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<document id="829AC4DDAF2C2D8DE0C8D7488787FE3D" ID-DOI="10.6620/ZS.2020.59-21" ID-ISSN="1810-522X" ID-PMC="PMC7807176" ID-PubMed="33456548" ID-Zenodo-Dep="12822787" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721923173720" checkinUser="felipe" docAuthor="Jouladeh-Roudbar, Arash, Ghanavi, Hamid Reza &amp; Doadrio, Ignacio" docDate="2020" docId="726C87BDE26C9C1AFCEF551D27F5FD01" docLanguage="en" docName="ZoolStud.59.21.1-303.pdf" docOrigin="Zoological Studies 59 (21)" docSource="http://dx.doi.org/10.5281/zenodo.12822535" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2019-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2019-.journal_article" docStyleVersion="1" docTitle="Capoeta capoeta" docType="treatment" docVersion="2" lastPageNumber="42" masterDocId="8E55FFC5E24B9C33FFDE5671272CFF97" masterDocTitle="Fig. 4 in Fig. 4 in Fig. 3 in Fig. 21. Sesarmops mora n in Paralbunea dayriti" masterLastPageNumber="303" masterPageNumber="1" pageNumber="40" updateTime="1721926655396" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B912564616850C8A8ED48B4BF7B8AA34">Fig. 4 in Fig. 4 in Fig. 3 in Fig. 21. Sesarmops mora n in Paralbunea dayriti</mods:title>
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<mods:name id="ADCD13A0EEC05727D829A86C6D94A3A2" type="personal">
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<mods:namePart id="2A40587DC8CEB4C78FE951784501BE25">Jouladeh-Roudbar, Arash</mods:namePart>
<mods:affiliation id="346BAA24192F64D7633E377FEE152BBC">Department of Fisheries, Faculty of Natural Resources, University of Tehran, Karaj, Alborz, Iran. E-mail: jouladehroudbar @ ut. ac. ir</mods:affiliation>
<mods:nameIdentifier id="5388EB30177068C2DAC177D8B16DF164" type="email">jouladehroudbar@ut.ac.ir</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="FDDF9ADC013589DAC6D02D54DDCAFBE7">Ghanavi, Hamid Reza</mods:namePart>
<mods:affiliation id="5AD9CFF726BC061E286FAE2FBB1FBC65">Department of Biology, Lund University, Lund, Sweden. * Correspondence: E-mail: hamid. ghanavi @ gmail. com (Ghanavi)</mods:affiliation>
<mods:nameIdentifier id="9FBBF9FCFB2EBD27FCF4AC0387445C96" type="email">hamid.ghanavi@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BCF0519F6DFA613369B837C66CB7CE5E">Doadrio, Ignacio</mods:namePart>
<mods:affiliation id="C22867BE72EF73FF194C3A311F5835AE">Biodiversity and Evolutionary Biology Department, Museo Nacional de Ciencias Naturales-CSIC, Madrid, Spain. E-mail: doadrio @ mncn. csic. es</mods:affiliation>
<mods:nameIdentifier id="5326243F47292282530FB7647EB4479E" type="email">doadrio@mncn.csic.es</mods:nameIdentifier>
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<mods:title id="1D4D3F9AD4847D00A52E6D994EE66300">Zoological Studies</mods:title>
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<mods:date id="57B1447B3D0707CC5DC8BEC42DEA4A86">2020</mods:date>
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<treatment id="726C87BDE26C9C1AFCEF551D27F5FD01" LSID="urn:lsid:plazi:treatment:726C87BDE26C9C1AFCEF551D27F5FD01" httpUri="http://treatment.plazi.org/id/726C87BDE26C9C1AFCEF551D27F5FD01" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40">
<subSubSection id="B2DF6520E26C9C14FCEF551D22ADFC11" box="[817,1409,876,902]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="FA7A36ABE26C9C14FCEF551D22ADFC11" blockId="39.[817,1409,876,902]" box="[817,1409,876,902]" pageId="39" pageNumber="40">
<heading id="A13281C7E26C9C14FCEF551D22ADFC11" bold="true" box="[817,1409,876,902]" centered="true" fontSize="10" level="2" pageId="39" pageNumber="40" reason="3">
<emphasis id="C8B1EAB9E26C9C14FCEF551D22ADFC11" bold="true" box="[817,1409,876,902]" pageId="39" pageNumber="40">
<taxonomicName id="3DC54D28E26C9C14FCEF551D2238FC11" ID-CoL="5X9LR" authority="(Guldenstaedt, 1773)" baseAuthorityName="Guldenstaedt" baseAuthorityYear="1773" box="[817,1300,876,902]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="capoeta">
<emphasis id="C8B1EAB9E26C9C14FCEF551D232AFC11" bold="true" box="[817,1030,876,902]" italics="true" pageId="39" pageNumber="40">Capoeta capoeta</emphasis>
(Güldenstaedt, 1773)
</taxonomicName>
Native
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2DF6520E26C9C1AFBF455FC27F5FD01" lastPageId="41" lastPageNumber="42" pageId="39" pageNumber="40" type="description">
<paragraph id="FA7A36ABE26C9C14FBF455FC23A4FC32" blockId="39.[1066,1160,909,933]" box="[1066,1160,909,933]" pageId="39" pageNumber="40">
(
<figureCitation id="62FE2A2EE26C9C14FBEC55FC23ACFC32" box="[1074,1152,909,933]" captionStart="Fig" captionStartId="41.[136,167,1953,1972]" captionText="Fig. 66. Capoeta capoeta. Photograph from Gedar chay River, Urmia Lake basin." pageId="39" pageNumber="40">Fig. 66</figureCitation>
)
</paragraph>
<paragraph id="FA7A36ABE26C9C14FCF355B923BBFC61" blockId="39.[813,1415,968,1174]" pageId="39" pageNumber="40">
<taxonomicName id="3DC54D28E26C9C14FCF355B92314FC4C" authority="Guldenstadt" authorityName="Guldenstadt" authorityYear="1773" box="[813,1080,968,987]" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="capoeta">
<emphasis id="C8B1EAB9E26C9C14FCF355B924EBFC4C" box="[813,967,968,987]" italics="true" pageId="39" pageNumber="40">Cyprinus capoeta</emphasis>
Güldenstädt
</taxonomicName>
[J. A. von] 1773: 508;
<typeStatus id="257E8809E26C9C14FAD255B8221BFC4B" box="[1292,1335,969,988]" pageId="39" pageNumber="40">Type</typeStatus>
locality:
<collectingRegion id="3801F849E26C9C14FC84559224A5FC61" box="[858,905,995,1014]" country="Georgia" name="Tbilisi" pageId="39" pageNumber="40">Tiflis</collectingRegion>
, Caspian Sea. No
<typeStatus id="257E8809E26C9C14FBFA5592237DFC61" box="[1060,1105,995,1014]" pageId="39" pageNumber="40">types</typeStatus>
known.
</paragraph>
<paragraph id="FA7A36ABE26C9C14FCF3558F2384FBBC" blockId="39.[813,1415,968,1174]" pageId="39" pageNumber="40">
<taxonomicName id="3DC54D28E26C9C14FCF3558F2310FB87" authority="Guldenstadt" authorityName="Guldenstadt" authorityYear="1787" box="[813,1084,1021,1041]" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="fundulus">
<emphasis id="C8B1EAB9E26C9C14FCF3558F24E1FB87" box="[813,973,1021,1041]" italics="true" pageId="39" pageNumber="40">Cyprinus fundulus</emphasis>
Güldenstädt
</taxonomicName>
[J. A. von] 1787: 222;
<typeStatus id="257E8809E26C9C14FAD3558F2214FB86" box="[1293,1336,1022,1041]" pageId="39" pageNumber="40">Type</typeStatus>
locality: Kura
<collectingRegion id="3801F849E26C9C14FC5452692495FBBC" box="[906,953,1048,1067]" country="Gambia" name="Upper River" pageId="39" pageNumber="40">River</collectingRegion>
at
<collectingRegion id="3801F849E26C9C14FC0D52692321FBBC" box="[979,1037,1048,1067]" country="Georgia" name="Tbilisi" pageId="39" pageNumber="40">Tbilisi</collectingRegion>
. No
<typeStatus id="257E8809E26C9C14FBEB5268234EFBBB" box="[1077,1122,1049,1068]" pageId="39" pageNumber="40">types</typeStatus>
known.
</paragraph>
<paragraph id="FA7A36ABE26C9C14FCF352422392FBF7" blockId="39.[813,1415,968,1174]" pageId="39" pageNumber="40">
<taxonomicName id="3DC54D28E26C9C14FCF35242236EFBD1" authority="Keyserling" authorityName="Keyserling" authorityYear="1861" box="[813,1090,1075,1094]" family="Cyprinidae" genus="Scaphiodon" kingdom="Animalia" order="Cypriniformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="asmussii">
<emphasis id="C8B1EAB9E26C9C14FCF3524224F3FBD2" box="[813,991,1075,1094]" italics="true" pageId="39" pageNumber="40">Scaphiodon asmussii</emphasis>
Keyserling
</taxonomicName>
[E. von] 1861: 14 [17];
<typeStatus id="257E8809E26C9C14FACE52422216FBD1" box="[1296,1338,1075,1094]" pageId="39" pageNumber="40">Type</typeStatus>
locality: Near
<collectingRegion id="3801F849E26C9C14FC57523F2497FBF6" box="[905,955,1102,1121]" country="Afghanistan" name="Herat" pageId="39" pageNumber="40">Herat</collectingRegion>
,
<collectingCountry id="82D2763BE26C9C14FC1C523C2301FBF7" box="[962,1069,1101,1120]" name="Afghanistan" pageId="39" pageNumber="40">Afghanistan</collectingCountry>
. No
<typeStatus id="257E8809E26C9C14FB8A523F23ADFBF6" box="[1108,1153,1102,1121]" pageId="39" pageNumber="40">types</typeStatus>
saved.
</paragraph>
<paragraph id="FA7A36ABE26C9C14FCF352192215FB01" blockId="39.[813,1415,968,1174]" pageId="39" pageNumber="40">
<taxonomicName id="3DC54D28E26C9C14FCF352192336FBEC" authority="De Filippi" authorityName="De Filippi" authorityYear="1865" box="[813,1050,1128,1148]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="sevangi">
<emphasis id="C8B1EAB9E26C9C14FCF352192490FBEC" box="[813,956,1128,1147]" italics="true" pageId="39" pageNumber="40">Capoeta sevangi</emphasis>
<materialsCitation id="4AAD3CF6E26C9C14FC1C52182336FBEC" box="[962,1050,1128,1148]" collectionCode="MZUT" country="Armenia" location="Lake Sevan" pageId="39" pageNumber="40" specimenCode="MZUT 695" specimenCount="1" typeStatus="holotype">De Filippi</materialsCitation>
</taxonomicName>
[F.] 1865: 312; Type locality:
<location id="FF1A6070E26C9C14FAFF521922A9FBEC" LSID="urn:lsid:plazi:treatment:726C87BDE26C9C1AFCEF551D27F5FD01:FF1A6070E26C9C14FAFF521922A9FBEC" box="[1313,1413,1128,1147]" country="Armenia" name="Lake Sevan" pageId="39" pageNumber="40">Lake Sevan</location>
(= Goktscha),
<collectingCountry id="82D2763BE26C9C14FC0A52F2230DFB01" box="[980,1057,1155,1174]" name="Armenia" pageId="39" pageNumber="40">Armenia</collectingCountry>
.
<typeStatus id="257E8809E26C9C14FBF752F22354FB01" box="[1065,1144,1155,1174]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
(unique):
<specimenCode id="AA639ED0E26C9C14FB0F52F2221AFB01" box="[1233,1334,1155,1174]" collectionCode="MZUT" country="Italy" httpUri="http://grbio.org/cool/yjj8-is06" name="Museo e Instituto DI Zoologia Sistematica dell' UniversitaDI Torino" pageId="39" pageNumber="40">MZUT 695</specimenCode>
.
</paragraph>
<caption id="AEBA6623E26C9C14FF5651D0251BF822" box="[136,567,1953,1973]" pageId="39" pageNumber="40" startId="39.[136,167,1953,1972]" targetType="figure">
<paragraph id="FA7A36ABE26C9C14FF5651D0251BF822" blockId="39.[136,567,1953,1973]" box="[136,567,1953,1973]" pageId="39" pageNumber="40">
<emphasis id="C8B1EAB9E26C9C14FF5651D027E7F823" bold="true" box="[136,203,1953,1972]" pageId="39" pageNumber="40">Fig. 62.</emphasis>
Distribution map of
<taxonomicName id="3DC54D28E26C9C14FE5D51D3251EF823" authorityName="Jouladeh-Roudbar, Eagderi, Ghanavi and Doadrio" authorityYear="2016" box="[387,562,1953,1973]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="alborzensis">
<emphasis id="C8B1EAB9E26C9C14FE5D51D3251EF823" box="[387,562,1953,1973]" italics="true" pageId="39" pageNumber="40">Capoeta alborzensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="FA7A36ABE2639C1BFF70568C25A5FED2" blockId="40.[174,775,253,459]" pageId="40" pageNumber="41">
<taxonomicName id="3DC54D28E2639C1BFF70568C254CFE87" authority="Kessler" authorityName="Kessler" authorityYear="1872" box="[174,608,253,272]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="steindachneri" subGenus="Scaphiodon">
<emphasis id="C8B1EAB9E2639C1BFF70568C27D1FE87" box="[174,253,253,272]" italics="true" pageId="40" pageNumber="41">Capoeta</emphasis>
(
<emphasis id="C8B1EAB9E2639C1BFED0568C2651FE87" box="[270,381,253,272]" italics="true" pageId="40" pageNumber="41">Scaphiodon</emphasis>
)
<emphasis id="C8B1EAB9E2639C1BFE50568C2523FE87" box="[398,527,253,272]" italics="true" pageId="40" pageNumber="41">steindachneri</emphasis>
Kessler
</taxonomicName>
[K. F.] 1872: 49 [5]; Type locality: Zeravshan River,
<collectingCountry id="82D2763BE2639C1BFDE95766258DFEBD" box="[567,673,279,298]" name="Uzbekistan" pageId="40" pageNumber="41">Uzbekistan</collectingCountry>
.
<typeStatus id="257E8809E2639C1BFD725766242FFEBD" box="[684,771,279,298]" pageId="40" pageNumber="41" type="syntype">Syntypes</typeStatus>
: whereabouts unknown, not ZIN 449395 (1, 2, 3).
</paragraph>
<paragraph id="FA7A36ABE2639C1BFF70573C2621FE02" blockId="40.[174,775,253,459]" pageId="40" pageNumber="41">
<taxonomicName id="3DC54D28E2639C1BFF70573C2685FEF7" authority="Kessler" authorityName="Kessler" authorityYear="1877" box="[174,425,333,352]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="hohenackeri">
<emphasis id="C8B1EAB9E2639C1BFF70573C2648FEF7" box="[174,356,333,352]" italics="true" pageId="40" pageNumber="41">Capoeta hohenackeri</emphasis>
<materialsCitation id="4AAD3CF6E2639C1BFEB7573C2685FEF7" box="[361,425,333,352]" collectionCode="ZIN" collectorName="Kaymaram F" country="Azerbaijan" location="Probably Kura-Araks river" pageId="40" pageNumber="41" specimenCode="ZIN 2864" specimenCount="1" typeStatus="holotype">Kessler</materialsCitation>
</taxonomicName>
[
<collectorName id="5730537DE2639C1BFE6A573C26CDFEF7" box="[436,481,333,352]" name="Kaymaram F" pageId="40" pageNumber="41">K. F.</collectorName>
] 1877: 89; Type locality:
<location id="FF1A6070E2639C1BFD64573C265FFEED" LSID="urn:lsid:plazi:treatment:726C87BDE26C9C1AFCEF551D27F5FD01:FF1A6070E2639C1BFD64573C265FFEED" country="Azerbaijan" name="Probably Kura-Araks river" pageId="40" pageNumber="41">Probably Kura-Araks river</location>
system,
<collectingCountry id="82D2763BE2639C1BFE1A5716250BFEED" box="[452,551,359,378]" name="Azerbaijan" pageId="40" pageNumber="41">Azerbaijan</collectingCountry>
.
<typeStatus id="257E8809E2639C1BFDEF571625AEFEED" box="[561,642,359,378]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
(unique):
<specimenCode id="AA639ED0E2639C1BFD3C57192624FE02" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="40" pageNumber="41" type="Museum">ZIN 2864</specimenCode>
.
</paragraph>
<paragraph id="FA7A36ABE2639C1BFF7057EC25ECFE5D" blockId="40.[174,775,253,459]" pageId="40" pageNumber="41">
<taxonomicName id="3DC54D28E2639C1BFF7057EC26BFFE27" authority="Nikolskii" authorityName="Nikolskii" authorityYear="1897" box="[174,403,413,432]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="gibbosa">
<emphasis id="C8B1EAB9E2639C1BFF7057EC2612FE27" box="[174,318,413,432]" italics="true" pageId="40" pageNumber="41">Capoeta gibbosa</emphasis>
Nikolskii
</taxonomicName>
[A. M.] 1897: 344; Type locality: Bochsani [Bukhsani], se
<collectingRegion id="3801F849E2639C1BFE8257C6269CFE5D" box="[348,432,439,458]" country="Iran" name="Khorasan-e Shomali" pageId="40" pageNumber="41">Khorasan</collectingRegion>
,
<collectingCountry id="82D2763BE2639C1BFE6657C926F1FE5C" box="[440,477,440,459]" name="Iran" pageId="40" pageNumber="41">Iran</collectingCountry>
.
<typeStatus id="257E8809E2639C1BFE3B57C6251BFE5D" box="[485,567,439,458]" pageId="40" pageNumber="41" type="syntype">Syntypes</typeStatus>
: ZIN 11104 (2).
</paragraph>
<paragraph id="FA7A36ABE2639C1BFF3457822510FDBD" blockId="40.[174,775,499,715]" pageId="40" pageNumber="41">
<emphasis id="C8B1EAB9E2639C1BFF34578226A3FD9C" box="[234,399,499,523]" italics="true" pageId="40" pageNumber="41">Common name</emphasis>
: Pr: Siyah mahi ghafghazi, Siyah mahi mamoli, En: Caucasian scraper.
</paragraph>
<paragraph id="FA7A36ABE2639C1BFF34544226C1FD3C" blockId="40.[174,775,499,715]" pageId="40" pageNumber="41">
<emphasis id="C8B1EAB9E2639C1BFF3454422674FDDD" box="[234,344,563,586]" italics="true" pageId="40" pageNumber="41">Diagnosis</emphasis>
: This species is similar to
<taxonomicName id="3DC54D28E2639C1BFD565442242BFDDC" baseAuthorityName="Valenciennes" baseAuthorityYear="1844" box="[648,775,563,587]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="aculeata">
<emphasis id="C8B1EAB9E2639C1BFD565442242BFDDC" box="[648,775,563,587]" italics="true" pageId="40" pageNumber="41">C. aculeata</emphasis>
</taxonomicName>
and
<taxonomicName id="3DC54D28E2639C1BFF01542226ADFDFC" authorityName="Jouladeh-Roudbar, Eagderi, Ghanavi and Doadrio" authorityYear="2016" box="[223,385,595,619]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="alborzensis">
<emphasis id="C8B1EAB9E2639C1BFF01542226ADFDFC" box="[223,385,595,619]" italics="true" pageId="40" pageNumber="41">C. alborzensis</emphasis>
</taxonomicName>
but distinguished from them by a smaller scale size and a higher number of total lateral line scales (4654 vs. 3944).
</paragraph>
<paragraph id="FA7A36ABE2639C1BFF3454C224B0FEA1" blockId="40.[174,775,499,715]" lastBlockId="40.[850,1452,253,726]" pageId="40" pageNumber="41">
<emphasis id="C8B1EAB9E2639C1BFF3454C226E8FD5C" box="[234,452,691,715]" italics="true" pageId="40" pageNumber="41">Meristic characters</emphasis>
: D: IIIIV 78 (7), A: III 5, P: 1620 (1819), V: 910, GR: 1921, LL: 5558, TV: 4344.
</paragraph>
<paragraph id="FA7A36ABE2639C1BFC51574C236CFE42" blockId="40.[850,1452,253,726]" pageId="40" pageNumber="41">
<emphasis id="C8B1EAB9E2639C1BFC51574C233DFEC2" box="[911,1041,317,341]" italics="true" pageId="40" pageNumber="41">Distribution</emphasis>
: Caspian Sea and Urmia Lake basins (
<figureCitation id="62FE2A2EE2639C1BFC84572C2487FEE2" box="[858,939,349,373]" captionStart="Fig" captionStartId="42.[174,205,1163,1182]" captionText="Fig. 67. Distribution map of Capoeta capoeta." pageId="40" pageNumber="41">Fig. 67</figureCitation>
). Found in Aras, Balekhlochay, Qarasu, Kargan
<collectingRegion id="3801F849E2639C1BFC8C570C24A2FE02" box="[850,910,381,405]" country="Gambia" name="Upper River" pageId="40" pageNumber="41">River</collectingRegion>
in the Caspian Sea, also Gedar, Baranduz chay, Nazlo chay, Zarineh, Simineh and Ghale chay Rivers in the Urmia Lake basin.
</paragraph>
<paragraph id="FA7A36ABE2639C1AFC5157AF26CBFDC2" blockId="40.[850,1452,253,726]" lastBlockId="41.[136,737,253,662]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<emphasis id="C8B1EAB9E2639C1BFC5157AF24D0FE62" box="[911,1020,478,501]" italics="true" pageId="40" pageNumber="41">Taxonomy</emphasis>
:
<bibRefCitation id="9E544B5AE2639C1BFBD457AF23A7FE61" author="Berg LS" box="[1034,1163,478,502]" pageId="40" pageNumber="41" refId="ref125108" refString="Berg LS. 1949. Israel program for scientific translations, Jerusalem Freshwater fishes of the USSR and adjacent countries, 496 pp." type="book" year="1949">Berg (1949)</bibRefCitation>
placed in
<taxonomicName id="3DC54D28E2639C1BFB2757AC22AEFE62" authorityName="Ruppell" authorityYear="1835" box="[1273,1410,477,501]" class="Actinopterygii" family="Cyprinidae" genus="Varicorhinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="C8B1EAB9E2639C1BFB2757AC22AEFE62" box="[1273,1410,477,501]" italics="true" pageId="40" pageNumber="41">Varicorhinus</emphasis>
</taxonomicName>
but
<bibRefCitation id="9E544B5AE2639C1BFC8C578C24FBFD81" author="Coad BW" box="[850,983,509,534]" pageId="40" pageNumber="41" pagination="1 - 49" refId="ref126054" refString="Coad BW. 1991. Fishes of the Tigris-Euphrates basin: A Critical Checklist. Syllogeus, Ottawa 68: 1 - 49." type="journal article" year="1991">Coad (1991)</bibRefCitation>
transferred to
<taxonomicName id="3DC54D28E2639C1BFBAE578F23E5FD82" authorityName="Valenciennes" authorityYear="1842" box="[1136,1225,510,533]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="C8B1EAB9E2639C1BFBAE578F23E5FD82" box="[1136,1225,510,533]" italics="true" pageId="40" pageNumber="41">Capoeta</emphasis>
</taxonomicName>
. Zareian et al. (2017) considered
<taxonomicName id="3DC54D28E2639C1BFC12546F2310FDA2" authorityName="De Filippi" authorityYear="1865" box="[972,1084,542,565]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="sevangi">
<emphasis id="C8B1EAB9E2639C1BFC12546F2310FDA2" box="[972,1084,542,565]" italics="true" pageId="40" pageNumber="41">C. sevangi</emphasis>
</taxonomicName>
as a full and valid species without any clear justification; on the other hand,
<bibRefCitation id="9E544B5AE2639C1BFAEF544C2497FDE2" author="Ghanavi HR &amp; Gonzalez EG &amp; Doadrio I." pageId="40" pageNumber="41" pagination="8205 - 8222" refId="ref130657" refString="Ghanavi HR, Gonzalez EG, Doadrio I. 2016. Phylogenetic relationships of freshwater fishes of the genus Capoeta (Actinopterygii, Cyprinidae) in Iran. Ecol Evol 6: 8205 - 8222. doi: 10.1002 / ece 3.2411." type="journal article" year="2016">Ghanavi et al. (2016)</bibRefCitation>
using Cyt
<emphasis id="C8B1EAB9E2639C1BFBF3542C2316FDE2" box="[1069,1082,605,629]" italics="true" pageId="40" pageNumber="41">b</emphasis>
genetic marker placed
<taxonomicName id="3DC54D28E2639C1BFAE7542F2287FDE2" authorityName="De Filippi" authorityYear="1865" box="[1337,1451,606,629]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="sevangi">
<emphasis id="C8B1EAB9E2639C1BFAE7542F2287FDE2" box="[1337,1451,606,629]" italics="true" pageId="40" pageNumber="41">C. sevangi</emphasis>
</taxonomicName>
samples from Sevan Lake (
<typeStatus id="257E8809E2639C1BFBA5540F239EFD01" box="[1147,1202,638,662]" pageId="40" pageNumber="41">Type</typeStatus>
locality of
<taxonomicName id="3DC54D28E2639C1BFAEE540F228EFD02" authorityName="De Filippi" authorityYear="1865" box="[1328,1442,638,661]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="sevangi">
<emphasis id="C8B1EAB9E2639C1BFAEE540F228EFD02" box="[1328,1442,638,661]" italics="true" pageId="40" pageNumber="41">C. sevangi</emphasis>
</taxonomicName>
) nested within
<taxonomicName id="3DC54D28E2639C1BFC3154EF2349FD22" authorityName="Guldenstadt" authorityYear="1773" box="[1007,1125,670,693]" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="capoeta">
<emphasis id="C8B1EAB9E2639C1BFC3154EF2349FD22" box="[1007,1125,670,693]" italics="true" pageId="40" pageNumber="41">C. capoeta</emphasis>
</taxonomicName>
samples from Arpa and Aras Rivers (see
<figureCitation id="62FE2A2EE2639C1BFC0A54CC233FFD41" box="[980,1043,701,726]" captionStart="Fig" captionStartId="1.[136,167,1953,1972]" captionTargetBox="[138,1414,886,1924]" captionTargetId="figure-6@1.[125,1422,871,1939]" captionTargetPageId="1" captionText="Fig. 1. Drainage basin map of Iran." figureDoi="http://doi.org/10.5281/zenodo.12822789" httpUri="https://zenodo.org/record/12822789/files/figure.png" pageId="40" pageNumber="41">
Fig.
<quantity id="3D3D9B4EE2639C1BFBD854CF233FFD41" box="[1030,1043,702,726]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="40" pageNumber="41" unit="in" value="1.0">1</quantity>
</figureCitation>
in
<bibRefCitation id="9E544B5AE2639C1BFBE754CC23DEFD41" author="Levin BA &amp; Freyhof J &amp; Lajbner Z &amp; Perea S &amp; Abdoli A &amp; Gaffaroglu M &amp; Ozulug M &amp; Rubenyan HR &amp; Salnikov VB &amp; Doadrio I &amp; Ozulug M &amp; Rubenyan HR &amp; Salnikov VB &amp; Doadrio I." box="[1081,1266,701,726]" pageId="40" pageNumber="41" pagination="542 - 549" refId="ref132668" refString="Levin BA, Freyhof J, Lajbner Z, Perea S, Abdoli A, Gaffaroglu M, Ozulug M, Rubenyan HR, Salnikov VB, Doadrio I, Ozulug M, Rubenyan HR, Salnikov VB, Doadrio I. 2012. Phylogenetic relationships of the algae scraping cyprinid genus Capoeta (Teleostei: Cyprinidae). Mol Phylogenet Evol 62: 542 - 549. doi: 10.1016 / j. ympev. 2011.09.004." type="journal article" year="2012">Levin et al. 2012</bibRefCitation>
and
<figureCitation id="62FE2A2EE2639C1BFAF754CC22A2FD42" box="[1321,1422,701,725]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[136,167,770,789]" captionStartId-1="9.[136,167,1359,1378]" captionTargetBox-0="[252,1288,249,745]" captionTargetBox-1="[252,1288,838,1333]" captionTargetId-0="figure-6@9.[261,1206,258,741]" captionTargetId-1="figure-8@9.[264,1281,847,1325]" captionTargetPageId-0="9" captionTargetPageId-1="9" captionText-0="Fig. 4. Orders of freshwater fishes of Iran (Number of species, Percent of total)." captionText-1="Fig. 5. Families of freshwater fishes of Iran (Number of species, Percent of total)." figureDoi-0="http://doi.org/10.5281/zenodo.12822797" figureDoi-1="http://doi.org/10.5281/zenodo.12822801" httpUri-0="https://zenodo.org/record/12822797/files/figure.png" httpUri-1="https://zenodo.org/record/12822801/files/figure.png" pageId="40" pageNumber="41">
Figs.
<quantity id="3D3D9B4EE2639C1BFAB854CC22A2FD42" box="[1382,1422,701,725]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="40" pageNumber="41" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">45</quantity>
</figureCitation>
in
<bibRefCitation id="9E544B5AE2629C1AFF56568C2644FE82" author="Ghanavi HR &amp; Gonzalez EG &amp; Doadrio I." box="[136,360,253,277]" pageId="41" pageNumber="42" pagination="8205 - 8222" refId="ref130657" refString="Ghanavi HR, Gonzalez EG, Doadrio I. 2016. Phylogenetic relationships of freshwater fishes of the genus Capoeta (Actinopterygii, Cyprinidae) in Iran. Ecol Evol 6: 8205 - 8222. doi: 10.1002 / ece 3.2411." type="journal article" year="2016">Ghanavi et al. 2016</bibRefCitation>
). In the mentioned study, samples from the Sevan Lake, apart from not forming any independent clade, differed from other samples of
<taxonomicName id="3DC54D28E2629C1AFD16574F27F2FEE2" authorityName="Guldenstadt" authorityYear="1773" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="capoeta">
<emphasis id="C8B1EAB9E2629C1AFD16574F27F2FEE2" italics="true" pageId="41" pageNumber="42">C. capoeta</emphasis>
</taxonomicName>
with less than 0.4% genetic distance (closely related species in the same study had more than 1% genetic distance). In addition, we could not separate this species using the morphological diagnostic characters suggested by Zareian et al. (2017). Therefore, we consider
<taxonomicName id="3DC54D28E2629C1AFF2F578F2646FD82" authorityName="De Filippi" authorityYear="1865" box="[241,362,510,533]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="sevangi">
<emphasis id="C8B1EAB9E2629C1AFF2F578F2646FD82" box="[241,362,510,533]" italics="true" pageId="41" pageNumber="42">C. sevangi</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="3DC54D28E2629C1AFDE3578F2594FD82" authorityName="Guldenstadt" authorityYear="1773" box="[573,696,510,533]" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="capoeta">
<emphasis id="C8B1EAB9E2629C1AFDE3578F2594FD82" box="[573,696,510,533]" italics="true" pageId="41" pageNumber="42">C. capoeta</emphasis>
</taxonomicName>
, as suggested by previous authors (
<bibRefCitation id="9E544B5AE2629C1AFE00546C25BBFDA1" author="Gabrielyan BK" box="[478,663,541,566]" pageId="41" pageNumber="42" pagination="23 - 29" refId="ref130539" refString="Gabrielyan BK. 2001. An annotated checklist of freshwater fishes of Armenia. Naga, The ICLARM Quarterly 24: 23 - 29." type="journal article" year="2001">Gabrielyan 2001</bibRefCitation>
;
<bibRefCitation id="9E544B5AE2629C1AFD7C546F27D0FDC2" author="Turan D &amp; Kottelat M &amp; Kirankaya SG &amp; Engin S." pageId="41" pageNumber="42" pagination="147 - 156" refId="ref135587" refString="Turan D, Kottelat M, Kirankaya SG, Engin S. 2006. Capoeta ekmekciae, a new species of cyprinid fish from northeastern Anatolia (Teleostei: Cyprinidae). Ichthyol Explor Freshw 17: 147 - 156." type="journal article" year="2006">Turan et al. 2006</bibRefCitation>
;
<bibRefCitation id="9E544B5AE2629C1AFED8544C26F7FDC2" author="Ghanavi HR &amp; Gonzalez EG &amp; Doadrio I." box="[262,475,573,597]" pageId="41" pageNumber="42" pagination="8205 - 8222" refId="ref130657" refString="Ghanavi HR, Gonzalez EG, Doadrio I. 2016. Phylogenetic relationships of freshwater fishes of the genus Capoeta (Actinopterygii, Cyprinidae) in Iran. Ecol Evol 6: 8205 - 8222. doi: 10.1002 / ece 3.2411." type="journal article" year="2016">Ghanavi et al. 2016</bibRefCitation>
).
</paragraph>
<caption id="AEBA6623E2639C1BFF7153962548FA6C" box="[175,612,1511,1531]" pageId="40" pageNumber="41" startId="40.[175,206,1511,1530]" targetType="figure">
<paragraph id="FA7A36ABE2639C1BFF7153962548FA6C" blockId="40.[175,612,1511,1531]" box="[175,612,1511,1531]" pageId="40" pageNumber="41">
<emphasis id="C8B1EAB9E2639C1BFF71539627DEFA6D" bold="true" box="[175,242,1511,1530]" pageId="40" pageNumber="41">Fig. 63.</emphasis>
Distribution map of
<taxonomicName id="3DC54D28E2639C1BFE7753992573FA6D" box="[425,607,1512,1531]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="anamisensis">
<emphasis id="C8B1EAB9E2639C1BFE7753992573FA6D" box="[425,607,1512,1531]" italics="true" pageId="40" pageNumber="41">Capoeta anamisensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="AEBA6623E2639C1BFF7051D0243AF823" box="[174,790,1953,1973]" pageId="40" pageNumber="41" startId="40.[174,205,1953,1972]" targetType="figure">
<paragraph id="FA7A36ABE2639C1BFF7051D0243AF823" blockId="40.[174,790,1953,1973]" box="[174,790,1953,1973]" pageId="40" pageNumber="41">
<emphasis id="C8B1EAB9E2639C1BFF7051D027DDF823" bold="true" box="[174,241,1953,1972]" pageId="40" pageNumber="41">Fig. 64.</emphasis>
<taxonomicName id="3DC54D28E2639C1BFF2551D326ADF823" authorityName="Kessler" authorityYear="1877" box="[251,385,1953,1973]" family="Cyprinidae" genus="Capoeta" kingdom="Animalia" order="Cypriniformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="buhsei">
<emphasis id="C8B1EAB9E2639C1BFF2551D326ADF823" box="[251,385,1953,1973]" italics="true" pageId="40" pageNumber="41">Capoeta buhsei</emphasis>
</taxonomicName>
. Photograph from Jajurud River, Namak basin.
</paragraph>
</caption>
<paragraph id="FA7A36ABE2629C1AFF1B542F27F5FD01" blockId="41.[136,737,253,662]" pageId="41" pageNumber="42">
<emphasis id="C8B1EAB9E2629C1AFF1B542F2673FDE2" box="[197,351,606,629]" italics="true" pageId="41" pageNumber="42">Conservation</emphasis>
: IUCN: Least Concern (
<bibRefCitation id="9E544B5AE2629C1AFD5A542C27E2FD01" author="Freyhof J." pageId="41" pageNumber="42" refId="ref128060" refString="Freyhof J. 2014 e. Capoeta capoeta. The IUCN Red List of Threatened Species 2014: e. T 19025492 A 19222863." type="journal volume" year="2014">Freyhof 2014e</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>