161 lines
19 KiB
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161 lines
19 KiB
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<document ID-CLB-Dataset="25973" ID-DOI="http://dx.doi.org/10.3897/phytokeys.128.34637" ID-GBIF-Dataset="3d9e0594-f67b-4482-948e-706bd3821b6a" ID-PMC="PMC6711933" ID-Pensoft-Pub="1314-2003-128-97" ID-Pensoft-UUID="CFE1800CC09C53E78AFCB0D1E6864E45" ID-PubMed="31523155" ID-Zenodo-Dep="3378135" ModsDocID="1314-2003-128-97" checkinTime="1566303206021" checkinUser="pensoft" docAuthor="Nobis, Marcin, Gudkova, Polina D. & Pendry, Colin A." docDate="2019" docId="72673E9169B852D2BBED1C13FC8CB9CD" docLanguage="en" docName="PhytoKeys 128: 97-119" docOrigin="PhytoKeys 128" docSource="http://dx.doi.org/10.3897/phytokeys.128.34637" docTitle="Trikeraia hookeri Bor, Kew Bull. 9 (4): 555 - 556 1954" docType="treatment" docVersion="8" id="CFE1800CC09C53E78AFCB0D1E6864E45" lastPageNumber="114" masterDocId="CFE1800CC09C53E78AFCB0D1E6864E45" masterDocTitle="Synopsis of the tribe Stipeae (Poaceae) in Nepal" masterLastPageNumber="119" masterPageNumber="97" pageNumber="114" updateTime="1732929356087" updateUser="ExternalLinkService">
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<mods:title id="185172AF707FBF9E7C867D9BCD6FF986">Synopsis of the tribe Stipeae (Poaceae) in Nepal</mods:title>
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<mods:namePart id="8BDE0517B2DE64C24CA30556C91C38D1">Nobis, Marcin</mods:namePart>
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<mods:namePart id="16C75C9883A4C4E6DB9099E5D1E19E1F">Gudkova, Polina D.</mods:namePart>
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<mods:namePart id="D46C67887A2EAF90253BFD820DA8FCC0">Pendry, Colin A.</mods:namePart>
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<mods:typeOfResource id="7297D3782DB2758A880775C3C7469EEC">text</mods:typeOfResource>
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<mods:title id="EDC12CAB0C47BF0B137959C6C367CAA6">PhytoKeys</mods:title>
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<mods:date id="4BAF0D4D9BD85FBDF806238C6A135333">2019</mods:date>
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<mods:detail id="7E04E7C9BF8E6F82E79392CE7180D949" type="volume">
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<mods:number id="609921B32F7DE7BC2FEF3C4D782212D7">128</mods:number>
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<mods:url id="B3A0FD056E753B3603D3F659BB0A55B0">http://dx.doi.org/10.3897/phytokeys.128.34637</mods:url>
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<mods:classification id="CDBD6A30D56614FFBC87FE22C3241251">journal article</mods:classification>
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<mods:identifier id="22E9B0E17432868B57A20E4611DDBFD0" type="DOI">http://dx.doi.org/10.3897/phytokeys.128.34637</mods:identifier>
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<treatment id="72673E9169B852D2BBED1C13FC8CB9CD" ID-GBIF-Taxon="159168977" LSID="urn:lsid:plazi:treatment:72673E9169B852D2BBED1C13FC8CB9CD" httpUri="http://treatment.plazi.org/id/72673E9169B852D2BBED1C13FC8CB9CD" lastPageNumber="114" pageId="17" pageNumber="114" scope_family="Poaceae">
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<subSubSection id="EA64E18B7BB961E3EECBA0C9777EBF4E" pageId="17" pageNumber="114" type="nomenclature">
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<paragraph id="A54D8EC8608894DAD0708AC38480DFAB" pageId="17" pageNumber="114">
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<taxonomicName id="AC032DA98678120249723BE7D9EA59E3" ID-CoL="58TTH" LSID="72673E91-69B8-52D2-BBED-1C13FC8CB9CD" authority="(Stapf) Bor, Kew Bull. 9 (4): 555 - 556 (1954)." authorityName="Bor, Kew Bull. 9 (4): 555 - 556" authorityYear="1954" baseAuthorityName="Stapf" class="Liliopsida" family="Poaceae" genus="Trikeraia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trikeraia hookeri" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="species" species="hookeri">
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<pageBreakToken id="0726F91E1F42757B4AB81267907C2888" pageId="17" pageNumber="114" start="start">Trikeraia</pageBreakToken>
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hookeri (Stapf) Bor, Kew Bull. 9(4): 555-556 (1954).
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C6B892C486C2366703441A85FF167A35" pageId="17" pageNumber="114" type="reference_group">
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<paragraph id="C803557AFF2022149CD94FC67F959467" pageId="17" pageNumber="114">
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<taxonomicName id="46E32D018EC8E817F58760030114A5C5" class="Liliopsida" family="Poaceae" genus="Stipa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Stipa hookeri" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="species" species="hookeri">Stipa hookeri</taxonomicName>
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Stapf, J. Linn. Soc., Bot. 30: 120 (1894) (Basionym).
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</paragraph>
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<paragraph id="30C88E25507EC34F73F4A326B9299402" pageId="17" pageNumber="114">
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≡
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<taxonomicName id="123AC25C301F4235124C56E4B95FB104" class="Liliopsida" family="Poaceae" genus="Achnatherum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Achnatherum hookeri" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="species" species="hookeri">Achnatherum hookeri</taxonomicName>
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(Stapf) Keng, Claves Gen. Sp. Gram. Prim. Sinic. 106, 213 (1957).
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</paragraph>
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<paragraph id="874763B33E3C82E5BF5E361060789FBC" pageId="17" pageNumber="114">
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=
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<taxonomicName id="728F302C613466F8C5A36C391E00A8F1" authorityName="Hitchc" authorityYear="1933" class="Liliopsida" family="Poaceae" genus="Timouria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Timouria aurita" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="species" species="aurita">Timouria aurita</taxonomicName>
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Hitchc., J. Wash. Acad. Sci. 23: 134 (1933).
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</paragraph>
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</subSubSection>
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<subSubSection id="1BBBEA1699DDE84382F54EC6DD27BF07" pageId="17" pageNumber="114" type="materials_examined">
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<paragraph id="64A982831AEB5B0AF3B0089C3C8B6A35" pageId="17" pageNumber="114">Type.</paragraph>
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<paragraph id="F9CD6F6E984641A6DDDFC83ED2A7D1F4" pageId="17" pageNumber="114">Tibet, 4500 m, sheltered nullahs near water, Jul-Sep. 1891, Thorold 124 (holotype K!, isotype C).</paragraph>
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</subSubSection>
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<subSubSection id="ABADA6FB560FE481621992B00D2F6F99" pageId="17" pageNumber="114" type="distribution">
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<paragraph id="461943EF4969E13410192EB6FDCE3923" pageId="17" pageNumber="114">General distribution.</paragraph>
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<paragraph id="07A890A286B1B58364234796532D1DE1" pageId="17" pageNumber="114">
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China (Tibet), India (Sikkim, Ladakh), Pakistan (
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<bibRefCitation id="0139C85DE568748AE936A426A673F88E" author="Freitag, H" journalOrPublisher="Notes from the Royal Botanic Garden Edinburgh" pageId="18" pageNumber="115" pagination="355 - 489" refId="B8" refString="Freitag, H, 1985. The genus Stipa (Gramineae) in southwest and south Asia. Notes from the Royal Botanic Garden Edinburgh 42: 355 - 489" title="The genus Stipa (Gramineae) in southwest and south Asia." volume="42" year="1985">Freitag 1985</bibRefCitation>
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;
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<bibRefCitation id="BE51A7FD17280837547A062187401922" author="Wu, ZL" editor="Wu, ZY" journalOrPublisher="22. Science Press Beijing and Missouri Botanical Garden Press St. Louis" pageId="20" pageNumber="117" pagination="188 - 212" refId="B43" refString="Wu, ZL, Phillips, SM, 2006. Tribe Stipeae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China (Poaceae), vol. 22. Science Press Beijing and Missouri Botanical Garden Press St. Louis: 188 - 212" title="Tribe Stipeae." volumeTitle="Flora of China (Poaceae), vol." year="2006">Wu and Phillips 2006</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="186F6EE1F260C37B43A811F0277FCEE4" pageId="17" pageNumber="114" type="distribution">
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<paragraph id="80A441875CAE73C5056DB355A77C2972" pageId="17" pageNumber="114">Distribution in Nepal.</paragraph>
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<paragraph id="5C26F758141692BDF7F3CFB2CA277DD0" pageId="17" pageNumber="114">
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Although we did not find any specimens of
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<taxonomicName id="463AA0E5619CC77A48F7CF8F124FFD6F" authorityName="Bor, Kew Bull. 9 (4): 555 - 556" authorityYear="1954" baseAuthorityName="Stapf" class="Liliopsida" family="Poaceae" genus="Trikeraia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trikeraia hookeri" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="species" species="hookeri">
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<emphasis id="49993D80916AB3519714BAD7487A1AAC" italics="true" pageId="17" pageNumber="114">Trikeraia hookeri</emphasis>
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</taxonomicName>
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during this study we include it here because it is known from nearby regions (
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<bibRefCitation id="FF3C44E2BE09EB9BA87B121045B40305" author="Freitag, H" journalOrPublisher="Notes from the Royal Botanic Garden Edinburgh" pageId="18" pageNumber="115" pagination="341 - 408" refId="B7" refString="Freitag, H, 1975. The genus Piptatherum (Gramineae) in southwest Asia. Notes from the Royal Botanic Garden Edinburgh 33: 341 - 408" title="The genus Piptatherum (Gramineae) in southwest Asia." volume="33" year="1975">Freitag 1975</bibRefCitation>
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,
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<bibRefCitation id="C0C7B69730427BD361CA9B3392E5CA03" author="Wu, ZL" editor="Wu, ZY" journalOrPublisher="22. Science Press Beijing and Missouri Botanical Garden Press St. Louis" pageId="20" pageNumber="117" pagination="188 - 212" refId="B43" refString="Wu, ZL, Phillips, SM, 2006. Tribe Stipeae. In: Wu, ZY, Raven, PH, Hong, DY, Eds., Flora of China (Poaceae), vol. 22. Science Press Beijing and Missouri Botanical Garden Press St. Louis: 188 - 212" title="Tribe Stipeae." volumeTitle="Flora of China (Poaceae), vol." year="2006">Wu and Phillips 2006</bibRefCitation>
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), so it is very likely to be present in Nepal.
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<bibRefCitation id="4F5A68C6316CB6F76C7A902B08244B4E" author="Freitag, H" journalOrPublisher="Notes from the Royal Botanic Garden Edinburgh" pageId="18" pageNumber="115" pagination="355 - 489" refId="B8" refString="Freitag, H, 1985. The genus Stipa (Gramineae) in southwest and south Asia. Notes from the Royal Botanic Garden Edinburgh 42: 355 - 489" title="The genus Stipa (Gramineae) in southwest and south Asia." volume="42" year="1985">Freitag (1985)</bibRefCitation>
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reported this species from Nepal based on
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<emphasis id="EE82F96019FC087A74DD1FD2BCF5D646" italics="true" pageId="17" pageNumber="114">Sufed 104</emphasis>
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(K), Mt. Everest, Tinkye palin, 4270 m, but this specimen appears to have been collected on the Tibetan side of Mt Everest.
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</paragraph>
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</subSubSection>
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<subSubSection id="CBCC550E9F3BC4432FF7354CB10480CD" pageId="17" pageNumber="114" type="habitat">
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<paragraph id="8886328D25A7B20933526772F0FF96B4" pageId="17" pageNumber="114">Habitat.</paragraph>
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<paragraph id="B37C12F8CF80071B33FC660BC7BB08B8" pageId="17" pageNumber="114">Scrublands, alpine mats.</paragraph>
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</subSubSection>
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<subSubSection id="4E69E3AC9D993A72A7EAF86B4A6DE314" pageId="17" pageNumber="114" type="altitude range">
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<paragraph id="93CB9E8AD3DEF939323EA579DE0711F2" pageId="17" pageNumber="114">Altitude range.</paragraph>
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<paragraph id="33654DA6D63C6424182369444AFF8030" pageId="17" pageNumber="114">4000-4300 m.</paragraph>
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</subSubSection>
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<subSubSection id="249EA7C4DDC8A833394570A6940DEB44" pageId="17" pageNumber="114" type="notes">
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<paragraph id="FF721328395FF7DBAC0DAF960BA29D41" pageId="17" pageNumber="114">Note.</paragraph>
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<paragraph id="85C43C8E4824E3942B2E4B2C132E9C38" pageId="17" pageNumber="114">
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The 2005 DNEP3 expedition to Solukhumbu collected several specimens which were identified as
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<taxonomicName id="DAAB1471996C6C5EABFE45C3147FB1A1" authorityName="Cope" authorityYear="1987" class="Liliopsida" family="Poaceae" genus="Trikeraia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trikeraia oreophila" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="species" species="oreophila">
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<emphasis id="A82F985FFF77134EF7313C7D8674C481" italics="true" pageId="17" pageNumber="114">Trikeraia oreophila</emphasis>
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</taxonomicName>
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Cope by H. Noltie [Dingboche, on trail south to the Lobuche Khola bridge, east facing valley side,
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<geoCoordinate id="A3C4B9278C45DCF0A46B51975626F069" degrees="27" direction="north" minutes="52" orientation="latitude" precision="15" seconds="50" value="27.880556">27°52'50"N</geoCoordinate>
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,
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<geoCoordinate id="2EE92AEBEE896CFE986F6B4590093036" degrees="86" direction="east" minutes="49" orientation="longitude" precision="15" seconds="7" value="86.81861">86°49'7"E</geoCoordinate>
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, 4230 m, 23 Sep. 2005,
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<emphasis id="6221F66E244A9050FEEEA4FA3198D3F2" italics="true" pageId="17" pageNumber="114">M.F. Watson et al. DNEP3 AX107</emphasis>
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(E, KATH)]. These specimens are characterized by having 4-6 mm long lemma lobes, the awn arising below the middle of the lemma and ovary with two stigmas. Recently,
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<taxonomicName id="13AD75E111F677D421ABC7D2D0812C62" authorityName="Cope" authorityYear="1987" class="Liliopsida" family="Poaceae" genus="Trikeraia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trikeraia oreophila" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="species" species="oreophila">
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<emphasis id="CE4E3784A0B3892BA10C05E7D3432A5A" italics="true" pageId="17" pageNumber="114">Trikeraia oreophila</emphasis>
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</taxonomicName>
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was found to be conspecific with
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<taxonomicName id="87D6E4AD8E805B11D84815E1F5451F80" authorityName="Keng" authorityYear="1958" class="Liliopsida" family="Poaceae" genus="Sinochasea" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Sinochasea trigyna" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="species" species="trigyna">
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<emphasis id="25CD5ECB357AEC3682A68A9C489DEDC7" italics="true" pageId="17" pageNumber="114">Sinochasea trigyna</emphasis>
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</taxonomicName>
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Keng (
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<bibRefCitation id="967AF8B53AB5AC4082C5E7E49EA76319" author="WCSP" journalOrPublisher="Academiya Nauk SSSR, Leningrad" pageId="20" pageNumber="117" publicationUrl="https://wcsp.science.kew.org/home.do" refId="B42" refString="WCSP, 2019. World Checklists of Selected Plant Families. https://wcsp.science.kew.org/home.do" title="World Checklists of Selected Plant Families." url="https://wcsp.science.kew.org/home.do" year="2019">WCSP 2019</bibRefCitation>
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). Morphological and molecular studies have shown that the genus
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<taxonomicName id="D3ACC163E31EABB1207EC0574F4A38EE" authorityName="Y.L.Keng" authorityYear="1958" class="Liliopsida" family="Poaceae" genus="Sinochasea" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Sinochasea" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="genus">
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<emphasis id="8DA40820C4F9BE266E242F1378EE7635" italics="true" pageId="17" pageNumber="114">Sinochasea</emphasis>
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</taxonomicName>
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is distinct not only from
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<taxonomicName id="F45FC3EB752888A183CC981C0A60E804" class="Liliopsida" family="Poaceae" genus="Trikeraia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trikeraia" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="genus">
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<emphasis id="A57B6721C272DD01F96B23317F761A68" italics="true" pageId="17" pageNumber="114">Trikeraia</emphasis>
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</taxonomicName>
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, but also from all the other genera of the tribe
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<taxonomicName id="7596DA027ED58C4B9A72E8E0A6CD7738" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Poales" pageId="17" pageNumber="114" phylum="Tracheophyta" rank="tribe" tribe="Stipeae">Stipeae</taxonomicName>
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, and therefore it was transferred to the tribe
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<taxonomicName id="4EE675470F973C6BF89B7FF5E6E33466" authorityName="Renvoize & Clayton" authorityYear="1985" genus="Phaenospermateae" lsidName="Phaenospermateae" pageId="17" pageNumber="114" rank="genus">
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<emphasis id="FAF911DC3125467F94553381A40C444F" italics="true" pageId="17" pageNumber="114">Phaenospermateae</emphasis>
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</taxonomicName>
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Renvoize & Clayton (
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<bibRefCitation id="44396A8E1ABFA53E2E704E6ECB50EDA4" DOI="https://doi.org/10.1080/14772000.2010.544339" author="Schneider, J" journalOrPublisher="Systematics and Biodiversity" pageId="20" pageNumber="117" pagination="27 - 44" refId="B33" refString="Schneider, J, Winterfeld, G, Hoffmann, MH, Roser, MR, 2011. Duthieeae, a new tribe of grasses (Poaceae) identified among the early diverging lineages of subfamily Pooideae: Molecular phylogenetics, morphological delineation, cytogenetics and biogeography. Systematics and Biodiversity 9 (1): 27 - 44, DOI: https://doi.org/10.1080/14772000.2010.544339" title="Duthieeae, a new tribe of grasses (Poaceae) identified among the early diverging lineages of subfamily Pooideae: Molecular phylogenetics, morphological delineation, cytogenetics and biogeography." url="https://doi.org/10.1080/14772000.2010.544339" volume="9" year="2011">Schneider et al. 2011</bibRefCitation>
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;
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<bibRefCitation id="B52469EFD8FE8B94520DD8C7D83B69D8" DOI="https://doi.org/10.1002/tax.611002" author="Romaschenko, K" journalOrPublisher="Taxon" pageId="20" pageNumber="117" pagination="18 - 44" refId="B30" refString="Romaschenko, K, Peterson, PM, Soreng, RJ, Garcia-Jacas, N, Futorna, O, Susanna, A, 2012. Systematics and evolution of the needle grasses (Poaceae: Pooideae: Stipeae) based on analysis of multiple chloroplast loci, ITS, and lemma micromorphology. Taxon 61 (1): 18 - 44, DOI: https://doi.org/10.1002/tax.611002" title="Systematics and evolution of the needle grasses (Poaceae: Pooideae: Stipeae) based on analysis of multiple chloroplast loci, ITS, and lemma micromorphology." url="https://doi.org/10.1002/tax.611002" volume="61" year="2012">Romaschenko et al. 2012</bibRefCitation>
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;
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<bibRefCitation id="64599D52302A4F27602D4A7E8A1238FC" author="Kellogg, EA" editor="Kubitski, K" journalOrPublisher="13. Springer International, Cham" pageId="18" pageNumber="115" pagination="1 - 416" refId="B15" refString="Kellogg, EA, 2015. Flowering plants. Monocots: Poaceae. In: Kubitski, K, Ed., The Families and Genera of Vascular Plants, vol. 13. Springer International, Cham: 1 - 416" title="Flowering plants. Monocots: Poaceae." volumeTitle="The Families and Genera of Vascular Plants, vol." year="2015">Kellogg 2015</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |