604 lines
75 KiB
XML
604 lines
75 KiB
XML
<document id="8853DC8950F5CFCD91EA5F5AE18C1788" ID-DOI="10.11646/zootaxa.3647.4.3" ID-GBIF-Dataset="bde365aa-bc27-44b2-9463-80ea385bd6a1" ID-ISSN="1175-5326" ID-Zenodo-Dep="217972" ID-ZooBank="7FD5A595-EE31-4D08-BEA9-BB838054B0C8" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460607203880" checkinUser="plazi" docAuthor="Takahashi, Akio, Ōki, Kimihiko, Ishido, Takahiro & Hirayama, Ren" docDate="2013" docId="726CA8361A1ED03DFF4BFEFE328BF9F8" docLanguage="en" docName="zt03647p540.pdf" docOrigin="Zootaxa 3647 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ocadia tanegashimensis Takahashi, Ōki, Ishido & Hirayama, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="534" masterDocId="8E55D04E1A1DD03AFFDCFF913030FFAC" masterDocTitle="A new species of the genus Ocadia (Testudines: Geoemydidae) from the middle Miocene of Tanegashima Island, southwestern Japan and its paleogeographic implications" masterLastPageNumber="540" masterPageNumber="527" pageNumber="530" updateTime="1698328003695" updateUser="plazi">
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<mods:title id="EF94B9C937FF0177B667FD264DBA4997">A new species of the genus Ocadia (Testudines: Geoemydidae) from the middle Miocene of Tanegashima Island, southwestern Japan and its paleogeographic implications</mods:title>
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<mods:namePart id="808298FE7CAB74FDD221004581AEB848">Takahashi, Akio</mods:namePart>
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<mods:namePart id="05B00B3AFD16E1DAD6674607EB21BBB1">Ōki, Kimihiko</mods:namePart>
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<mods:namePart id="5E3174ABEE8B61E8DE1297505EEE8F02">Ishido, Takahiro</mods:namePart>
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<mods:namePart id="AC55B743F7F6C22665EB2577D0661BB8">Hirayama, Ren</mods:namePart>
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<mods:title id="B382048B344EC5DA6E276730D05D0CDC">Zootaxa</mods:title>
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<mods:date id="0366108644F59DAFCBE80DE7F26EDCFB">2013</mods:date>
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<mods:number id="2B100070B1FDA2B3E0E156875934AB45">3647</mods:number>
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<mods:classification id="6AEFD10F46E3487B250EF4CAC7A186DB">journal article</mods:classification>
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<mods:identifier id="8F154E9CE00CB27BEF754D59E27B16D2" type="DOI">10.11646/zootaxa.3647.4.3</mods:identifier>
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<treatment id="726CA8361A1ED03DFF4BFEFE328BF9F8" ID-DOI="http://doi.org/10.5281/zenodo.6152205" ID-GBIF-Taxon="119567885" ID-Zenodo-Dep="6152205" LSID="urn:lsid:plazi:treatment:726CA8361A1ED03DFF4BFEFE328BF9F8" httpUri="http://treatment.plazi.org/id/726CA8361A1ED03DFF4BFEFE328BF9F8" lastPageId="7" lastPageNumber="534" pageId="3" pageNumber="530">
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<subSubSection id="B2DF4AAB1A1ED039FF4BFEFE3143FE62" pageId="3" pageNumber="530" type="nomenclature">
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<paragraph id="FA7A19201A1ED039FF4BFEFE3211FE25" blockId="3.[151,741,367,462]" box="[151,545,367,393]" pageId="3" pageNumber="530">
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<heading id="A132AE4C1A1ED039FF4BFEFE3211FE25" bold="true" box="[151,545,367,393]" fontSize="11" level="1" pageId="3" pageNumber="530" reason="1">
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<emphasis id="C8B1C5321A1ED039FF4BFEFE3211FE25" bold="true" box="[151,545,367,393]" pageId="3" pageNumber="530">
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<taxonomicName id="3DC562A31A1ED039FF4BFEFE318DFE25" box="[151,445,367,393]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="530" phylum="Chordata" rank="species" species="tanegashimensis" status="sp. nov.">
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<emphasis id="C8B1C5321A1ED039FF4BFEFE318DFE25" bold="true" box="[151,445,367,393]" italics="true" pageId="3" pageNumber="530">Ocadia tanegashimensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D38278491A1ED039FE18FEFE3211FE25" box="[452,545,367,393]" pageId="3" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="FA7A19201A1ED039FF4BFE033143FE62" blockId="3.[151,741,367,462]" pageId="3" pageNumber="530">
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(Suggested Japanese name: Tanegashima hanagame) (
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<figureCitation id="62FE05A51A1ED039FF43FE27313CFE62" box="[159,268,438,462]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,250,1414,1437]" captionStartId-1="6.[151,250,1635,1658]" captionStartId-2="7.[151,250,1049,1072]" captionTargetBox-0="[281,1305,625,1392]" captionTargetBox-1="[193,1420,222,1608]" captionTargetBox-2="[151,1436,200,1011]" captionTargetId-0="figure@4.[281,1305,625,1393]" captionTargetId-1="figure@6.[158,1428,195,1614]" captionTargetId-2="figure@7.[151,1436,193,1028]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 2. Picture showing occurrence of Ocadia tanegashimensis (MTE 1) from the Kawachi Formation in Kawachi, Minamitane – cho, southern part of Tanegashima Island." captionText-1="FIGURE 3. Ocadia tanegashimensis (MTE 1) from Tanegashima Island. A – C, the carapace in dorsal (A), ventral (B) and anterior (C) views. D, left fifth to seventh costals in ventral view. E – F, the plastron in dorsal (E) and ventral (F) views. Arrows indicate locations of upper ends of the plastral buttresses. Scale bar: 5 cm." captionText-2="FIGURE 4. Illustration of Ocadia tanegashimensis (MTE 1) from Tanegashima Island. A, the carapace in dorsal and B, the plastron in ventral views. Abbreviations: abd, abdominal; an, anal; ce, cervical; co, costal; ent, entoplastron; epi, epiplastron; fe, femoral; gu, gular; hu, humeral; hyo, hyoplastron; hyp, hypoplastron; LE, length of the epiplastron; LHS, length of the hyoplastral suture; m, marginal; ne, neural; nu, nuchal; pec, pectoral; pl, pleural; v, vertebral; xip, xiphiplastron. Symbol +: locations of upper ends of the plastral buttresses in ventral view. Scale bar: 5 cm." httpUri-0="https://zenodo.org/record/217974/files/figure.png" httpUri-1="https://zenodo.org/record/217975/files/figure.png" httpUri-2="https://zenodo.org/record/217976/files/figure.png" pageId="3" pageNumber="530">Figs. 2–4</figureCitation>
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;
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<tableCitation id="B7472C9B1A1ED039FECBFE273158FE62" box="[279,360,438,462]" captionStart="TABLE 1" captionStartId="5.[151,239,511,535]" captionText="TABLE 1. Comparisons of the Tanegashima geoemydid (MTE 1) with hingeless geoemydids (see Appendix) having the entoplastron intersected by the humero – pectoral sulcus and the neurals short – sided anteriorly. Symbols, +, –, and?, indicate presence, absence, and undetermined status of a given character state. Character state (1): second and third vertebrals rectangular and as long as wide, character state (2): posterior margin of the carapace unserrated, character state (3): medial length of the gular shorter than interhumeral sulcus, character state (4): plastral buttresses developed, extending halfway of the costals, character state (5): medial and two lateral obtuse discoutinuous keels on the carapace present, character state (6): first vertebral scute expanded anterolaterally, reaching the second marginal, character state (7): second and third vertebral scutes much narrower than the first anteroposteriorly, character state (8): length of entoplastron nearly as long as interhyoplastral suture, character state (9): costals dovetailed with one another in outline, and character state (10): third pleural overlapping only sixth and seventh peripherals." pageId="3" pageNumber="530">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B2DF4AAB1A1ED039FF4BFE6E3491FD2E" pageId="3" pageNumber="530" type="materials_examined">
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<paragraph id="FA7A19201A1ED039FF4BFE6E34DFFDBA" blockId="3.[151,1436,510,2010]" box="[151,1263,510,535]" pageId="3" pageNumber="530">
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<emphasis id="C8B1C5321A1ED039FF4BFE6E313BFDBB" bold="true" box="[151,267,511,535]" pageId="3" pageNumber="530">
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<typeStatus id="257EA7821A1ED039FF4BFE6E3136FDBB" box="[151,262,511,535]" pageId="3" pageNumber="530" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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A shell, deposited in the collection of the Minamitane–cho Board of Education (MTE1).
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</paragraph>
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<paragraph id="FA7A19201A1ED039FF15FDB2342FFDF2" blockId="3.[151,1436,510,2010]" pageId="3" pageNumber="530">
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<emphasis id="C8B1C5321A1ED039FF15FDB2315DFD97" bold="true" box="[201,365,546,571]" pageId="3" pageNumber="530">
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<typeStatus id="257EA7821A1ED039FF15FDB23132FD97" box="[201,258,547,571]" pageId="3" pageNumber="530">Type</typeStatus>
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locality.
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</emphasis>
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South slope located approximately
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<quantity id="3D3DB4C51A1ED039FCF5FDB3334FFD96" box="[809,895,546,570]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="530" unit="km" value="1.5">1.5 km</quantity>
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southwest from the Kaminaka crossroad on Prefectural Road 588, Kawachi, Minamitane–cho, Kagoshima Prefecture,
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<collectingCountry id="82D259B01A1ED039FC0BFDD6342BFDF2" box="[983,1051,583,606]" name="Japan" pageId="3" pageNumber="530">Japan</collectingCountry>
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.
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</paragraph>
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<paragraph id="FA7A19201A1ED039FF15FDFB3491FD2E" blockId="3.[151,1436,510,2010]" box="[201,1185,618,643]" pageId="3" pageNumber="530">
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<emphasis id="C8B1C5321A1ED039FF15FDFB3102FD2F" bold="true" box="[201,306,618,643]" pageId="3" pageNumber="530">Horizon.</emphasis>
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Middle part of Kawachi Formation (early middle Miocene), Kukinaga Group.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2DF4AAB1A1ED039FF15FD1E317AFD66" pageId="3" pageNumber="530" type="etymology">
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<paragraph id="FA7A19201A1ED039FF15FD1E317AFD66" blockId="3.[151,1436,510,2010]" pageId="3" pageNumber="530">
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<emphasis id="C8B1C5321A1ED039FF15FD1E317FFD0B" bold="true" box="[201,335,655,679]" pageId="3" pageNumber="530">Etymology.</emphasis>
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The specific epithet refers to the name of the island (Tanegashima Island) from which the fossil was discovered.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2DF4AAB1A1ED039FF15FD473575FCD2" pageId="3" pageNumber="530" type="diagnosis">
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<paragraph id="FA7A19201A1ED039FF15FD473575FCD2" blockId="3.[151,1436,510,2010]" pageId="3" pageNumber="530">
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<emphasis id="C8B1C5321A1ED039FF15FD473173FD43" bold="true" box="[201,323,726,751]" pageId="3" pageNumber="530">Diagnosis.</emphasis>
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A medium–sized geoemydid species larger than
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<taxonomicName id="3DC562A31A1ED039FCB7FD4633D7FD42" box="[875,999,727,750]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="3" pageNumber="530" phylum="Chordata" rank="species" species="sinensis">
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<emphasis id="C8B1C5321A1ED039FCB7FD46334FFD42" box="[875,895,727,750]" italics="true" pageId="3" pageNumber="530">O</emphasis>
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.
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<emphasis id="C8B1C5321A1ED039FC52FD4633D7FD42" box="[910,999,727,750]" italics="true" pageId="3" pageNumber="530">sinensis</emphasis>
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</taxonomicName>
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, characterized by a combination of the following character states: the first vertebral expanded anterolaterally, reaching the second marginal; the second and third vertebral scutes much narrower than the first vertebral anteroposteriorly; a medial and two lateral obtuse discontinuous keels on carapace; length of the entoplastron as long as the interhyoplastral suture; the costals dovetailed with one another in outline; the third pleural overlapping only the sixth and seventh peripherals.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2DF4AAB1A1ED03DFF15FC1B328BF9F8" lastPageId="7" lastPageNumber="534" pageId="3" pageNumber="530" type="description">
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<paragraph id="FA7A19201A1ED039FF15FC1B3113FB6E" blockId="3.[151,1436,510,2010]" pageId="3" pageNumber="530">
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<emphasis id="C8B1C5321A1ED039FF15FC1B3197FC0F" bold="true" box="[201,423,906,931]" pageId="3" pageNumber="530">Description. Shell.</emphasis>
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The shell is incompletely preserved and strongly depressed dorsoventrally. It lacks the eighth neural; the first and second suprapygals; the right fifth, eighth and eleventh peripherals; the left sixth and eleventh peripherals; and the pygal. The shell margin is not emarginated anterolaterally and posterolaterally. The shell surface is smooth, lacks growth annuli, and has an obtuse medial and two lateral discontinuous knob–like longitudinal keels. The carapace and plastron are moderately thick (carapace:
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<quantity id="3D3DB4C51A1ED039FBD2FB8B3443FB9E" box="[1038,1139,1050,1074]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="530" unit="mm" value="10.3">10.3 mm</quantity>
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thick in the middle of the thickened part of the nuchal,
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<quantity id="3D3DB4C51A1ED039FE3CFBAF3207FBFA" box="[480,567,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="3" pageNumber="530" unit="mm" value="9.4">9.4 mm</quantity>
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in the posterior part of the fifth neural, and
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<quantity id="3D3DB4C51A1ED039FBF4FBAF344FFBFA" box="[1064,1151,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="530" unit="mm" value="8.5">8.5 mm</quantity>
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in the second peripheral; plastron:
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<quantity id="3D3DB4C51A1ED039FEDDFBF33155FBD6" box="[257,357,1122,1146]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="530" unit="mm" value="11.5">11.5 mm</quantity>
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thick in the left lateral part of the gular lip, 8.0 mm in the posterior corner of the entoplastron,
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<quantity id="3D3DB4C51A1ED039FF4BFB1730C0FB32" box="[151,240,1158,1182]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="530" unit="mm" value="4.9">4.9 mm</quantity>
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in the posterolateral corner of the right hyoplastron, and
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<quantity id="3D3DB4C51A1ED039FC4DFB1733C8FB32" box="[913,1016,1158,1182]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="3" pageNumber="530" unit="mm" value="12.8">12.8 mm</quantity>
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in the posteriormost part of the left femoral lip).
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</paragraph>
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<paragraph id="FA7A19201A1ED039FF15FB5F33E7FADA" blockId="3.[151,1436,510,2010]" pageId="3" pageNumber="530">The nuchal is hexagonal and slightly short–sided posteriorly. The first neural is barrel–shaped and the second to seventh neurals are hexagonal and short–sided anteriorly. Widths of the costals are variable; the second, fourth, sixth, and eighth costals are distally expanded, whereas the third and fifth costals are distally contracted. The first costal contacts the first to third peripherals and the second costal connects the third and fourth peripherals laterally. The remaining costals articulate with the adjacent two peripherals distally.</paragraph>
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<paragraph id="FA7A19201A1ED039FF15FA133210F972" blockId="3.[151,1436,510,2010]" pageId="3" pageNumber="530">
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The cervical scute is rectangular and longer than wide in dorsal view and trapezoidal and much wider than long in ventral view. The first vertebral is slightly longer than wide (
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<quantity id="3D3DB4C51A1ED039FC53FA3733C4FA12" box="[911,1012,1446,1470]" metricMagnitude="-2" metricUnit="m" metricValue="8.37" pageId="3" pageNumber="530" unit="mm" value="83.7">83.7 mm</quantity>
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long and
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<quantity id="3D3DB4C51A1ED039FBBDFA3734F5FA12" box="[1121,1221,1446,1470]" metricMagnitude="-2" metricUnit="m" metricValue="8.12" pageId="3" pageNumber="530" unit="mm" value="81.2">81.2 mm</quantity>
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wide) and laterally notched, reaching the posteromedial corner of the second marginal. The second and third vertebrals are nearly rectangular in shape and narrower than the first vertebral (the second,
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<quantity id="3D3DB4C51A1ED039FC73FA7F3424F9AA" box="[943,1044,1518,1542]" metricMagnitude="-2" metricUnit="m" metricValue="6.74" pageId="3" pageNumber="530" unit="mm" value="67.4">67.4 mm</quantity>
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long and
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<quantity id="3D3DB4C51A1ED039FB58FA7F34D9F9AA" box="[1156,1257,1518,1542]" metricMagnitude="-2" metricUnit="m" metricValue="6.76" pageId="3" pageNumber="530" unit="mm" value="67.6">67.6 mm</quantity>
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wide; the third,
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<quantity id="3D3DB4C51A1ED039FF4BF98330CDF986" box="[151,253,1554,1578]" metricMagnitude="-2" metricUnit="m" metricValue="7.029999999999999" pageId="3" pageNumber="530" unit="mm" value="70.3">70.3 mm</quantity>
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long and
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<quantity id="3D3DB4C51A1ED039FEAEF98331E9F986" box="[370,473,1554,1578]" metricMagnitude="-2" metricUnit="m" metricValue="7.32" pageId="3" pageNumber="530" unit="mm" value="73.2">73.2 mm</quantity>
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wide). The fourth is obviously wider than the second and third vertebrals. The first pleural contacts the posterolateral corner of the first marginal anteromedially and the second to fifth marginals anterolaterally. The second pleural appears to contact the fifth and sixth marginals laterally, and the third pleural contacts the sixth to eighth marginals. The fourth pleural contacts the eighth to tenth marginals. Of the four pleural scutes, the first covers four and the second and fourth cover three peripherals, whereas the third covers only two peripherals (the sixth and seventh).
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</paragraph>
|
||
<paragraph id="FA7A19201A1ED039FF15F97A3511F8C2" blockId="3.[151,1436,510,2010]" pageId="3" pageNumber="530">
|
||
<emphasis id="C8B1C5321A1ED039FF15F97A3105F8AF" bold="true" box="[201,309,1771,1795]" pageId="3" pageNumber="530">Plastron.</emphasis>
|
||
The plastron preserves all components: the epiplastra, the entoplastron, the hyoplastra, the hypoplastra and the xiphiplastra. The maximum plastron length is estimated to be approximately
|
||
<quantity id="3D3DB4C51A1ED039FADCF89F3553F88A" box="[1280,1379,1806,1830]" metricMagnitude="-1" metricUnit="m" metricValue="3.25" pageId="3" pageNumber="530" unit="mm" value="325.0">325 mm</quantity>
|
||
. The presence of a shallow but distinct ventral concavity on the plastron in ventral view and the absence of the laterally rounded posterior lobe suggest that this individual is male (Ernst and Barbour 1989; Zhang
|
||
<emphasis id="C8B1C5321A1ED039FB45F8C934FAF8C3" box="[1177,1226,1879,1903]" italics="true" pageId="3" pageNumber="530">et al</emphasis>
|
||
. 1998).
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A1ED03EFF15F8EB3132FD90" blockId="3.[151,1436,510,2010]" lastBlockId="4.[151,1436,151,572]" lastPageId="4" lastPageNumber="531" pageId="3" pageNumber="530">
|
||
The epiplastra are mostly preserved, lacking only the lateral margin of the left epiplastron. Of these, the right is
|
||
<quantity id="3D3DB4C51A1ED039FF4BF80F30CBF81A" box="[151,251,1950,1974]" metricMagnitude="-2" metricUnit="m" metricValue="5.67" pageId="3" pageNumber="530" unit="mm" value="56.7">56.7 mm</quantity>
|
||
long and
|
||
<quantity id="3D3DB4C51A1ED039FEB0F80F31E1F81A" box="[364,465,1950,1974]" metricMagnitude="-2" metricUnit="m" metricValue="5.72" pageId="3" pageNumber="530" unit="mm" value="57.2">57.2 mm</quantity>
|
||
wide. They have a relatively wide epiplastral lip represented by dorsal overlapping of the gular scute (
|
||
<quantity id="3D3DB4C51A1ED039FE97F8533181F876" box="[331,433,1986,2010]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="3" pageNumber="530" unit="mm" value="15.7">15.7 mm</quantity>
|
||
in the right) and lack the epiplastral excavation. The interepiplastral suture is short (
|
||
<quantity id="3D3DB4C51A1ED03EFAB5F85330F4FF03" lastPageId="4" lastPageNumber="531" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="3" pageNumber="530" unit="mm" value="24.3">24.3 mm</quantity>
|
||
) and approximately two–fifths of the median length of the entoplastron (
|
||
<quantity id="3D3DB4C51A19D03EFBDBFF063440FF03" box="[1031,1136,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="5.74" pageId="4" pageNumber="531" unit="mm" value="57.4">57.4 mm</quantity>
|
||
) in ventral view. It is also short (
|
||
<quantity id="3D3DB4C51A19D03EFF38FF2D3161FF78" box="[228,337,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="4" pageNumber="531" unit="mm" value="25.3">25.3 mm</quantity>
|
||
) dorsally and approximately half of the median length of the entoplastron (
|
||
<quantity id="3D3DB4C51A19D03EFB3BFF2D3567FF78" box="[1255,1367,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="4.37" pageId="4" pageNumber="531" unit="mm" value="43.7">43.7 mm</quantity>
|
||
). The hyoplastra lack most of the axillary buttresses, the anterolateral fringes corresponding to a portion of the bridge to the carapace, and the posterolateral part of the left forelobe. The left hyoplastron (
|
||
<quantity id="3D3DB4C51A19D03EFB9CFE953485FEB0" box="[1088,1205,260,284]" metricMagnitude="-1" metricUnit="m" metricValue="1.0959999999999999" pageId="4" pageNumber="531" unit="mm" value="109.6">109.6 mm</quantity>
|
||
long and
|
||
<quantity id="3D3DB4C51A19D03EFAF5FE9535ACFEB0" box="[1321,1436,260,284]" metricMagnitude="-1" metricUnit="m" metricValue="1.087" pageId="4" pageNumber="531" unit="mm" value="108.7">108.7 mm</quantity>
|
||
wide) preserves the upper part of the axillary buttress, suturing with the middle parts of the left first costal in ventral view. The interhyoplastral suture is
|
||
<quantity id="3D3DB4C51A19D03EFD5EFEDD32DAFEC8" box="[642,746,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="6.26" pageId="4" pageNumber="531" unit="mm" value="62.6">62.6 mm</quantity>
|
||
, slightly longer than the median length of the entoplastron in ventral view. The hypoplastra are also preserved but their anterolateral and anteromedial corners and the ascending processes of the inguinal buttresses are missing. However, the left fifth costal preserves the sutural facet in ventral view, indicating that the inguinal buttresses extend to the middle part of the fifth costals. The xiphiplastra are nearly completely preserved, missing just small portions in their anteromedial parts and the posteromedial part of the left xiphiplastron. The left xiphiplastron is
|
||
<quantity id="3D3DB4C51A19D03EFD93FE6E3284FDBB" box="[591,692,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="7.79" pageId="4" pageNumber="531" unit="mm" value="77.9">77.9 mm</quantity>
|
||
long and
|
||
<quantity id="3D3DB4C51A19D03EFCF9FE6E33A8FDBB" box="[805,920,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="6.8629999999999995" pageId="4" pageNumber="531" unit="mm" value="68.63">68.63 mm</quantity>
|
||
wide. Length of the interhypoplstral suture is
|
||
<quantity id="3D3DB4C51A19D03EFF4BFDB530CEFD90" box="[151,254,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="5.81" pageId="4" pageNumber="531" unit="mm" value="58.1">58.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="AEBA49A81A19D03EFF4BFA1732F6FA17" httpUri="https://zenodo.org/record/217974/files/figure.png" pageId="4" pageNumber="531" targetBox="[281,1305,625,1392]" targetPageId="4">
|
||
<paragraph id="FA7A19201A19D03EFF4BFA1732F6FA17" blockId="4.[151,1436,1414,1467]" pageId="4" pageNumber="531">
|
||
<emphasis id="C8B1C5321A19D03EFF4BFA173128FA30" bold="true" box="[151,280,1414,1437]" pageId="4" pageNumber="531">FIGURE 2.</emphasis>
|
||
Picture showing occurrence of
|
||
<taxonomicName id="3DC562A31A19D03EFDAFFA163340FA31" box="[627,880,1415,1437]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="531" phylum="Chordata" rank="species" species="tanegashimensis">
|
||
<emphasis id="C8B1C5321A19D03EFDAFFA163340FA31" box="[627,880,1415,1437]" italics="true" pageId="4" pageNumber="531">Ocadia tanegashimensis</emphasis>
|
||
</taxonomicName>
|
||
(MTE1) from the Kawachi Formation in Kawachi, Minamitane–cho, southern part of Tanegashima Island.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA7A19201A19D03EFF1BFA733368F926" blockId="4.[151,1436,1506,2035]" pageId="4" pageNumber="531">
|
||
The gular scutes reach the anterior tip of the entoplastron. Length of the intergular sulcus (
|
||
<quantity id="3D3DB4C51A19D03EFB65FA733510FA56" box="[1209,1312,1506,1530]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="4" pageNumber="531" unit="mm" value="35.6">35.6 mm</quantity>
|
||
) is slightly shorter than that of the interhumeral sulcus (
|
||
<quantity id="3D3DB4C51A19D03EFD55F99732C4F9B2" box="[649,756,1542,1566]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="4" pageNumber="531" unit="mm" value="36.2">36.2 mm</quantity>
|
||
). The humero–pectoral sulcus intersects the posterior part of the entoplastron. It slightly bends backward on the hyoplastra. The pectoro–abdominal sulcus arches over the lateral part of the hyoplastra. The axillary and right inguinal scutes are invisible, although the left inguinal is present and is isolated from the femoral scute as in
|
||
<taxonomicName id="3DC562A31A19D03EFD0AF9E23361F927" box="[726,849,1651,1675]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="531" phylum="Chordata" rank="species" species="sinensis">
|
||
<emphasis id="C8B1C5321A19D03EFD0AF9E232DAF926" box="[726,746,1651,1674]" italics="true" pageId="4" pageNumber="531">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A19D03EFD24F9E53361F927" box="[760,849,1652,1675]" italics="true" pageId="4" pageNumber="531">sinensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A19D03FFF15F90730C0FE63" blockId="4.[151,1436,1506,2035]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="532" pageId="4" pageNumber="531">
|
||
<emphasis id="C8B1C5321A19D03EFF15F907315FF903" bold="true" box="[201,367,1686,1711]" pageId="4" pageNumber="531">Comparisons.</emphasis>
|
||
The musk duct foramen in the right third peripheral of the present material (MTE1) indicates that this taxon belongs to the family
|
||
<taxonomicName id="3DC562A31A19D03EFD9AF92B32D2F97E" box="[582,738,1722,1746]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="531" phylum="Chordata" rank="family">Geoemydidae</taxonomicName>
|
||
(Hirayama 1985; Joyce and Bell 2004). MTE1 is similar to
|
||
<taxonomicName id="3DC562A31A19D03EFF4BF94F3180F95A" authority="Cuvierichelys" authorityName="Cuvierichelys" box="[151,432,1758,1782]" class="Testudines" family="Geoemydidae" genus="Chinemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A19D03EFF4BF94F3135F95A" box="[151,261,1758,1782]" italics="true" pageId="4" pageNumber="531">Chinemys</emphasis>
|
||
,
|
||
<emphasis id="C8B1C5321A19D03EFECAF94F3180F95A" box="[278,432,1758,1782]" italics="true" pageId="4" pageNumber="531">Cuvierichelys</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A19D03EFE1DF9713206F95B" box="[449,566,1760,1783]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A19D03EFE1DF9713206F95B" box="[449,566,1760,1783]" italics="true" pageId="4" pageNumber="531">Mauremys</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A19D03EFD94F94F32AAF95A" box="[584,666,1758,1782]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A19D03EFD94F94F32AAF95A" box="[584,666,1758,1782]" italics="true" pageId="4" pageNumber="531">Ocadia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A19D03EFD77F94F3370F95A" box="[683,832,1758,1782]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A19D03EFD77F94F3370F95A" box="[683,832,1758,1782]" italics="true" pageId="4" pageNumber="531">Palaeochelys</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A19D03EFC8EF94F3396F95A" box="[850,934,1758,1782]" class="Reptilia" family="Geoemydidae" genus="Sacalia" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A19D03EFC8EF94F3396F95A" box="[850,934,1758,1782]" italics="true" pageId="4" pageNumber="531">Sacalia</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3DC562A31A19D03EFC35F94F34A6F95A" box="[1001,1174,1758,1782]" class="Reptilia" family="Geoemydidae" genus="Siebenrockiella" kingdom="Animalia" order="Testudines" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A19D03EFC35F94F34A6F95A" box="[1001,1174,1758,1782]" italics="true" pageId="4" pageNumber="531">Siebenrockiella</emphasis>
|
||
</taxonomicName>
|
||
in lacking the plastral hinge and in having the entoplastron intersected by the humero–pectoral sulcus and the neurals short–sided anteriorly. In addition, MTE1 has four informative character states for genus level taxonomy (
|
||
<tableCitation id="B7472C9B1A19D03EFB64F8B7353DF892" box="[1208,1293,1830,1854]" captionStart="TABLE 1" captionStartId="5.[151,239,511,535]" captionText="TABLE 1. Comparisons of the Tanegashima geoemydid (MTE 1) with hingeless geoemydids (see Appendix) having the entoplastron intersected by the humero – pectoral sulcus and the neurals short – sided anteriorly. Symbols, +, –, and?, indicate presence, absence, and undetermined status of a given character state. Character state (1): second and third vertebrals rectangular and as long as wide, character state (2): posterior margin of the carapace unserrated, character state (3): medial length of the gular shorter than interhumeral sulcus, character state (4): plastral buttresses developed, extending halfway of the costals, character state (5): medial and two lateral obtuse discoutinuous keels on the carapace present, character state (6): first vertebral scute expanded anterolaterally, reaching the second marginal, character state (7): second and third vertebral scutes much narrower than the first anteroposteriorly, character state (8): length of entoplastron nearly as long as interhyoplastral suture, character state (9): costals dovetailed with one another in outline, and character state (10): third pleural overlapping only sixth and seventh peripherals." pageId="4" pageNumber="531">Table 1</tableCitation>
|
||
): (1) second and third vertebrals nearly rectangular, as long as wide (modified from Hirayama
|
||
<emphasis id="C8B1C5321A19D03EFB87F8DD34A0F8CF" box="[1115,1168,1867,1891]" italics="true" pageId="4" pageNumber="531">et al</emphasis>
|
||
. 2007), (2) absence of serration on the posterior margin of the carapace (e.g., Ernst and Barbour 1989; Yasukawa
|
||
<emphasis id="C8B1C5321A19D03EFB44F8E134F9F82A" box="[1176,1225,1902,1927]" italics="true" pageId="4" pageNumber="531">et al</emphasis>
|
||
. 2001), (3) medial length of the gular shorter than the interhumeral sulcus (Hirayama
|
||
<emphasis id="C8B1C5321A19D03EFC76F80533EEF807" box="[938,990,1939,1963]" italics="true" pageId="4" pageNumber="531">et al</emphasis>
|
||
. 2007), and (4) the plastral buttresses moderately developed, extending to half way of the costals (Hirayama
|
||
<emphasis id="C8B1C5321A19D03EFC6AF82933D8F862" box="[950,1000,1974,1999]" italics="true" pageId="4" pageNumber="531">et al</emphasis>
|
||
. 2007). Of these, the character (3) and (4) are known as diagnostic characters for the genus
|
||
<taxonomicName id="3DC562A31A19D03EFD28F84A3466F85E" authority="Hirayama et al. 2007" authorityName="Hirayama et al." authorityYear="2007" box="[756,1110,2010,2035]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A19D03EFD28F84A3377F85F" box="[756,839,2011,2035]" italics="true" pageId="4" pageNumber="531">Ocadia</emphasis>
|
||
(Hirayama
|
||
<emphasis id="C8B1C5321A19D03EFC13F84D3432F85F" box="[975,1026,2011,2035]" italics="true" pageId="4" pageNumber="531">et al</emphasis>
|
||
. 2007)
|
||
</taxonomicName>
|
||
, but variably seen in several comparative taxa (
|
||
<tableCitation id="B7472C9B1A18D03FFEB7FF0631F2FF03" box="[363,450,151,175]" captionStart="TABLE 1" captionStartId="5.[151,239,511,535]" captionText="TABLE 1. Comparisons of the Tanegashima geoemydid (MTE 1) with hingeless geoemydids (see Appendix) having the entoplastron intersected by the humero – pectoral sulcus and the neurals short – sided anteriorly. Symbols, +, –, and?, indicate presence, absence, and undetermined status of a given character state. Character state (1): second and third vertebrals rectangular and as long as wide, character state (2): posterior margin of the carapace unserrated, character state (3): medial length of the gular shorter than interhumeral sulcus, character state (4): plastral buttresses developed, extending halfway of the costals, character state (5): medial and two lateral obtuse discoutinuous keels on the carapace present, character state (6): first vertebral scute expanded anterolaterally, reaching the second marginal, character state (7): second and third vertebral scutes much narrower than the first anteroposteriorly, character state (8): length of entoplastron nearly as long as interhyoplastral suture, character state (9): costals dovetailed with one another in outline, and character state (10): third pleural overlapping only sixth and seventh peripherals." pageId="5" pageNumber="532">Table 1</tableCitation>
|
||
). Character (1) is shared exclusively with
|
||
<taxonomicName id="3DC562A31A18D03FFC68FF093479FF1C" box="[948,1097,152,176]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="nipponica">
|
||
<emphasis id="C8B1C5321A18D03FFC68FF0933F8FF03" box="[948,968,152,175]" italics="true" pageId="5" pageNumber="532">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A18D03FFC05FF083479FF1C" box="[985,1097,153,176]" italics="true" pageId="5" pageNumber="532">nipponica</emphasis>
|
||
</taxonomicName>
|
||
, some
|
||
<taxonomicName id="3DC562A31A18D03FFB43FF09352DFF1C" box="[1183,1309,152,176]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="sinensis">
|
||
<emphasis id="C8B1C5321A18D03FFB43FF09352DFF1C" box="[1183,1309,152,176]" italics="true" pageId="5" pageNumber="532">O. sinensis</emphasis>
|
||
</taxonomicName>
|
||
, and some
|
||
<taxonomicName id="3DC562A31A18D03FFF4BFF2D311DFF78" box="[151,301,188,212]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A18D03FFF4BFF2D311DFF78" box="[151,301,188,212]" italics="true" pageId="5" pageNumber="532">Palaeochelys</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="3DC562A31A18D03FFEE0FF2C3182FF78" box="[316,434,188,212]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="crocheti">
|
||
<emphasis id="C8B1C5321A18D03FFEE0FF2C3182FF78" box="[316,434,188,212]" italics="true" pageId="5" pageNumber="532">P. crocheti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3DC562A31A18D03FFE3BFF2C326BFF78" box="[487,603,188,212]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="C8B1C5321A18D03FFE3BFF2C326BFF78" box="[487,603,188,212]" italics="true" pageId="5" pageNumber="532">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
). A similar state is seen in
|
||
<taxonomicName id="3DC562A31A18D03FFC55FF2C3434FF78" box="[905,1028,188,212]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="gaudryi">
|
||
<emphasis id="C8B1C5321A18D03FFC55FF2C3390FF78" box="[905,928,189,212]" italics="true" pageId="5" pageNumber="532">M</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A18D03FFC71FF2D3434FF78" box="[941,1028,188,212]" italics="true" pageId="5" pageNumber="532">gaudryi</emphasis>
|
||
</taxonomicName>
|
||
, some
|
||
<taxonomicName id="3DC562A31A18D03FFB88FF2C34EBFF78" box="[1108,1243,189,212]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="japonica">
|
||
<emphasis id="C8B1C5321A18D03FFB88FF2C345BFF78" box="[1108,1131,189,212]" italics="true" pageId="5" pageNumber="532">M</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A18D03FFBA5FF2C34EBFF78" box="[1145,1243,189,212]" italics="true" pageId="5" pageNumber="532">japonica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A18D03FFB35FF2C35A4FF78" box="[1257,1428,188,212]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="massiliensis">
|
||
<emphasis id="C8B1C5321A18D03FFB35FF2C3530FF78" box="[1257,1280,189,212]" italics="true" pageId="5" pageNumber="532">M</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A18D03FFAD1FF2D35A4FF78" box="[1293,1428,188,212]" italics="true" pageId="5" pageNumber="532">massiliensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3DC562A31A18D03FFF1AFF70316BFF54" box="[198,347,224,248]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="sarmatica">
|
||
<emphasis id="C8B1C5321A18D03FFF1AFF7030EDFF54" box="[198,221,225,248]" italics="true" pageId="5" pageNumber="532">M</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A18D03FFF37FF70316BFF54" box="[235,347,225,248]" italics="true" pageId="5" pageNumber="532">sarmatica</emphasis>
|
||
</taxonomicName>
|
||
, in which the second and third vertebrals are much wider than long (de Broin 1977; Hervet 2004; Takahashi per. obs. on
|
||
<taxonomicName id="3DC562A31A18D03FFE79FE943200FEB0" box="[421,560,261,284]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="japonica">
|
||
<emphasis id="C8B1C5321A18D03FFE79FE943200FEB0" box="[421,560,261,284]" italics="true" pageId="5" pageNumber="532">M. japonica</emphasis>
|
||
</taxonomicName>
|
||
). Character state (2) is informative for discriminating
|
||
<taxonomicName id="3DC562A31A18D03FFB72FE953531FEB0" box="[1198,1281,260,284]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A18D03FFB72FE953531FEB0" box="[1198,1281,260,284]" italics="true" pageId="5" pageNumber="532">Ocadia</emphasis>
|
||
</taxonomicName>
|
||
from several species of
|
||
<taxonomicName id="3DC562A31A18D03FFEC9FEB831BAFEEC" box="[277,394,297,320]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A18D03FFEC9FEB831BAFEEC" box="[277,394,297,320]" italics="true" pageId="5" pageNumber="532">Mauremys</emphasis>
|
||
</taxonomicName>
|
||
as well as a few
|
||
<taxonomicName id="3DC562A31A18D03FFDBFFEB932C8FEEC" box="[611,760,296,320]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A18D03FFDBFFEB932C8FEEC" box="[611,760,296,320]" italics="true" pageId="5" pageNumber="532">Palaeochelys</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="3DC562A31A18D03FFCD0FEB833B4FEEC" box="[780,900,296,320]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="C8B1C5321A18D03FFCD0FEB833B4FEEC" box="[780,900,296,320]" italics="true" pageId="5" pageNumber="532">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
), and
|
||
<taxonomicName id="3DC562A31A18D03FFC0FFEB934B0FEEC" box="[979,1152,296,320]" class="Reptilia" family="Geoemydidae" genus="Siebenrockiella" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A18D03FFC0FFEB934B0FEEC" box="[979,1152,296,320]" italics="true" pageId="5" pageNumber="532">Siebenrockiella</emphasis>
|
||
</taxonomicName>
|
||
(Sacco 1889; Ernst and Barbour 1989; Hervet 2004; Claude
|
||
<emphasis id="C8B1C5321A18D03FFDE0FEDC3246FEC8" box="[572,630,332,356]" italics="true" pageId="5" pageNumber="532">et al.</emphasis>
|
||
2007; Chesi
|
||
<emphasis id="C8B1C5321A18D03FFCCDFEDC337BFEC8" box="[785,843,332,356]" italics="true" pageId="5" pageNumber="532">et al.</emphasis>
|
||
2009). Among the comparative genera, all of these four character states seen in MTE1 are exclusively shared with the genus
|
||
<taxonomicName id="3DC562A31A18D03FFC36FEE1340DFE24" box="[1002,1085,368,392]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A18D03FFC36FEE1340DFE24" box="[1002,1085,368,392]" italics="true" pageId="5" pageNumber="532">Ocadia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="B7472C9B1A18D03FFB91FEFE3493FE2B" box="[1101,1187,367,391]" captionStart="TABLE 1" captionStartId="5.[151,239,511,535]" captionText="TABLE 1. Comparisons of the Tanegashima geoemydid (MTE 1) with hingeless geoemydids (see Appendix) having the entoplastron intersected by the humero – pectoral sulcus and the neurals short – sided anteriorly. Symbols, +, –, and?, indicate presence, absence, and undetermined status of a given character state. Character state (1): second and third vertebrals rectangular and as long as wide, character state (2): posterior margin of the carapace unserrated, character state (3): medial length of the gular shorter than interhumeral sulcus, character state (4): plastral buttresses developed, extending halfway of the costals, character state (5): medial and two lateral obtuse discoutinuous keels on the carapace present, character state (6): first vertebral scute expanded anterolaterally, reaching the second marginal, character state (7): second and third vertebral scutes much narrower than the first anteroposteriorly, character state (8): length of entoplastron nearly as long as interhyoplastral suture, character state (9): costals dovetailed with one another in outline, and character state (10): third pleural overlapping only sixth and seventh peripherals." pageId="5" pageNumber="532">Table 1</tableCitation>
|
||
). The other character states (5–10) are not informative for genus level identification. Thus, in the following, we compare MTE1 within
|
||
<taxonomicName id="3DC562A31A18D03FFF4BFE2930DAFE7C" box="[151,234,440,464]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A18D03FFF4BFE2930DAFE7C" box="[151,234,440,464]" italics="true" pageId="5" pageNumber="532">Ocadia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="AEBA49A81A18D03FFF4BFE6E3412FC9B" pageId="5" pageNumber="532">
|
||
<paragraph id="FA7A19201A18D03FFF4BFE6E3412FC9B" blockId="5.[151,1436,511,823]" pageId="5" pageNumber="532">
|
||
<emphasis id="C8B1C5321A18D03FFF4BFE6E313AFDBB" bold="true" box="[151,266,511,535]" pageId="5" pageNumber="532">TABLE 1.</emphasis>
|
||
Comparisons of the Tanegashima geoemydid (MTE1) with hingeless geoemydids (see Appendix) having the entoplastron intersected by the humero–pectoral sulcus and the neurals short–sided anteriorly. Symbols, +, –, and?, indicate presence, absence, and undetermined status of a given character state. Character state (1): second and third vertebrals rectangular and as long as wide, character state (2): posterior margin of the carapace unserrated, character state (3): medial length of the gular shorter than interhumeral sulcus, character state (4): plastral buttresses developed, extending halfway of the costals, character state (5): medial and two lateral obtuse discoutinuous keels on the carapace present, character state (6): first vertebral scute expanded anterolaterally, reaching the second marginal, character state (7): second and third vertebral scutes much narrower than the first anteroposteriorly, character state (8): length of entoplastron nearly as long as interhyoplastral suture, character state (9): costals dovetailed with one another in outline, and character state (10): third pleural overlapping only sixth and seventh peripherals.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA7A19201A18D03FFF43FCC23320FCC4" blockId="5.[159,1388,850,2003]" box="[159,784,850,872]" pageId="5" pageNumber="532">Genus Species Character state</paragraph>
|
||
<paragraph id="FA7A19201A18D03FFDA5FCED355CFC3D" blockId="5.[159,1388,850,2003]" box="[633,1388,891,914]" pageId="5" pageNumber="532">
|
||
1 2 3 4 5 6 7
|
||
<date id="8E7B3FE01A18D03FFB72FCED355CFC3D" box="[1198,1388,892,913]" pageId="5" pageNumber="532" value="2010-09-08">8 9 10</date>
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A18D03FFF43FC343550FC17" blockId="5.[159,1388,850,2003]" box="[159,1376,933,955]" pageId="5" pageNumber="532">MTE1 + + + + + + + + + +</paragraph>
|
||
<paragraph id="FA7A19201A18D03FFF43FC5E356FFB9B" blockId="5.[159,1388,850,2003]" pageId="5" pageNumber="532">
|
||
<emphasis id="C8B1C5321A18D03FFF43FC5E3230FC49" box="[159,512,975,997]" italics="true" pageId="5" pageNumber="532">
|
||
<taxonomicName id="3DC562A31A18D03FFF43FC5E3133FC49" box="[159,259,975,997]" class="Testudines" family="Geoemydidae" genus="Chinemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">Chinemys</taxonomicName>
|
||
<taxonomicName id="3DC562A31A18D03FFEACFC5E3230FC49" box="[368,512,975,997]" class="Testudines" family="Geoemydidae" genus="Chinemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="fenhoense">Ch. fenhoense</taxonomicName>
|
||
</emphasis>
|
||
– +?? – – – – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFC6931CBFBA2" box="[368,507,1016,1038]" class="Testudines" family="Geoemydidae" genus="Chinemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="nigricans">
|
||
<emphasis id="C8B1C5321A18D03FFEACFC6931CBFBA2" box="[368,507,1016,1038]" italics="true" pageId="5" pageNumber="532">Ch. nigricans</emphasis>
|
||
</taxonomicName>
|
||
– + – – – – – – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFBB031D8FB9B" box="[368,488,1057,1079]" class="Testudines" family="Geoemydidae" genus="Chinemys" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="reevesii">
|
||
<emphasis id="C8B1C5321A18D03FFEACFBB031D8FB9B" box="[368,488,1057,1079]" italics="true" pageId="5" pageNumber="532">Ch. reevesii</emphasis>
|
||
</taxonomicName>
|
||
– + +/– – + +/– +/– – – –
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A18D03FFF43FBDA356DFBCD" blockId="5.[159,1388,850,2003]" box="[159,1373,1099,1121]" pageId="5" pageNumber="532">
|
||
<emphasis id="C8B1C5321A18D03FFF43FBDA3238FBCD" box="[159,520,1099,1121]" italics="true" pageId="5" pageNumber="532">
|
||
Cuvierichelys
|
||
<taxonomicName id="3DC562A31A18D03FFEACFBDD3238FBCD" box="[368,520,1100,1121]" pageId="5" pageNumber="532" rank="species" species="parisiensis">Cu. parisiensis</taxonomicName>
|
||
</emphasis>
|
||
? + +? –?? – –?
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A18D03FFF43FBE7356FF92F" blockId="5.[159,1388,850,2003]" pageId="5" pageNumber="532">
|
||
<emphasis id="C8B1C5321A18D03FFF43FBE73239FB27" box="[159,521,1142,1163]" italics="true" pageId="5" pageNumber="532">
|
||
<taxonomicName id="3DC562A31A18D03FFF43FBE73139FB27" box="[159,265,1142,1163]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">Mauremys</taxonomicName>
|
||
<taxonomicName id="3DC562A31A18D03FFEACFBE73239FB27" box="[368,521,1142,1163]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="annamensis">M. annamensis</taxonomicName>
|
||
</emphasis>
|
||
– + +/– + – +/– + – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFB3131C0FB19" box="[368,496,1184,1205]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="campanii">
|
||
<emphasis id="C8B1C5321A18D03FFEACFB3131C0FB19" box="[368,496,1184,1205]" italics="true" pageId="5" pageNumber="532">M. campanii</emphasis>
|
||
</taxonomicName>
|
||
– – –?? + + + – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFB5B31D1FB73" box="[368,481,1225,1247]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="gaudryi">
|
||
<emphasis id="C8B1C5321A18D03FFEACFB5B31D1FB73" box="[368,481,1225,1247]" italics="true" pageId="5" pageNumber="532">M. gaudryi</emphasis>
|
||
</taxonomicName>
|
||
–? –? – + + – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFB6531DBFAA5" box="[368,491,1268,1289]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="japonica">
|
||
<emphasis id="C8B1C5321A18D03FFEACFB6531DBFAA5" box="[368,491,1268,1289]" italics="true" pageId="5" pageNumber="532">M. japonica</emphasis>
|
||
</taxonomicName>
|
||
– – +/– – – +/– + – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFA8F31EDFA9F" box="[368,477,1309,1331]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="leprosa">
|
||
<emphasis id="C8B1C5321A18D03FFEACFA8F31EDFA9F" box="[368,477,1309,1331]" italics="true" pageId="5" pageNumber="532">M. leprosa</emphasis>
|
||
</taxonomicName>
|
||
– + – – + + – – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFAD9323CFAF1" box="[368,524,1351,1373]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="massiliensis">
|
||
<emphasis id="C8B1C5321A18D03FFEACFAD9323CFAF1" box="[368,524,1351,1373]" italics="true" pageId="5" pageNumber="532">M. massiliensis</emphasis>
|
||
</taxonomicName>
|
||
– + –? + + – –? –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFAE331E7FA2B" box="[368,471,1394,1415]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="mutica">
|
||
<emphasis id="C8B1C5321A18D03FFEACFAE331E7FA2B" box="[368,471,1394,1415]" italics="true" pageId="5" pageNumber="532">M. mutica</emphasis>
|
||
</taxonomicName>
|
||
– – +/– – – +/– + + – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFA0D31E4FA1D" box="[368,468,1436,1457]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="portisi">
|
||
<emphasis id="C8B1C5321A18D03FFEACFA0D31E4FA1D" box="[368,468,1436,1457]" italics="true" pageId="5" pageNumber="532">M. portisi</emphasis>
|
||
</taxonomicName>
|
||
– – –? – + – – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFA5731CDFA77" box="[368,509,1477,1499]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="pygolopha">
|
||
<emphasis id="C8B1C5321A18D03FFEACFA5731CDFA77" box="[368,509,1477,1499]" italics="true" pageId="5" pageNumber="532">M. pygolopha</emphasis>
|
||
</taxonomicName>
|
||
– – – – – – – + – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACFA6131D2F9A9" box="[368,482,1519,1541]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="rivulata">
|
||
<emphasis id="C8B1C5321A18D03FFEACFA6131D2F9A9" box="[368,482,1519,1541]" italics="true" pageId="5" pageNumber="532">M. rivulata</emphasis>
|
||
</taxonomicName>
|
||
– + – – + – – – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACF98B31C7F983" box="[368,503,1562,1583]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="sarmatica">
|
||
<emphasis id="C8B1C5321A18D03FFEACF98B31C7F983" box="[368,503,1562,1583]" italics="true" pageId="5" pageNumber="532">M. sarmatica</emphasis>
|
||
</taxonomicName>
|
||
– + – – – + + + – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACF9D53222F9F5" box="[368,530,1603,1625]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="thanhinensis">
|
||
<emphasis id="C8B1C5321A18D03FFEACF9D53222F9F5" box="[368,530,1603,1625]" italics="true" pageId="5" pageNumber="532">M. thanhinensis</emphasis>
|
||
</taxonomicName>
|
||
– +/– +/– – – +/– + + – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACF9FF31F9F92F" box="[368,457,1645,1667]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="yabei">
|
||
<emphasis id="C8B1C5321A18D03FFEACF9FF31F9F92F" box="[368,457,1645,1667]" italics="true" pageId="5" pageNumber="532">M. yabei</emphasis>
|
||
</taxonomicName>
|
||
–? – – – + + + – –
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A18D03FFF43F906356FF97B" blockId="5.[159,1388,850,2003]" pageId="5" pageNumber="532">
|
||
<emphasis id="C8B1C5321A18D03FFF43F90631C4F901" box="[159,500,1687,1709]" italics="true" pageId="5" pageNumber="532">
|
||
<taxonomicName id="3DC562A31A18D03FFF43F90630DAF901" box="[159,234,1687,1709]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">Ocadia</taxonomicName>
|
||
<taxonomicName id="3DC562A31A18D03FFEACF90931C4F901" box="[368,500,1688,1709]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="nipponica">O. nipponica</taxonomicName>
|
||
</emphasis>
|
||
+ + + + – + + – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACF95331EFF97B" box="[368,479,1730,1751]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="sinensis">
|
||
<emphasis id="C8B1C5321A18D03FFEACF95331EFF97B" box="[368,479,1730,1751]" italics="true" pageId="5" pageNumber="532">O. sinensis</emphasis>
|
||
</taxonomicName>
|
||
+/– + +/– + + – – – – –
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A18D03FFF43F97A356FF8F9" blockId="5.[159,1388,850,2003]" pageId="5" pageNumber="532">
|
||
<emphasis id="C8B1C5321A18D03FFF43F97A314DF8AD" box="[159,381,1771,1793]" italics="true" pageId="5" pageNumber="532">
|
||
<taxonomicName id="3DC562A31A18D03FFF43F97A3116F8AD" box="[159,294,1771,1793]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">Palaeochelys</taxonomicName>
|
||
<taxonomicName id="3DC562A31A18D03FFEACF97D314DF8AD" box="[368,381,1772,1793]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="crocheti">P</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A18D03FFE56F97A31ECF8AD" box="[394,476,1771,1793]" italics="true" pageId="5" pageNumber="532">crocheti</emphasis>
|
||
+ + + – – – + + – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACF88731E8F887" box="[368,472,1813,1835]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="laurenti">
|
||
<emphasis id="C8B1C5321A18D03FFEACF88731E8F887" box="[368,472,1813,1835]" italics="true" pageId="5" pageNumber="532">P. laurenti</emphasis>
|
||
</taxonomicName>
|
||
+ – + – – +/– + – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACF8D131C0F8F9" box="[368,496,1855,1877]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="vallisnerii">
|
||
<emphasis id="C8B1C5321A18D03FFEACF8D131C0F8F9" box="[368,496,1855,1877]" italics="true" pageId="5" pageNumber="532">P. vallisnerii</emphasis>
|
||
</taxonomicName>
|
||
– + + – – + + – – –
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A18D03FFF43F8F8356FF8D3" blockId="5.[159,1388,850,2003]" box="[159,1375,1897,1919]" pageId="5" pageNumber="532">
|
||
<taxonomicName id="3DC562A31A18D03FFF43F8F8318CF8D3" box="[159,444,1897,1919]" class="Reptilia" family="Geoemydidae" genus="Sacalia" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A18D03FFF43F8F8318CF8D3" box="[159,444,1897,1919]" italics="true" pageId="5" pageNumber="532">Sacalia Sacalia</emphasis>
|
||
</taxonomicName>
|
||
spp. – + +/– – – – +/– – – –
|
||
</paragraph>
|
||
<paragraph id="FA7A19201A18D03CFF43F8023550F87C" blockId="5.[159,1388,850,2003]" lastBlockId="6.[151,1436,1760,2001]" lastPageId="6" lastPageNumber="533" pageId="5" pageNumber="532">
|
||
<emphasis id="C8B1C5321A18D03FFF43F80231CDF805" box="[159,509,1939,1961]" italics="true" pageId="5" pageNumber="532">
|
||
<taxonomicName id="3DC562A31A18D03FFF43F802310CF805" box="[159,316,1939,1961]" class="Reptilia" family="Geoemydidae" genus="Siebenrockiella" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="genus">Siebenrockiella</taxonomicName>
|
||
<taxonomicName id="3DC562A31A18D03FFEACF80531CDF805" box="[368,509,1939,1961]" class="Reptilia" family="Geoemydidae" genus="Siebenrockiella" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="crassicollis">S. crassicollis</taxonomicName>
|
||
</emphasis>
|
||
– – – + – – +/– – – –
|
||
<taxonomicName id="3DC562A31A18D03FFEACF82F31D1F87F" box="[368,481,1981,2003]" class="Reptilia" family="Geoemydidae" genus="Siebenrockiella" kingdom="Animalia" order="Testudines" pageId="5" pageNumber="532" phylum="Chordata" rank="species" species="leytensis">
|
||
<emphasis id="C8B1C5321A18D03FFEACF82F31D1F87F" box="[368,481,1981,2003]" italics="true" pageId="5" pageNumber="532">S. leytensis</emphasis>
|
||
</taxonomicName>
|
||
– – – + – – – – – – Within the genus
|
||
<taxonomicName id="3DC562A31A1BD03CFE51F97031EFF955" box="[397,479,1761,1785]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="533" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A1BD03CFE51F97031EFF955" box="[397,479,1761,1785]" italics="true" pageId="6" pageNumber="533">Ocadia</emphasis>
|
||
</taxonomicName>
|
||
, MTE1 exclusively shares the first vertebral expanded anterolaterally and reaching to the second marginal (character 6) with
|
||
<taxonomicName id="3DC562A31A1BD03CFDBAF8973427F8B1" authority="Hirayama et al. 2007" authorityName="Hirayama et al." authorityYear="2007" box="[614,1047,1797,1821]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="533" phylum="Chordata" rank="species" species="nipponica">
|
||
<emphasis id="C8B1C5321A1BD03CFDBAF897324AF8B1" box="[614,634,1798,1821]" italics="true" pageId="6" pageNumber="533">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A1BD03CFD50F89732CCF8B1" box="[652,764,1798,1821]" italics="true" pageId="6" pageNumber="533">nipponica</emphasis>
|
||
(Hirayama
|
||
<emphasis id="C8B1C5321A1BD03CFC57F89733F7F8B1" box="[907,967,1797,1821]" italics="true" pageId="6" pageNumber="533">et al.</emphasis>
|
||
2007)
|
||
</taxonomicName>
|
||
. On the other hand, it shares the tricarinate discontinuous knob–like keels (character state 5) only with
|
||
<taxonomicName id="3DC562A31A1BD03CFC6CF8B8341CF8ED" box="[944,1068,1833,1857]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="533" phylum="Chordata" rank="species" species="sinensis">
|
||
<emphasis id="C8B1C5321A1BD03CFC6CF8B833F4F8EC" box="[944,964,1833,1856]" italics="true" pageId="6" pageNumber="533">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A1BD03CFC0FF8BB341CF8ED" box="[979,1068,1834,1857]" italics="true" pageId="6" pageNumber="533">sinensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="B7472C9B1A1BD03CFBE6F8B934A0F8EC" box="[1082,1168,1832,1856]" captionStart="TABLE 1" captionStartId="5.[151,239,511,535]" captionText="TABLE 1. Comparisons of the Tanegashima geoemydid (MTE 1) with hingeless geoemydids (see Appendix) having the entoplastron intersected by the humero – pectoral sulcus and the neurals short – sided anteriorly. Symbols, +, –, and?, indicate presence, absence, and undetermined status of a given character state. Character state (1): second and third vertebrals rectangular and as long as wide, character state (2): posterior margin of the carapace unserrated, character state (3): medial length of the gular shorter than interhumeral sulcus, character state (4): plastral buttresses developed, extending halfway of the costals, character state (5): medial and two lateral obtuse discoutinuous keels on the carapace present, character state (6): first vertebral scute expanded anterolaterally, reaching the second marginal, character state (7): second and third vertebral scutes much narrower than the first anteroposteriorly, character state (8): length of entoplastron nearly as long as interhyoplastral suture, character state (9): costals dovetailed with one another in outline, and character state (10): third pleural overlapping only sixth and seventh peripherals." pageId="6" pageNumber="533">Table 1</tableCitation>
|
||
, Smith 1931; Ernst and Barbour 1989; Hirayama
|
||
<emphasis id="C8B1C5321A1BD03CFE68F8DF31D5F8C9" box="[436,485,1869,1893]" italics="true" pageId="6" pageNumber="533">et al</emphasis>
|
||
. 2007). Ernst and Barbour (1989) also described that the dorsal keels on the carapace disappear in aged individuals of
|
||
<taxonomicName id="3DC562A31A1BD03CFDCAF8E032A5F825" box="[534,661,1905,1929]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="533" phylum="Chordata" rank="species" species="sinensis">
|
||
<emphasis id="C8B1C5321A1BD03CFDCAF8E0321AF824" box="[534,554,1905,1928]" italics="true" pageId="6" pageNumber="533">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A1BD03CFDE0F8E332A5F825" box="[572,661,1906,1929]" italics="true" pageId="6" pageNumber="533">sinensis</emphasis>
|
||
</taxonomicName>
|
||
. This suggests that absence of the keels in
|
||
<taxonomicName id="3DC562A31A1BD03CFB44F8E0351EF825" box="[1176,1326,1905,1929]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="533" phylum="Chordata" rank="species" species="nipponica">
|
||
<emphasis id="C8B1C5321A1BD03CFB44F8E0349CF824" box="[1176,1196,1905,1928]" italics="true" pageId="6" pageNumber="533">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A1BD03CFB62F8E3351EF825" box="[1214,1326,1906,1929]" italics="true" pageId="6" pageNumber="533">nipponica</emphasis>
|
||
</taxonomicName>
|
||
could be attributed to loss resulting from aging. Moreover, the second and third vertebrals in MTE1 are much narrower than the first (character state 7), which is one of the diagnostic characters for
|
||
<taxonomicName id="3DC562A31A1BD03CFC1DF828356CF87C" authority="Hirayama et al. 2007" authorityName="Hirayama et al." authorityYear="2007" box="[961,1372,1976,2001]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="533" phylum="Chordata" rank="species" species="nipponica">
|
||
<emphasis id="C8B1C5321A1BD03CFC1DF82833E5F87C" box="[961,981,1977,2000]" italics="true" pageId="6" pageNumber="533">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A1BD03CFC3FF82B3463F87D" box="[995,1107,1978,2001]" italics="true" pageId="6" pageNumber="533">nipponica</emphasis>
|
||
(Hirayama
|
||
<emphasis id="C8B1C5321A1BD03CFB05F82B353AF87D" box="[1241,1290,1977,2001]" italics="true" pageId="6" pageNumber="533">et al</emphasis>
|
||
. 2007)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="AEBA49A81A1BD03CFF4BF9F233B5F91B" httpUri="https://zenodo.org/record/217975/files/figure.png" pageId="6" pageNumber="533" targetBox="[193,1420,222,1608]" targetPageId="6">
|
||
<paragraph id="FA7A19201A1BD03CFF4BF9F233B5F91B" blockId="6.[151,1435,1635,1719]" pageId="6" pageNumber="533">
|
||
<emphasis id="C8B1C5321A1BD03CFF4BF9F23126F9D4" bold="true" box="[151,278,1635,1658]" pageId="6" pageNumber="533">FIGURE 3.</emphasis>
|
||
<taxonomicName id="3DC562A31A1BD03CFEFCF9F2322BF9D5" box="[288,539,1635,1657]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="6" pageNumber="533" phylum="Chordata" rank="species" species="tanegashimensis">
|
||
<emphasis id="C8B1C5321A1BD03CFEFCF9F2322BF9D5" box="[288,539,1635,1657]" italics="true" pageId="6" pageNumber="533">Ocadia tanegashimensis</emphasis>
|
||
</taxonomicName>
|
||
(MTE1) from Tanegashima Island. A–C, the carapace in dorsal (A), ventral (B) and anterior (C) views. D, left fifth to seventh costals in ventral view. E–F, the plastron in dorsal (E) and ventral (F) views. Arrows indicate locations of upper ends of the plastral buttresses. Scale bar: 5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AEBA49A81A1AD03DFF4BFB8833FBFB06" httpUri="https://zenodo.org/record/217976/files/figure.png" pageId="7" pageNumber="534" targetBox="[151,1436,200,1011]" targetPageId="7">
|
||
<paragraph id="FA7A19201A1AD03DFF4BFB8833FBFB06" blockId="7.[151,1436,1049,1194]" pageId="7" pageNumber="534">
|
||
<emphasis id="C8B1C5321A1AD03DFF4BFB883125FB82" bold="true" box="[151,277,1049,1072]" pageId="7" pageNumber="534">FIGURE 4.</emphasis>
|
||
Illustration of
|
||
<taxonomicName id="3DC562A31A1AD03DFE6EFB88329CFB83" box="[434,684,1049,1071]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="tanegashimensis">
|
||
<emphasis id="C8B1C5321A1AD03DFE6EFB88329CFB83" box="[434,684,1049,1071]" italics="true" pageId="7" pageNumber="534">Ocadia tanegashimensis</emphasis>
|
||
</taxonomicName>
|
||
(MTE1) from Tanegashima Island. A, the carapace in dorsal and B, the plastron in ventral views. Abbreviations: abd, abdominal; an, anal; ce, cervical; co, costal; ent, entoplastron; epi, epiplastron; fe, femoral; gu, gular; hu, humeral; hyo, hyoplastron; hyp, hypoplastron; LE, length of the epiplastron; LHS, length of the hyoplastral suture; m, marginal; ne, neural; nu, nuchal; pec, pectoral; pl, pleural; v, vertebral; xip, xiphiplastron. Symbol +: locations of upper ends of the plastral buttresses in ventral view. Scale bar: 5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA7A19201A1AD03DFF15FB45328BF9F8" blockId="7.[151,1437,1236,1620]" pageId="7" pageNumber="534">
|
||
Furthermore, MTE1 has three peculiar character states: length of the entoplastron nearly as long as the interhyoplastral suture (character 8), the second to sixth costal bones showing dovetailed shapes (character 9), and the third pleural overlapping only the sixth and seventh peripherals (character 10). Character 8 is also shared with several species of
|
||
<taxonomicName id="3DC562A31A1AD03DFEB6FAD031EFFAF4" box="[362,479,1345,1368]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="534" phylum="Chordata" rank="genus">
|
||
<emphasis id="C8B1C5321A1AD03DFEB6FAD031EFFAF4" box="[362,479,1345,1368]" italics="true" pageId="7" pageNumber="534">Mauremys</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="3DC562A31A1AD03DFE2EFAD032B1FAF4" box="[498,641,1345,1368]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="campanii">
|
||
<emphasis id="C8B1C5321A1AD03DFE2EFAD032B1FAF4" box="[498,641,1345,1368]" italics="true" pageId="7" pageNumber="534">M. campanii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A1AD03DFD4EFAD03335FAF4" box="[658,773,1345,1368]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="mutica">
|
||
<emphasis id="C8B1C5321A1AD03DFD4EFAD03335FAF4" box="[658,773,1345,1368]" italics="true" pageId="7" pageNumber="534">M. mutica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A1AD03DFCC9FAD03383FAF4" box="[789,947,1344,1368]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="pygolopha">
|
||
<emphasis id="C8B1C5321A1AD03DFCC9FAD03383FAF4" box="[789,947,1344,1368]" italics="true" pageId="7" pageNumber="534">M. pygolopha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A1AD03DFC18FAD0346BFAF4" box="[964,1115,1345,1368]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="sarmatica">
|
||
<emphasis id="C8B1C5321A1AD03DFC18FAD0346BFAF4" box="[964,1115,1345,1368]" italics="true" pageId="7" pageNumber="534">M. sarmatica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3DC562A31A1AD03DFBB7FAD03525FAF4" box="[1131,1301,1344,1368]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="thanhinensi">
|
||
<emphasis id="C8B1C5321A1AD03DFBB7FAD03525FAF4" box="[1131,1301,1344,1368]" italics="true" pageId="7" pageNumber="534">M. thanhinensi</emphasis>
|
||
</taxonomicName>
|
||
s,
|
||
<taxonomicName id="3DC562A31A1AD03DFAEDFAD035A4FAF4" box="[1329,1428,1344,1368]" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="yabei">
|
||
<emphasis id="C8B1C5321A1AD03DFAEDFAD035A4FAF4" box="[1329,1428,1344,1368]" italics="true" pageId="7" pageNumber="534">M. yabei</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3DC562A31A1AD03DFF17FAF43176FAD0" box="[203,326,1380,1404]" class="Reptilia" family="Geoemydidae" genus="Palaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="crocheti">
|
||
<emphasis id="C8B1C5321A1AD03DFF17FAF43176FAD0" box="[203,326,1380,1404]" italics="true" pageId="7" pageNumber="534">P. crocheti</emphasis>
|
||
</taxonomicName>
|
||
; Shikama 1949; de Broin 1977; Hervet 2004; Claude
|
||
<emphasis id="C8B1C5321A1AD03DFC10FAF43439FAD0" box="[972,1033,1380,1404]" italics="true" pageId="7" pageNumber="534">et al.</emphasis>
|
||
2007; Takahashi pers. obs. on
|
||
<taxonomicName id="3DC562A31A1AD03DFAA1FAF430D3FA0C" class="Reptilia" family="Geoemydidae" genus="Mauremys" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="mutica">
|
||
<emphasis id="C8B1C5321A1AD03DFAA1FAF430D3FA0C" italics="true" pageId="7" pageNumber="534">M. mutica</emphasis>
|
||
</taxonomicName>
|
||
). In MTE1, the even–numbered costal plates are expanded whereas the odd–numbered plates exclusive of the first one are narrower distally (character 9). In contrast, the costals in the other comparative taxa are nearly rectangular. The third pleural covers only two peripherals, the seventh and eighth (character 10) in MTE1, but overlaps three peripherals in the two congeneric species as well as the other comparative species (the sixth to eighth, generally, but very rarely the fifth to seventh in
|
||
<taxonomicName id="3DC562A31A1AD03DFCFFF9893392F99C" box="[803,930,1560,1584]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="sinensis">
|
||
<emphasis id="C8B1C5321A1AD03DFCFFF9893307F983" box="[803,823,1560,1583]" italics="true" pageId="7" pageNumber="534">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A1AD03DFC95F9883392F99C" box="[841,930,1561,1584]" italics="true" pageId="7" pageNumber="534">sinensis</emphasis>
|
||
</taxonomicName>
|
||
). Based on these features, MTE1 is clearly distinguished from
|
||
<taxonomicName id="3DC562A31A1AD03DFEADF9AC31DCF9F8" box="[369,492,1597,1620]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="sinensis">
|
||
<emphasis id="C8B1C5321A1AD03DFEADF9AC31B5F9F8" box="[369,389,1597,1620]" italics="true" pageId="7" pageNumber="534">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A1AD03DFE4FF9AC31DCF9F8" box="[403,492,1597,1620]" italics="true" pageId="7" pageNumber="534">sinensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3DC562A31A1AD03DFDFEF9AC3284F9F8" box="[546,692,1597,1620]" class="Testudines" family="Geoemydidae" genus="Ocadia" higherTaxonomySource="CoL" kingdom="Animalia" order="Cryptodira" pageId="7" pageNumber="534" phylum="Chordata" rank="species" species="nipponica">
|
||
<emphasis id="C8B1C5321A1AD03DFDFEF9AC3206F9F8" box="[546,566,1597,1620]" italics="true" pageId="7" pageNumber="534">O</emphasis>
|
||
.
|
||
<emphasis id="C8B1C5321A1AD03DFD98F9AC3284F9F8" box="[580,692,1597,1620]" italics="true" pageId="7" pageNumber="534">nipponica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |