111,649,1040,1319]" pageId="28" pageNumber="449">
Six main ridges; first ridge convex and lingually bent forming a “crochet” (
<emphasis id="C8AEEA81FFBDFFD4BACCAE31FE17177C" box="[389,441,1104,1129]" italics="true" pageId="28" pageNumber="449">sensu</emphasis>
<bibRefCitation id="9E4B4B62FFBDFFD4BAF7AE2EFF04179C" author="HUGUENEY M. &amp; MEIN P." pageId="28" pageNumber="449" pagination="109 - 123" refId="ref32228" refString="HUGUENEY M. &amp; MEIN P. 1965. - Lagomorphes et rongeurs du Neogene de Lissieu (Rhone). Travaux du Laboratoire de Geologie de la Faculte des Sciences de Lyon Nouvelle Serie 12: 109 - 123." type="journal article" year="1965">Hugueney &amp; Mein 1965</bibRefCitation>
); second ridge lingually inclined; all ridges but the first one connected by an elongate endoloph; a single specimen possesses small labial and lingual portions of an extra ridge situated between the third and fourth crests; four roots, three specimens with a supplementary anterolabial rootlet.
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38AF2DFF361673" blockId="28.[111,647,1356,1510]" box="[113,152,1356,1382]" pageId="28" pageNumber="449">
<emphasis id="C8AEEA81FFBDFFD4BB38AF2DFF361673" box="[113,152,1356,1382]" italics="true" pageId="28" pageNumber="449">M2</emphasis>
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38AF0DFD2F16F3" blockId="28.[111,647,1356,1510]" pageId="28" pageNumber="449">Eight main low ridges; a complete endoloph connects all the ridges; half of the specimens exhibit a small labial portion of an extra ridge; usually arising as a bifurcation of the sixth ridge; three or four roots.</paragraph>
<paragraph id="FA653693FFBDFFD4BB38AC6BFF361531" blockId="28.[111,648,1546,1700]" box="[113,152,1546,1572]" pageId="28" pageNumber="449">
<emphasis id="C8AEEA81FFBDFFD4BB38AC6BFF361531" box="[113,152,1546,1572]" italics="true" pageId="28" pageNumber="449">M3</emphasis>
</paragraph>
<paragraph id="FA653693FFBDFFD4BB38AC4BFD2915B6" blockId="28.[111,648,1546,1700]" pageId="28" pageNumber="449">Trapezoid or triangular in outline; eight main ridges connected by a complete endoloph; half of the specimens with a small labial portion of an extra ridge; usually as a bifurcation of the fifth or the</paragraph>
<paragraph id="FA653693FFBDFFD4B9EFAAB9FC6B1207" blockId="28.[677,1211,216,274]" pageId="28" pageNumber="449">sixth ridge; rare specimens with two labial portion of extra ridges; three roots.</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECAB56FD111244" blockId="28.[677,1210,311,401]" box="[677,703,311,337]" pageId="28" pageNumber="449">
<emphasis id="C8AEEA81FFBDFFD4B9ECAB56FD111244" box="[677,703,311,337]" italics="true" pageId="28" pageNumber="449">p4</emphasis>
</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECAB37FC1F1285" blockId="28.[677,1210,311,401]" pageId="28" pageNumber="449">Ovoid in outline; three main ridges weakly connected on the labial side.</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECABD7FD6912C5" blockId="28.[675,1210,438,623]" box="[677,711,438,464]" pageId="28" pageNumber="449">
<emphasis id="C8AEEA81FFBDFFD4B9ECABD7FD6912C5" box="[677,711,438,464]" italics="true" pageId="28" pageNumber="449">m1</emphasis>
</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECABB4FBFF117A" blockId="28.[675,1210,438,623]" pageId="28" pageNumber="449">Six main ridges; first and second ridges connected both on labial and lingual sides; third ridge convex or straight; forth, fifth and sixth ridges slightly convex; fifth and sixth ridges sometimes connected on the labial or lingual side; three roots.</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECA8F2FD6911B8" blockId="28.[675,1211,659,1003]" box="[677,711,659,685]" pageId="28" pageNumber="449">
<emphasis id="C8AEEA81FFBDFFD4B9ECA8F2FD6911B8" box="[677,711,659,685]" italics="true" pageId="28" pageNumber="449">m2</emphasis>
</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECA8D2FBEA10FE" blockId="28.[675,1211,659,1003]" pageId="28" pageNumber="449">Six main ridges; lingual side of the molar slightly longer than labial one; a swelling of the enamel occasionally occurs on the anterolabial border of the molar; ridges frequently connected on the labial side, most notably between first and second and between fifth and sixth ridges; a lingual portion of an extra ridge regularly present between the third and fourth ridges; in half of the specimens a smaller labial portion of an extra ridge is present between the third and fourth ridges; four roots.</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECAE71FD69173F" blockId="28.[675,1212,1040,1255]" box="[677,711,1040,1066]" pageId="28" pageNumber="449">
<emphasis id="C8AEEA81FFBDFFD4B9ECAE71FD69173F" box="[677,711,1040,1066]" italics="true" pageId="28" pageNumber="449">m3</emphasis>
</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECAE4EFC0D17F2" blockId="28.[675,1212,1040,1255]" pageId="28" pageNumber="449">Trapezoid in outline; lingual side longer than labial one; six main ridges; first and second ridges and fifth and sixth ridges occasionally connected on the labial and lingual side; 60% of specimens with a small lingual portion of an extra ridge between third and fourth ridges.</paragraph>
</subSubSection>
<subSubSection id="B2C06518FFBDFFD5B9ECAF6CFC2015B6" lastPageId="29" lastPageNumber="450" pageId="28" pageNumber="449" type="discussion">
<paragraph id="FA653693FFBDFFD4B9ECAF6CFCA71630" blockId="28.[675,1211,1293,1700]" box="[677,777,1293,1319]" pageId="28" pageNumber="449">REMARKS</paragraph>
<paragraph id="FA653693FFBDFFD4B9ECAF4CFC1B16B3" blockId="28.[675,1211,1293,1700]" pageId="28" pageNumber="449">
The flattened molars bearing a high number of roughly parallel ridges clearly support the attribution of the material documented herein to the genus
<taxonomicName id="3DDA4D10FFBDFFD4B9EAAFEDFC1F16B3" authority="Kaup, 1829" authorityName="Kaup" authorityYear="1829" box="[675,945,1420,1446]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="449" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBDFFD4B9EAAFEDFC8916B3" box="[675,807,1420,1446]" italics="true" pageId="28" pageNumber="449">Muscardinus</emphasis>
Kaup, 1829
</taxonomicName>
.
</paragraph>
<paragraph id="FA653693FFBDFFD5B9F2AFCAFD2812A5" blockId="28.[675,1211,1293,1700]" lastBlockId="29.[111,649,216,1699]" lastPageId="29" lastPageNumber="450" pageId="28" pageNumber="449">
The measurements of the studied material fit well with those of
<taxonomicName id="3DDA4D10FFBDFFD4B87CAFAAFD751510" authority="Fejfar &amp; Storch, 1990" authorityName="Fejfar &amp; Storch" authorityYear="1990" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="449" phylum="Chordata" rank="species" species="helleri">
<emphasis id="C8AEEA81FFBDFFD4B87CAFAAFBAD16F0" box="[821,1027,1483,1509]" italics="true" pageId="28" pageNumber="449">Muscardinus helleri</emphasis>
<bibRefCitation id="9E4B4B62FFBDFFD4BF41AFAAFD751510" author="FEJFAR O. &amp; STORCH G." pageId="28" pageNumber="449" pagination="139 - 184" refId="ref31419" refString="FEJFAR O. &amp; STORCH G. 1990. - Eine Pliozane (ober- Ruscinische) Kleinsaugerfauna aus Gundersheim, Rheinhessen- 1. Nagetiere: Mammalia, Rodentia. Senckenbergiana Lethaea 71: 139 - 184." type="journal article" year="1990">Fejfar &amp; Storch, 1990</bibRefCitation>
</taxonomicName>
from the Ruscinian locality of Gundersheim 4, even if the width of this latter species is generally larger. The morphology of the teeth from MCC is roughly similar to that of
<taxonomicName id="3DDA4D10FFBDFFD4B8A8AC2AFBE51571" authorityName="Fejfar &amp; Storch" authorityYear="1990" box="[993,1099,1610,1636]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="28" pageNumber="449" phylum="Chordata" rank="species" species="helleri">
<emphasis id="C8AEEA81FFBDFFD4B8A8AC2AFBE51571" box="[993,1099,1610,1636]" italics="true" pageId="28" pageNumber="449">M. helleri</emphasis>
</taxonomicName>
primarily concerning the presence of six main ridges on the lower molars and M1, eight ridges on M2 and M3, as well as small lingual and labial extra ridges on the m2. However, some minor differences can be recognized, including the presence of a short endoloph in the M1 and M
<quantity id="3D229B76FFBCFFD5BA37AB56FE091244" box="[382,423,311,337]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="29" pageNumber="450" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="3DDA4D10FFBCFFD5BAE4AB59FDBD1244" authorityName="Fejfar &amp; Storch" authorityYear="1990" box="[429,531,311,337]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="helleri">
<emphasis id="C8AEEA81FFBCFFD5BAE4AB59FDBD1244" box="[429,531,311,337]" italics="true" pageId="29" pageNumber="450">M. helleri</emphasis>
</taxonomicName>
not reaching the last two ridges, the absence of extra ridges in the M2, and the presence in fewer specimens with a lingual “crochet” on the first ridge of M1.
</paragraph>
<paragraph id="FA653693FFBCFFD5BBC1ABD4FD2911B8" blockId="29.[111,649,216,1699]" pageId="29" pageNumber="450">
The European Plio-Pleistocene
<taxonomicName id="3DDA4D10FFBCFFD5BAB0ABD4FE3712FA" authority="Kowalski, 1963" authorityName="Kowalski" authorityYear="1963" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="pliocaenicus">
<emphasis id="C8AEEA81FFBCFFD5BAB0ABD4FF4212FA" italics="true" pageId="29" pageNumber="450">Muscardinus pliocaenicus</emphasis>
<bibRefCitation id="9E4B4B62FFBCFFD5BBBBABB4FE3712FA" author="KOWALSKI K." box="[242,409,469,495]" pageId="29" pageNumber="450" pagination="533 - 567" refId="ref32461" refString="KOWALSKI K. 1963. - The Pliocene and Pleistocene Gliridae (Mammalia, Rodentia) from Poland. Acta Zoologica Cracoviensia 8: 533 - 567." type="journal article" year="1963">Kowalski, 1963</bibRefCitation>
</taxonomicName>
and the Late Miocene
<taxonomicName id="3DDA4D10FFBCFFD5BB39AB94FF52113A" authority="Bachmayer &amp; Wilson, 1970" authorityName="Bachmayer &amp; Wilson" authorityYear="1970" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="subSpecies" species="pliocaenicus" subSpecies="austriacus">
<emphasis id="C8AEEA81FFBCFFD5BB39AB94FE46111A" box="[112,488,501,527]" italics="true" pageId="29" pageNumber="450">Muscardinus pliocaenicus austriacus</emphasis>
<bibRefCitation id="9E4B4B62FFBCFFD5BAA7AB94FF52113A" author="BACHMAYER W. &amp; WILSON R. W." pageId="29" pageNumber="450" pagination="533 - 587" refId="ref29281" refString="BACHMAYER W. &amp; WILSON R. W. 1970. - Die Fauna der altpliozanen Hohlen und Spaltenfullungen bei Kohfidisch, Burgenland Osterreich). Annalen des Naturhistorischen Museums in Wien 74: 533 - 587." type="journal article" year="1970">Bachmayer &amp; Wilson, 1970</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="82CD7603FFBCFFD5BA71A874FE2F113A" box="[312,385,533,559]" name="Austria" pageId="29" pageNumber="450">Austria</collectingCountry>
(see also Daxner-Höck &amp;
<bibRefCitation id="9E4B4B62FFBCFFD5BB26A855FF47115B" author="HOCK G. &amp; HOCK E." box="[111,233,564,590]" pageId="29" pageNumber="450" pagination="375 - 444" refId="ref30576" refString="DAXNER- HOCK G. &amp; HOCK E. 2009. - New data on Eomyidae and Gliridae (Rodentia, Mammalia) from the Late Miocene of Austria. Annalen des Naturhistorischen Museums in Wien 111 A: 375 - 444." type="journal article" year="2009">Höck 2009</bibRefCitation>
) are slightly smaller than
<taxonomicName id="3DDA4D10FFBCFFD5BAB8A854FDE5115B" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[497,587,565,590]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBCFFD5BAB8A854FDE5115B" box="[497,587,565,590]" italics="true" pageId="29" pageNumber="450">M. vireti</emphasis>
</taxonomicName>
from MCC. Moreover, these taxa are characterized by a minor number of ridges in the M1 and M2 and by the absence of extra ridges in the lower molars.
</paragraph>
<paragraph id="FA653693FFBCFFD5BBC1A8D2FDA71079" blockId="29.[111,649,216,1699]" pageId="29" pageNumber="450">
<taxonomicName id="3DDA4D10FFBCFFD5BBC1A8D2FDB611D8" authority="Kormos, 1930" authorityName="Kormos" authorityYear="1930" box="[136,536,691,718]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="dacicus">
<emphasis id="C8AEEA81FFBCFFD5BBC1A8D2FEC711D8" box="[136,361,691,717]" italics="true" pageId="29" pageNumber="450">Muscardinus dacicus</emphasis>
Kormos, 1930
</taxonomicName>
from the Plio-Pleistocene of Europe shows a larger size and can be easily distinguished from the material of MCC by its simplified dental pattern without extra ridges on the m2 and with less developed ridges separated by larger valleys in the M1.
</paragraph>
<paragraph id="FA653693FFBCFFD5BBC1A910FF44171F" blockId="29.[111,649,216,1699]" pageId="29" pageNumber="450">
The Late Miocene species
<taxonomicName id="3DDA4D10FFBCFFD5BAF8A910FE3810B9" authority="Hugueney &amp; Mein, 1965" authorityName="Hugueney &amp; Mein" authorityYear="1965" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="davidi">
<emphasis id="C8AEEA81FFBCFFD5BAF8A910FD28109E" box="[433,646,881,907]" italics="true" pageId="29" pageNumber="450">Muscardinus davidi</emphasis>
<bibRefCitation id="9E4B4B62FFBCFFD5BB3AA9F0FE3810B9" author="HUGUENEY M. &amp; MEIN P." box="[115,406,913,940]" pageId="29" pageNumber="450" pagination="109 - 123" refId="ref32228" refString="HUGUENEY M. &amp; MEIN P. 1965. - Lagomorphes et rongeurs du Neogene de Lissieu (Rhone). Travaux du Laboratoire de Geologie de la Faculte des Sciences de Lyon Nouvelle Serie 12: 109 - 123." type="journal article" year="1965">Hugueney &amp; Mein, 1965</bibRefCitation>
</taxonomicName>
from Lissieu is characterized by a size comparable to that of MCC from which it differs in having less seven ridges in the M2.
</paragraph>
<paragraph id="FA653693FFBCFFD5BBC1AE6EFE4E16F0" blockId="29.[111,649,216,1699]" pageId="29" pageNumber="450">
The measurements of
<taxonomicName id="3DDA4D10FFBCFFD5BA3EAE6EFEEF177C" authority="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal, 2008" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="C8AEEA81FFBCFFD5BA3EAE6EFD28173C" box="[375,646,1039,1065]" italics="true" pageId="29" pageNumber="450">Muscardinus meridionalis</emphasis>
García-Alix, Minwer-Barakat, Martín-Suárez &amp; Freudenthal, 2008
</taxonomicName>
from the localities Purcal 24 (Late Turolian) and Purcal 4 (Early Ruscinian) in the
<collectingCountry id="82CD7603FFBCFFD5BBD5AEEFFF5417BD" box="[156,250,1166,1192]" name="Grenada" pageId="29" pageNumber="450">Granada</collectingCountry>
Basin (García-Alix
<emphasis id="C8AEEA81FFBCFFD5BA9DAEEEFDA417BD" box="[468,522,1166,1192]" italics="true" pageId="29" pageNumber="450">et al.</emphasis>
2008c) are only slightly larger than those of the studied material from MCC. The general pattern of the molars is rather similar in displaying a similar number of main ridges in each dental element. However, in
<taxonomicName id="3DDA4D10FFBCFFD5BB3BAF4CFEB61652" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" box="[114,280,1325,1351]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="C8AEEA81FFBCFFD5BB3BAF4CFEB61652" box="[114,280,1325,1351]" italics="true" pageId="29" pageNumber="450">M. meridionalis</emphasis>
</taxonomicName>
extra ridges are not present on M2 and M3, the endoloph of upper molars seems to be weaker, the “crochet” on the first ridge of the M1 only rarely occurs and the extra ridges on the lower molars are extremely infrequent in the m2 and completely absent in the m3.
</paragraph>
<paragraph id="FA653693FFBCFFD5BBC1AF8AFC6412A5" blockId="29.[111,649,216,1699]" lastBlockId="29.[675,1211,216,1699]" pageId="29" pageNumber="450">
Compared with
<taxonomicName id="3DDA4D10FFBCFFD5BA7FAF8AFF561530" authority="Hugueney &amp; Mein, 1965" authorityName="Hugueney &amp; Mein" authorityYear="1965" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBCFFD5BA7FAF8AFE5B1511" box="[310,501,1515,1541]" italics="true" pageId="29" pageNumber="450">Muscardinus vireti</emphasis>
<bibRefCitation id="9E4B4B62FFBCFFD5BAB2AF8AFF561530" author="HUGUENEY M. &amp; MEIN P." pageId="29" pageNumber="450" pagination="109 - 123" refId="ref32228" refString="HUGUENEY M. &amp; MEIN P. 1965. - Lagomorphes et rongeurs du Neogene de Lissieu (Rhone). Travaux du Laboratoire de Geologie de la Faculte des Sciences de Lyon Nouvelle Serie 12: 109 - 123." type="journal article" year="1965">Hugueney &amp; Mein, 1965</bibRefCitation>
</taxonomicName>
from Lissieu, the specimens from MCC display slightly larger mean lengths and widths even if the size ranges partially overlap and also show a very similar morphology. In
<taxonomicName id="3DDA4D10FFBCFFD5B96EAC0BFD2B1596" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[551,645,1642,1667]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBCFFD5B96EAC0BFD2B1596" box="[551,645,1642,1667]" italics="true" pageId="29" pageNumber="450">M. vireti</emphasis>
</taxonomicName>
the number of ridges is identical for each dental element (six in M1 and lower molars, eight in M2 and M3), the endoloph is well-developed and connects the last five ridges of M1, lingual and labial portions of extra ridges occur in many m2 and, finally, some M2 and M3 exhibit incomplete extra ridges very similar in shape to those observed in the specimens from MCC.
</paragraph>
<paragraph id="FA653693FFBCFFD5B9F2ABD7FC8511D8" blockId="29.[675,1211,216,1699]" pageId="29" pageNumber="450">
A few specimens assigned to
<taxonomicName id="3DDA4D10FFBCFFD5BF4CABD4FD7212FA" authorityName="Hugueney &amp; Mein" authorityYear="1965" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBCFFD5BF4CABD4FB2312DA" box="[1029,1165,437,463]" italics="true" pageId="29" pageNumber="450">Muscardinus</emphasis>
aff.
<emphasis id="C8AEEA81FFBCFFD5B9ECABB7FD7212FA" box="[677,732,470,495]" italics="true" pageId="29" pageNumber="450">vireti</emphasis>
</taxonomicName>
were reported from Baccinello V3 (
<bibRefCitation id="9E4B4B62FFBCFFD5BF10ABB4FD4C111A" author="ENGESSER B." pageId="29" pageNumber="450" pagination="763 - 780" refId="ref31261" refString="ENGESSER B. 1983. - Die jungtertiaren Kleinsauger des Gebietes der Maremma (Toskana, Italien). 1. Teil: Gliridae (Rodentia, Mammalia). Eclogae Geologicae Helvetiae 76: 763 - 780." type="journal article" year="1983">Engesser 1983</bibRefCitation>
). The measurements of that material are intermediate between those from MCC and Lissieu except for the m3 that are smaller. From the morphological point of view, the arrangement of the ridges of the molars is remarkably similar to that from MCC and no reliable differences can be detected.
</paragraph>
<paragraph id="FA653693FFBCFFD5B9F2A8B2FC7910FE" blockId="29.[675,1211,216,1699]" pageId="29" pageNumber="450">
The
<taxonomicName id="3DDA4D10FFBCFFD5B9A1A8B2FCC711F8" authorityName="Kaup" authorityYear="1829" box="[744,873,723,749]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBCFFD5B9A1A8B2FCC711F8" box="[744,873,723,749]" italics="true" pageId="29" pageNumber="450">Muscardinus</emphasis>
</taxonomicName>
material from MCC is therefore assigned to
<taxonomicName id="3DDA4D10FFBCFFD5B850A892FCDC1019" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[793,882,755,780]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBCFFD5B850A892FCDC1019" box="[793,882,755,780]" italics="true" pageId="29" pageNumber="450">M. vireti</emphasis>
</taxonomicName>
. The slight size differences with the material from Lissieu can be explained by the different age of the two localities, since Lissieu is probably slightly older than MCC (
<bibRefCitation id="9E4B4B62FFBCFFD5BF6EA930FCBA109E" author="GOMEZ CANO A. R. &amp; HERNANDEZ FERNANDEZ M. &amp; SIERRA M. A." pageId="29" pageNumber="450" pagination="193 - 204" refId="ref32052" refString="GOMEZ CANO A. R., HERNANDEZ FERNANDEZ M. &amp; ALVAREZ- SIERRA M. A. 2011. - Biogeographic provincialism in rodent faunas from the Iberoccitanian Region (southwestern Europe) generates severe diachrony within the Mammalian Neogene (MN) biochronologic scale during the Late Miocene. Palaeogeography, Palaeoclimatology, Palaeoecology 307: 193 - 204." type="journal article" year="2011">
Gómez Cano
<emphasis id="C8AEEA81FFBCFFD5B9EDA913FD7B109E" box="[676,725,881,907]" italics="true" pageId="29" pageNumber="450">et al.</emphasis>
2011
</bibRefCitation>
). The intermediate size of the teeth from Baccinello V3 confirms that slight size differences are to be expected in assemblages of different age and geographic provenance.
</paragraph>
<paragraph id="FA653693FFBCFFD5B9F2A991FC2015B6" blockId="29.[675,1211,216,1699]" pageId="29" pageNumber="450">
According to García-Alix
<emphasis id="C8AEEA81FFBCFFD5B880A990FC54171F" box="[969,1018,1008,1034]" italics="true" pageId="29" pageNumber="450">et al.</emphasis>
(2008c),
<taxonomicName id="3DDA4D10FFBCFFD5BF16A990FB14171F" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[1119,1210,1009,1034]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBCFFD5BF16A990FB14171F" box="[1119,1210,1009,1034]" italics="true" pageId="29" pageNumber="450">M. vireti</emphasis>
</taxonomicName>
should be considered the ancestor of
<taxonomicName id="3DDA4D10FFBCFFD5BF6BAE71FD15175C" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="C8AEEA81FFBCFFD5BF6BAE71FD15175C" italics="true" pageId="29" pageNumber="450">M. meridionalis</emphasis>
</taxonomicName>
. This interpretation is followed herein, justified by several relevant morphological features that are shared by these two taxa, including the number of crests in upper and lower molars. Actually,
<taxonomicName id="3DDA4D10FFBCFFD5BF22AEEEFCA617DD" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="C8AEEA81FFBCFFD5BF22AEEEFCA617DD" italics="true" pageId="29" pageNumber="450">M. meridionalis</emphasis>
</taxonomicName>
only differs in having a slightly modified dental pattern with very uncommon extra ridges, a much rarer occurrence of the “crochet” on the M1, and the weaker endoloph. García-Alix
<emphasis id="C8AEEA81FFBCFFD5BFCCAF6FFB141632" box="[1157,1210,1293,1319]" italics="true" pageId="29" pageNumber="450">et al.</emphasis>
(2008c) considered
<taxonomicName id="3DDA4D10FFBCFFD5B8CAAF4CFC431652" authorityName="Fejfar &amp; Storch" authorityYear="1990" box="[899,1005,1325,1351]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="helleri">
<emphasis id="C8AEEA81FFBCFFD5B8CAAF4CFC431652" box="[899,1005,1325,1351]" italics="true" pageId="29" pageNumber="450">M. helleri</emphasis>
</taxonomicName>
as the descendant of
<taxonomicName id="3DDA4D10FFBCFFD5B988AF2CFCC61673" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" box="[705,872,1356,1382]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="C8AEEA81FFBCFFD5B988AF2CFCC61673" box="[705,872,1356,1382]" italics="true" pageId="29" pageNumber="450">M. meridionalis</emphasis>
</taxonomicName>
. Such hypothesis appears to be weak considering that
<taxonomicName id="3DDA4D10FFBCFFD5B8DDAF0CFC571693" authorityName="Fejfar &amp; Storch" authorityYear="1990" box="[916,1017,1388,1414]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="helleri">
<emphasis id="C8AEEA81FFBCFFD5B8DDAF0CFC571693" box="[916,1017,1388,1414]" italics="true" pageId="29" pageNumber="450">M. helleri</emphasis>
</taxonomicName>
displays a slightly more complicated dental pattern than
<taxonomicName id="3DDA4D10FFBCFFD5BF0AAFEDFD4E16D0" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="C8AEEA81FFBCFFD5BF0AAFEDFD4E16D0" italics="true" pageId="29" pageNumber="450">M. meridionalis</emphasis>
</taxonomicName>
, with more developed extra ridges in the m2. Therefore, the lineage
<emphasis id="C8AEEA81FFBCFFD5B8DEAFADFCBD1510" italics="true" pageId="29" pageNumber="450">
<taxonomicName id="3DDA4D10FFBCFFD5B8DEAFADFC5A16F0" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[919,1012,1484,1509]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="vireti">M. vireti</taxonomicName>
<taxonomicName id="3DDA4D10FFBCFFD5BF4BAFADFB0516F0" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" box="[1026,1195,1483,1509]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">M. meridionalis</taxonomicName>
<taxonomicName id="3DDA4D10FFBCFFD5B9ECAF8AFCBD1510" authorityName="Fejfar &amp; Storch" authorityYear="1990" box="[677,787,1515,1541]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="helleri">M. helleri</taxonomicName>
</emphasis>
seems to be dubious since it would imply a simplification of the dental pattern (
<emphasis id="C8AEEA81FFBCFFD5BFD3AC6AFC381551" italics="true" pageId="29" pageNumber="450">
<taxonomicName id="3DDA4D10FFBCFFD5BFD3AC6AFD721551" authorityName="Hugueney &amp; Mein" authorityYear="1965" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="vireti">M. vireti</taxonomicName>
<taxonomicName id="3DDA4D10FFBCFFD5B9A3AC4AFC381551" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" box="[746,918,1578,1604]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">M. meridionalis</taxonomicName>
</emphasis>
) followed by a reversal of this trend (
<emphasis id="C8AEEA81FFBCFFD5B85EAC2AFB8B1571" box="[791,1061,1610,1636]" italics="true" pageId="29" pageNumber="450">
<taxonomicName id="3DDA4D10FFBCFFD5B85EAC2AFC161571" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" box="[791,952,1610,1636]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">M. meridionalis</taxonomicName>
<taxonomicName id="3DDA4D10FFBCFFD5B88AAC2AFB8B1571" authorityName="Fejfar &amp; Storch" authorityYear="1990" box="[963,1061,1610,1636]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="helleri">M. helleri</taxonomicName>
</emphasis>
).
<taxonomicName id="3DDA4D10FFBCFFD5BF70AC2BFC891596" authorityName="Garcia-Alix, Minwer-Barakat, Martin-Suarez &amp; Freudenthal" authorityYear="2008" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="29" pageNumber="450" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="C8AEEA81FFBCFFD5BF70AC2BFC891596" italics="true" pageId="29" pageNumber="450">Muscardinus meridionalis</emphasis>
</taxonomicName>
possibly represents a taxon exclusive from southern
<collectingCountry id="82CD7603FFBCFFD5B800ACE8FC2415B6" box="[841,906,1673,1699]" name="Spain" pageId="29" pageNumber="450">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>