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<document id="A16D7338F521A85D43CB4755945FE458" ID-DOI="10.6620/ZS.2020.59-21" ID-ISSN="1810-522X" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721923173720" checkinUser="felipe" docAuthor="Jouladeh-Roudbar, Arash, Ghanavi, Hamid Reza &amp; Doadrio, Ignacio" docDate="2020" docId="726C87BDE25C9C24FF19568C24F7FD21" docLanguage="en" docName="ZoolStud.59.21.1-303.pdf" docOrigin="Zoological Studies 59 (21)" docSource="http://dx.doi.org/10.5281/zenodo.12822535" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2019-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2019-.journal_article" docStyleVersion="1" docTitle="Alosa kessleri" docType="treatment" docVersion="1" lastPageNumber="24" masterDocId="8E55FFC5E24B9C33FFDE5671272CFF97" masterDocTitle="Fig. 4 in Fig. 4 in Fig. 3 in Fig. 21. Sesarmops mora n in Paralbunea dayriti" masterLastPageNumber="303" masterPageNumber="1" pageNumber="24" updateTime="1721925126478" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="84616C87F0B21477E6F9F6634688259F">Fig. 4 in Fig. 4 in Fig. 3 in Fig. 21. Sesarmops mora n in Paralbunea dayriti</mods:title>
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<mods:name id="8636F02AFE9D89645027F85388B818BF" type="personal">
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<mods:namePart id="FF5CBBF79A1D8CB1FBE74C0FF29E69AA">Jouladeh-Roudbar, Arash</mods:namePart>
<mods:affiliation id="2977DF925154B9EA286D0105B751E1F4">Department of Fisheries, Faculty of Natural Resources, University of Tehran, Karaj, Alborz, Iran. E-mail: jouladehroudbar @ ut. ac. ir</mods:affiliation>
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<mods:namePart id="643DFA20CEDB0E43FC1EB59CB8A42F1C">Ghanavi, Hamid Reza</mods:namePart>
<mods:affiliation id="043E0C4D63D65D9A360BFFEA2CA7EB20">Department of Biology, Lund University, Lund, Sweden. * Correspondence: E-mail: hamid. ghanavi @ gmail. com (Ghanavi)</mods:affiliation>
<mods:nameIdentifier id="F5EFB4A7F86C87A66E8EF07593D0F908" type="email">hamid.ghanavi@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BDB3739583C516D4F2D91E08DCC2D4CB">Doadrio, Ignacio</mods:namePart>
<mods:affiliation id="9B34304F5B7D13F1AC214ECB37DD3068">Biodiversity and Evolutionary Biology Department, Museo Nacional de Ciencias Naturales-CSIC, Madrid, Spain. E-mail: doadrio @ mncn. csic. es</mods:affiliation>
<mods:nameIdentifier id="56E16B8892053F1571A95CA690908A16" type="email">doadrio@mncn.csic.es</mods:nameIdentifier>
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<mods:title id="225DF99473846C02553129AD4F891A3C">Zoological Studies</mods:title>
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<mods:date id="DB66EBE39AF4A1AD81BEF21D7BCB35E2">2020</mods:date>
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<treatment id="726C87BDE25C9C24FF19568C24F7FD21" LSID="urn:lsid:plazi:treatment:726C87BDE25C9C24FF19568C24F7FD21" httpUri="http://treatment.plazi.org/id/726C87BDE25C9C24FF19568C24F7FD21" lastPageNumber="24" pageId="23" pageNumber="24">
<subSubSection id="B2DF6520E25C9C24FF19568C26C8FEA1" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="FA7A36ABE25C9C24FF19568C26C8FEA1" blockId="23.[199,674,252,310]" pageId="23" pageNumber="24">
<emphasis id="C8B1EAB9E25C9C24FF19568C258EFE81" bold="true" box="[199,674,252,278]" pageId="23" pageNumber="24">
<taxonomicName id="3DC54D28E25C9C24FF19568C2518FE81" ID-CoL="5TWJV" authority="(Grimm, 1887)" baseAuthorityName="Grimm" baseAuthorityYear="1887" box="[199,564,252,278]" family="Clupeidae" genus="Alosa" kingdom="Animalia" order="Clupeiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="kessleri">
<emphasis id="C8B1EAB9E25C9C24FF19568C2656FE81" bold="true" box="[199,378,253,278]" italics="true" pageId="23" pageNumber="24">Alosa kessleri</emphasis>
(Grimm, 1887)
</taxonomicName>
Native
</emphasis>
(
<figureCitation id="62FE2A2EE25C9C24FE50576C26F0FEA1" box="[398,476,285,310]" captionStart="Fig" captionStartId="23.[136,167,1953,1972]" captionText="Fig. 34. Alosa kessleri. Photograph from Fereydunkenar shore, Caspian Sea." pageId="23" pageNumber="24">Fig. 34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B2DF6520E25C9C24FF56572924F7FD21" pageId="23" pageNumber="24" type="description">
<paragraph id="FA7A36ABE25C9C24FF565729261BFE36" blockId="23.[136,739,344,524]" pageId="23" pageNumber="24">
<taxonomicName id="3DC54D28E25C9C24FF5657292676FEFC" authority="Grimm" authorityName="Grimm" authorityYear="1887" box="[136,346,344,363]" family="Clupeidae" genus="Clupea" kingdom="Animalia" order="Clupeiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="kessleri">
<emphasis id="C8B1EAB9E25C9C24FF565729263FFEFC" box="[136,275,344,363]" italics="true" pageId="23" pageNumber="24">Clupea kessleri</emphasis>
Grimm
</taxonomicName>
[O. von] 1887: 7, 16; Type locality: Volga
<collectingRegion id="3801F849E25C9C24FF6B570227C9FE11" box="[181,229,371,390]" country="Gambia" name="Upper River" pageId="23" pageNumber="24">River</collectingRegion>
delta,
<collectingRegion id="3801F849E25C9C24FEFD57022653FE11" box="[291,383,371,390]" country="Russia" name="Astrakhan'" pageId="23" pageNumber="24">Astrakhan</collectingRegion>
.
<typeStatus id="257E8809E25C9C24FE57570226CFFE11" box="[393,483,371,390]" pageId="23" pageNumber="24" type="lectotype">Lectotype</typeStatus>
: ZIN 15925.
<typeStatus id="257E8809E25C9C24FD82570225F2FE11" box="[604,734,371,390]" pageId="23" pageNumber="24" type="paralectotype">Paralectotypes</typeStatus>
: ZIN 15922 (1).
</paragraph>
<paragraph id="FA7A36ABE25C9C24FF5657D9261BFD9C" blockId="23.[136,739,344,524]" pageId="23" pageNumber="24">
<taxonomicName id="3DC54D28E25C9C24FF5657D926D3FE2C" authority="Tanassiychuk" authorityName="Tanassiychuk" authorityYear="1940" box="[136,511,424,443]" family="Clupeidae" genus="Caspialosa" kingdom="Animalia" order="Clupeiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="subSpecies" species="volgensis" subSpecies="bergi">
<emphasis id="C8B1EAB9E25C9C24FF5657D92652FE2C" box="[136,382,424,443]" italics="true" pageId="23" pageNumber="24">Caspialosa volgensis bergi</emphasis>
Tanassiychuk
</taxonomicName>
[N. P.] 1940: 103; Type locality: Near villiage of Nikolskoye,
<quantity id="3D3D9B4EE25C9C24FDDC57B22568FE41" box="[514,580,451,470]" metricMagnitude="5" metricUnit="m" metricValue="2.4" pageId="23" pageNumber="24" unit="km" value="240.0">240 km</quantity>
above
<collectingRegion id="3801F849E25C9C24FD5A57B225F2FE41" box="[644,734,451,470]" country="Russia" name="Astrakhan'" pageId="23" pageNumber="24">Astrakhan</collectingRegion>
, Volga Delta, northern Caspian Sea,
<collectingCountry id="82D2763BE25C9C24FE2D57AF2503FE66" box="[499,559,478,497]" name="Russia" pageId="23" pageNumber="24">Russia</collectingCountry>
.
<typeStatus id="257E8809E25C9C24FDE757AF25A7FE66" box="[569,651,478,497]" pageId="23" pageNumber="24" type="syntype">Syntypes</typeStatus>
: (several) ZIN 29212 (1).
</paragraph>
<paragraph id="FA7A36ABE25C9C24FF1B544526E6FD1B" blockId="23.[136,737,564,844]" pageId="23" pageNumber="24">
<emphasis id="C8B1EAB9E25C9C24FF1B544526AEFDDB" box="[197,386,564,588]" italics="true" pageId="23" pageNumber="24">Common name</emphasis>
: Pr: Shagmahi-e-mohajer, Puzanok, Zalom, En: Caspian anadromous Shad, Volga herring, Black-spined herring.
</paragraph>
<paragraph id="FA7A36ABE25C9C24FF1B54E42334FEA2" blockId="23.[136,737,564,844]" lastBlockId="23.[813,1414,253,694]" pageId="23" pageNumber="24">
<emphasis id="C8B1EAB9E25C9C24FF1B54E42618FD3B" box="[197,308,661,684]" italics="true" pageId="23" pageNumber="24">Diagnosis</emphasis>
: Body fairly elongate, more “herring-like” than “shad-like”. Total gill rakers 57 to 95 (as in
<taxonomicName id="3DC54D28E25C9C24FF5654A527C2FD7B" baseAuthorityName="Eichwald" baseAuthorityYear="1838" box="[136,238,724,748]" family="Clupeidae" genus="Alosa" kingdom="Animalia" order="Clupeiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caspia">
<emphasis id="C8B1EAB9E25C9C24FF5654A527C2FD7B" box="[136,238,724,748]" italics="true" pageId="23" pageNumber="24">A. caspia</emphasis>
</taxonomicName>
), thick, coarse and shorter than gill filaments in some, long, thin and equal to or longer than gill filaments in others (
<emphasis id="C8B1EAB9E25C9C24FE8055642652FCBB" box="[350,382,789,812]" italics="true" pageId="23" pageNumber="24">i.e.</emphasis>
,
<taxonomicName id="3DC54D28E25C9C24FE5255652521FCBB" baseAuthorityName="Berg" baseAuthorityYear="1913" box="[396,525,788,812]" family="Clupeidae" genus="Caspialosa" kingdom="Animalia" order="Clupeiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="volgensis">
<emphasis id="C8B1EAB9E25C9C24FE5255652521FCBB" box="[396,525,788,812]" italics="true" pageId="23" pageNumber="24">A. volgensis</emphasis>
</taxonomicName>
75128 GR). Teeth found in both jaws and well-developed. Other Caspian shads have less than 50 gill rakers, except
<taxonomicName id="3DC54D28E25C9C24FAC2568F22A9FE82" baseAuthorityName="Eichwald" baseAuthorityYear="1838" box="[1308,1413,254,277]" family="Clupeidae" genus="Alosa" kingdom="Animalia" order="Clupeiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caspia">
<emphasis id="C8B1EAB9E25C9C24FAC2568F22A9FE82" box="[1308,1413,254,277]" italics="true" pageId="23" pageNumber="24">A. caspia</emphasis>
</taxonomicName>
which is deep-bodied.
</paragraph>
<paragraph id="FA7A36ABE25C9C24FCB7574F23D1FE02" blockId="23.[813,1414,253,694]" pageId="23" pageNumber="24">
<emphasis id="C8B1EAB9E25C9C24FCB7574F236CFEC2" box="[873,1088,317,341]" italics="true" pageId="23" pageNumber="24">Meristic characters</emphasis>
: D: IIIV 1216 (14), A: II IV (III) 1521 (18), P: 89, GR: 5795, LL: 5356, TV: 4750 but
<bibRefCitation id="9E544B5AE25C9C24FC41570C234AFE02" author="Svetovidov AN" box="[927,1126,381,405]" pageId="23" pageNumber="24" refId="ref135119" refString="Svetovidov AN. 1952. Israel Program for Scientific Translations, Jerusalem Fauna of the U. S. S. R. (Fauna SSSR). Fishes (Ryby)." type="book" year="1952">Svetovidov (1952)</bibRefCitation>
report 5054.
</paragraph>
<paragraph id="FA7A36ABE25C9C24FCB757EC2375FDA2" blockId="23.[813,1414,253,694]" pageId="23" pageNumber="24">
<emphasis id="C8B1EAB9E25C9C24FCB757EC24C2FE22" box="[873,1006,413,437]" italics="true" pageId="23" pageNumber="24">Distribution</emphasis>
: Caspian Sea basin (
<figureCitation id="62FE2A2EE25C9C24FB0457EC2201FE22" box="[1242,1325,413,437]" captionStart="Fig" captionStartId="24.[174,205,1628,1647]" captionText="Fig. 35. Distribution map of Alosa kessleri." pageId="23" pageNumber="24">Fig. 35</figureCitation>
). In sea and along shores of the central and northern parts, but in south and especially southeast in winter. Enters northern rivers to spawn. In Iranian waters it is found in the southern Caspian Sea basin.
</paragraph>
<paragraph id="FA7A36ABE25C9C24FCB7544F245CFDE2" blockId="23.[813,1414,253,694]" pageId="23" pageNumber="24">
<emphasis id="C8B1EAB9E25C9C24FCB7544F24F2FDC2" box="[873,990,574,597]" italics="true" pageId="23" pageNumber="24">Taxonomy</emphasis>
:
<bibRefCitation id="9E544B5AE25C9C24FC2F544C2221FDC1" author="Whitehead P &amp; Nelson G &amp; Wongratana T." box="[1009,1293,573,598]" pageId="23" pageNumber="24" pagination="460 - 475" refId="ref135840" refString="Whitehead P, Nelson G, Wongratana T. 1988. FAO species catalogue. Clupeoid fishes of the world (Suborder Clupeoidei). Part 2. Engraulididae. FAO Fisheries Synopsis 125: 460 - 475." type="journal article" year="1988">Whitehead et al. (1988)</bibRefCitation>
placed in
<taxonomicName id="3DC54D28E25C9C24FCF3542C2446FDE2" authorityName="Linck" authorityYear="1790" box="[813,874,605,629]" family="Clupeidae" genus="Alosa" kingdom="Animalia" order="Clupeiformes" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="C8B1EAB9E25C9C24FCF3542C2446FDE2" box="[813,874,605,629]" italics="true" pageId="23" pageNumber="24">Alosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FA7A36ABE25C9C24FCB7540F24F7FD21" blockId="23.[813,1414,253,694]" pageId="23" pageNumber="24">
<emphasis id="C8B1EAB9E25C9C24FCB7540F24D5FD02" box="[873,1017,638,661]" italics="true" pageId="23" pageNumber="24">Conservation</emphasis>
: IUCN: Least Concern (
<bibRefCitation id="9E544B5AE25C9C24FADF540C24E3FD21" author="Freyhof J &amp; Kottelat" pageId="23" pageNumber="24" refId="ref129384" refString="Freyhof J, Kottelat. 2008 e. Alosa kessleri. The IUCN Red List of Threatened Species 2008: e. T 135537 A 4140730." type="journal volume" year="2008">Freyhof and Kottelat 2008e</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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