168 lines
16 KiB
XML
168 lines
16 KiB
XML
<document id="C85BE20014ED8341EEEA255CC591DCA1" ID-DOI="10.11646/zootaxa.3647.4.3" ID-GBIF-Dataset="bde365aa-bc27-44b2-9463-80ea385bd6a1" ID-ISSN="1175-5326" ID-Zenodo-Dep="217972" ID-ZooBank="7FD5A595-EE31-4D08-BEA9-BB838054B0C8" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460607203880" checkinUser="plazi" docAuthor="Takahashi, Akio, Ōki, Kimihiko, Ishido, Takahiro & Hirayama, Ren" docDate="2013" docId="726CA8361A1FD039FF4BFA2F33AAFEB0" docLanguage="en" docName="zt03647p540.pdf" docOrigin="Zootaxa 3647 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ocadia Gray 1870" docType="treatment" docVersion="7" lastPageNumber="529" masterDocId="8E55D04E1A1DD03AFFDCFF913030FFAC" masterDocTitle="A new species of the genus Ocadia (Testudines: Geoemydidae) from the middle Miocene of Tanegashima Island, southwestern Japan and its paleogeographic implications" masterLastPageNumber="540" masterPageNumber="527" pageNumber="529" updateTime="1698328003695" updateUser="plazi">
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<mods:title id="E893D86521F94F42466ADF43177D9CE5">A new species of the genus Ocadia (Testudines: Geoemydidae) from the middle Miocene of Tanegashima Island, southwestern Japan and its paleogeographic implications</mods:title>
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<mods:namePart id="52756EFAC30F08AEF75D3A705EB9BB71">Takahashi, Akio</mods:namePart>
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<mods:roleTerm id="6CF401F224F7A26D9A455BD9F646F51B">Author</mods:roleTerm>
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<mods:namePart id="62199213ECA2512A44A3E09C75673A24">Ōki, Kimihiko</mods:namePart>
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</mods:role>
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<mods:namePart id="8759258DA87F362048C3D329988E0CB6">Ishido, Takahiro</mods:namePart>
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</mods:name>
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<mods:name id="18434BF0035A147ED6DEA44EE81D6084" type="personal">
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<mods:roleTerm id="995EBAA91E1DBC2766EBACA5070E29A3">Author</mods:roleTerm>
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<mods:namePart id="2126462391BA2F5FECDF2F8F78A01F7D">Hirayama, Ren</mods:namePart>
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<mods:typeOfResource id="2E599A454CF059310FFC2211D89A885F">text</mods:typeOfResource>
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<mods:title id="0DE59BBD8D9516BDE4B614D25708E72B">Zootaxa</mods:title>
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<mods:part id="8BF7D7376ED59E8211B9571319CCE3DD">
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<mods:date id="9FA3EA9BAAC12FC6238BA65CA9343133">2013</mods:date>
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<mods:detail id="C765296922C70460E98FA4FEC6F1B08D" type="volume">
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<mods:number id="70C93CC3416D98BC2F9629F2B0B0F993">3647</mods:number>
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</mods:detail>
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<mods:detail id="1A00C401DDEE3D91347569E2429B4F35" type="issue">
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<mods:number id="CBAFF6FDDFC3310DF9ABF3CB5CA2FB5D">4</mods:number>
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<mods:start id="2C2EA26FC5A19E269E5D455CF58F2F6A">527</mods:start>
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<mods:end id="5E5CE36C3366041DEF66EE814FE38658">540</mods:end>
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<mods:classification id="0C52E231148AA95A5371015E1CCE03E8">journal article</mods:classification>
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<mods:identifier id="F8E9F5C0849F0FD0BEED7C6E01848B97" type="DOI">10.11646/zootaxa.3647.4.3</mods:identifier>
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<mods:identifier id="90EF2E2015812EB5DE6DB55ECD2FCF09" type="GBIF-Dataset">bde365aa-bc27-44b2-9463-80ea385bd6a1</mods:identifier>
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<mods:identifier id="FC949763EFDB096288CF0B14303F25F4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="726CA8361A1FD039FF4BFA2F33AAFEB0" ID-DOI="http://doi.org/10.5281/zenodo.6152203" ID-GBIF-Taxon="119567878" ID-Zenodo-Dep="6152203" LSID="urn:lsid:plazi:treatment:726CA8361A1FD039FF4BFA2F33AAFEB0" httpUri="http://treatment.plazi.org/id/726CA8361A1FD039FF4BFA2F33AAFEB0" lastPageId="3" lastPageNumber="529" pageId="2" pageNumber="529">
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<subSubSection id="B2DF4AAB1A1FD038FF4BFA2F314DFA74" box="[151,381,1470,1496]" pageId="2" pageNumber="529" type="nomenclature">
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<paragraph id="FA7A19201A1FD038FF4BFA2F314DFA74" blockId="2.[151,381,1470,1496]" box="[151,381,1470,1496]" pageId="2" pageNumber="529">
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<heading id="A132AE4C1A1FD038FF4BFA2F314DFA74" bold="true" box="[151,381,1470,1496]" fontSize="11" level="1" pageId="2" pageNumber="529" reason="1">
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<taxonomicName id="3DC562A31A1FD038FF4BFA2F314DFA74" authority="Gray, 1870" authorityName="Gray" authorityYear="1870" box="[151,381,1470,1496]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="529" phylum="Chordata" rank="genus">
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<emphasis id="C8B1C5321A1FD038FF4BFA2F314DFA74" bold="true" box="[151,381,1470,1496]" pageId="2" pageNumber="529">
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<emphasis id="C8B1C5321A1FD038FF4BFA2F30DDFA74" bold="true" box="[151,237,1470,1496]" italics="true" pageId="2" pageNumber="529">Ocadia</emphasis>
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Gray, 1870
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B2DF4AAB1A1FD038FF4BF99434B5F9ED" pageId="2" pageNumber="529" type="materials_examined">
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<paragraph id="FA7A19201A1FD038FF4BF99432B8F9B1" blockId="2.[151,1436,1541,2033]" box="[151,648,1541,1566]" pageId="2" pageNumber="529">
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<emphasis id="C8B1C5321A1FD038FF4BF9943102F9B2" bold="true" box="[151,306,1541,1566]" pageId="2" pageNumber="529">
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<typeStatus id="257EA7821A1FD038FF4BF99430E2F9B1" box="[151,210,1541,1565]" pageId="2" pageNumber="529">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="3DC562A31A1FD038FEE5F99432B4F9B1" authority="Gray, 1834" authorityName="Gray" authorityYear="1834" box="[313,644,1541,1565]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="529" phylum="Chordata" rank="species" species="sinensis">
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<emphasis id="C8B1C5321A1FD038FEE5F99431DBF9B1" box="[313,491,1541,1565]" italics="true" pageId="2" pageNumber="529">Ocadia sinensis</emphasis>
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(Gray, 1834)
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="FA7A19201A1FD038FF15F9B834B5F9ED" blockId="2.[151,1436,1541,2033]" box="[201,1157,1576,1601]" pageId="2" pageNumber="529">
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<emphasis id="C8B1C5321A1FD038FF15F9B831A3F9ED" bold="true" box="[201,403,1576,1601]" pageId="2" pageNumber="529">Included species.</emphasis>
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<taxonomicName id="3DC562A31A1FD038FE46F9B832D5F9ED" authority="Gray, 1834" authorityName="Gray" authorityYear="1834" box="[410,741,1577,1601]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="529" phylum="Chordata" rank="species" species="sinensis">
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<emphasis id="C8B1C5321A1FD038FE46F9B8327DF9ED" box="[410,589,1577,1601]" italics="true" pageId="2" pageNumber="529">Ocadia sinensis</emphasis>
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(Gray, 1834)
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</taxonomicName>
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;
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<taxonomicName id="3DC562A31A1FD038FD2DF9BB34B1F9ED" authority="Hirayama et al., 2007" authorityName="Hirayama et al." authorityYear="2007" box="[753,1153,1577,1601]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="529" phylum="Chordata" rank="species" species="nipponica">
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<emphasis id="C8B1C5321A1FD038FD2DF9BB33B3F9ED" box="[753,899,1578,1601]" italics="true" pageId="2" pageNumber="529">O. nipponica</emphasis>
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Hirayama
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<emphasis id="C8B1C5321A1FD038FC23F9BB341FF9ED" box="[1023,1071,1577,1601]" italics="true" pageId="2" pageNumber="529">et al</emphasis>
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., 2007
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2DF4AAB1A1FD038FF15F9DC3473F8CD" pageId="2" pageNumber="529" type="description">
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<paragraph id="FA7A19201A1FD038FF15F9DC3473F8CD" blockId="2.[151,1436,1541,2033]" pageId="2" pageNumber="529">
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<emphasis id="C8B1C5321A1FD038FF15F9DC3199F9CA" bold="true" box="[201,425,1613,1638]" pageId="2" pageNumber="529">
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<taxonomicName id="3DC562A31A1FD038FF15F9DC3116F9CA" box="[201,294,1613,1638]" pageId="2" pageNumber="538" rank="species" species="revised">Revised</taxonomicName>
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diagnosis.
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</emphasis>
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Modified on the basis of Smith (1931), Bourret (1941), Ernst and Barbour (1989), Hirayama
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<emphasis id="C8B1C5321A1FD038FED1F9E33170F925" box="[269,320,1649,1673]" italics="true" pageId="2" pageNumber="529">et al</emphasis>
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. (2007), and this study: medium to large–sized hingeless geoemydids with following combination of character states: hexagonal neural plates short–sided in front; posterior margin of the carapace unserrated; second and third vertebral scutes frequently showing nearly rectangular in shape, as long as wide; the entoplastron intersected by the humero–pectoral sulcus; the plastral buttresses moderately developed, extending to half way of the costal plates; median length of the gular shorter than the interhumeral sulcus and the gular often isolated from the entoplastron; enlarged upper and lower triturating surface decorated by the lingual ridges; the upper triturating surface with a denticulate median ridge; and the foramen palatinum posterius small.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2DF4AAB1A1FD039FF15F8FC33AAFEB0" lastPageId="3" lastPageNumber="530" pageId="2" pageNumber="529" type="discussion">
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<paragraph id="FA7A19201A1FD039FF15F8FC33AAFEB0" blockId="2.[151,1436,1541,2033]" lastBlockId="3.[151,1436,151,284]" lastPageId="3" lastPageNumber="530" pageId="2" pageNumber="529">
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<emphasis id="C8B1C5321A1FD038FF15F8FC310DF829" bold="true" box="[201,317,1901,1925]" pageId="2" pageNumber="529">Remarks.</emphasis>
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Recent molecular phylogenetic analyses have shown that the genus
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<taxonomicName id="3DC562A31A1FD038FB91F8FC3490F829" box="[1101,1184,1901,1925]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="529" phylum="Chordata" rank="genus">
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<emphasis id="C8B1C5321A1FD038FB91F8FC3490F829" box="[1101,1184,1901,1925]" italics="true" pageId="2" pageNumber="529">Ocadia</emphasis>
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</taxonomicName>
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is included in a clade with
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<taxonomicName id="3DC562A31A1FD038FF09F8003173F805" box="[213,323,1937,1961]" class="Testudines" family="Geoemydidae" genus="Chinemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="2" pageNumber="529" phylum="Chordata" rank="genus">
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<emphasis id="C8B1C5321A1FD038FF09F8003173F805" box="[213,323,1937,1961]" italics="true" pageId="2" pageNumber="529">Chinemys</emphasis>
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</taxonomicName>
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spp. and
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<taxonomicName id="3DC562A31A1FD038FE62F8033292F805" box="[446,674,1938,1961]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="2" pageNumber="529" phylum="Chordata" rank="species" species="japonica">
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<emphasis id="C8B1C5321A1FD038FE62F8033292F805" box="[446,674,1938,1961]" italics="true" pageId="2" pageNumber="529">Mauremys japonica</emphasis>
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</taxonomicName>
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and that these taxa form another more inclusive clade with the remaining living species of
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<taxonomicName id="3DC562A31A1FD038FE08F8273279F861" box="[468,585,1974,1997]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="2" pageNumber="529" phylum="Chordata" rank="genus">
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<emphasis id="C8B1C5321A1FD038FE08F8273279F861" box="[468,585,1974,1997]" italics="true" pageId="2" pageNumber="529">Mauremys</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="3DC562A31A1FD038FD87F82732E0F861" box="[603,720,1974,1997]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="2" pageNumber="529" phylum="Chordata" rank="genus" sensu="lato">
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<emphasis id="C8B1C5321A1FD038FD87F82732E0F861" box="[603,720,1974,1997]" italics="true" pageId="2" pageNumber="529">Mauremys</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D38278491A1FD038FD05F827337DF861" box="[729,845,1973,1997]" pageId="2" pageNumber="529" sensu="lato">sensu lato</taxonomicNameLabel>
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, Honda
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<emphasis id="C8B1C5321A1FD038FC71F82733D7F861" box="[941,999,1973,1997]" italics="true" pageId="2" pageNumber="529">et al.</emphasis>
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2002; Barth
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<emphasis id="C8B1C5321A1FD038FBA2F8273488F861" box="[1150,1208,1973,1997]" italics="true" pageId="2" pageNumber="529">et al.</emphasis>
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2004; Feldman and Parham 2004; Spinks
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<emphasis id="C8B1C5321A1FD038FE50F84B31F3F85D" box="[396,451,2009,2033]" italics="true" pageId="2" pageNumber="529">et al.</emphasis>
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2004; Sasaki
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<emphasis id="C8B1C5321A1FD038FD83F84B32A7F85D" box="[607,663,2009,2033]" italics="true" pageId="2" pageNumber="529">et al.</emphasis>
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2006; Jiang
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<emphasis id="C8B1C5321A1FD038FCF8F84B336CF85D" box="[804,860,2009,2033]" italics="true" pageId="2" pageNumber="529">et al.</emphasis>
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2011). However, no synapomorphic morphological character states are known for the two clades (e.g., Honda
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<emphasis id="C8B1C5321A1ED039FCE0FF083346FF1C" box="[828,886,152,176]" italics="true" pageId="3" pageNumber="530">et al.</emphasis>
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2002; Hirayama
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<emphasis id="C8B1C5321A1ED039FB9CFF08344AFF1C" box="[1088,1146,152,176]" italics="true" pageId="3" pageNumber="530">et al.</emphasis>
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2007). Actually, because
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<taxonomicName id="3DC562A31A1ED039FF4BFF2D30DAFF78" box="[151,234,188,212]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="530" phylum="Chordata" rank="genus">
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<emphasis id="C8B1C5321A1ED039FF4BFF2D30DAFF78" box="[151,234,188,212]" italics="true" pageId="3" pageNumber="530">Ocadia</emphasis>
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</taxonomicName>
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differs much from
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<taxonomicName id="3DC562A31A1ED039FE10FF2D320AFF78" box="[460,570,188,212]" class="Testudines" family="Geoemydidae" genus="Chinemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="530" phylum="Chordata" rank="genus">
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<emphasis id="C8B1C5321A1ED039FE10FF2D320AFF78" box="[460,570,188,212]" italics="true" pageId="3" pageNumber="530">Chinemys</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="3DC562A31A1ED039FDAAFF2C32DBFF78" box="[630,747,189,212]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="3" pageNumber="530" phylum="Chordata" rank="genus">
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<emphasis id="C8B1C5321A1ED039FDAAFF2C32DBFF78" box="[630,747,189,212]" italics="true" pageId="3" pageNumber="530">Mauremys</emphasis>
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</taxonomicName>
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in several skull and shell characters (e.g., McDowell 1964; Hirayama 1985; Gaffney and Meylan 1988; Yasukawa
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<emphasis id="C8B1C5321A1ED039FCD7FF703375FF54" box="[779,837,224,248]" italics="true" pageId="3" pageNumber="530">et al.</emphasis>
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2001), we retain the genus
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<taxonomicName id="3DC562A31A1ED039FB5FFF7134E5FF54" box="[1155,1237,224,248]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="530" phylum="Chordata" rank="genus">
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<emphasis id="C8B1C5321A1ED039FB5FFF7134E5FF54" box="[1155,1237,224,248]" italics="true" pageId="3" pageNumber="530">Ocadia</emphasis>
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</taxonomicName>
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for disclosing its past species diversity from paleontological view in the present study.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |