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<document id="54FF91FD6652CC0370471958DF070D1A" ID-CLB-Dataset="27316" ID-DOI="10.11646/zootaxa.4583.1.1" ID-GBIF-Dataset="027ed27d-5035-4646-93a6-c23b565f3d20" ID-ISSN="1175-5326" ID-Zenodo-Dep="2637603" ID-ZooBank="27F24995-359E-46F6-AB22-75568BACFDCF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555054242242" checkinUser="plazi" docAuthor="Zhang, - H." docDate="2019" docId="724987F6FFAA2433FF7EBB96FF1A38F8" docLanguage="en" docName="zootaxa.4583.1.1.pdf" docOrigin="Zootaxa 4583 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Edgbastonia Ponder, W.-H. Zhang, Hallan &amp; Shea 2019" docType="treatment" docVersion="16" lastPageNumber="47" masterDocId="8E70FF8EFF86241DFFE9BE79FFE73928" masterDocTitle="New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages" masterLastPageNumber="67" masterPageNumber="1" pageNumber="45" updateTime="1698720421338" updateUser="ExternalLinkService">
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<mods:title id="9E608377F01E9637D407AA807D1A50A8">New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages</mods:title>
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<mods:namePart id="F0BE851213BF4D944FA0D8C3832179E3">Zhang, - H.</mods:namePart>
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<mods:date id="7F5B4245679DDFACB7567759D7903FAC">2019</mods:date>
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<mods:number id="23AC08515B1C6178B48A580495E7A6EB">2019-04-10</mods:number>
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<mods:identifier id="D5BF710A848DF1693756BD58A73017BB" type="CLB-Dataset">27316</mods:identifier>
<mods:identifier id="C3FA3A55D83044C4AB426EB49D4C9C51" type="DOI">10.11646/zootaxa.4583.1.1</mods:identifier>
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<treatment id="724987F6FFAA2433FF7EBB96FF1A38F8" ID-DOI="http://doi.org/10.5281/zenodo.5582738" ID-GBIF-Taxon="156198415" ID-Zenodo-Dep="5582738" LSID="urn:lsid:plazi:treatment:724987F6FFAA2433FF7EBB96FF1A38F8" httpUri="http://treatment.plazi.org/id/724987F6FFAA2433FF7EBB96FF1A38F8" lastPageId="46" lastPageNumber="47" pageId="44" pageNumber="45">
<subSubSection id="B2FA656BFFAA2431FF7EBB96FDDE3F22" box="[151,569,1519,1546]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="FA5F36E0FFAA2431FF7EBB96FDDE3F22" blockId="44.[151,569,1519,1546]" box="[151,569,1519,1546]" pageId="44" pageNumber="45">
<heading id="A117818CFFAA2431FF7EBB96FDDE3F22" bold="true" box="[151,569,1519,1546]" fontSize="11" level="1" pageId="44" pageNumber="45" reason="1">
<emphasis id="C894EAF2FFAA2431FF7EBB96FDDE3F22" bold="true" box="[151,569,1519,1546]" pageId="44" pageNumber="45">
<taxonomicName id="3DE04D63FFAA2431FF7EBB96FE083F21" authority="Ponder, W.-H. Zhang, Hallan &amp; Shea, 2019" authorityName="Ponder, W.-H. Zhang, Hallan &amp; Shea" authorityYear="2019" box="[151,495,1519,1546]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="44" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis id="C894EAF2FFAA2431FF7EBB96FEC93F21" bold="true" box="[151,302,1519,1545]" italics="true" pageId="44" pageNumber="45">Edgbastonia</emphasis>
?
<emphasis id="C894EAF2FFAA2431FEADBB96FE083F21" bold="true" box="[324,495,1519,1545]" italics="true" pageId="44" pageNumber="45">conjuboyensis</emphasis>
</taxonomicName>
n. sp.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFFAA2431FF7EB84FFE6C3EBB" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="FA5F36E0FFAA2431FF7EB84FFBBA3FBF" blockId="44.[151,1437,1590,1939]" pageId="44" pageNumber="45">
<emphasis id="C894EAF2FFAA2431FF7EB84FFE663F67" bold="true" box="[151,385,1590,1615]" pageId="44" pageNumber="45">Material examined.</emphasis>
<materialsCitation id="4A883CBDFFAA2431FE60B84FFEEF3FBF" ID-GBIF-Occurrence="2235592650" collectingDate="2004-08-23" collectorName="W. F. Ponder &amp; J. B. Studdert &amp; C. Slatyer &amp; J. M. Ponder" county="Einasleigh" latitude="-18.667221" location="Basalt Spring" longLatPrecision="21" longitude="144.74333" municipality="Conjuboy Station" pageId="44" pageNumber="45" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="255B8842FFAA2431FE60B84FFE123F67" box="[393,501,1590,1615]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
:
<collectingRegion id="3824F802FFAA2431FDE9B84FFD603F67" box="[512,647,1590,1615]" country="Australia" name="Queensland" pageId="44" pageNumber="45">Queensland</collectingRegion>
,
<locationDeviation id="EBED2FC2FFAA2431FD7AB84EFC863F67" box="[659,865,1590,1615]" location="Georgetown" pageId="44" pageNumber="45">W of Georgetown</locationDeviation>
, S of
<collectingCounty id="133E4E6CFFAA2431FC4DB84FFBFA3F67" box="[932,1053,1590,1615]" pageId="44" pageNumber="45">Einasleigh</collectingCounty>
,
<collectingMunicipality id="1A3BAC9AFFAA2431FBC0B84FFB163F67" box="[1065,1265,1590,1615]" pageId="44" pageNumber="45">Conjuboy Station</collectingMunicipality>
,
<location id="FF3F603BFFAA2431FB14B84FFA703F67" LSID="urn:lsid:plazi:treatment:724987F6FFAA2433FF7EBB96FF1A38F8:FF3F603BFFAA2431FB14B84FFA703F67" box="[1277,1431,1590,1615]" county="Einasleigh" latitude="-18.667221" longLatPrecision="21" longitude="144.74333" municipality="Conjuboy Station" name="Basalt Spring" pageId="44" pageNumber="45" stateProvince="Queensland">Basalt Spring</location>
,
<geoCoordinate id="9FD45027FFAA2431FF7EB822FED03F5B" box="[151,311,1627,1651]" direction="south" orientation="latitude" pageId="44" pageNumber="45" precision="15" value="-18.667221">18° 40' 02&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FFAA2431FEA1B822FE1F3F5A" box="[328,504,1626,1651]" direction="east" orientation="longitude" pageId="44" pageNumber="45" precision="15" value="144.74333">144° 44' 36&quot; E</geoCoordinate>
, on rocks,
<collectorName id="57155336FFAA2431FD94B822FCEB3F5B" box="[637,780,1626,1651]" pageId="44" pageNumber="45">W.F. Ponder</collectorName>
,
<collectorName id="57155336FFAA2431FCF3B822FC553F5B" box="[794,946,1626,1651]" pageId="44" pageNumber="45">J.B. Studdert</collectorName>
,
<collectorName id="57155336FFAA2431FC28B822FBD33F5B" box="[961,1076,1626,1651]" pageId="44" pageNumber="45">C. Slatyer</collectorName>
&amp;
<collectorName id="57155336FFAA2431FBB6B822FB0A3F5B" box="[1119,1261,1626,1651]" pageId="44" pageNumber="45">J.M. Ponder</collectorName>
,
<collectingDate id="9E1AE9C8FFAA2431FB15B822FA7F3F5B" box="[1276,1432,1627,1651]" pageId="44" pageNumber="45" value="2004-08-23">
<date id="8E5E1020FFAA2431FB15B822FA7F3F5B" box="[1276,1432,1627,1651]" pageId="44" pageNumber="45" value="2004-08-23">23 Aug 2004</date>
</collectingDate>
, C.479956
</materialsCitation>
.
<typeStatus id="255B8842FFAA2431FEFBB806FE643FBF" box="[274,387,1663,1687]" pageId="44" pageNumber="45" type="paratype">Paratypes</typeStatus>
: Same data,
<date id="8E5E1020FFAA2431FDF9B806FD423FBF" box="[528,677,1663,1687]" pageId="44" pageNumber="45" value="2004-08-23">23 Aug 2004</date>
, C.445416, 20+ dry;
<materialsCitation id="4A883CBDFFAA2431FC7AB806FBBF3FBF" ID-GBIF-Occurrence="2235592665" box="[915,1112,1662,1687]" collectionCode="QM" pageId="44" pageNumber="45" specimenCode="MO85771, 5" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="9CF1AE25FFAA2431FC7AB806FC273FBF" box="[915,960,1663,1687]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="44" pageNumber="45" type="Museum">QM</collectionCode>
<specimenCode id="AA469E9BFFAA2431FC2EB807FBDA3FBF" box="[967,1085,1662,1687]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="44" pageNumber="45" type="Museum">MO85771</specimenCode>
,
<specimenCode id="AA469E9BFFAA2431FBA1B807FBBF3FBF" box="[1096,1112,1662,1687]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="44" pageNumber="45" type="Museum">5</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="FA5F36E0FFAA2431FF2EB8DBFE6C3EBB" blockId="44.[151,1437,1590,1939]" pageId="44" pageNumber="45">
<emphasis id="C894EAF2FFAA2431FF2EB8DBFEE43F93" bold="true" box="[199,259,1698,1723]" pageId="44" pageNumber="45">Shell</emphasis>
(
<figureCitation id="62DB2A65FFAA2431FEFDB8DBFE983F93" box="[276,383,1698,1723]" captionStart="FIGURE 13" captionStartId="28.[151,250,1847,1869]" captionTargetBox="[275,1320,252,1754]" captionTargetId="figure@28.[214,1372,223,1786]" captionTargetPageId="28" captionText="FIGURE 13. Shells of species of Edgbastonia (Barcaldonia) n. subgen. A. Edgbastonia (B.) hufferensis n. sp., holotype (C.479951). B. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990) (C. 479947). C, D. Edgbastonia (B.) routhensis n. sp., Routh Stn, C, holotype (C.479960). D. paratype (C.448850). E, Edgbastonia (B.) pagoda n. sp., holotype, Myross Stn (C.479948). F. Edgbastonia (B.) rugosa n. sp., holotype, Myross Stn (C.479949). G. Edgbastonia (B.) corrugata umbilicata n. sp., holotype, Myross Stn (C.479946). H. Edgbastonia? conjuboyensis, holotype, Conjuboy Stn (C.479956). I. Edgbastonia (B.) chillagoensis, holotype, Chillagoe Stn (C.479958). Scale bar 1 mm." httpUri="https://zenodo.org/record/2637629/files/figure.png" pageId="44" pageNumber="45">Fig. 13H</figureCitation>
). Spire outline slightly convex, normally coiled, translucent. Length
<quantity id="3D189B05FFAA2431FB7EB8DAFAC63F92" box="[1175,1313,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="44" pageNumber="45" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.41.8 mm</quantity>
(mean
<quantity id="3D189B05FFAA2431FA90B8DBFF203FF7" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="44" pageNumber="45" unit="mm" value="1.6">1.6 mm</quantity>
), width
<quantity id="3D189B05FFAA2431FEC1B8BEFE543FF6" box="[296,435,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="44" pageNumber="45" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.91.1 mm</quantity>
(mean 1.0 mm). Protoconch of about 1.3 whorls, lacking punctate sculpture but with initial part with an irregular surface resembling uneven plastering, and the latter part with faint transverse wrinkles and extremely fine spiral striae. Teleoconch whorls moderately convex, evenly rounded, total number 3.54.1 (mean 3.7). Umbilicus represented by chink only. Aperture ovate or pyriform, inner lip narrow, thin or medium, slightly separated along whole length of parietal wall, outer lip thin or medium. Periostracum very thin and inconspicuous, white.
</paragraph>
</subSubSection>
<caption id="AE9F6668FFAB2430FF7EBEE1FD9A39E7" ID-Table-UUID="AE9F6668FFAB2430FF7EBEE1FD9A39E7" httpUri="http://table.plazi.org/id/AE9F6668FFAB2430FF7EBEE1FD9A39E7" pageId="45" pageNumber="46" targetBox="[159,1346,234,504]" targetIsTable="true" targetPageId="45">
<paragraph id="FA5F36E0FFAB2430FF7EBEE1FD9A39E7" blockId="45.[151,1436,151,207]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF7EBEE1FEF03987" bold="true" box="[151,279,152,175]" pageId="45" pageNumber="46">TABLE 16.</emphasis>
Shell measurements (in mm) of type material of
<taxonomicName id="3DE04D63FFAB2430FCC2BEEEFC553987" box="[811,946,151,175]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="45" pageNumber="46" phylum="Mollusca" rank="genus">
<emphasis id="C894EAF2FFAB2430FCC2BEEEFC553987" box="[811,946,151,175]" italics="true" pageId="45" pageNumber="46">Edgbastonia</emphasis>
</taxonomicName>
?
<emphasis id="C894EAF2FFAB2430FC2DBEEEFBBC3987" box="[964,1115,151,175]" italics="true" pageId="45" pageNumber="46">conjuboyensis</emphasis>
<emphasis id="C894EAF2FFAB2430FB8ABEE1FB473987" bold="true" box="[1123,1184,151,175]" pageId="45" pageNumber="46">n. sp.</emphasis>
(paratypes, C.445416). See Materials and Methods for abbreviations.
</paragraph>
</caption>
<paragraph id="FA5F36E0FFAB2430FE2BBE93FAA538D0" pageId="45" pageNumber="46">
<table id="88E0C440FFABDBE2FF76BE93FAA538D0" box="[159,1346,234,504]" gridcols="7" gridrows="7" pageId="45" pageNumber="46">
<tr id="44D034A2FFABDBE2FF76BE93FAA53828" box="[159,1346,234,256]" gridrow="0" pageId="45" pageNumber="46" rowspan-0="1">
<th id="07015DDEFFABDBE2FE2BBE93FE093828" box="[450,494,234,256]" gridcol="1" gridrow="0" pageId="45" pageNumber="46">SH</th>
<th id="07015DDEFFABDBE2FDBEBE93FD643828" box="[599,643,234,256]" gridcol="2" gridrow="0" pageId="45" pageNumber="46">SW</th>
<th id="07015DDEFFABDBE2FCE9BE93FCCB3828" box="[768,812,234,256]" gridcol="3" gridrow="0" pageId="45" pageNumber="46">AH</th>
<th id="07015DDEFFABDBE2FC43BE93FC313828" box="[938,982,234,256]" gridcol="4" gridrow="0" pageId="45" pageNumber="46">AW</th>
<th id="07015DDEFFABDBE2FBBDBE93FB673828" box="[1108,1152,234,256]" gridcol="5" gridrow="0" pageId="45" pageNumber="46">BW</th>
<th id="07015DDEFFABDBE2FAFFBE93FAA53828" box="[1302,1346,234,256]" gridcol="6" gridrow="0" pageId="45" pageNumber="46">TW</th>
</tr>
<tr id="44D034A2FFABDBE2FF76BF6DFAA53802" box="[159,1346,276,298]" gridrow="1" pageId="45" pageNumber="46">
<th id="07015DDEFFABDBE2FF76BF6DFF1A3802" box="[159,253,276,298]" gridcol="0" gridrow="1" pageId="45" pageNumber="46">Holotype</th>
<td id="07015DDEFFABDBE2FE2BBF6DFE093802" box="[450,494,276,298]" gridcol="1" gridrow="1" pageId="45" pageNumber="46">1.74</td>
<td id="07015DDEFFABDBE2FDBEBF6DFD643802" box="[599,643,276,298]" gridcol="2" gridrow="1" pageId="45" pageNumber="46">1.20</td>
<td id="07015DDEFFABDBE2FCE9BF6DFCCB3802" box="[768,812,276,298]" gridcol="3" gridrow="1" pageId="45" pageNumber="46">0.77</td>
<td id="07015DDEFFABDBE2FC43BF6DFC313802" box="[938,982,276,298]" gridcol="4" gridrow="1" pageId="45" pageNumber="46">0.62</td>
<td id="07015DDEFFABDBE2FBBDBF6DFB673802" box="[1108,1152,276,298]" gridcol="5" gridrow="1" pageId="45" pageNumber="46">1.31</td>
<td id="07015DDEFFABDBE2FAFFBF6DFAA53802" box="[1302,1346,276,298]" gridcol="6" gridrow="1" pageId="45" pageNumber="46">3</td>
</tr>
<tr id="44D034A2FFABDBE2FF76BF44FAA5387B" box="[159,1346,317,339]" gridrow="2" pageId="45" pageNumber="46">
<th id="07015DDEFFABDBE2FF76BF44FAA5387B" box="[159,1346,317,339]" colspan="7" colspanRight="6" gridcol="0" gridrow="2" pageId="45" pageNumber="46">Paratypes (20)</th>
</tr>
<tr id="44D034A2FFABDBE2FF76BF1FFAA53854" box="[159,1346,358,380]" gridrow="3" pageId="45" pageNumber="46">
<th id="07015DDEFFABDBE2FF76BF1FFF1A3854" box="[159,253,358,380]" gridcol="0" gridrow="3" pageId="45" pageNumber="46">Min</th>
<td id="07015DDEFFABDBE2FE2BBF1FFE093854" box="[450,494,358,380]" gridcol="1" gridrow="3" pageId="45" pageNumber="46">1.40</td>
<td id="07015DDEFFABDBE2FDBEBF1FFD643854" box="[599,643,358,380]" gridcol="2" gridrow="3" pageId="45" pageNumber="46">0.88</td>
<td id="07015DDEFFABDBE2FCE9BF1FFCCB3854" box="[768,812,358,380]" gridcol="3" gridrow="3" pageId="45" pageNumber="46">0.60</td>
<td id="07015DDEFFABDBE2FC43BF1FFC313854" box="[938,982,358,380]" gridcol="4" gridrow="3" pageId="45" pageNumber="46">0.60</td>
<td id="07015DDEFFABDBE2FBBDBF1FFB673854" box="[1108,1152,358,380]" gridcol="5" gridrow="3" pageId="45" pageNumber="46">1.15</td>
<td id="07015DDEFFABDBE2FAFFBF1FFAA53854" box="[1302,1346,358,380]" gridcol="6" gridrow="3" pageId="45" pageNumber="46">3.45</td>
</tr>
<tr id="44D034A2FFABDBE2FF76BFE9FAA5388E" box="[159,1346,400,422]" gridrow="4" pageId="45" pageNumber="46">
<th id="07015DDEFFABDBE2FF76BFE9FF1A388E" box="[159,253,400,422]" gridcol="0" gridrow="4" pageId="45" pageNumber="46">Max</th>
<td id="07015DDEFFABDBE2FE2BBFE9FE09388E" box="[450,494,400,422]" gridcol="1" gridrow="4" pageId="45" pageNumber="46">1.75</td>
<td id="07015DDEFFABDBE2FDBEBFE9FD64388E" box="[599,643,400,422]" gridcol="2" gridrow="4" pageId="45" pageNumber="46">1.09</td>
<td id="07015DDEFFABDBE2FCE9BFE9FCCB388E" box="[768,812,400,422]" gridcol="3" gridrow="4" pageId="45" pageNumber="46">0.78</td>
<td id="07015DDEFFABDBE2FC43BFE9FC31388E" box="[938,982,400,422]" gridcol="4" gridrow="4" pageId="45" pageNumber="46">0.76</td>
<td id="07015DDEFFABDBE2FBBDBFE9FB67388E" box="[1108,1152,400,422]" gridcol="5" gridrow="4" pageId="45" pageNumber="46">1.41</td>
<td id="07015DDEFFABDBE2FAFFBFE9FAA5388E" box="[1302,1346,400,422]" gridcol="6" gridrow="4" pageId="45" pageNumber="46">4.10</td>
</tr>
<tr id="44D034A2FFABDBE2FF76BFC0FAA538E7" box="[159,1346,441,463]" gridrow="5" pageId="45" pageNumber="46">
<th id="07015DDEFFABDBE2FF76BFC0FF1A38E7" box="[159,253,441,463]" gridcol="0" gridrow="5" pageId="45" pageNumber="46">Mean</th>
<td id="07015DDEFFABDBE2FE2BBFC0FE0938E7" box="[450,494,441,463]" gridcol="1" gridrow="5" pageId="45" pageNumber="46">1.58</td>
<td id="07015DDEFFABDBE2FDBEBFC0FD6438E7" box="[599,643,441,463]" gridcol="2" gridrow="5" pageId="45" pageNumber="46">1.00</td>
<td id="07015DDEFFABDBE2FCE9BFC0FCCB38E7" box="[768,812,441,463]" gridcol="3" gridrow="5" pageId="45" pageNumber="46">0.68</td>
<td id="07015DDEFFABDBE2FC43BFC0FC3138E7" box="[938,982,441,463]" gridcol="4" gridrow="5" pageId="45" pageNumber="46">0.69</td>
<td id="07015DDEFFABDBE2FBBDBFC0FB6738E7" box="[1108,1152,441,463]" gridcol="5" gridrow="5" pageId="45" pageNumber="46">1.27</td>
<td id="07015DDEFFABDBE2FAFFBFC0FAA538E7" box="[1302,1346,441,463]" gridcol="6" gridrow="5" pageId="45" pageNumber="46">3.70</td>
</tr>
<tr id="44D034A2FFABDBE2FF76BF9BFAA538D0" box="[159,1346,482,504]" gridrow="6" pageId="45" pageNumber="46">
<th id="07015DDEFFABDBE2FF76BF9BFF1A38D0" box="[159,253,482,504]" gridcol="0" gridrow="6" pageId="45" pageNumber="46">SD</th>
<td id="07015DDEFFABDBE2FE2BBF9BFE0938D0" box="[450,494,482,504]" gridcol="1" gridrow="6" pageId="45" pageNumber="46">0.12</td>
<td id="07015DDEFFABDBE2FDBEBF9BFD6438D0" box="[599,643,482,504]" gridcol="2" gridrow="6" pageId="45" pageNumber="46">0.06</td>
<td id="07015DDEFFABDBE2FCE9BF9BFCCB38D0" box="[768,812,482,504]" gridcol="3" gridrow="6" pageId="45" pageNumber="46">0.05</td>
<td id="07015DDEFFABDBE2FC43BF9BFC3138D0" box="[938,982,482,504]" gridcol="4" gridrow="6" pageId="45" pageNumber="46">0.05</td>
<td id="07015DDEFFABDBE2FBBDBF9BFB6738D0" box="[1108,1152,482,504]" gridcol="5" gridrow="6" pageId="45" pageNumber="46">0.08</td>
<td id="07015DDEFFABDBE2FAFFBF9BFAA538D0" box="[1302,1346,482,504]" gridcol="6" gridrow="6" pageId="45" pageNumber="46">0.18</td>
</tr>
</table>
</paragraph>
<subSubSection id="B2FA656BFFAB2430FF2EBC40FB063CA6" pageId="45" pageNumber="46" type="description">
<paragraph id="FA5F36E0FFAB2430FF2EBC40FD183B5E" blockId="45.[151,1437,569,2034]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EBC40FEA83B7A" bold="true" box="[199,335,569,594]" pageId="45" pageNumber="46">Operculum</emphasis>
(
<figureCitation id="62DB2A65FFAB2430FE88BC40FE133B79" box="[353,500,569,594]" captionStart="FIGURE 14" captionStartId="29.[151,250,1810,1832]" captionTargetBox="[218,1365,209,1793]" captionTargetId="figure@29.[200,1397,193,1801]" captionTargetPageId="29" captionText="FIGURE 14. Opercula of species of Edgbastonia (Barcaldonia) n. subgen. A. Edgbastonia (B.) hufferensis n. sp., internal view (C.423238). B, C. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990). B. External view. C. Internal view (C. 457755). D, E. Edgbastonia (B.) routhensis n. sp. D. External view. E. Internal view (C438551). F, G. Edgbastonia (B.) pagoda n. sp. F. External view. G. Internal view (C.457756). H. Edgbastonia (B.) rugosa n. sp., internal view (C.457753). I, J. Edgbastonia (B.) corrugata umbilicata n. subsp. I. External view. J. Internal view (C.415845). K, L. Edgbastonia? conjuboyensis n. sp. K. External view. L. Internal view (C.445416). M, N. Edgbastonia (B.) chillagoensis n. sp. M. External view. N. Internal view (C.438557). Scale bar 200 µm." httpUri="https://zenodo.org/record/2637631/files/figure.png" pageId="45" pageNumber="46">Fig. 14M, N</figureCitation>
). Transparent, pale yellow, slightly concave. Inner side lacking white smear, with weak, smooth ridge near columellar edge of inner side.
</paragraph>
<paragraph id="FA5F36E0FFAB2430FF2EBCF8FB463B96" blockId="45.[151,1437,569,2034]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EBCF8FDCB3BB2" bold="true" box="[199,556,641,666]" pageId="45" pageNumber="46">Head-foot and external body.</emphasis>
Snout, neck and opercular lobes pigmented, tentacles and dorsal and lateral foot unpigmented, mantle roof densely pigmented or black, visceral coil densely pigmented.
</paragraph>
<paragraph id="FA5F36E0FFAB2430FF2EBCB0FC383ABE" blockId="45.[151,1437,569,2034]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EBCB0FE953BCA" bold="true" box="[199,370,713,738]" pageId="45" pageNumber="46">Mantle cavity.</emphasis>
Ctenidium well-developed, filaments 1315, broadly triangular, apex right edge. Osphradium narrowly oval, towards anterior end of ctenidium, length relative to gill 0.210.31. Hypobranchial gland not distinguishable (absent). Rectum with U-shaped bend, faecal pellets longitudinally orientated or orientated obliquely and longitudinally, anus at or very near mantle collar. Kidney extends for about third of length into mantle cavity roof, or half or more in roof of mantle cavity. Renal gland transverse. Pericardium extends for about third of length into mantle cavity roof, abutting posterior end of ctenidium.
</paragraph>
<paragraph id="FA5F36E0FFAB2430FF2EBDD8FF3A3D0D" blockId="45.[151,1437,569,2034]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EBDD8FEF93A92" bold="true" box="[199,286,929,954]" pageId="45" pageNumber="46">Radula</emphasis>
(
<figureCitation id="62DB2A65FFAB2430FED8BDD8FE5C3A92" box="[305,443,929,954]" captionStart="FIGURE 19" captionStartId="36.[151,250,1281,1303]" captionTargetBox="[168,1402,212,1252]" captionTargetId="figure@36.[151,1435,193,1280]" captionTargetPageId="36" captionText="FIGURE 19. Radulae of species of Edgbastonia (Barcaldonia) n. subgen. A, B. Edgbastonia (B.) rugosa n. sp. A. Half row. B. Lateral teeth (C.457753). C, D. Edgbastonia (B.) corrugata umbilicata n. subsp. C. Half row. D. Lateral and marginal teeth (C.415845). E, F. Edgbastonia? conjuboyensis n. sp. E. Half row. F. Marginal teeth (C.445416). G, H, I. Edgbastonia (B.) chillagoensis n. sp. G. Half row. H. Lateral and marginal teeth. I. Outer marginal teeth (C.438557). Scale bars 10 µm." httpUri="https://zenodo.org/record/2637641/files/figure.png" pageId="45" pageNumber="46">Fig. 19E, F</figureCitation>
). Central teeth with cusp formula 45+1+45, basal cusps 2+2; median cusp narrow, sharp about twice as long as adjacent cusps. Lateral teeth with cusp formula 34+1+34; main cusp about twice as long as adjacent cusps, sharp to blunt. Inner marginal teeth with 1921 cusps. Outer marginal teeth with 1721 cusps.
</paragraph>
<paragraph id="FA5F36E0FFAB2430FF2EBA48FD793C0A" blockId="45.[151,1437,569,2034]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EBA48FDFC3D62" bold="true" box="[199,539,1073,1098]" pageId="45" pageNumber="46">Female reproductive system</emphasis>
(
<figureCitation id="62DB2A65FFAB2430FDC4BA48FD263D62" box="[557,705,1073,1098]" captionStart="FIGURE 16" captionStartId="33.[151,250,1801,1823]" captionTargetBox="[150,1389,213,1728]" captionTargetId="figure@33.[150,1436,193,1779]" captionTargetPageId="33" captionText="FIGURE 16. Female reproductive system of species of Edgbastonia (Barcaldonia) n. subgen., shown from left side. A, B. Edgbastonia (B.) hufferensis n. sp. (C.423238). C, F. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990) (C.457755). D, E. Edgbastonia (B.) routhensis n. sp. (C.438551). G, H, Edgbastonia (B.) pagoda n. sp. (C.457756). I, L. Edgbastonia (B.) rugosa n. sp. (C.457753). J, K. Edgbastonia (B.) corrugata umbilicata n. subsp., (C.415845). M, N. Edgbastonia? conjuboyensis n. sp. (C.445416). O, P. Edgbastonia (B.) chillagoensis n. sp. (C.438557). Abbreviations: ag—albumen gland; bc—bursa copulatrix; cg—capsule gland; o—opening of oviduct; pw—posterior wall of mantle cavity; ro—renal oviduct; sr—seminal receptacle; vc—ventral channel. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637635/files/figure.png" pageId="45" pageNumber="46">Fig. 16M, N</figureCitation>
). Ovary simple sac. Renal oviduct makes tight, near vertical to anteriorly orientated loop, distinctly muscular. Seminal receptacle absent; orientated sperm located in renal oviduct. Bursa copulatrix behind albumen gland, elongately-oval or triangular, shorter than albumen gland, bursal duct enters bursa mid anteriorly, bursal duct joins coiled oviduct little behind posterior mantle cavity wall. Albumen gland partly in mantle cavity. Capsule gland with two distinct glandular zones, medium thickness in cross section, markedly indented by rectum. Anterior vestibule not defined, opening subterminal, short, cowl and/or gutter associated with oviduct opening absent.
</paragraph>
<paragraph id="FA5F36E0FFAB2430FF2EBB54FB063CA6" blockId="45.[151,1437,569,2034]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EBB54FDE63C6E" bold="true" box="[199,513,1325,1350]" pageId="45" pageNumber="46">Male reproductive system</emphasis>
(
<figureCitation id="62DB2A65FFAB2430FDFABB54FD9D3C6E" box="[531,634,1325,1350]" captionStart="FIGURE 17" captionStartId="34.[151,250,1813,1835]" captionTargetBox="[183,1412,762,1756]" captionTargetId="figure@34.[151,1436,741,1775]" captionTargetPageId="34" captionText="FIGURE 17. Prostate and vas deferens of species of Edgbastonia (Barcaldonia) n. subgen. A. Edgbastonia (B.) hufferensis n. sp. (C.423238). B. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990) (C.457755). C. Edgbastonia (B.) routhensis n. sp. (C.438551). D. Edgbastonia (B.) pagoda n. sp. (C.457756). E. Edgbastonia (B.) rugosa n. sp. (C.457753). F. Edgbastonia (B.) corrugata umbilicata n. subsp. (C.415845). G. Edgbastonia? conjuboyensis n. sp. (C. 445416). H. Edgbastonia (B.) chillagoensis n. sp. (C.438557). Abbreviations: avd—anterior vas deferens; pg—prostate gland; pvd—posterior vas deferens; pw—position of posterior wall of pallial cavity. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637637/files/figure.png" pageId="45" pageNumber="46">Fig. 17G</figureCitation>
,
<figureCitation id="62DB2A65FFAB2430FD61BB57FD333C6E" box="[648,724,1326,1350]" captionStart="FIGURE 18" captionStartId="35.[151,250,1854,1876]" captionTargetBox="[200,1347,213,1826]" captionTargetId="figure@35.[185,1406,193,1841]" captionTargetPageId="35" captionText="FIGURE 18. Penis and vas deferens of species of Edgbastonia (Barcaldonia) n. subgen. A. Edgbastonia (B.) hufferensis n. sp. (C.423238). B, C. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990) (C.457755). D, Edgbastonia (B.) routhensis n. sp. (C.438551). E. Edgbastonia (B.) pagoda n. sp. (C.457756). F. Edgbastonia (B.) rugosa n. sp. (C.457753). G. Edgbastonia (B.) corrugata umbilicata n. subsp. H, I. Edgbastonia? conjuboyensis n. sp. (C.445416). J, K. Edgbastonia (B.) chilagoensis n. sp. (C.438557). Abbreviations: avd—anterior vas deferens; ngl—non-glandular lobe; p—papilla; pd—penial duct. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637639/files/figure.png" pageId="45" pageNumber="46">18H, I</figureCitation>
). Prostate gland half in mantle roof, bean-shaped, medium in cross section. Posterior pallial vas deferens slightly undulating, anteriorly strongly and tightly undulating near base of penis. Penis towards middle of head, distal end narrow, elongate, not tapering, blunt terminally.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFFAB2430FF2EBBE0FD473C9A" box="[199,672,1433,1458]" pageId="45" pageNumber="46" type="etymology">
<paragraph id="FA5F36E0FFAB2430FF2EBBE0FD473C9A" blockId="45.[151,1437,569,2034]" box="[199,672,1433,1458]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EBBE0FEA93C9A" bold="true" box="[199,334,1433,1458]" pageId="45" pageNumber="46">Etymology.</emphasis>
Named for Conjuboy Station.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFFAB2430FF2EBBC4FB333CD2" pageId="45" pageNumber="46" type="distribution">
<paragraph id="FA5F36E0FFAB2430FF2EBBC4FB333CD2" blockId="45.[151,1437,569,2034]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EBBC4FE163CFE" bold="true" box="[199,497,1469,1494]" pageId="45" pageNumber="46">Distribution and habitat.</emphasis>
Known only from a non-GAB spring on Conjuboy Station (
<figureCitation id="62DB2A65FFAB2430FB7DBBC4FB333CFE" box="[1172,1236,1469,1494]" captionStart="FIGURE 1" captionStartId="4.[151,250,1844,1866]" captionTargetBox="[151,1435,498,1802]" captionTargetId="figure@4.[151,1435,498,1802]" captionTargetPageId="4" captionText="FIGURE 1. Map of the spring supergroups associated with the Great Artesian Basin (GAB) in Queensland, showing the main non-town locations mentioned in the text (squares) and some of the towns in the area (ovals). The pale grey shading is the discharge part of the GAB, the dark grey the recharge part. The inset map of Australia shows the area illustrated. The approximate location of Conondale is indicated by a triangle and Nundle by a star. The spring supergroups are outlined with dotted lines and are as follows: A, Mulligan River; B, Springvale; C, Flinders River; D, Hughenden; E, Barcaldine (see inset at upper right for details); F, Springsure; G, Eulo." httpUri="https://zenodo.org/record/2637605/files/figure.png" pageId="45" pageNumber="46">Fig. 1</figureCitation>
) in north western
<collectingRegion id="3824F802FFAB2430FF7EBB98FEF93CD2" box="[151,286,1505,1530]" country="Australia" name="Queensland" pageId="45" pageNumber="46">Queensland</collectingRegion>
, about
<quantity id="3D189B05FFAB2430FE87BB9BFE243CD2" box="[366,451,1505,1530]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="45" pageNumber="46" unit="km" value="120.0">120 km</quantity>
south of Mt Garnet in the mid-eastern part of the Einasleigh Uplands.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFFAB2433FF2EB87CFF1A38F8" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="discussion">
<paragraph id="FA5F36E0FFAB2430FF2EB87CFF383E32" blockId="45.[151,1437,569,2034]" pageId="45" pageNumber="46">
<emphasis id="C894EAF2FFAB2430FF2EB87CFEDC3F36" bold="true" box="[199,315,1541,1566]" pageId="45" pageNumber="46">Remarks.</emphasis>
This species was found in a spring emanating from basalt rocks on Conjuboy Station and thus it is not from the Great Artesian Basin aquifers. It is the only tateid known to date from a basal Tertiary spring (
<bibRefCitation id="9E714B11FFAB2430FF77B834FEBA3F4E" author="Habermehl, M. A." box="[158,349,1613,1638]" pageId="45" pageNumber="46" refId="ref34878" refString="Habermehl, M. A. (1982) Springs of the Great Artesian Basin, Australia-their origin and nature. Australian Bureau of Mineral Resources, Geology and Geophysics. Report 235. Australian Government Publishing Service for the Bureau of Mineral Resources, Geology and Geophysics, Canberra, 50 leaves." type="book" year="1982">Habermehl 1982</bibRefCitation>
) within the area occupied by the
<collectingRegion id="3824F802FFAB2430FD3FB834FCBD3F4E" box="[726,858,1613,1638]" country="Australia" name="Queensland" pageId="45" pageNumber="46">Queensland</collectingRegion>
part of the GAB. Some of the basalts on Conjuboy Station are Murronga basalts formed around 120,000 to 180,000 years ago (
<bibRefCitation id="9E714B11FFAB2430FC02B808FAF93FA2" author="Griffin, T. J. &amp; McDougall, I." box="[1003,1310,1649,1674]" pageId="45" pageNumber="46" pagination="387 - 396" refId="ref34563" refString="Griffin, T. J. &amp; McDougall, I. (1975) Geochronology of the Cainozoic McBride Volcanic Province, Northern Queensland. Journal of the Geological Society of Australia, 22, 387 - 396. https: // doi. org / 10.1080 / 00167617508728905" type="journal article" year="1975">Griffin &amp; McDougall 1975</bibRefCitation>
) which are slightly younger than the famous Undara Lava Tubes to the south which were formed a little earlier, around 190,000 years ago. However, other basalts on Conjuboy Station are much older, around 400,000 years and some 2.27 million years old, while others in the same region are in excess of 7 million years (
<bibRefCitation id="9E714B11FFAB2430FB4AB8A4FF343E32" author="Griffin, T. J. &amp; McDougall, I." pageId="45" pageNumber="46" pagination="387 - 396" refId="ref34563" refString="Griffin, T. J. &amp; McDougall, I. (1975) Geochronology of the Cainozoic McBride Volcanic Province, Northern Queensland. Journal of the Geological Society of Australia, 22, 387 - 396. https: // doi. org / 10.1080 / 00167617508728905" type="journal article" year="1975">Griffin &amp; McDougall 1975</bibRefCitation>
).
</paragraph>
<paragraph id="FA5F36E0FFAB2433FF2EB95CFDEC3835" blockId="45.[151,1437,569,2034]" lastBlockId="46.[151,1436,151,464]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
Regrettably our available material of this species was not able to be sequenced. It occurs well to the north of the Barcaldine Supergroup springs, in the same general area as
<emphasis id="C894EAF2FFAB2430FC8AB930FC6D3E4A" box="[867,906,1865,1890]" italics="true" pageId="45" pageNumber="46">Ed.</emphasis>
<taxonomicName id="3DE04D63FFAB2430FC78B933FBBF3E4A" authorityName="Zhang" authorityYear="2019" box="[913,1112,1865,1890]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="chillagoensis" subGenus="Barcaldinia">
(
<emphasis id="C894EAF2FFAB2430FC72B933FC4B3E49" box="[923,940,1866,1889]" italics="true" pageId="45" pageNumber="46">B</emphasis>
.)
<emphasis id="C894EAF2FFAB2430FC2DB930FBBF3E4A" box="[964,1112,1865,1890]" italics="true" pageId="45" pageNumber="46">chillagoensis</emphasis>
</taxonomicName>
and
<emphasis id="C894EAF2FFAB2430FB66B930FB513E4A" box="[1167,1206,1865,1890]" italics="true" pageId="45" pageNumber="46">Ed.</emphasis>
<taxonomicName id="3DE04D63FFAB2430FB54B933FA8E3E4A" authorityName="Zhang" authorityYear="2019" box="[1213,1385,1865,1890]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="hufferensis" subGenus="Barcaldinia">
(
<emphasis id="C894EAF2FFAB2430FB2FB933FB303E49" box="[1222,1239,1866,1889]" italics="true" pageId="45" pageNumber="46">B</emphasis>
.)
<emphasis id="C894EAF2FFAB2430FB06B930FA8E3E4A" box="[1263,1385,1865,1890]" italics="true" pageId="45" pageNumber="46">hufferensis</emphasis>
</taxonomicName>
. All the springs in this area (the Einasleigh Uplands) are not fed from water from aquifers making up the Great Artesian Basin (
<bibRefCitation id="9E714B11FFAB2430FF0FB9E8FE4F3E82" author="Habermehl, M. A." box="[230,424,1937,1962]" pageId="45" pageNumber="46" refId="ref34878" refString="Habermehl, M. A. (1982) Springs of the Great Artesian Basin, Australia-their origin and nature. Australian Bureau of Mineral Resources, Geology and Geophysics. Report 235. Australian Government Publishing Service for the Bureau of Mineral Resources, Geology and Geophysics, Canberra, 50 leaves." type="book" year="1982">Habermehl 1982</bibRefCitation>
). While the molecular data for the other two species appears to confirm their relationship with
<taxonomicName id="3DE04D63FFAB2430FF3BB9CCFE873EE6" box="[210,352,1973,1998]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="45" pageNumber="46" phylum="Mollusca" rank="genus">
<emphasis id="C894EAF2FFAB2430FF3BB9CCFE873EE6" box="[210,352,1973,1998]" italics="true" pageId="45" pageNumber="46">Edgbastonia</emphasis>
</taxonomicName>
, we include the present species in that genus only tentatively, pending confirmation. It differs from species included in any of the genera dealt with here in its more regularly conical shell. Anatomically, it mainly differs in the female lacking a bursal duct. Another unusual feature is its peculiar protoconch microsculpture, which is unlike that seen in any other Australian tateid. Despite the differences, the general anatomy, radula, and operculum do not show other substantive differences from many of the other species included here in the
<emphasis id="C894EAF2FFA82433FEF4BF7DFE683835" box="[285,399,260,285]" italics="true" pageId="46" pageNumber="47">Jardinella</emphasis>
radiation.
</paragraph>
<paragraph id="FA5F36E0FFA82433FF2EBF5EFF1A38F8" blockId="46.[151,1436,151,464]" pageId="46" pageNumber="47">
Species of the SE Australian/Tasmanian genus
<emphasis id="C894EAF2FFA82433FD3DBF51FC8D3817" box="[724,874,296,319]" italics="true" pageId="46" pageNumber="47">Austropygrus</emphasis>
Cotton have a similar shell morphology, but those all have a pegged operculum, more than two pairs of basal cusps and differ in some anatomical details (e.g., possessing a gastric caecum) and in protoconch microsculpture (see
<bibRefCitation id="9E714B11FFA82433FC09BF16FB5D38A0" author="Clark, S. A. &amp; Miller, A. C. &amp; Ponder, W. F." box="[992,1210,367,392]" pageId="46" pageNumber="47" pagination="1 - 109" refId="ref33317" refString="Clark, S. A., Miller, A. C. &amp; Ponder, W. F. (2003) A revision of the snail genus Austropyrgus (Gastropoda: Hydrobiidae): a morphostatic radiation of freshwater gastropods in southeastern Australia. Records of the Australian Museum, 28 (Supplement), 1 - 109. https: // doi. org / 10.3853 / j. 0812 - 7387.28.2003.1377" type="journal article" year="2003">
Clark
<emphasis id="C894EAF2FFA82433FBC7BF08FB8938A0" box="[1070,1134,367,392]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2003
</bibRefCitation>
). Some species of
<emphasis id="C894EAF2FFA82433FF7EBFEDFEC13885" box="[151,294,404,429]" italics="true" pageId="46" pageNumber="47">Fonscochlea</emphasis>
<bibRefCitation id="9E714B11FFA82433FEDBBFEDFDF03884" author="Ponder, W. F. &amp; Hershler, R. &amp; Jenkins, B." box="[306,535,404,429]" pageId="46" pageNumber="47" pagination="1 - 140" refId="ref36212" refString="Ponder, W. F., Hershler, R. &amp; Jenkins, B. (1989) An endemic radiation of Hydrobiidae from artesian springs in northern South Australia: their taxonomy, physiology, distribution and anatomy. Malacologia, 31, 1 - 140." type="journal article" year="1989">
Ponder
<emphasis id="C894EAF2FFA82433FE67BFECFE2B3885" box="[398,460,404,429]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
, 1989
</bibRefCitation>
are generally similar but members of that genus differ markedly in female anatomy.
</paragraph>
</subSubSection>
</treatment>
</document>