338 lines
36 KiB
XML
338 lines
36 KiB
XML
<document id="33678CC3E3AC888E8C246DFA394403D7" ID-CLB-Dataset="46225" ID-DOI="10.5281/zenodo.202819" ID-GBIF-Dataset="172f8751-669a-4ec6-9832-f824818c3887" ID-ISSN="1175-5326" ID-Zenodo-Dep="202819" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460397122923" checkinUser="plazi" docAuthor="Clark, Roger N. & Jewett, Stephen C." docDate="2011" docId="724AC7727328377C86DBFBC788A8F94D" docLanguage="en" docName="zt03051p013.pdf" docOrigin="Zootaxa 3051" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pteraster willsi Clark & Jewett, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="11" masterDocId="8E73BF0A73203776864CFFA98B62FF93" masterDocTitle="Three new sea stars (Asteroidea: Solasteridae & Pterasteridae) from the Aleutian Islands" masterLastPageNumber="13" masterPageNumber="1" pageNumber="9" updateTime="1698298806485" updateUser="ExternalLinkService">
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<mods:mods id="B96ECEC58DF8936E7B1BCCB9159981B3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="36CA513DBA2A1829BF5E32F3919FE340">
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<mods:title id="45C0EC61523EB23C37DCF7AE1B3F870D">Three new sea stars (Asteroidea: Solasteridae & Pterasteridae) from the Aleutian Islands</mods:title>
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</mods:titleInfo>
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<mods:name id="E1A6939F2DC967FF5CD7682C9D528083" type="personal">
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<mods:namePart id="D8DB8E320173D5A4209E9C527528D64D">Clark, Roger N.</mods:namePart>
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<mods:name id="3214FA2198A1DCC98C1E864BE36BA1D3" type="personal">
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<mods:roleTerm id="244C024118C9A6930E939D70A6DC15B4">Author</mods:roleTerm>
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<mods:namePart id="5F65601ECD6E2EBAE2D5DE14C559FDFB">Jewett, Stephen C.</mods:namePart>
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<mods:typeOfResource id="30D015A6E19FE713E11C2F1F2DF45EA6">text</mods:typeOfResource>
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<mods:title id="CB2AE02324DACE9006513CD47DC25906">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="72E9ABBFD16CF014C254EDA621E21724">
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<mods:date id="754AF894B9A082BB5E3D84C42A4BA755">2011</mods:date>
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<mods:detail id="9C587CA9F814BCA4D181423BA9D48062" type="volume">
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<mods:number id="5A2491C6E806C653BDB301D7C8423254">3051</mods:number>
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<mods:start id="33B620ECCFE96E57A258DED489D355C7">1</mods:start>
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<mods:end id="2F679FEFB0500CED13870C6E19E2BC90">13</mods:end>
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<mods:classification id="17D2ED66FAD383F7E36492340D300986">journal article</mods:classification>
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<mods:identifier id="08B4C7440E5E0CA874A75FF398475F55" type="CLB-Dataset">46225</mods:identifier>
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<mods:identifier id="06119B2FFBE31F1C3BD09C84D37CC722" type="DOI">10.5281/zenodo.202819</mods:identifier>
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<mods:identifier id="1393608813A127CEFB95E24986E660B1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="869EC8E47F9221E00FA23EFD398ECCEC" type="Zenodo-Dep">202819</mods:identifier>
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<treatment id="724AC7727328377C86DBFBC788A8F94D" ID-DOI="http://doi.org/10.5281/zenodo.5611037" ID-GBIF-Taxon="119418362" ID-Zenodo-Dep="5611037" LSID="urn:lsid:plazi:treatment:724AC7727328377C86DBFBC788A8F94D" httpUri="http://treatment.plazi.org/id/724AC7727328377C86DBFBC788A8F94D" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
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<subSubSection id="B2F925EF7328377E86DBFBC78A36FB3A" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="FA5C76647328377E86DBFBC78AD2FB1B" blockId="8.[151,432,1134,1193]" box="[151,432,1134,1160]" pageId="8" pageNumber="9">
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<heading id="A114C1087328377E86DBFBC78AD2FB1B" bold="true" box="[151,432,1134,1160]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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<emphasis id="C897AA767328377E86DBFBC78AD2FB1B" bold="true" box="[151,432,1134,1160]" pageId="8" pageNumber="9">
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<taxonomicName id="3DE30DE77328377E86DBFBC78A28FB1B" ID-CoL="4PTKY" box="[151,330,1134,1160]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="willsi" status="sp. nov.">
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<emphasis id="C897AA767328377E86DBFBC78A28FB1B" bold="true" box="[151,330,1134,1160]" italics="true" pageId="8" pageNumber="9">Pteraster willsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D3A4170D7328377E871EFBC78AD2FB1B" box="[338,432,1134,1160]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="FA5C76647328377E86DBFB388A36FB3A" blockId="8.[151,432,1134,1193]" box="[151,340,1169,1193]" pageId="8" pageNumber="9">Figures 7, 14–20</paragraph>
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</subSubSection>
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<subSubSection id="B2F925EF7328377E86DBFB738AB3FA86" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="FA5C76647328377E86DBFB738E12FB62" blockId="8.[151,1436,1241,2020]" box="[151,1392,1241,1266]" pageId="8" pageNumber="9">
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<emphasis id="C897AA767328377E86DBFB738A57FB61" bold="true" box="[151,309,1241,1266]" pageId="8" pageNumber="9">
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<typeStatus id="2558C8C67328377E86DBFB738BB0FB61" box="[151,210,1242,1266]" pageId="8" pageNumber="9">Type</typeStatus>
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locality.
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</emphasis>
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Alaska, Aleutian Islands, Rat Islands, West of Kiska Island (52°
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<geoCoordinate id="9FD710A37328377E8248FB708F11FB62" box="[1028,1139,1241,1265]" direction="north" orientation="latitude" pageId="8" pageNumber="9" precision="55" value="2.066">02.066 N</geoCoordinate>
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, 176°
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<geoCoordinate id="9FD710A37328377E82FCFB708E6CFB62" box="[1200,1294,1241,1265]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="555" value="27.04">27.04 E</geoCoordinate>
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),
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<specimenCount id="ECE5BDED7328377E836EFB708E12FB62" box="[1314,1392,1241,1265]" pageId="8" pageNumber="9" type="generic">
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<quantity id="3D1BDB817328377E836EFB708E0EFB62" box="[1314,1388,1241,1265]" metricMagnitude="2" metricUnit="m" metricValue="1.6199999999999999" pageId="8" pageNumber="9" unit="m" value="162.0">162 m</quantity>
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.
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</specimenCount>
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</paragraph>
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<paragraph id="FA5C76647328377E8689FB548AB3FA86" blockId="8.[151,1436,1241,2020]" box="[197,465,1277,1302]" pageId="8" pageNumber="9">
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<emphasis id="C897AA767328377E8689FB548A11FA85" bold="true" box="[197,371,1277,1302]" pageId="8" pageNumber="9">
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<typeStatus id="2558C8C67328377E8689FB548A62FA86" box="[197,256,1277,1301]" pageId="8" pageNumber="9">Type</typeStatus>
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material.
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</emphasis>
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<tableCitation id="B76143DF7328377E8735FB548AA8FA86" box="[377,458,1277,1301]" captionStart="TABLE 3" captionStartId="10.[151,235,152,173]" captionTargetBox="[151,1381,201,1383]" captionTargetPageId="10" captionText="TABLE 3. Type material: Pteraster willsi sp. nov." httpUri="http://table.plazi.org/id/AE9C26EC732A377C86DBFF3189F4FF3E" pageId="8" pageNumber="9" tableUuid="AE9C26EC732A377C86DBFF3189F4FF3E">Table 3</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2F925EF7328377E86DBFA888858F97A" pageId="8" pageNumber="9" type="description">
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<paragraph id="FA5C76647328377E86DBFA8889F6FA7C" blockId="8.[151,1436,1241,2020]" pageId="8" pageNumber="9">
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<emphasis id="C897AA767328377E86DBFA888A78FAAB" bold="true" box="[151,282,1313,1336]" pageId="8" pageNumber="9">Description.</emphasis>
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Small, stellate form,
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<typeStatus id="2558C8C67328377E87B0FA888938FAA4" box="[508,602,1313,1335]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="62D86AE17328377E8426FA8889D5FAAB" box="[618,695,1313,1336]" captionStart="FIGURE 14" captionStartId="9.[151,250,1762,1785]" captionTargetBox="[257,1274,193,1716]" captionTargetId="figure@9.[257,1274,193,1718]" captionTargetPageId="9" captionText="FIGURE 14. Pteraster willsi sp. nov. Holotype (LACM 2002 - 101.001). Whole animal, aboral view; bar = 1.0 cm. FIGURES 15 – 18. Pteraster willsi sp. nov. Paratypes (LACM 1997 - 222.001). Figure 15, Close-up, aboral surface; bar = 2.0 mm. Figure 16, Close-up, nidamental chamber, pseudopaxillae / spines and supradorsal membrane; Bar = 1.5 mm. Figure 17, Close-up, adambulacral area (ad = adambulacral spines; al = actinolateral spine); Bar = 1.0 mm. Figure 18, Close-up, oral region; bar = 3.0 mm." httpUri="https://zenodo.org/record/202822/files/figure.png" pageId="8" pageNumber="9">Fig. 14</figureCitation>
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); disc broad, rays relatively long, tapering, disc plump, inflated R =
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<quantity id="3D1BDB817328377E83CFFA8B8BA3FAC5" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="mm" value="30.0">30 mm</quantity>
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, r =
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<quantity id="3D1BDB817328377E86B9FAE98A5CFAC5" box="[245,318,1344,1366]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="9" unit="mm" value="22.0">22 mm</quantity>
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R:r = 1.36, (range 1:1.3–1:1.9). Supradorsal membrane (Fig. 15), relatively thin, firm, papillose. Abactinal plates 4–5 lobed; paxillar columns (Fig. 16), short (height less than half the length of paxillar spines), relatively thick, crowned with 5–7 long, slender spines, the tips of which protrude through the supradorsal membrane, and are covered by a fleshy papilla. Madreporite small, spherical, located beneath the supradorsal membrane, near the osculum. Osculum conspicuous, closed by five palmate valves, each formed by 4 large robust spines, along the facing edge of a large stout paxillae bearing about 16 spines. Spiraculae small, very numerous.
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</paragraph>
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<paragraph id="FA5C76647328377E8689FA5E8F46F933" blockId="8.[151,1436,1241,2020]" pageId="8" pageNumber="9">Adambulacral plates (Fig. 17), with L-shaped series of 5–7 webbed spines, distal 3–4 sub-equal, arranged transversely to groove, proximal 1–3 much smaller, set along groove, and a long actinolateral spine, flattened and sometimes bifurcated at the distal end; immersed in actinal membrane. Actinolateral fringe relatively wide, tapering. Each half of jaw plate (Fig. 18), bears 6–7 slender, webbed, grading spines, not continuously webbed to the adjacent plate, and a single large, triangular suboral spine. Tube feet in two rows.</paragraph>
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<paragraph id="FA5C76647328377E8689F9058858F97A" blockId="8.[151,1436,1241,2020]" pageId="8" pageNumber="9">
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Color of living specimens: from deep water uniformly white, sometimes with orange-tipped ocular spine tips, shallow water specimens (
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<figureCitation id="62D86AE17328377E87F2F9668973F974" box="[446,529,1743,1767]" captionStart="FIGURE 19" captionStartId="9.[151,250,1915,1938]" captionText="FIGURE 19. Pteraster willsi sp. nov. Paratype (LACM 2007 - 120.001). Live, in situ, Rat Island, 14 m. Image: Héloïse Chenelot, 1 July, 2007. R = approximately 1.7 cm." pageId="8" pageNumber="9">Fig. 19</figureCitation>
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), uniformly bright orange.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2F925EF7328377E8689F95A8A7EF8E4" pageId="8" pageNumber="9" type="distribution">
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<paragraph id="FA5C76647328377E8689F95A8A07F8BC" blockId="8.[151,1436,1241,2020]" pageId="8" pageNumber="9">
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<emphasis id="C897AA767328377E8689F95A8A3CF89F" bold="true" box="[197,350,1779,1804]" pageId="8" pageNumber="9">Distribution.</emphasis>
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Known from the central and western Aleutians (Fig. 7), Kanaga Island (
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<geoCoordinate id="9FD710A37328377E82C6F95D8E6AF89F" box="[1162,1288,1780,1804]" direction="west" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-178.43333">178°26’ W</geoCoordinate>
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) to near Attu Island (172°57 E).
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</paragraph>
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<paragraph id="FA5C76647328377E8689F8928A7EF8E4" blockId="8.[151,1436,1241,2020]" pageId="8" pageNumber="9">
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<emphasis id="C897AA767328377E8689F8928A4AF8C7" bold="true" box="[197,296,1851,1876]" pageId="8" pageNumber="9">Habitat.</emphasis>
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Rocky and pebbly bottoms, rich in sponges and hydrocorals,
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<quantity id="3D1BDB817328377E8591F8958F2FF8C7" box="[989,1101,1852,1876]" metricMagnitude="1" metricUnit="m" metricValue="8.85" metricValueMax="16.6" metricValueMin="1.1" pageId="8" pageNumber="9" unit="m" value="88.5" valueMax="166.0" valueMin="11.0">11–166 m</quantity>
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with a bottom temperature of 3.7° to 6°C.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2F925EF7328377C868BF82D8800F9B9" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="etymology">
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<paragraph id="FA5C76647328377E868BF82D8E06F877" blockId="8.[151,1436,1241,2020]" pageId="8" pageNumber="9">
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<emphasis id="C897AA767328377E868BF82D8A2CF80F" bold="true" box="[199,334,1924,1948]" pageId="8" pageNumber="9">Etymology.</emphasis>
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It is with great pleasure that we name this species in honor of Dr. Irvin A. Wills (deceased) of John Brown University, Siloam Springs, Arkansas. For nearly four decades Dr. Wills served JBU as Professor and Head of the Department of Biology where he mentored thousands of biology students, including Stephen C. Jewett.
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</paragraph>
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<caption id="AE9C26EC7329377F86DBF94B8A17F8E0" httpUri="https://zenodo.org/record/202822/files/figure.png" pageId="9" pageNumber="10" targetBox="[257,1274,193,1716]" targetPageId="9">
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<paragraph id="FA5C76647329377F86DBF94B8A17F8E0" blockId="9.[151,1437,1762,2030]" pageId="9" pageNumber="10">
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<emphasis id="C897AA767329377F86DBF94B8A7DF96B" bold="true" box="[151,287,1762,1785]" pageId="9" pageNumber="10">FIGURE 14.</emphasis>
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<taxonomicName id="3DE30DE77329377F8769F94A8AA3F96B" box="[293,449,1762,1784]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="willsi" status="sp. nov.">
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<emphasis id="C897AA767329377F8769F94A8AA3F96B" box="[293,449,1762,1784]" italics="true" pageId="9" pageNumber="10">Pteraster willsi</emphasis>
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</taxonomicName>
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<emphasis id="C897AA767329377F878BF94A8975F96B" bold="true" box="[455,535,1763,1784]" pageId="9" pageNumber="10">
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<taxonomicNameLabel id="D3A4170D7329377F878BF94A8975F96B" box="[455,535,1763,1784]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Holotype (LACM 2002-101.001). Whole animal, aboral view; bar = 1.0 cm.
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<emphasis id="C897AA767329377F86DBF8A88A37F885" bold="true" box="[151,341,1793,1816]" pageId="9" pageNumber="10">FIGURES 15–18.</emphasis>
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<taxonomicName id="3DE30DE77329377F8710F8AB8A98F884" box="[348,506,1793,1815]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="willsi" status="sp. nov.">
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<emphasis id="C897AA767329377F8710F8AB8A98F884" box="[348,506,1793,1815]" italics="true" pageId="9" pageNumber="10">Pteraster willsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D3A4170D7329377F844EF8AB8931F884" box="[514,595,1794,1815]" pageId="9" pageNumber="10" rank="species">
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<emphasis id="C897AA767329377F844EF8AB8931F884" bold="true" box="[514,595,1794,1815]" pageId="9" pageNumber="10">sp. nov.</emphasis>
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</taxonomicNameLabel>
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Paratypes (LACM 1997-222.001). Figure 15, Close-up, aboral surface; bar = 2.0 mm. Figure 16, Close-up, nidamental chamber, pseudopaxillae/spines and supradorsal membrane; Bar = 1.5 mm. Figure 17, Close-up, adambulacral area (ad = adambulacral spines; al = actinolateral spine); Bar = 1.0 mm. Figure 18, Close-up, oral region; bar = 3.0 mm.
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</paragraph>
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</caption>
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<caption id="AE9C26EC7329377F86DBF8D289F5F823" pageId="9" pageNumber="10">
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<paragraph id="FA5C76647329377F86DBF8D289F5F823" blockId="9.[151,1437,1762,2030]" pageId="9" pageNumber="10">
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<emphasis id="C897AA767329377F86DBF8D28A46F802" bold="true" box="[151,292,1915,1938]" pageId="9" pageNumber="10">FIGURE 19.</emphasis>
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<taxonomicName id="3DE30DE77329377F8763F8D48AADF801" box="[303,463,1916,1938]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="willsi" status="sp. nov.">
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<emphasis id="C897AA767329377F8763F8D48AADF801" box="[303,463,1916,1938]" italics="true" pageId="9" pageNumber="10">Pteraster willsi</emphasis>
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</taxonomicName>
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<emphasis id="C897AA767329377F8796F8D5894DF802" bold="true" box="[474,559,1916,1937]" pageId="9" pageNumber="10">
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<taxonomicNameLabel id="D3A4170D7329377F8796F8D5894DF802" box="[474,559,1916,1937]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Paratype (LACM 2007-120.001). Live,
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<emphasis id="C897AA767329377F85AEF8D48F47F801" box="[994,1061,1917,1938]" italics="true" pageId="9" pageNumber="10">in situ</emphasis>
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, Rat Island, 14 m. Image: Héloïse Chenelot, 1 July, 2007. R = approximately 1.7 cm.
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</paragraph>
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</caption>
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<caption id="AE9C26EC7329377F86DBF81089C7F87D" pageId="9" pageNumber="10">
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<paragraph id="FA5C76647329377F86DBF81089C7F87D" blockId="9.[151,1437,1762,2030]" pageId="9" pageNumber="10">
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<emphasis id="C897AA767329377F86DBF8108A7DF85D" bold="true" box="[151,287,1977,2000]" pageId="9" pageNumber="10">FIGURE 20.</emphasis>
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<taxonomicName id="3DE30DE77329377F8769F8138AA2F85C" box="[293,448,1977,1999]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="willsi" status="sp. nov.">
|
||
<emphasis id="C897AA767329377F8769F8138AA2F85C" box="[293,448,1977,1999]" italics="true" pageId="9" pageNumber="10">Pteraster willsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C897AA767329377F878AF8138974F85C" bold="true" box="[454,534,1978,1999]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="D3A4170D7329377F878AF8138974F85C" box="[454,534,1978,1999]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Paratypes (LACM 2007-118.002). Live,
|
||
<emphasis id="C897AA767329377F85F4F8138897F85C" box="[952,1013,1978,1999]" italics="true" pageId="9" pageNumber="10">in situ</emphasis>
|
||
, Vega Bay, Kiska Island, 14 m. Image: R. N. Clark, 29 June, 2007. R = approximately 2.0 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AE9C26EC732A377C86DBFF3189F4FF3E" ID-Table-UUID="AE9C26EC732A377C86DBFF3189F4FF3E" box="[151,662,152,174]" httpUri="http://table.plazi.org/id/AE9C26EC732A377C86DBFF3189F4FF3E" pageId="10" pageNumber="11" targetBox="[151,1381,201,1383]" targetIsTable="true" targetPageId="10">
|
||
<paragraph id="FA5C7664732A377C86DBFF3189F4FF3E" blockId="10.[151,662,152,174]" box="[151,662,152,174]" pageId="10" pageNumber="11">
|
||
<emphasis id="C897AA76732A377C86DBFF318A66FF3E" bold="true" box="[151,260,152,173]" pageId="10" pageNumber="11">TABLE 3.</emphasis>
|
||
Type material:
|
||
<taxonomicName id="3DE30DE7732A377C87EFFF30895DFF3D" box="[419,575,152,174]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="willsi" status="sp. nov.">
|
||
<emphasis id="C897AA76732A377C87EFFF30895DFF3D" box="[419,575,152,174]" italics="true" pageId="10" pageNumber="11">Pteraster willsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D3A4170D732A377C8409FF3189F4FF3E" box="[581,662,152,173]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA5C7664732A377C86D3FF638E7EFADB" pageId="10" pageNumber="11">
|
||
<table id="88E384C4732AC88986DBFF608E07FAF4" box="[151,1381,201,1383]" gridcols="5" gridrows="12" pageId="10" pageNumber="11">
|
||
<tr id="44D37426732AC88986DBFF608E07FF6D" box="[151,1381,201,254]" gridrow="0" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFF608A60FF6D" box="[151,258,201,254]" gridcol="0" gridrow="0" pageId="10" pageNumber="11">Type</th>
|
||
<th id="07021D5A732AC8898706FF608822FF6D" box="[330,832,201,254]" gridcol="1" gridrow="0" pageId="10" pageNumber="11">Deposition & No. of Collection Site Specimens</th>
|
||
<th id="07021D5A732AC889853BFF608883FF6D" box="[887,993,201,254]" gridcol="2" gridrow="0" pageId="10" pageNumber="11">Collection Depth, m</th>
|
||
<th id="07021D5A732AC8898258FF608FD6FF6D" box="[1044,1204,201,254]" gridcol="3" gridrow="0" pageId="10" pageNumber="11">Collection Date</th>
|
||
<th id="07021D5A732AC88982ADFF608E07FF6D" box="[1249,1381,201,254]" gridcol="4" gridrow="0" pageId="10" pageNumber="11">Collector & Method</th>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFEB88E07FEF6" box="[151,1381,273,357]" gridrow="1" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFEB88A60FEF6" box="[151,258,273,357]" gridcol="0" gridrow="1" pageId="10" pageNumber="11">Holotype</th>
|
||
<td id="07021D5A732AC8898706FEB88822FEF6" box="[330,832,273,357]" gridcol="1" gridrow="1" pageId="10" pageNumber="11">LACM 2002- W of Kiska Island 101.001, dry 52°02.06 N, 176°27.97 E; NMFS 94-200201-77</td>
|
||
<td id="07021D5A732AC889853BFEB88883FEF6" box="[887,993,273,357]" gridcol="2" gridrow="1" pageId="10" pageNumber="11">162 m</td>
|
||
<td id="07021D5A732AC8898258FEB88FD6FEF6" box="[1044,1204,273,357]" gridcol="3" gridrow="1" pageId="10" pageNumber="11">16 June, 2002</td>
|
||
<td id="07021D5A732AC88982ADFEB88E07FEF6" box="[1249,1381,273,357]" gridcol="4" gridrow="1" pageId="10" pageNumber="11">
|
||
R. N. Clark, trawled, R/V
|
||
<emphasis id="C897AA76732A377C82AEFEE68E34FEF6" box="[1250,1366,335,357]" italics="true" pageId="10" pageNumber="11">Vesteraalen</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFED18E07FE58" box="[151,1381,376,459]" gridrow="2" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFED18A60FE58" box="[151,258,376,459]" gridcol="0" gridrow="2" pageId="10" pageNumber="11">Paratype</th>
|
||
<td id="07021D5A732AC8898706FED18822FE58" box="[330,832,376,459]" gridcol="1" gridrow="2" pageId="10" pageNumber="11">LACM 2002- W of Kiska Island 101.002, 52°02.06 N, 176°27.97 E; 1 specimen, dry NMFS 94-200201-77</td>
|
||
<td id="07021D5A732AC889853BFED18883FE58" box="[887,993,376,459]" gridcol="2" gridrow="2" pageId="10" pageNumber="11">162 m</td>
|
||
<td id="07021D5A732AC8898258FED18FD6FE58" box="[1044,1204,376,459]" gridcol="3" gridrow="2" pageId="10" pageNumber="11">16 June, 2002</td>
|
||
<td id="07021D5A732AC88982ADFED18E07FE58" box="[1249,1381,376,459]" gridcol="4" gridrow="2" pageId="10" pageNumber="11">
|
||
R. N. Clark, trawled, R/V
|
||
<emphasis id="C897AA76732A377C82AEFE1C8E34FE58" box="[1250,1366,437,459]" italics="true" pageId="10" pageNumber="11">Vesteraalen</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFE778E07FDA1" box="[151,1381,478,562]" gridrow="3" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFE778A60FDA1" box="[151,258,478,562]" gridcol="0" gridrow="3" pageId="10" pageNumber="11">Paratype</th>
|
||
<td id="07021D5A732AC8898706FE778822FDA1" box="[330,832,478,562]" gridcol="1" gridrow="3" pageId="10" pageNumber="11">USNM 1153701, N of Kiska Island 1 specimen, dry 52°04.29 N, 177°15.33 E; NMFS 23-199701-199</td>
|
||
<td id="07021D5A732AC889853BFE778883FDA1" box="[887,993,478,562]" gridcol="2" gridrow="3" pageId="10" pageNumber="11">130 m</td>
|
||
<td id="07021D5A732AC8898258FE778FD6FDA1" box="[1044,1204,478,562]" gridcol="3" gridrow="3" pageId="10" pageNumber="11">31 July, 1997</td>
|
||
<td id="07021D5A732AC88982ADFE778E07FDA1" box="[1249,1381,478,562]" gridcol="4" gridrow="3" pageId="10" pageNumber="11">
|
||
R. N. Clark, trawled, R/V
|
||
<emphasis id="C897AA76732A377C82AEFDB48E33FDA1" box="[1250,1361,541,562]" italics="true" pageId="10" pageNumber="11">Dominator</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFDEC8E07FD0B" box="[151,1381,581,664]" gridrow="4" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFDEC8A60FD0B" box="[151,258,581,664]" gridcol="0" gridrow="4" pageId="10" pageNumber="11">Paratype</th>
|
||
<td id="07021D5A732AC8898706FDEC8822FD0B" box="[330,832,581,664]" gridcol="1" gridrow="4" pageId="10" pageNumber="11">USNM 1153702, N of Agattu Island 1 specimen, dry 52°20.58 N, 173°31.64 E; NMFS 23-199701-231</td>
|
||
<td id="07021D5A732AC889853BFDEC8883FD0B" box="[887,993,581,664]" gridcol="2" gridrow="4" pageId="10" pageNumber="11">86 m</td>
|
||
<td id="07021D5A732AC8898258FDEC8FD6FD0B" box="[1044,1204,581,664]" gridcol="3" gridrow="4" pageId="10" pageNumber="11">6 August, 1997</td>
|
||
<td id="07021D5A732AC88982ADFDEC8E07FD0B" box="[1249,1381,581,664]" gridcol="4" gridrow="4" pageId="10" pageNumber="11">
|
||
R. N. Clark, trawled, R/V
|
||
<emphasis id="C897AA76732A377C82AEFD2A8E33FD0B" box="[1250,1361,643,664]" italics="true" pageId="10" pageNumber="11">Dominator</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFD028E07FD6C" box="[151,1381,683,767]" gridrow="5" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFD028A60FD6C" box="[151,258,683,767]" gridcol="0" gridrow="5" pageId="10" pageNumber="11">Paratypes</th>
|
||
<td id="07021D5A732AC8898706FD028822FD6C" box="[330,832,683,767]" gridcol="1" gridrow="5" pageId="10" pageNumber="11">CASIZ 184726, E of Kiska Island 2 specimens, dry 52°01.81 N, 177°43.57 E; NMFS 23-199701-196</td>
|
||
<td id="07021D5A732AC889853BFD028883FD6C" box="[887,993,683,767]" gridcol="2" gridrow="5" pageId="10" pageNumber="11">110 m</td>
|
||
<td id="07021D5A732AC8898258FD028FD6FD6C" box="[1044,1204,683,767]" gridcol="3" gridrow="5" pageId="10" pageNumber="11">31 July, 1997</td>
|
||
<td id="07021D5A732AC88982ADFD028E07FD6C" box="[1249,1381,683,767]" gridcol="4" gridrow="5" pageId="10" pageNumber="11">
|
||
R. N. Clark, trawled, R/V
|
||
<emphasis id="C897AA76732A377C82AEFD438E33FD6C" box="[1250,1361,746,767]" italics="true" pageId="10" pageNumber="11">Dominator</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFCBB8E07FCF5" box="[151,1381,786,870]" gridrow="6" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFCBB8A60FCF5" box="[151,258,786,870]" gridcol="0" gridrow="6" pageId="10" pageNumber="11">Paratypes</th>
|
||
<td id="07021D5A732AC8898706FCBB8822FCF5" box="[330,832,786,870]" gridcol="1" gridrow="6" pageId="10" pageNumber="11">CASIZ 184727, S of Attu Island 2 specimens, dry 52°29.30 N, 172°57.50 E; NMFS 23-199701-210</td>
|
||
<td id="07021D5A732AC889853BFCBB8883FCF5" box="[887,993,786,870]" gridcol="2" gridrow="6" pageId="10" pageNumber="11">166 m</td>
|
||
<td id="07021D5A732AC8898258FCBB8FD6FCF5" box="[1044,1204,786,870]" gridcol="3" gridrow="6" pageId="10" pageNumber="11">2 August, 1997</td>
|
||
<td id="07021D5A732AC88982ADFCBB8E07FCF5" box="[1249,1381,786,870]" gridcol="4" gridrow="6" pageId="10" pageNumber="11">
|
||
R. N. Clark, trawled, R/V
|
||
<emphasis id="C897AA76732A377C82AEFCF88E33FCF5" box="[1250,1361,849,870]" italics="true" pageId="10" pageNumber="11">Dominator</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFCD08E07FC5E" box="[151,1381,889,973]" gridrow="7" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFCD08A60FC5E" box="[151,258,889,973]" gridcol="0" gridrow="7" pageId="10" pageNumber="11">Paratypes</th>
|
||
<td id="07021D5A732AC8898706FCD08822FC5E" box="[330,832,889,973]" gridcol="1" gridrow="7" pageId="10" pageNumber="11">LACM 1997- SW of Buldir Island 222.001, 52°14.94 N, 175°14.90 E; 6 specimens, dry NMFS 23-199701-239</td>
|
||
<td id="07021D5A732AC889853BFCD08883FC5E" box="[887,993,889,973]" gridcol="2" gridrow="7" pageId="10" pageNumber="11">141 m</td>
|
||
<td id="07021D5A732AC8898258FCD08FD6FC5E" box="[1044,1204,889,973]" gridcol="3" gridrow="7" pageId="10" pageNumber="11">8 August, 1997</td>
|
||
<td id="07021D5A732AC88982ADFCD08E07FC5E" box="[1249,1381,889,973]" gridcol="4" gridrow="7" pageId="10" pageNumber="11">
|
||
R. N. Clark, trawled, R/V
|
||
<emphasis id="C897AA76732A377C82AEFC118E33FC5E" box="[1250,1361,952,973]" italics="true" pageId="10" pageNumber="11">Dominator</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFC498E07FBA0" box="[151,1381,992,1075]" gridrow="8" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFC498A60FBA0" box="[151,258,992,1075]" gridcol="0" gridrow="8" pageId="10" pageNumber="11">Paratypes</th>
|
||
<td id="07021D5A732AC8898706FC498822FBA0" box="[330,832,992,1075]" gridcol="1" gridrow="8" pageId="10" pageNumber="11">LACM 2007- Vega Bay, Kiska Island 118.002, 51°54.86 N, 177°26.86 E; 3 specimens, dry AKALE07-0017</td>
|
||
<td id="07021D5A732AC889853BFC498883FBA0" box="[887,993,992,1075]" gridcol="2" gridrow="8" pageId="10" pageNumber="11">14 m</td>
|
||
<td id="07021D5A732AC8898258FC498FD6FBA0" box="[1044,1204,992,1075]" gridcol="3" gridrow="8" pageId="10" pageNumber="11">29 June, 2007</td>
|
||
<td id="07021D5A732AC88982ADFC498E07FBA0" box="[1249,1381,992,1075]" gridcol="4" gridrow="8" pageId="10" pageNumber="11">R. N. Clark, scuba</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFBEF8E07FB09" box="[151,1381,1094,1178]" gridrow="9" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFBEF8A60FB09" box="[151,258,1094,1178]" gridcol="0" gridrow="9" pageId="10" pageNumber="11">Paratype</th>
|
||
<td id="07021D5A732AC8898706FBEF8822FB09" box="[330,832,1094,1178]" gridcol="1" gridrow="9" pageId="10" pageNumber="11">LACM 1997- S of Tanaga Island 179.003, 51°37.36 N, 178°26.46 W; 1 specimen NMFS 23-199701-135</td>
|
||
<td id="07021D5A732AC889853BFBEF8883FB09" box="[887,993,1094,1178]" gridcol="2" gridrow="9" pageId="10" pageNumber="11">163 m</td>
|
||
<td id="07021D5A732AC8898258FBEF8FD6FB09" box="[1044,1204,1094,1178]" gridcol="3" gridrow="9" pageId="10" pageNumber="11">14 July, 1997</td>
|
||
<td id="07021D5A732AC88982ADFBEF8E07FB09" box="[1249,1381,1094,1178]" gridcol="4" gridrow="9" pageId="10" pageNumber="11">
|
||
R. N. Clark, trawled, R/V
|
||
<emphasis id="C897AA76732A377C82AEFB2C8E33FB09" box="[1250,1361,1157,1178]" italics="true" pageId="10" pageNumber="11">Dominator</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFB048E07FA93" box="[151,1381,1197,1280]" gridrow="10" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFB048A60FA93" box="[151,258,1197,1280]" gridcol="0" gridrow="10" pageId="10" pageNumber="11">Paratype</th>
|
||
<td id="07021D5A732AC8898706FB048822FA93" box="[330,832,1197,1280]" gridcol="1" gridrow="10" pageId="10" pageNumber="11">LACM 2007- Rat Island 120.001, 51°49.59 N, 178°16.67 E; 1 specimen, dry AKALE07-A0031</td>
|
||
<td id="07021D5A732AC889853BFB048883FA93" box="[887,993,1197,1280]" gridcol="2" gridrow="10" pageId="10" pageNumber="11">14 m</td>
|
||
<td id="07021D5A732AC8898258FB048FD6FA93" box="[1044,1204,1197,1280]" gridcol="3" gridrow="10" pageId="10" pageNumber="11">1 July, 2007</td>
|
||
<td id="07021D5A732AC88982ADFB048E07FA93" box="[1249,1381,1197,1280]" gridcol="4" gridrow="10" pageId="10" pageNumber="11">H. Chenelot, scuba</td>
|
||
</tr>
|
||
<tr id="44D37426732AC88986DBFABA8E07FAF4" box="[151,1381,1299,1383]" gridrow="11" pageId="10" pageNumber="11">
|
||
<th id="07021D5A732AC88986DBFABA8A60FAF4" box="[151,258,1299,1383]" gridcol="0" gridrow="11" pageId="10" pageNumber="11">Paratype</th>
|
||
<td id="07021D5A732AC8898706FABA8822FAF4" box="[330,832,1299,1383]" gridcol="1" gridrow="11" pageId="10" pageNumber="11">UAF 8139, Little Sitkin Island 1 specimen, dry 51°58.20 N, 178°27.18 E; AKALE07-0045</td>
|
||
<td id="07021D5A732AC889853BFABA8883FAF4" box="[887,993,1299,1383]" gridcol="2" gridrow="11" pageId="10" pageNumber="11">11 m</td>
|
||
<td id="07021D5A732AC8898258FABA8FD6FAF4" box="[1044,1204,1299,1383]" gridcol="3" gridrow="11" pageId="10" pageNumber="11">1 July, 2007</td>
|
||
<td id="07021D5A732AC88982ADFABA8E07FAF4" box="[1249,1381,1299,1383]" gridcol="4" gridrow="11" pageId="10" pageNumber="11">R. N. Clark, scuba</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="FA5C7664732A377C868BFA0F8800F9B9" blockId="10.[151,1436,1446,1759]" pageId="10" pageNumber="11">
|
||
<emphasis id="C897AA76732A377C868BFA0F8A11FA2C" bold="true" box="[199,371,1446,1471]" pageId="10" pageNumber="11">Reproduction.</emphasis>
|
||
Like many
|
||
<taxonomicName id="3DE30DE7732A377C87B0FA018906FA2C" box="[508,612,1448,1471]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="C897AA76732A377C87B0FA018906FA2C" box="[508,612,1448,1471]" italics="true" pageId="10" pageNumber="11">Pteraster</emphasis>
|
||
</taxonomicName>
|
||
spp.,
|
||
<emphasis id="C897AA76732A377C84E6FA01899DFA2D" box="[682,767,1446,1471]" italics="true" pageId="10" pageNumber="11">P. w i l l s i</emphasis>
|
||
<emphasis id="C897AA76732A377C8547FA0E8804FA2C" bold="true" box="[779,870,1447,1471]" pageId="10" pageNumber="11">sp. nov.</emphasis>
|
||
broods its young in the nidamental chamber until they are developed enough to break through the supradorsal membrane, crawl away and fend for themselves (R =
|
||
<quantity id="3D1BDB81732A377C86DBFA478B96F995" box="[151,244,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="10" pageNumber="11" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
) (
|
||
<figureCitation id="62D86AE1732A377C8747FA478A39F995" box="[267,347,1518,1542]" captionStart="FIGURE 20" captionStartId="9.[151,250,1977,2000]" captionText="FIGURE 20. Pteraster willsi sp. nov. Paratypes (LACM 2007 - 118.002). Live, in situ, Vega Bay, Kiska Island, 14 m. Image: R. N. Clark, 29 June, 2007. R = approximately 2.0 cm." pageId="10" pageNumber="11">Fig. 20</figureCitation>
|
||
) (LACM 2007-118.002). This has been observed in many specimens, including several
|
||
<typeStatus id="2558C8C6732A377C8363FA468EFEF994" box="[1327,1436,1519,1543]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
|
||
(LACM 2007-118-002, LACM 1997-222.001, CASIZ 184727).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2F925EF732A377C868BF99E88A8F94D" pageId="10" pageNumber="11" type="discussion">
|
||
<paragraph id="FA5C7664732A377C868BF99E8E0AF905" blockId="10.[151,1436,1446,1759]" pageId="10" pageNumber="11">
|
||
<emphasis id="C897AA76732A377C868BF99E8A52F9DC" bold="true" box="[199,304,1591,1615]" pageId="10" pageNumber="11">Remark.</emphasis>
|
||
<taxonomicName id="3DE30DE7732A377C877BF9918A81F9DD" box="[311,483,1590,1615]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="willsi" status="sp. nov.">
|
||
<emphasis id="C897AA76732A377C877BF9918A81F9DD" box="[311,483,1590,1615]" italics="true" pageId="10" pageNumber="11">Pteraster willsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C897AA76732A377C87A7F99E8926F9DC" bold="true" box="[491,580,1591,1615]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="D3A4170D732A377C87A7F99E8926F9DC" box="[491,580,1591,1615]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="3DE30DE7732A377C849BF9918830F9DD" box="[727,850,1590,1615]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="militaris">
|
||
<emphasis id="C897AA76732A377C849BF9918830F9DD" box="[727,850,1590,1615]" italics="true" pageId="10" pageNumber="11">P. militaris</emphasis>
|
||
</taxonomicName>
|
||
, but differs in 1) smaller size and firmer, more rigid supradorsal membrane which lacks the convolutions, 2) large, trigonal suboral spines, and 3) fewer adambulacral spines, 3–4 compared to
|
||
<quantity id="3D1BDB81732A377C87E2F9D78A97F905" box="[430,501,1662,1686]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="10" pageNumber="11" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">6–9 in</quantity>
|
||
<taxonomicName id="3DE30DE7732A377C87B0F9298914F905" box="[508,630,1662,1687]" class="Asteroidea" family="Pterasteridae" genus="Pteraster" kingdom="Animalia" order="Velatida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="militaris">
|
||
<emphasis id="C897AA76732A377C87B0F9298914F905" box="[508,630,1662,1687]" italics="true" pageId="10" pageNumber="11">P. militaris</emphasis>
|
||
</taxonomicName>
|
||
. The two species are frequently found together in the same habitat.
|
||
</paragraph>
|
||
<paragraph id="FA5C7664732A377C868BF90A88A8F94D" blockId="10.[151,1436,1446,1759]" pageId="10" pageNumber="11">
|
||
A
|
||
<typeStatus id="2558C8C6732A377C86AAF90A8A2AF928" box="[230,328,1699,1723]" pageId="10" pageNumber="11" type="paratype">Paratype</typeStatus>
|
||
specimen from station AKALE07-0017 (LACM 2007-118.002) was apparently feeding on the (scavenged?) chela of the small lithode crab
|
||
<taxonomicName id="3DE30DE7732A377C84C7F96188A8F94D" authority="Brandt." authorityName="Brandt." box="[651,970,1734,1759]" class="Malacostraca" family="Lithodidae" genus="Dermaturus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mandtii">
|
||
<emphasis id="C897AA76732A377C84C7F961880DF94D" box="[651,879,1734,1759]" italics="true" pageId="10" pageNumber="11">Dermaturus mandtii</emphasis>
|
||
Brandt.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |