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<document id="58CDA5290739FC18908457DA2516DBBC" ID-CLB-Dataset="6393" ID-DOI="10.5252/g2014n3a4" ID-GBIF-Dataset="08337548-044b-4fce-8702-fe17f207d9a2" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613150662339" checkinUser="felipe" docAuthor="Colombero, Simone, Pavia, Giulio &amp; Carnevale, Giorgio" docDate="2014" docId="72738785FFB8FFD3BBFBAE0FFBEC15B6" docLanguage="en" docName="Geodiversitas.36.3.421-475.pdf" docOrigin="Geodiversitas 36 (3)" docSource="http://dx.doi.org/10.5252/g2014n3a4" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Eliomys intermedius Friant 1953" docType="treatment" docVersion="6" lastPageNumber="447" masterDocId="8E4AFFFDFFA1FFC8BB49AA61FFAE1315" masterDocTitle="Messinian rodents from Moncucco Torinese, NW Italy: palaeobiodiversity and biochronology" masterLastPageNumber="475" masterPageNumber="421" pageNumber="446" updateTime="1698929471688" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="C66EDCA95731DEAE6A410F51713EB843">Messinian rodents from Moncucco Torinese, NW Italy: palaeobiodiversity and biochronology</mods:title>
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<treatment id="72738785FFB8FFD3BBFBAE0FFBEC15B6" ID-DOI="http://doi.org/10.5281/zenodo.4836533" ID-GBIF-Taxon="182404781" ID-Zenodo-Dep="4836533" LSID="urn:lsid:plazi:treatment:72738785FFB8FFD3BBFBAE0FFBEC15B6" httpUri="http://treatment.plazi.org/id/72738785FFB8FFD3BBFBAE0FFBEC15B6" lastPageId="27" lastPageNumber="447" pageId="25" pageNumber="446">
<subSubSection id="B2C06518FFB8FFD1BBFBAE0FFE1717BC" pageId="25" pageNumber="446" type="nomenclature">
<paragraph id="FA653693FFB8FFD1BBFBAE0FFE1717BC" blockId="25.[178,582,1134,1193]" pageId="25" pageNumber="446">
<taxonomicName id="3DDA4D10FFB8FFD1BBFBAE0FFDE8179C" authority="Friant, 1953" authorityName="Friant" authorityYear="1953" box="[178,582,1134,1161]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="446" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFB8FFD1BBFBAE0FFEA9179C" bold="true" box="[178,263,1134,1161]" italics="true" pageId="25" pageNumber="446">Eliomys</emphasis>
aff.
<emphasis id="C8AEEA81FFB8FFD1BA7EAE0FFE19179C" bold="true" box="[311,439,1134,1161]" italics="true" pageId="25" pageNumber="446">intermedius</emphasis>
Friant, 1953
</taxonomicName>
(
<figureCitation id="62E12A16FFB8FFD1BA0FAEEFFE2A17BC" box="[326,388,1166,1193]" captionStart="FIG" captionStartId="26.[113,124,913,931]" captionTargetBox="[122,1206,227,849]" captionTargetId="figure-331@26.[661,1211,213,777]" captionTargetPageId="26" captionText="FIG. 6. — Isolated teeth of Eliomys Wagner, 1840 and Muscardinus Kaup, 1829 from Moncucco Torinese:A-E, Eliomys aff.intermedius Friant, 1953; A, MGPT-PU12128348, P4 dex.; B, MGPT-PU128270 M1 sin.; C, MGPT-PU127495, M2 sin.; D, MGPT-PU128269, m1 dex.; E, MGPT-PU127127368, m2 dex.; F-P, Muscardinus vireti Hugueney &amp; Mein, 1965; F, MGPT-PU127604, P4 dex.; G, MGPT- PU12392, M1 dex.; H, MGPT-PU127606, M2 sin.; I, MGPT-PU128178, M2 dex.; J, MGPT-PU127607, M3 dex.; K, MGPT-PU128182, M3 dex.; L, MGPT-PU127608, p4 sin.; M, MGPT-PU127609, m1 dex.; N, MGPT-PU127610, m2 sin.; O, MGPT-PU127611, m3 sin.; P, MGPT-PU127513, m3 dex. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4538575" httpUri="https://zenodo.org/record/4538575/files/figure.png" pageId="25" pageNumber="446">Fig. 6</figureCitation>
A-E)
</paragraph>
</subSubSection>
<subSubSection id="B2C06518FFB8FFD1BB38AEA6FD2917CA" box="[113,647,1223,1249]" pageId="25" pageNumber="446" type="reference_group">
<paragraph id="FA653693FFB8FFD1BB38AEA6FD2917CA" blockId="25.[113,647,1223,1249]" box="[113,647,1223,1249]" pageId="25" pageNumber="446">
<treatmentCitationGroup id="DACA11BDFFB8FFD1BB38AEA6FD2917CA" box="[113,647,1223,1249]" pageId="25" pageNumber="446">
<taxonomicName id="3DDA4D10FFB8FFD1BB38AEA6FEA717CA" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[113,265,1223,1249]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="446" phylum="Chordata" rank="species" species="truci">
<emphasis id="C8AEEA81FFB8FFD1BB38AEA6FF1717CA" box="[113,185,1223,1247]" italics="true" pageId="25" pageNumber="446">Eliomys</emphasis>
cf.
<emphasis id="C8AEEA81FFB8FFD1BB94AEA9FEA717CA" box="[221,265,1224,1247]" italics="true" pageId="25" pageNumber="446">truci</emphasis>
</taxonomicName>
–
<treatmentCitation id="7B7B1082FFB8FFD1BA6AAEA6FDBB17CA" author="ANGELONE C. &amp; COLOMBERO S. &amp; ESU D. &amp; GIUNTELLI P. &amp; MARCOLINI F. &amp; PAVIA M. &amp; TRENKWALDER S. &amp; VAN DEN HOEK OSTENDE L. W. &amp; ZUNINO M. &amp; PAVIA G." box="[291,533,1223,1247]" page="99" pageId="25" pageNumber="446" year="2011">
<bibRefCitation id="9E4B4B62FFB8FFD1BA6AAEA6FDBB17CA" author="ANGELONE C. &amp; COLOMBERO S. &amp; ESU D. &amp; GIUNTELLI P. &amp; MARCOLINI F. &amp; PAVIA M. &amp; TRENKWALDER S. &amp; VAN DEN HOEK OSTENDE L. W. &amp; ZUNINO M. &amp; PAVIA G." box="[291,533,1223,1247]" pageId="25" pageNumber="446" pagination="89 - 104" refId="ref29204" refString="ANGELONE C., COLOMBERO S., ESU D., GIUNTELLI P., MARCOLINI F., PAVIA M., TRENKWALDER S., VAN DEN HOEK OSTENDE L. W., ZUNINO M. &amp; PAVIA G. 2011. - Moncucco Torinese, a new post-evaporitic Messinian fossiliferous site from Piedmont (NW Italy). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 259: 89 - 104." type="journal article" year="2011">
Angelone
<emphasis id="C8AEEA81FFB8FFD1BACCAEA9FE1A17CA" box="[389,436,1223,1247]" italics="true" pageId="25" pageNumber="446">et al.</emphasis>
2011: 99
</bibRefCitation>
</treatmentCitation>
, fig. 6 (17).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B2C06518FFB8FFD1BB38AE96FEBD164C" pageId="25" pageNumber="446" type="materials_examined">
<paragraph id="FA653693FFB8FFD1BB38AE96FE03161B" blockId="25.[113,429,1271,1295]" box="[113,429,1271,1295]" pageId="25" pageNumber="446">
<collectionCode id="9CCBAE56FFB8FFD1BB38AE96FEAA1618" box="[113,260,1271,1294]" pageId="25" pageNumber="446">
<typeStatus id="25618831FFB8FFD1BB38AE96FF0C1618" box="[113,162,1271,1294]" pageId="25" pageNumber="446">TYPE</typeStatus>
LOCALITY
</collectionCode>
. — Sète,
<collectingCountry id="82CD7603FFB8FFD1BA2CAE96FE07161B" box="[357,425,1271,1294]" name="France" pageId="25" pageNumber="446">France</collectingCountry>
.
</paragraph>
<paragraph id="FA653693FFB8FFD1BB38AF49FEBD164C" blockId="25.[113,646,1319,1369]" pageId="25" pageNumber="446">OCCURRENCE IN THE STUDIED LAYERS. — MCC4; MCC5; MCC7.</paragraph>
</subSubSection>
<subSubSection id="B2C06518FFB8FFD1BB38AF10FB151510" pageId="25" pageNumber="446" type="description">
<paragraph id="FA653693FFB8FFD1BB38AF10FEC716B6" blockId="25.[112,646,1392,1443]" pageId="25" pageNumber="446">REFERRED MATERIAL. — A single P4, three M1, two M2, two m1, and a single m2.</paragraph>
<paragraph id="FA653693FFB8FFD1BB38AFDAFE2716C6" blockId="25.[113,393,1467,1491]" box="[113,393,1467,1491]" pageId="25" pageNumber="446">
MEASUREMENTS. —
<tableCitation id="B7580328FFB8FFD1BA73AFDAFE2716C6" box="[314,393,1467,1491]" captionStart="TABLE" captionStartId="25.[117,124,219,237]" captionTargetPageId="25" captionText="TABLE 8. — Dental measurements (in mm) of Eliomys aff. intermedius Friant,1953 from Moncucco Torinese. Abbreviations: see Table 1." pageId="25" pageNumber="446">Table 8.</tableCitation>
</paragraph>
<paragraph id="FA653693FFB8FFD1BB38AF8AFEAC1516" blockId="25.[111,647,1515,1700]" box="[113,258,1515,1541]" pageId="25" pageNumber="446">DESCRIPTION</paragraph>
<paragraph id="FA653693FFB8FFD1BB38AC6AFF211530" blockId="25.[111,647,1515,1700]" box="[113,143,1547,1573]" pageId="25" pageNumber="446">
<emphasis id="C8AEEA81FFB8FFD1BB38AC6AFF211530" box="[113,143,1547,1573]" italics="true" pageId="25" pageNumber="446">P4</emphasis>
</paragraph>
<paragraph id="FA653693FFB8FFD1BB38AC4BFEB715B1" blockId="25.[111,647,1515,1700]" pageId="25" pageNumber="446">Triangular outline; anteroloph short and connected to the paracone; the anterior centroloph extends to the midpoint of the tooth; posteroloph not fused with metacone.</paragraph>
<paragraph id="FA653693FFB8FFD1B9ECA815FD4C119B" blockId="25.[675,1212,628,1034]" box="[677,738,628,654]" pageId="25" pageNumber="446">
<emphasis id="C8AEEA81FFB8FFD1B9ECA815FD4C119B" box="[677,738,628,654]" italics="true" pageId="25" pageNumber="446">M1-2</emphasis>
</paragraph>
<paragraph id="FA653693FFB8FFD1B9ECA8F2FC9C171F" blockId="25.[675,1212,628,1034]" pageId="25" pageNumber="446">M1 and M2 display a very similar dental pattern; the M2 usually presents a less concave anteroloph; molars are shallow basined with trapezoidal outline, the lingual side being slightly shorter than the labial one; endoloph strong and continuous; paracone slightly stouter and higher than the metacone; the anterior centroloph reaches the midpoint of the molar; posterior centroloph always present; in both M2 and in one of three M1 the posterior centroloph fuses with the anterior centroloph forming a “Y-shaped” ridge that extends to the midpoint of the molar.</paragraph>
<paragraph id="FA653693FFB8FFD1B9ECAE4EFD69175C" blockId="25.[674,1211,1071,1351]" box="[677,711,1071,1097]" pageId="25" pageNumber="446">
<emphasis id="C8AEEA81FFB8FFD1B9ECAE4EFD69175C" box="[677,711,1071,1097]" italics="true" pageId="25" pageNumber="446">m1</emphasis>
</paragraph>
<paragraph id="FA653693FFB8FFD1B9ECAE2EFC5C1652" blockId="25.[674,1211,1071,1351]" pageId="25" pageNumber="446">Trapezoid in outline; the endolophid is discontinuous; anterolophid connected to protoconid in one of the two available specimens; metalophid connected to metaconid; centrolophid extending beyond the midpoint of the molar; well-developed posterior extra ridge; in both specimens, the connection between hypoconid and posterolophid marked by a slight narrowing of the ridge.</paragraph>
<paragraph id="FA653693FFB8FFD1B9ECAF0DFD691693" blockId="25.[675,1212,1388,1541]" box="[677,711,1388,1414]" pageId="25" pageNumber="446">
<emphasis id="C8AEEA81FFB8FFD1B9ECAF0DFD691693" box="[677,711,1388,1414]" italics="true" pageId="25" pageNumber="446">m2</emphasis>
</paragraph>
<paragraph id="FA653693FFB8FFD1B9ECAFEDFB151510" blockId="25.[675,1212,1388,1541]" pageId="25" pageNumber="446">Broader than long; the endolophid is discontinous; anterolophid not connected to the protoconid; metalophid not connected with the metaconid; centrolophid short; very reduced posterior extra ridge.</paragraph>
</subSubSection>
<subSubSection id="B2C06518FFB8FFD3B9ECAC4AFBEC15B6" lastPageId="27" lastPageNumber="448" pageId="25" pageNumber="446" type="discussion">
<paragraph id="FA653693FFB8FFD1B9ECAC4AFCA71556" blockId="25.[675,1210,1579,1700]" box="[677,777,1579,1605]" pageId="25" pageNumber="446">REMARKS</paragraph>
<paragraph id="FA653693FFB8FFD2B9ECAC2BFE781672" blockId="25.[675,1210,1579,1700]" lastBlockId="26.[111,648,1103,1699]" lastPageId="26" lastPageNumber="447" pageId="25" pageNumber="446">
The presence of a continuous and strong endoloph in the upper molars excludes any possible assignment of the material from MCC to the genus
<taxonomicName id="3DDA4D10FFBBFFD2BB38AE31FE2D177C" authority="Ognev, 1924" authorityName="Ognev" authorityYear="1924" box="[113,387,1103,1129]" class="Mammalia" family="Gliridae" genus="Myomimus" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBBFFD2BB38AE31FF48177C" box="[113,230,1104,1129]" italics="true" pageId="26" pageNumber="447">Myomimus</emphasis>
Ognev, 1924
</taxonomicName>
. The presence of wider than long lower molars and continuous, strong endolophs in upper molars are characteristic of the genera
<taxonomicName id="3DDA4D10FFBBFFD2BBF3AECEFE0317DD" authority="Thomas, 1906" authorityName="Thomas" authorityYear="1906" box="[186,429,1198,1224]" class="Mammalia" family="Gliridae" genus="Dryomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBBFFD2BBF3AECEFEBC17DD" box="[186,274,1199,1224]" italics="true" pageId="26" pageNumber="447">Dryomys</emphasis>
Thomas, 1906
</taxonomicName>
and
<taxonomicName id="3DDA4D10FFBBFFD2BA97AECFFF0717FD" authority="Wagner, 1840" authorityName="Wagner" authorityYear="1840" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBBFFD2BA97AECFFD8517DD" box="[478,555,1198,1224]" italics="true" pageId="26" pageNumber="447">Eliomys</emphasis>
Wagner, 1840
</taxonomicName>
. According to Freudenthal &amp; Martín-Suárez (2006), the presence of an endolophid in the lower molars might be used to separate
<taxonomicName id="3DDA4D10FFBBFFD2BAA3AF6FFDE81632" authorityName="Thomas" authorityYear="1906" box="[490,582,1294,1319]" class="Mammalia" family="Gliridae" genus="Dryomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBBFFD2BAA3AF6FFDE81632" box="[490,582,1294,1319]" italics="true" pageId="26" pageNumber="447">Dryomys</emphasis>
</taxonomicName>
from
<taxonomicName id="3DDA4D10FFBBFFD2BB38AF4CFF6E1652" authorityName="Wagner" authorityYear="1840" box="[113,192,1325,1351]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBBFFD2BB38AF4CFF6E1652" box="[113,192,1325,1351]" italics="true" pageId="26" pageNumber="447">Eliomys</emphasis>
</taxonomicName>
. For this reason, we assign the specimens from MCC to the genus
<taxonomicName id="3DDA4D10FFBBFFD2BAC9AF2DFE611673" authorityName="Wagner" authorityYear="1840" box="[384,463,1356,1382]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBBFFD2BAC9AF2DFE611673" box="[384,463,1356,1382]" italics="true" pageId="26" pageNumber="447">Eliomys</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="AEA5661BFFBBFFD2BB38A9F0FE531701" ID-DOI="http://doi.org/10.5281/zenodo.4538575" ID-Zenodo-Dep="4538575" httpUri="https://zenodo.org/record/4538575/files/figure.png" pageId="26" pageNumber="447" startId="26.[113,124,913,931]" targetBox="[122,1206,227,849]" targetPageId="26">
<paragraph id="FA653693FFBBFFD2BB38A9F0FE531701" blockId="26.[113,1211,913,1044]" pageId="26" pageNumber="447">
FIG. 6. — Isolated teeth of
<taxonomicName id="3DDA4D10FFBBFFD2BA0EA9F0FE5010B6" authority="Wagner, 1840" authorityName="Wagner" authorityYear="1840" box="[327,510,913,931]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBBFFD2BA0EA9F0FE2910B6" box="[327,391,913,931]" italics="true" pageId="26" pageNumber="447">Eliomys</emphasis>
Wagner, 1840
</taxonomicName>
and
<taxonomicName id="3DDA4D10FFBBFFD2B96CA9F0FD5D10B6" authority="Kaup, 1829" authorityName="Kaup" authorityYear="1829" box="[549,755,913,931]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBBFFD2B96CA9F0FD2110B6" box="[549,655,913,931]" italics="true" pageId="26" pageNumber="447">Muscardinus</emphasis>
Kaup, 1829
</taxonomicName>
from Moncucco Torinese:
<emphasis id="C8AEEA81FFBBFFD2B885A9F0FC4210B6" bold="true" box="[972,1004,913,931]" pageId="26" pageNumber="447">A -E</emphasis>
,
<taxonomicName id="3DDA4D10FFBBFFD2B8BFA9F0FF7610AC" authority="Friant, 1953" authorityName="Friant" authorityYear="1953" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBBFFD2B8BFA9F0FB9810B6" box="[1014,1078,913,931]" italics="true" pageId="26" pageNumber="447">Eliomys</emphasis>
aff.
<emphasis id="C8AEEA81FFBBFFD2BF11A9F0FB1710B6" box="[1112,1209,913,931]" italics="true" pageId="26" pageNumber="447">intermedius</emphasis>
Friant, 1953
</taxonomicName>
;
<emphasis id="C8AEEA81FFBBFFD2BBA9A9C6FF4310AC" bold="true" box="[224,237,935,953]" pageId="26" pageNumber="447">A</emphasis>
, MGPT-PU12128348, P4 dex.;
<emphasis id="C8AEEA81FFBBFFD2BABDA9C6FDAF10AC" bold="true" box="[500,513,935,953]" pageId="26" pageNumber="447">B</emphasis>
, MGPT-PU128270 M1 sin.;
<emphasis id="C8AEEA81FFBBFFD2B9A5A9C6FD5410AC" bold="true" box="[748,762,935,953]" pageId="26" pageNumber="447">C</emphasis>
, MGPT-PU127495, M2 sin.;
<emphasis id="C8AEEA81FFBBFFD2B8A3A9C6FC5610AC" bold="true" box="[1002,1016,935,953]" pageId="26" pageNumber="447">D</emphasis>
, MGPT-PU128269, m1 dex.;
<emphasis id="C8AEEA81FFBBFFD2BBE8A9DFFF0310C5" bold="true" box="[161,173,958,976]" pageId="26" pageNumber="447">E</emphasis>
, MGPT-PU127127368, m2 dex.;
<emphasis id="C8AEEA81FFBBFFD2BA8FA9DFFE4A10C5" bold="true" box="[454,484,958,976]" pageId="26" pageNumber="447">F -P</emphasis>
,
<taxonomicName id="3DDA4D10FFBBFFD2BAB9A9DFFCF210C5" authority="Hugueney &amp; Mein, 1965" authorityName="Hugueney &amp; Mein" authorityYear="1965" box="[496,860,958,976]" class="Mammalia" family="Gliridae" genus="Muscardinus" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="species" species="vireti">
<emphasis id="C8AEEA81FFBBFFD2BAB9A9DFFD2610C5" box="[496,648,958,976]" italics="true" pageId="26" pageNumber="447">Muscardinus vireti</emphasis>
<bibRefCitation id="9E4B4B62FFBBFFD2B9C7A9DFFCF210C5" author="HUGUENEY M. &amp; MEIN P." box="[654,860,958,976]" pageId="26" pageNumber="447" pagination="109 - 123" refId="ref32228" refString="HUGUENEY M. &amp; MEIN P. 1965. - Lagomorphes et rongeurs du Neogene de Lissieu (Rhone). Travaux du Laboratoire de Geologie de la Faculte des Sciences de Lyon Nouvelle Serie 12: 109 - 123." type="journal article" year="1965">Hugueney &amp; Mein, 1965</bibRefCitation>
</taxonomicName>
;
<emphasis id="C8AEEA81FFBBFFD2B82CA9DFFCDE10C5" bold="true" box="[869,880,958,976]" pageId="26" pageNumber="447">F</emphasis>
, MGPT-PU127604, P4 dex.;
<emphasis id="C8AEEA81FFBBFFD2BF2FA9DFFBDA10C5" bold="true" box="[1126,1140,958,976]" pageId="26" pageNumber="447">G</emphasis>
, MGPT- PU12392, M1 dex.;
<emphasis id="C8AEEA81FFBBFFD2BA5EA9B4FE8B10F2" bold="true" box="[279,293,981,999]" pageId="26" pageNumber="447">H</emphasis>
, MGPT-PU127606, M2 sin.;
<emphasis id="C8AEEA81FFBBFFD2B95BA9B4FDB610F2" bold="true" box="[530,536,981,999]" pageId="26" pageNumber="447">I</emphasis>
, MGPT-PU128178, M2 dex.;
<emphasis id="C8AEEA81FFBBFFD2B845A9B4FCB810F2" bold="true" box="[780,790,981,999]" pageId="26" pageNumber="447">J</emphasis>
, MGPT-PU127607, M3 dex.;
<emphasis id="C8AEEA81FFBBFFD2BF42A9B4FBB610F2" bold="true" box="[1035,1048,981,999]" pageId="26" pageNumber="447">K</emphasis>
, MGPT-PU128182, M3 dex.;
<emphasis id="C8AEEA81FFBBFFD2BB88A98AFF6210E8" bold="true" box="[193,204,1003,1021]" pageId="26" pageNumber="447">L</emphasis>
, MGPT-PU127608, p4 sin.;
<emphasis id="C8AEEA81FFBBFFD2BAF1A98AFE6710E8" bold="true" box="[440,457,1003,1021]" pageId="26" pageNumber="447">M</emphasis>
, MGPT-PU127609, m1 dex.;
<emphasis id="C8AEEA81FFBBFFD2B988A98AFD6110E8" bold="true" box="[705,719,1003,1021]" pageId="26" pageNumber="447">N</emphasis>
, MGPT-PU127610, m2 sin.;
<emphasis id="C8AEEA81FFBBFFD2B889A98AFC6110E8" bold="true" box="[960,975,1003,1021]" pageId="26" pageNumber="447">O</emphasis>
, MGPT-PU127611, m3 sin.;
<emphasis id="C8AEEA81FFBBFFD2BB38AE63FFD31701" bold="true" box="[113,125,1026,1044]" pageId="26" pageNumber="447">P</emphasis>
, MGPT-PU127513, m3 dex. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="FA653693FFBBFFD2BBC1AF0DFC221510" blockId="26.[111,648,1103,1699]" lastBlockId="26.[674,1212,1103,1700]" pageId="26" pageNumber="447">
Direct comparisons with
<taxonomicName id="3DDA4D10FFBBFFD2BAD4AF0DFE8F16B3" authority="Mein &amp; Michaux, 1970" authorityName="Mein &amp; Michaux" authorityYear="1970" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="species" species="truci">
<emphasis id="C8AEEA81FFBBFFD2BAD4AF0DFD8B1693" box="[413,549,1388,1414]" italics="true" pageId="26" pageNumber="447">Eliomys truci</emphasis>
<bibRefCitation id="9E4B4B62FFBBFFD2B964AF0DFE8F16B3" author="MEIN P. &amp; MICHAUX J." pageId="26" pageNumber="447" pagination="2780 - 2783" refId="ref33062" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sudoccidentale. Comptes rendus de l'Academie des Sciences D 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux, 1970</bibRefCitation>
</taxonomicName>
, a species distributed in Europe between the Late Miocene and the Late Pliocene, from the Early Ruscinian locality of Hautimagne, evidenced that the specimens from MCC mainly differ in the presence of a posterior centroloph in the upper molars, a shorter centrolophid in the m2 and in the strong reduction of the secondary accessory ridge in the m2. Moreover, in the m1-2 of
<taxonomicName id="3DDA4D10FFBBFFD2BA90AC0BFD8B1596" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[473,549,1642,1667]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="species" species="truci">
<emphasis id="C8AEEA81FFBBFFD2BA90AC0BFD8B1596" box="[473,549,1642,1667]" italics="true" pageId="26" pageNumber="447">E. truci</emphasis>
</taxonomicName>
from the
<collectingCountry id="82CD7603FFBBFFD2BB38ACE8FF6215B6" box="[113,204,1673,1699]" name="Grenada" pageId="26" pageNumber="447">Granada</collectingCountry>
and Guadix basin, the metalophid-meta- conid connection and the anterolophid-protoconid connection are rarer compared to the material from MCC (García-Alix
<emphasis id="C8AEEA81FFBBFFD2B825AEEEFC3217BD" box="[876,924,1166,1192]" italics="true" pageId="26" pageNumber="447">et al.</emphasis>
2008a). The measurements of the teeth from MCC are slightly larger than those of
<taxonomicName id="3DDA4D10FFBBFFD2B98AAEAFFCBA17F2" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[707,788,1230,1255]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="species" species="truci">
<emphasis id="C8AEEA81FFBBFFD2B98AAEAFFCBA17F2" box="[707,788,1230,1255]" italics="true" pageId="26" pageNumber="447">E. truci</emphasis>
</taxonomicName>
from Hautimagne (
<bibRefCitation id="9E4B4B62FFBBFFD2B8BDAEACFD4C1612" author="MEIN P. &amp; MICHAUX J." pageId="26" pageNumber="447" pagination="2780 - 2783" refId="ref33062" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sudoccidentale. Comptes rendus de l'Academie des Sciences D 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux 1970</bibRefCitation>
), Los Mansuetos, Concud 2 and 3 (
<bibRefCitation id="9E4B4B62FFBBFFD2BF38AE8CFC85163D" author="VAN DE WEERD A." pageId="26" pageNumber="447" pagination="1 - 217" refId="ref35291" refString="VAN DE WEERD A. 1976. - Rodent faunas of the Mio-Pliocene continental sediments of the Teruel-Alfambra region, Spain. Utrecht Micropaleontological Bulletins 2: 1 - 217." type="journal article" year="1976">van de Weerd 1976</bibRefCitation>
), Tollo de Chiclana 13 and 1B, Purcal 23 and 25a, La Dehesa 1 and 16, and Otura 1 and 4 (García-Alix
<emphasis id="C8AEEA81FFBBFFD2B874AF2CFCC01673" box="[829,878,1356,1382]" italics="true" pageId="26" pageNumber="447">et al.</emphasis>
2008a), especially as regards the width, whereas the length of the specimens of some other assemblages such as Masada del Valle 2 and 5, Concud Barranco de las Calaveras (
<bibRefCitation id="9E4B4B62FFBBFFD2BF64AFCAFD7316F3" author="VAN DE WEERD A." pageId="26" pageNumber="447" pagination="1 - 217" refId="ref35291" refString="VAN DE WEERD A. 1976. - Rodent faunas of the Mio-Pliocene continental sediments of the Teruel-Alfambra region, Spain. Utrecht Micropaleontological Bulletins 2: 1 - 217." type="journal article" year="1976">van de Weerd 1976</bibRefCitation>
), and La Gloria 4 (
<bibRefCitation id="9E4B4B62FFBBFFD2B8D3AFAAFBC516F0" author="ADROVER R. &amp; MEIN P. &amp; MOISSENET E." box="[922,1131,1483,1509]" pageId="26" pageNumber="447" pagination="47 - 84" refId="ref28616" refString="ADROVER R., MEIN P. &amp; MOISSENET E. 1993 a. - Roedores de la transicion Mio-Plioceno de la region de Teruel. Paleontologia i Evolucio 26 - 27: 47 - 84." type="journal article" year="1993">
Adrover
<emphasis id="C8AEEA81FFBBFFD2B8BDAFADFB8A16F0" box="[1012,1060,1483,1509]" italics="true" pageId="26" pageNumber="447">et al.</emphasis>
1993a
</bibRefCitation>
) is very close to that of MCC.
</paragraph>
<paragraph id="FA653693FFBBFFD3B9F2AC6BFBEC15B6" blockId="26.[674,1212,1103,1700]" lastBlockId="27.[674,1212,1039,1700]" lastPageId="27" lastPageNumber="448" pageId="26" pageNumber="447">
The measurements of
<taxonomicName id="3DDA4D10FFBBFFD2B8FBAC6BFCA31550" authority="Friant, 1953" authorityName="Friant" authorityYear="1953" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="447" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBBFFD2B8FBAC6BFB261531" box="[946,1160,1546,1572]" italics="true" pageId="26" pageNumber="447">Eliomys intermedius</emphasis>
Friant, 1953
</taxonomicName>
, widespread in Europe between the Late Miocene and the Pleistocene, are usually larger than those of the sample from MCC (
<bibRefCitation id="9E4B4B62FFBBFFD2B8A2AC0BFB1D1596" author="VAN DE WEERD A." box="[1003,1203,1641,1668]" pageId="26" pageNumber="447" pagination="1 - 217" refId="ref35291" refString="VAN DE WEERD A. 1976. - Rodent faunas of the Mio-Pliocene continental sediments of the Teruel-Alfambra region, Spain. Utrecht Micropaleontological Bulletins 2: 1 - 217." type="journal article" year="1976">van de Weerd 1976</bibRefCitation>
; García-Alix
<emphasis id="C8AEEA81FFBBFFD2B86EACEBFCF215B6" box="[807,860,1673,1699]" italics="true" pageId="26" pageNumber="447">et al.</emphasis>
2008a). Recent observations on some teeth from the
<typeStatus id="25618831FFBAFFD3BA1CAE71FE2D173F" box="[341,387,1040,1066]" pageId="27" pageNumber="448">type</typeStatus>
locality of Sète revealed that the remains of
<taxonomicName id="3DDA4D10FFBAFFD3BA76AE4EFE23175C" authorityName="Wagner" authorityYear="1840" box="[319,397,1071,1097]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBAFFD3BA76AE4EFE23175C" box="[319,397,1071,1097]" italics="true" pageId="27" pageNumber="448">Eliomys</emphasis>
</taxonomicName>
from MCC share some morphological characteristics with this species, such as the presence of a posterior centroloph in the upper molars and the rounded outline of some dental elements, particularly the m2. Moreover,
<taxonomicName id="3DDA4D10FFBAFFD3BB3AAEAFFEA617F2" authorityName="Friant" authorityYear="1953" box="[115,264,1229,1255]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3BB3AAEAFFEA617F2" box="[115,264,1229,1255]" italics="true" pageId="27" pageNumber="448">E. intermedius</emphasis>
</taxonomicName>
displays metalophid-metaconid and anterolophid-protoconid connections as observed in the small sample of MCC. The morphology of the lower molars from MCC is also similar to that of
<taxonomicName id="3DDA4D10FFBAFFD3BB3AAF2CFEAB1673" authorityName="Friant" authorityYear="1953" box="[115,261,1356,1382]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3BB3AAF2CFEAB1673" box="[115,261,1356,1382]" italics="true" pageId="27" pageNumber="448">E. intermedius</emphasis>
</taxonomicName>
since the single m2 exhibits a strong reduction of the secondary posterior ridge even if in the latter species the reduction is stronger and frequently reported in the lower molars (García-Alix
<emphasis id="C8AEEA81FFBAFFD3B910AFCDFD2916D0" box="[601,647,1451,1477]" italics="true" pageId="27" pageNumber="448">et al.</emphasis>
2008a). In a previous paper,
<bibRefCitation id="9E4B4B62FFBAFFD3BAD4AFAAFD2916F0" author="ANGELONE C. &amp; COLOMBERO S. &amp; ESU D. &amp; GIUNTELLI P. &amp; MARCOLINI F. &amp; PAVIA M. &amp; TRENKWALDER S. &amp; VAN DEN HOEK OSTENDE L. W. &amp; ZUNINO M. &amp; PAVIA G." box="[413,647,1483,1509]" pageId="27" pageNumber="448" pagination="89 - 104" refId="ref29204" refString="ANGELONE C., COLOMBERO S., ESU D., GIUNTELLI P., MARCOLINI F., PAVIA M., TRENKWALDER S., VAN DEN HOEK OSTENDE L. W., ZUNINO M. &amp; PAVIA G. 2011. - Moncucco Torinese, a new post-evaporitic Messinian fossiliferous site from Piedmont (NW Italy). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 259: 89 - 104." type="journal article" year="2011">
Angelone
<emphasis id="C8AEEA81FFBAFFD3B94EAFADFD9616F0" box="[519,568,1483,1509]" italics="true" pageId="27" pageNumber="448">et al.</emphasis>
(2011)
</bibRefCitation>
assigned a single left M1 from MCC (MGPT-PU 127) (erroneously identified as an M3) to
<taxonomicName id="3DDA4D10FFBAFFD3B973AC6BFF6F1551" authorityName="Mein &amp; Michaux" authorityYear="1970" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="truci">
<emphasis id="C8AEEA81FFBAFFD3B973AC6BFD261531" box="[570,648,1546,1572]" italics="true" pageId="27" pageNumber="448">Eliomys</emphasis>
cf.
<emphasis id="C8AEEA81FFBAFFD3BBDBAC4AFF6F1551" box="[146,193,1579,1604]" italics="true" pageId="27" pageNumber="448">truci</emphasis>
</taxonomicName>
; this molar is referred herein to as
<taxonomicName id="3DDA4D10FFBAFFD3B971AC4BFEBC1571" authorityName="Friant" authorityYear="1953" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3B971AC4BFD281551" box="[568,646,1578,1604]" italics="true" pageId="27" pageNumber="448">Eliomys</emphasis>
aff.
<emphasis id="C8AEEA81FFBAFFD3BBD2AC2BFEBC1571" box="[155,274,1610,1636]" italics="true" pageId="27" pageNumber="448">intermedius</emphasis>
</taxonomicName>
because its morphological features fit well with
<taxonomicName id="3DDA4D10FFBAFFD3BBBBAC0BFE2B1596" authorityName="Friant" authorityYear="1953" box="[242,389,1641,1667]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3BBBBAC0BFE2B1596" box="[242,389,1641,1667]" italics="true" pageId="27" pageNumber="448">E. intermedius</emphasis>
</taxonomicName>
,even if the size is smaller and closer to that of
<taxonomicName id="3DDA4D10FFBAFFD3BA06ACEBFE3215B6" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[335,412,1674,1699]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="truci">
<emphasis id="C8AEEA81FFBAFFD3BA06ACEBFE3215B6" box="[335,412,1674,1699]" italics="true" pageId="27" pageNumber="448">E. truci</emphasis>
</taxonomicName>
. Some assemblages of
<taxonomicName id="3DDA4D10FFBAFFD3B9ECAE6EFD5D173C" authorityName="Wagner" authorityYear="1840" box="[677,755,1039,1065]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="genus">
<emphasis id="C8AEEA81FFBAFFD3B9ECAE6EFD5D173C" box="[677,755,1039,1065]" italics="true" pageId="27" pageNumber="448">Eliomys</emphasis>
</taxonomicName>
from Greek and Spanish localities of Late Miocene/Early Pliocene age are referred to
<taxonomicName id="3DDA4D10FFBAFFD3BF24AE4EFCEC177C" authorityName="Friant" authorityYear="1953" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3BF24AE4EFB15175C" box="[1133,1211,1071,1097]" italics="true" pageId="27" pageNumber="448">Eliomys</emphasis>
aff.
<emphasis id="C8AEEA81FFBAFFD3B982AE2EFCEC177C" box="[715,834,1103,1129]" italics="true" pageId="27" pageNumber="448">intermedius</emphasis>
</taxonomicName>
(
<bibRefCitation id="9E4B4B62FFBAFFD3B806AE2EFB89177C" author="DE BRUIJN H. &amp; DAWSON M. R. &amp; MEIN P." box="[847,1063,1103,1129]" pageId="27" pageNumber="448" pagination="535 - 584" refId="ref30906" refString="DE BRUIJN H., DAWSON M. R. &amp; MEIN P. 1970. - Upper Pliocene Rodentia, Lagomorpha and Insectivora (Mammalia) from the Isle of Rhodes (Greece). I-III. Proceedings Koninklijke Nederlandse Akademie van Wetenschappen B 73: 535 - 584." type="journal article" year="1970">
de Bruijn
<emphasis id="C8AEEA81FFBAFFD3B8FEAE31FC46177C" box="[951,1000,1103,1129]" italics="true" pageId="27" pageNumber="448">et al.</emphasis>
1970
</bibRefCitation>
;
<bibRefCitation id="9E4B4B62FFBAFFD3BF79AE2EFD48179C" author="ADROVER R. &amp; MEIN P. &amp; MOISSENET E." pageId="27" pageNumber="448" pagination="47 - 84" refId="ref28616" refString="ADROVER R., MEIN P. &amp; MOISSENET E. 1993 a. - Roedores de la transicion Mio-Plioceno de la region de Teruel. Paleontologia i Evolucio 26 - 27: 47 - 84." type="journal article" year="1993">
Adrover
<emphasis id="C8AEEA81FFBAFFD3BFC0AE31FB14177C" box="[1161,1210,1103,1129]" italics="true" pageId="27" pageNumber="448">et al.</emphasis>
1993a
</bibRefCitation>
; García-Alix
<emphasis id="C8AEEA81FFBAFFD3B82CAE0EFC3D179D" box="[869,915,1134,1160]" italics="true" pageId="27" pageNumber="448">et al.</emphasis>
2008a),because they share an intermediate size between
<taxonomicName id="3DDA4D10FFBAFFD3B8E4AEEEFC5917BD" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[941,1015,1167,1192]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="truci">
<emphasis id="C8AEEA81FFBAFFD3B8E4AEEEFC5917BD" box="[941,1015,1167,1192]" italics="true" pageId="27" pageNumber="448">E.truci</emphasis>
</taxonomicName>
and
<taxonomicName id="3DDA4D10FFBAFFD3BF61AEEEFB1917BD" authorityName="Friant" authorityYear="1953" box="[1064,1207,1166,1192]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3BF61AEEEFB1917BD" box="[1064,1207,1166,1192]" italics="true" pageId="27" pageNumber="448">E.intermedius</emphasis>
</taxonomicName>
and display morphological traits that fit better with
<taxonomicName id="3DDA4D10FFBAFFD3B9EAAEAFFC9717F2" authorityName="Friant" authorityYear="1953" box="[675,825,1229,1255]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3B9EAAEAFFC9717F2" box="[675,825,1229,1255]" italics="true" pageId="27" pageNumber="448">E. intermedius</emphasis>
</taxonomicName>
, such as the presence of a posterior centroloph, more developed metalophid-metaconid and anterolophid-protoconid connections, and reduction of the posterior extra ridge. The assemblage of MCC exhibits some peculiar morphological features, such as the common occurrence of fused centrolophs developing into a Y-shaped ridge and a strong reduction of the posterior extra ridge in the m2. These assemblages are referred to
<taxonomicName id="3DDA4D10FFBAFFD3B8E0AFADFBCA16F0" authorityName="Friant" authorityYear="1953" box="[937,1124,1483,1510]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3B8E0AFADFC1716F0" box="[937,953,1484,1509]" italics="true" pageId="27" pageNumber="448">E</emphasis>
. aff.
<emphasis id="C8AEEA81FFBAFFD3B8A4AFAAFBCA16F0" box="[1005,1124,1483,1509]" italics="true" pageId="27" pageNumber="448">intermedius</emphasis>
</taxonomicName>
because the material is not rich enough to determine a clear attribution. Moreover even if the studied specimens display similar morphological features to those of
<taxonomicName id="3DDA4D10FFBAFFD3B9EFAC2AFC931571" authorityName="Friant" authorityYear="1953" box="[678,829,1610,1636]" class="Mammalia" family="Gliridae" genus="Eliomys" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="448" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="C8AEEA81FFBAFFD3B9EFAC2AFC931571" box="[678,829,1610,1636]" italics="true" pageId="27" pageNumber="448">E. intermedius</emphasis>
</taxonomicName>
, some reliable differences with the
<typeStatus id="25618831FFBAFFD3B9EFAC0BFD7D1591" box="[678,723,1642,1668]" pageId="27" pageNumber="448">type</typeStatus>
locality assemblage, prevent us to securely refer the material from MCC to this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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