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<document id="1E32CC18BCDC94B543DEC25CE9A300D7" ID-CLB-Dataset="27316" ID-DOI="10.11646/zootaxa.4583.1.1" ID-GBIF-Dataset="027ed27d-5035-4646-93a6-c23b565f3d20" ID-ISSN="1175-5326" ID-Zenodo-Dep="2637603" ID-ZooBank="27F24995-359E-46F6-AB22-75568BACFDCF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555054242242" checkinUser="plazi" docAuthor="Zhang, - H." docDate="2019" docId="724987F6FF9D2403FF7EBA25FEEC3EFA" docLanguage="en" docName="zootaxa.4583.1.1.pdf" docOrigin="Zootaxa 4583 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Edgbastonia (Barcaldinia) corrugata subsp. umbilicata Zhang 2019, n. subsp." docType="treatment" docVersion="16" lastPageNumber="31" masterDocId="8E70FF8EFF86241DFFE9BE79FFE73928" masterDocTitle="New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages" masterLastPageNumber="67" masterPageNumber="1" pageNumber="28" updateTime="1698720421338" updateUser="ExternalLinkService">
<mods:mods id="CB49ED775FFE569A59CFA90A324412F4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EEC80A4CBD586D48AE0913145A60381A">
<mods:title id="752BD5324507FA5FD350E12032E1D3C4">New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages</mods:title>
</mods:titleInfo>
<mods:name id="C3E70F5E5E5D441138DE4202AB6BBA08" type="personal">
<mods:role id="CABC5E2F5B2644E4FC8411F15F6F4DDB">
<mods:roleTerm id="499F44417D0987F684027133D33C7D8E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC42FD92310E3F2C631C6A1A8AA0AFFC">Zhang, - H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4F4890AA6A73FD037B35F441FBF8B556">text</mods:typeOfResource>
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<mods:titleInfo id="3FC830221EDE098647EAC5CC25F9AEA0">
<mods:title id="E586A8E346060FD5E0BCFEC0FC2FD1A5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7ED53B07DCFEB3A803028B19A79B64E6">
<mods:date id="ACE510FE3CE0CAA92C730E2FC0DD3499">2019</mods:date>
<mods:detail id="D022A465725278889BC7F6CE87678346" type="pubDate">
<mods:number id="4720C3D7FF0EC6A39A17F49DC45A477C">2019-04-10</mods:number>
</mods:detail>
<mods:detail id="2D87ADC1819565DEA9159BD8ED8ACCA8" type="volume">
<mods:number id="C47A1E7263820AA1AC2995A2F6EDF420">4583</mods:number>
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<mods:detail id="B11811B784391F2119B4817329A623B0" type="issue">
<mods:number id="6909E2A712E31C78E10DFA6329698742">1</mods:number>
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<mods:classification id="3C31A9EF281501863B48ECE5F8FADF8C">journal article</mods:classification>
<mods:identifier id="E004A49D4DBC6DCCCFE7B730D51E94F5" type="CLB-Dataset">27316</mods:identifier>
<mods:identifier id="EA0A3A91FB16DFDD834612880FA58734" type="DOI">10.11646/zootaxa.4583.1.1</mods:identifier>
<mods:identifier id="08B95DFFA825194C54BC0E82DC323402" type="GBIF-Dataset">027ed27d-5035-4646-93a6-c23b565f3d20</mods:identifier>
<mods:identifier id="4EAB50962E46B6FA3F02551DFADE3A97" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E9CA84D9D45E50704904061FF77B7D07" type="Zenodo-Dep">2637603</mods:identifier>
<mods:identifier id="2275BBBF10D81C2B9F95E1B081E1A22F" type="ZooBank">27F24995-359E-46F6-AB22-75568BACFDCF</mods:identifier>
</mods:mods>
<treatment id="724987F6FF9D2403FF7EBA25FEEC3EFA" ID-DOI="http://doi.org/10.5281/zenodo.5582690" ID-GBIF-Taxon="156198429" ID-Zenodo-Dep="5582690" LSID="urn:lsid:plazi:treatment:724987F6FF9D2403FF7EBA25FEEC3EFA" httpUri="http://treatment.plazi.org/id/724987F6FF9D2403FF7EBA25FEEC3EFA" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
<subSubSection id="B2FA656BFF9D2406FF7EBA25FCB43D5E" box="[151,851,1116,1142]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="FA5F36E0FF9D2406FF7EBA25FCB43D5E" blockId="27.[151,851,1116,1142]" box="[151,851,1116,1142]" pageId="27" pageNumber="28">
<heading id="A117818CFF9D2406FF7EBA25FCB43D5E" bold="true" box="[151,851,1116,1142]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="C894EAF2FF9D2406FF7EBA25FCB43D5E" bold="true" box="[151,851,1116,1142]" pageId="27" pageNumber="28">
<taxonomicName id="3DE04D63FF9D2406FF7EBA25FD3A3D5E" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[151,733,1116,1142]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="28" phylum="Mollusca" rank="subSpecies" species="corrugata" status="subsp. nov." subGenus="Barcaldinia" subSpecies="umbilicata">
<emphasis id="C894EAF2FF9D2406FF7EBA25FEC83D5E" bold="true" box="[151,303,1116,1142]" italics="true" pageId="27" pageNumber="28">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9D2406FED6BA25FE293D5E" bold="true" box="[319,462,1116,1142]" italics="true" pageId="27" pageNumber="28">Barcaldinia</emphasis>
)
<emphasis id="C894EAF2FF9D2406FE09BA25FD3A3D5E" bold="true" box="[480,733,1116,1142]" italics="true" pageId="27" pageNumber="28">corrugata umbilicata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9D2406FD0CBA25FCB43D5E" box="[741,851,1116,1142]" pageId="27" pageNumber="28" rank="subSpecies">n. subsp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF9D2406FF7EBADBFC0A3FA7" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="FA5F36E0FF9D2406FF7EBADBFD803CF3" blockId="27.[151,1437,1186,1967]" pageId="27" pageNumber="28">
<emphasis id="C894EAF2FF9D2406FF7EBADBFE663D93" bold="true" box="[151,385,1186,1211]" pageId="27" pageNumber="28">Material examined.</emphasis>
<materialsCitation id="4A883CBDFF9D2406FE61BADBFB6F3DF7" ID-GBIF-Occurrence="2235592658" collectingDate="2002-10-06" collectorName="W. F. Ponder &amp; A. Davis" latitude="-22.785" location="Myross Station" longLatPrecision="21" longitude="145.43333" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="255B8842FF9D2406FE61BADBFE123D93" box="[392,501,1186,1211]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
:
<collectingRegion id="3824F802FF9D2406FDE9BADBFD603D93" box="[512,647,1186,1211]" country="Australia" name="Queensland" pageId="27" pageNumber="28">Queensland</collectingRegion>
,
<location id="FF3F603BFF9D2406FD7ABADAFCA63D93" LSID="urn:lsid:plazi:treatment:724987F6FF9D2403FF7EBA25FEEC3EFA:FF3F603BFF9D2406FD7ABADAFCA63D93" box="[659,833,1186,1211]" latitude="-22.785" longLatPrecision="21" longitude="145.43333" name="Myross Station" pageId="27" pageNumber="28" stateProvince="Queensland">Myross Station</location>
, main spring,
<geoCoordinate id="9FD45027FF9D2406FC08BADAFB9E3D93" box="[993,1145,1186,1211]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="-22.785">22° 47' 06&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF9D2406FB6FBADBFACA3D92" box="[1158,1325,1186,1211]" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="15" value="145.43333">145° 26' 00&quot; E</geoCoordinate>
, at spring edge, amongst grass and on mud,
<collectorName id="57155336FF9D2406FDD0BABEFD2E3DF7" box="[569,713,1222,1247]" pageId="27" pageNumber="28">W.F. Ponder</collectorName>
&amp;
<collectorName id="57155336FF9D2406FD13BABEFC803DF7" box="[762,871,1222,1247]" pageId="27" pageNumber="28">A. Davis</collectorName>
,
<collectingDate id="9E1AE9C8FF9D2406FC90BABFFBE23DF7" box="[889,1029,1222,1247]" pageId="27" pageNumber="28" value="2002-10-06">
<date id="8E5E1020FF9D2406FC90BABFFBE23DF7" box="[889,1029,1222,1247]" pageId="27" pageNumber="28" value="2002-10-06">6 Oct 2002</date>
</collectingDate>
, C.479946
</materialsCitation>
.
<materialsCitation id="4A883CBDFF9D2406FB70BABEFC763C47" ID-GBIF-Occurrence="2235592659" collectingDate="2006-09-13" collectingDateMax="2008-07-01" collectingDateMin="2006-09-13" collectionCode="QM" collectorName="W. F. Ponder &amp; R. Fairfax &amp; R. Fensham &amp; R. J. Fensham &amp; Other" county="Same" latitude="-22.790833" location="Myross Station" longLatPrecision="21" longitude="145.44084" municipality="Other" pageId="27" pageNumber="28" specimenCode="MO85774, 5" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<typeStatus id="255B8842FF9D2406FB70BABEFAED3DF7" box="[1177,1290,1223,1247]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
:
<collectingCounty id="133E4E6CFF9D2406FAF2BABEFABE3DF7" box="[1307,1369,1223,1247]" pageId="27" pageNumber="28">Same</collectingCounty>
data, C.415845, 20+;
<collectionCode id="9CF1AE25FF9D2406FEA4BA92FE9D3C2B" box="[333,378,1259,1283]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="27" pageNumber="28" type="Museum">QM</collectionCode>
<specimenCode id="AA469E9BFF9D2406FE68BA93FE103C2B" box="[385,503,1258,1283]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="27" pageNumber="28" type="Museum">MO85774</specimenCode>
,
<specimenCode id="AA469E9BFF9D2406FDEBBA93FDF43C2B" box="[514,531,1258,1283]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="27" pageNumber="28" type="Museum">5</specimenCode>
.
<collectingMunicipality id="1A3BAC9AFF9D2406FDF7BA93FDB93C2B" box="[542,606,1258,1283]" pageId="27" pageNumber="28">Other</collectingMunicipality>
material examined:
<collectingRegion id="3824F802FF9D2406FCABBA93FC2E3C2B" box="[834,969,1258,1283]" country="Australia" name="Queensland" pageId="27" pageNumber="28">Queensland</collectingRegion>
,
<location id="FF3F603BFF9D2406FC3DBA92FB663C2B" LSID="urn:lsid:plazi:treatment:724987F6FF9D2403FF7EBA25FEEC3EFA:FF3F603BFF9D2406FC3DBA92FB663C2B" box="[980,1153,1258,1283]" county="Same" latitude="-22.790833" longLatPrecision="21" longitude="145.44084" municipality="Other" name="Myross Station" pageId="27" pageNumber="28" stateProvince="Queensland">Myross Station</location>
, main spring,
<geoCoordinate id="9FD45027FF9D2406FAF6BA92FF4F3C0F" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="-22.785276">22° 47' 07&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF9D2406FF5DBB77FEBE3C0E" box="[180,345,1294,1319]" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="15" value="145.43472">145° 26' 05&quot; E</geoCoordinate>
,
<collectorName id="57155336FF9D2406FE8DBB76FE083C0F" box="[356,495,1294,1319]" pageId="27" pageNumber="28">W.F. Ponder</collectorName>
,
<collectorName id="57155336FF9D2406FE13BB76FD8B3C0F" box="[506,620,1294,1319]" pageId="27" pageNumber="28">R. Fairfax</collectorName>
&amp;
<collectorName id="57155336FF9D2406FD67BB76FCF03C0F" box="[654,791,1294,1319]" pageId="27" pageNumber="28">R. Fensham</collectorName>
,
<collectingDate id="9E1AE9C8FF9D2406FCCBBB76FC573C0F" box="[802,944,1294,1319]" pageId="27" pageNumber="28" value="2006-09-13">
<date id="8E5E1020FF9D2406FCCBBB76FC573C0F" box="[802,944,1294,1319]" pageId="27" pageNumber="28" value="2006-09-13">13 Sep 2006</date>
</collectingDate>
, C.458219, 20+;
<geoCoordinate id="9FD45027FF9D2406FB99BB76FAE23C0F" box="[1136,1285,1295,1319]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="-22.795">22° 47' 42&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF9D2406FAF8BB77FF4C3C62" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="15" value="145.44055">145° 26' 26&quot; E</geoCoordinate>
,
<collectingDate id="9E1AE9C8FF9D2406FF5FBB4AFECA3C63" box="[182,301,1330,1355]" pageId="27" pageNumber="28" value="2008-07-01">
<date id="8E5E1020FF9D2406FF5FBB4AFECA3C63" box="[182,301,1330,1355]" pageId="27" pageNumber="28" value="2008-07-01">1 Jul 2008</date>
</collectingDate>
,
<collectorName id="57155336FF9D2406FED0BB4AFE363C63" box="[313,465,1330,1355]" pageId="27" pageNumber="28">R.J. Fensham</collectorName>
&amp;
<collectorName id="57155336FF9D2406FE1CBB4AFD8D3C63" box="[501,618,1330,1355]" pageId="27" pageNumber="28">R. Fairfax</collectorName>
, C.462094, 10+;
<geoCoordinate id="9FD45027FF9D2406FCC5BB4AFC253C63" box="[812,962,1331,1355]" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="15" value="-22.790833">22° 47' 27&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF9D2406FC26BB4BFB923C62" box="[975,1141,1330,1355]" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="15" value="145.44084">145° 26' 27&quot; E</geoCoordinate>
,
<collectingDate id="9E1AE9C8FF9D2406FB68BB4AFB1E3C63" box="[1153,1273,1330,1355]" pageId="27" pageNumber="28" value="2008-07-01">
<date id="8E5E1020FF9D2406FB68BB4AFB1E3C63" box="[1153,1273,1330,1355]" pageId="27" pageNumber="28" value="2008-07-01">1 Jul 2008</date>
</collectingDate>
,
<collectorName id="57155336FF9D2406FAEDBB4AFA7B3C63" box="[1284,1436,1330,1355]" pageId="27" pageNumber="28">R.J. Fensham</collectorName>
&amp;
<collectorName id="57155336FF9D2406FF5DBB2EFECF3C47" box="[180,296,1366,1391]" pageId="27" pageNumber="28">R. Fairfax</collectorName>
, C.462092, 20+.
<collectorName id="57155336FF9D2406FE01BB2FFDCF3C47" box="[488,552,1366,1391]" pageId="27" pageNumber="28">Other</collectorName>
material from the same locality
</materialsCitation>
: C.483045 12; C.483114, 20+; C.483112, 20+; C.483107, 20+; C.483110, 5; C.483113, 20+; C.483111, 5; C.483044, 20+; C.483108, 20+; C.483109, 3, C.505944,20+; C.505941,1; C.505949, 20+; C.505945, 20+, C.505951, 20+, C.505943, 1; C.505950, 12; C.505948, 5; C.505942, 13; C.505946, 5.
</paragraph>
<paragraph id="FA5F36E0FF9D2406FF2EBB9FFC0A3FA7" blockId="27.[151,1437,1186,1967]" pageId="27" pageNumber="28">
<emphasis id="C894EAF2FF9D2406FF2EBB9FFEE43CD7" bold="true" box="[199,259,1510,1535]" pageId="27" pageNumber="28">Shell</emphasis>
(
<figureCitation id="62DB2A65FF9D2406FEFDBB9FFE993CD7" box="[276,382,1510,1535]" captionStart="FIGURE 13" captionStartId="28.[151,250,1847,1869]" captionTargetBox="[275,1320,252,1754]" captionTargetId="figure@28.[214,1372,223,1786]" captionTargetPageId="28" captionText="FIGURE 13. Shells of species of Edgbastonia (Barcaldonia) n. subgen. A. Edgbastonia (B.) hufferensis n. sp., holotype (C.479951). B. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990) (C. 479947). C, D. Edgbastonia (B.) routhensis n. sp., Routh Stn, C, holotype (C.479960). D. paratype (C.448850). E, Edgbastonia (B.) pagoda n. sp., holotype, Myross Stn (C.479948). F. Edgbastonia (B.) rugosa n. sp., holotype, Myross Stn (C.479949). G. Edgbastonia (B.) corrugata umbilicata n. sp., holotype, Myross Stn (C.479946). H. Edgbastonia? conjuboyensis, holotype, Conjuboy Stn (C.479956). I. Edgbastonia (B.) chillagoensis, holotype, Chillagoe Stn (C.479958). Scale bar 1 mm." httpUri="https://zenodo.org/record/2637629/files/figure.png" pageId="27" pageNumber="28">Fig. 13G</figureCitation>
). Ovate, spire outline moderately convex, normally coiled, translucent. Length
<quantity id="3D189B05FF9D2406FAFABB9FFA7A3CD6" box="[1299,1437,1510,1535]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="27" pageNumber="28" unit="mm" value="2.4" valueMax="2.6" valueMin="2.2">2.2–2.6 mm</quantity>
(mean
<quantity id="3D189B05FF9D2406FF0DB872FEDA3F0B" box="[228,317,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="27" pageNumber="28" unit="mm" value="2.4">2.4 mm</quantity>
), width
<quantity id="3D189B05FF9D2406FE7EB873FDF93F0A" box="[407,542,1546,1571]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="27" pageNumber="28" unit="mm" value="2.15" valueMax="2.5" valueMin="1.8">1.8–2.5 mm</quantity>
(mean
<quantity id="3D189B05FF9D2406FD98B872FD2C3F0B" box="[625,715,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="27" pageNumber="28" unit="mm" value="2.1">2.1 mm</quantity>
). Protoconch about 1.2–1.3 whorls, distinctly pitted. Teleoconch whorls weakly convex, evenly rounded, total number 3.2–4.3 (mean 3.8). Umbilicus moderately wide. Aperture ovate, inner lip narrow, thin or medium, in partial contact to slightly separated from parietal wall, outer lip thin or medium. Periostracum moderately developed, pale yellow or yellow-brown.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF9D2403FF2EB8E3FB363D65" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28" type="description">
<paragraph id="FA5F36E0FF9D2406FF2EB8E3FA913F9B" blockId="27.[151,1437,1186,1967]" box="[199,1398,1690,1715]" pageId="27" pageNumber="28">
<emphasis id="C894EAF2FF9D2406FF2EB8E3FEA83F9B" bold="true" box="[199,335,1690,1715]" pageId="27" pageNumber="28">Operculum</emphasis>
(
<figureCitation id="62DB2A65FF9D2406FEB7B8E3FE343F9B" box="[350,467,1690,1715]" captionStart="FIGURE 14" captionStartId="29.[151,250,1810,1832]" captionTargetBox="[218,1365,209,1793]" captionTargetId="figure@29.[200,1397,193,1801]" captionTargetPageId="29" captionText="FIGURE 14. Opercula of species of Edgbastonia (Barcaldonia) n. subgen. A. Edgbastonia (B.) hufferensis n. sp., internal view (C.423238). B, C. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990). B. External view. C. Internal view (C. 457755). D, E. Edgbastonia (B.) routhensis n. sp. D. External view. E. Internal view (C438551). F, G. Edgbastonia (B.) pagoda n. sp. F. External view. G. Internal view (C.457756). H. Edgbastonia (B.) rugosa n. sp., internal view (C.457753). I, J. Edgbastonia (B.) corrugata umbilicata n. subsp. I. External view. J. Internal view (C.415845). K, L. Edgbastonia? conjuboyensis n. sp. K. External view. L. Internal view (C.445416). M, N. Edgbastonia (B.) chillagoensis n. sp. M. External view. N. Internal view (C.438557). Scale bar 200 µm." httpUri="https://zenodo.org/record/2637631/files/figure.png" pageId="27" pageNumber="28">
Fig.
<date id="8E5E1020FF9D2406FE7AB8E2FE5D3F9B" box="[403,442,1691,1715]" pageId="27" pageNumber="28" value="2014-01">14I</date>
, J
</figureCitation>
). Transparent, colourless, slightly concave. Inner side lacking white smear, simple.
</paragraph>
<paragraph id="FA5F36E0FF9D2406FF2EB8C7FA8B3FD3" blockId="27.[151,1437,1186,1967]" pageId="27" pageNumber="28">
<emphasis id="C894EAF2FF9D2406FF2EB8C7FDCF3FFF" bold="true" box="[199,552,1726,1751]" pageId="27" pageNumber="28">Head-foot and external body.</emphasis>
Snout, tentacles unpigmented or pigmented, neck, dorsal and lateral foot and opercular lobes unpigmented, mantle roof unpigmented or weakly pigmented, visceral coil weakly pigmented.
</paragraph>
<paragraph id="FA5F36E0FF9D2406FF2EB97FFAEE3E87" blockId="27.[151,1437,1186,1967]" pageId="27" pageNumber="28">
<emphasis id="C894EAF2FF9D2406FF2EB97FFE953E37" bold="true" box="[199,370,1798,1823]" pageId="27" pageNumber="28">Mantle cavity.</emphasis>
Ctenidium well-developed, filaments 24–28, broadly triangular, apex right edge. Osphradium narrowly oval, towards posterior end of ctenidium, length relative to gill 0.22–0.31. Hypobranchial gland moderately developed. Rectum with U-shaped bend, faecal pellets longitudinally orientated or orientated obliquely or sideways, anus at mantle collar or a little behind it. Kidney half or more in roof of mantle cavity. Renal gland transverse. Pericardium half or more in roof of mantle cavity, overlapping posterior end of ctenidium.
</paragraph>
<caption id="AE9F6668FF9A2401FF7EB94EFC963ECF" httpUri="https://zenodo.org/record/2637629/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1847,1869]" targetBox="[275,1320,252,1754]" targetPageId="28">
<paragraph id="FA5F36E0FF9A2401FF7EB94EFC963ECF" blockId="28.[151,1436,1847,2023]" pageId="28" pageNumber="29">
<emphasis id="C894EAF2FF9A2401FF7EB94EFEC43E65" bold="true" box="[151,291,1847,1869]" pageId="28" pageNumber="29">FIGURE 13.</emphasis>
Shells of species of
<taxonomicName id="3DE04D63FF9A2401FDECB94EFCFA3E65" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[517,797,1847,1869]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="subGenus" status="subgen. nov." subGenus="Barcaldonia">
<emphasis id="C894EAF2FF9A2401FDECB94EFD623E65" box="[517,645,1847,1869]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9A2401FD7EB94EFCF23E65" box="[663,789,1847,1869]" italics="true" pageId="28" pageNumber="29">Barcaldonia</emphasis>
)
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9A2401FCCEB94EFC7F3E65" box="[807,920,1847,1869]" pageId="28" pageNumber="29" rank="subGenus">
<emphasis id="C894EAF2FF9A2401FCCEB94EFC7F3E65" bold="true" box="[807,920,1847,1869]" pageId="28" pageNumber="29">n. subgen.</emphasis>
</taxonomicNameLabel>
A.
<taxonomicName id="3DE04D63FF9A2401FC2DB94EFB0A3E65" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[964,1261,1847,1870]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="hufferensis" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9A2401FC2DB94EFBA33E65" box="[964,1092,1847,1869]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9A2401FBBFB940FB8A3E66" box="[1110,1133,1849,1870]" italics="true" pageId="28" pageNumber="29">B.</emphasis>
)
<emphasis id="C894EAF2FF9A2401FB96B94EFB0A3E65" box="[1151,1261,1847,1869]" italics="true" pageId="28" pageNumber="29">hufferensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9A2401FB1FB94EFAD43E65" box="[1270,1331,1847,1869]" pageId="28" pageNumber="29" rank="species">
<emphasis id="C894EAF2FF9A2401FB1FB94EFAD43E65" bold="true" box="[1270,1331,1847,1869]" pageId="28" pageNumber="29">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype (C.479951). B.
<taxonomicName id="3DE04D63FF9A2401FED3B92FFCBB3E44" authority="(Ponder &amp; Clark, 1990)" authorityName="Ponder &amp; Clark" authorityYear="1990" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[314,860,1878,1900]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="pallida" subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9A2401FED3B92FFE5D3E44" box="[314,442,1878,1900]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9A2401FE22B92EFE063E44" box="[459,481,1879,1900]" italics="true" pageId="28" pageNumber="29">B.</emphasis>
) cf.
<emphasis id="C894EAF2FF9A2401FDFDB92FFDBB3E44" box="[532,604,1878,1900]" italics="true" pageId="28" pageNumber="29">pallida</emphasis>
(Ponder &amp; Clark, 1990)
</taxonomicName>
(C. 479947). C, D.
<taxonomicName id="3DE04D63FF9A2401FBD8B92FFAB53E44" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[1073,1362,1878,1900]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="routhensis" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9A2401FBD8B92FFB563E44" box="[1073,1201,1878,1900]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9A2401FB2BB92EFB3E3E44" box="[1218,1241,1879,1900]" italics="true" pageId="28" pageNumber="29">B.</emphasis>
)
<emphasis id="C894EAF2FF9A2401FB00B92FFAB53E44" box="[1257,1362,1878,1900]" italics="true" pageId="28" pageNumber="29">routhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9A2401FAB3B92FFA713E44" box="[1370,1430,1878,1900]" pageId="28" pageNumber="29" rank="species">
<emphasis id="C894EAF2FF9A2401FAB3B92FFA713E44" bold="true" box="[1370,1430,1878,1900]" pageId="28" pageNumber="29">n. sp.</emphasis>
</taxonomicNameLabel>
, Routh Stn, C, holotype (C.479960). D. paratype (C.448850). E,
<taxonomicName id="3DE04D63FF9A2401FCB8B90CFBBC3EA3" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[849,1115,1909,1931]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="pagoda" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9A2401FCB8B90CFC363EA3" box="[849,977,1909,1931]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9A2401FC0CB90FFC1B3EA3" box="[997,1020,1910,1931]" italics="true" pageId="28" pageNumber="29">B.</emphasis>
)
<emphasis id="C894EAF2FF9A2401FBE6B90CFBBC3EA3" box="[1039,1115,1909,1931]" italics="true" pageId="28" pageNumber="29">pagoda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9A2401FB8EB90CFB413EA3" box="[1127,1190,1909,1931]" pageId="28" pageNumber="29" rank="species">
<emphasis id="C894EAF2FF9A2401FB8EB90CFB413EA3" bold="true" box="[1127,1190,1909,1931]" pageId="28" pageNumber="29">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, Myross Stn (C.479948). F.
<taxonomicName id="3DE04D63FF9A2401FED9B9EDFDCD3E82" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[304,554,1940,1962]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="rugosa" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9A2401FED9B9EDFE573E82" box="[304,432,1940,1962]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9A2401FE29B9ECFE313E82" box="[448,470,1941,1962]" italics="true" pageId="28" pageNumber="29">B.</emphasis>
)
<emphasis id="C894EAF2FF9A2401FE0DB9ECFDCD3E82" box="[484,554,1941,1962]" italics="true" pageId="28" pageNumber="29">rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9A2401FDDBB9EDFD8C3E82" box="[562,619,1940,1962]" pageId="28" pageNumber="29" rank="species">
<emphasis id="C894EAF2FF9A2401FDDBB9EDFD8C3E82" bold="true" box="[562,619,1940,1962]" pageId="28" pageNumber="29">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, Myross Stn (C.479949). G.
<taxonomicName id="3DE04D63FF9A2401FC1CB9EDFA653E82" authorityName="Zhang" authorityYear="2019" box="[1013,1410,1940,1962]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="subSpecies" species="corrugata" subGenus="Barcaldinia" subSpecies="umbilicata">
<emphasis id="C894EAF2FF9A2401FC1CB9EDFB923E82" box="[1013,1141,1940,1962]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9A2401FB6DB9ECFB7D3E82" box="[1156,1178,1941,1962]" italics="true" pageId="28" pageNumber="29">B.</emphasis>
)
<emphasis id="C894EAF2FF9A2401FB40B9ECFA653E82" box="[1193,1410,1940,1962]" italics="true" pageId="28" pageNumber="29">corrugata umbilicata</emphasis>
</taxonomicName>
<emphasis id="C894EAF2FF9A2401FA6EB9EDFF523EE0" bold="true" pageId="28" pageNumber="29">n. sp.</emphasis>
, holotype, Myross Stn (C.479946). H.
<taxonomicName id="3DE04D63FF9A2401FDB9B9CBFD373EE0" box="[592,720,1970,1992]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus">
<emphasis id="C894EAF2FF9A2401FDB9B9CBFD373EE0" box="[592,720,1970,1992]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
</taxonomicName>
?
<emphasis id="C894EAF2FF9A2401FD0DB9CBFC943EE0" box="[740,883,1970,1992]" italics="true" pageId="28" pageNumber="29">conjuboyensis</emphasis>
, holotype, Conjuboy Stn (C.479956). I.
<taxonomicName id="3DE04D63FF9A2401FAF5B9CBFEAE3ECF" authorityName="Zhang" authorityYear="2019" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="chillagoensis" subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9A2401FAF5B9CBFA7B3EE0" box="[1308,1436,1970,1992]" italics="true" pageId="28" pageNumber="29">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9A2401FF76B9ABFF523ECF" box="[159,181,2002,2023]" italics="true" pageId="28" pageNumber="29">B.</emphasis>
)
<emphasis id="C894EAF2FF9A2401FF2DB9A8FEAE3ECF" box="[196,329,2001,2023]" italics="true" pageId="28" pageNumber="29">chillagoensis</emphasis>
</taxonomicName>
, holotype, Chillagoe Stn (C.479958). Scale bar 1 mm.
</paragraph>
</caption>
<caption id="AE9F6668FF9B2400FF7EB96BFE283ED9" httpUri="https://zenodo.org/record/2637631/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1810,1832]" targetBox="[218,1365,209,1793]" targetPageId="29">
<paragraph id="FA5F36E0FF9B2400FF7EB96BFE283ED9" blockId="29.[151,1437,1810,2033]" pageId="29" pageNumber="30">
<emphasis id="C894EAF2FF9B2400FF7EB96BFEC53E00" bold="true" box="[151,290,1810,1832]" pageId="29" pageNumber="30">FIGURE 14.</emphasis>
Opercula of species of
<taxonomicName id="3DE04D63FF9B2400FDF4B96BFCD43E00" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[541,819,1810,1832]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="subGenus" status="subgen. nov." subGenus="Barcaldonia">
<emphasis id="C894EAF2FF9B2400FDF4B96BFD7A3E00" box="[541,669,1810,1832]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9B2400FD47B96BFCCC3E00" box="[686,811,1810,1832]" italics="true" pageId="29" pageNumber="30">Barcaldonia</emphasis>
)
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9B2400FCD5B96BFC4C3E00" box="[828,939,1810,1832]" pageId="29" pageNumber="30" rank="subGenus">
<emphasis id="C894EAF2FF9B2400FCD5B96BFC4C3E00" bold="true" box="[828,939,1810,1832]" pageId="29" pageNumber="30">n. subgen.</emphasis>
</taxonomicNameLabel>
A.
<taxonomicName id="3DE04D63FF9B2400FC3CB96BFB1C3E00" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[981,1275,1810,1832]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="hufferensis" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9B2400FC3CB96BFBB23E00" box="[981,1109,1810,1832]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9B2400FB8EB96AFB9A3E00" box="[1127,1149,1811,1832]" italics="true" pageId="29" pageNumber="30">B.</emphasis>
)
<emphasis id="C894EAF2FF9B2400FB64B96BFB1C3E00" box="[1165,1275,1810,1832]" italics="true" pageId="29" pageNumber="30">hufferensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9B2400FAEDB96BFAA73E00" box="[1284,1344,1810,1832]" pageId="29" pageNumber="30" rank="species">
<emphasis id="C894EAF2FF9B2400FAEDB96BFAA73E00" bold="true" box="[1284,1344,1810,1832]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
, internal view (C.423238). B, C.
<taxonomicName id="3DE04D63FF9B2400FE6EB94AFC7F3E61" authority="(Ponder &amp; Clark, 1990)" authorityName="Ponder &amp; Clark" authorityYear="1990" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[391,920,1843,1865]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="pallida" subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9B2400FE6EB94AFDE03E61" box="[391,519,1843,1865]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9B2400FDFFB94DFDCB3E61" box="[534,556,1844,1865]" italics="true" pageId="29" pageNumber="30">B.</emphasis>
) cf.
<emphasis id="C894EAF2FF9B2400FDB0B94AFD463E61" box="[601,673,1843,1865]" italics="true" pageId="29" pageNumber="30">pallida</emphasis>
(Ponder &amp; Clark, 1990)
</taxonomicName>
. B. External view. C. Internal view (C. 457755). D, E.
<taxonomicName id="3DE04D63FF9B2400FF5DB92DFE333E42" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[180,468,1876,1898]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="routhensis" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9B2400FF5DB92DFED33E42" box="[180,308,1876,1898]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9B2400FEACB92CFEBC3E42" box="[325,347,1877,1898]" italics="true" pageId="29" pageNumber="30">B.</emphasis>
)
<emphasis id="C894EAF2FF9B2400FE82B92DFE333E42" box="[363,468,1876,1898]" italics="true" pageId="29" pageNumber="30">routhensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9B2400FE35B92DFDF03E42" box="[476,535,1876,1898]" pageId="29" pageNumber="30" rank="species">
<emphasis id="C894EAF2FF9B2400FE35B92DFDF03E42" bold="true" box="[476,535,1876,1898]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
D. External view. E. Internal view (C438551). F, G.
<taxonomicName id="3DE04D63FF9B2400FBD5B92DFAD83E42" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[1084,1343,1876,1898]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="pagoda" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9B2400FBD5B92DFB5B3E42" box="[1084,1212,1876,1898]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9B2400FB24B92CFB043E42" box="[1229,1251,1877,1898]" italics="true" pageId="29" pageNumber="30">B.</emphasis>
)
<emphasis id="C894EAF2FF9B2400FB1AB92DFAD83E42" box="[1267,1343,1876,1898]" italics="true" pageId="29" pageNumber="30">pagoda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9B2400FAAEB92DFA653E42" box="[1351,1410,1876,1898]" pageId="29" pageNumber="30" rank="species">
<emphasis id="C894EAF2FF9B2400FAAEB92DFA653E42" bold="true" box="[1351,1410,1876,1898]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
F. External view. G. Internal view (C.457756). H.
<taxonomicName id="3DE04D63FF9B2400FD9CB90CFC893EA4" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[629,878,1909,1932]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="rugosa" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9B2400FD9CB90CFD123EA3" box="[629,757,1909,1931]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9B2400FCEDB90EFCFD3EA4" box="[772,794,1911,1932]" italics="true" pageId="29" pageNumber="30">B.</emphasis>
)
<emphasis id="C894EAF2FF9B2400FCC1B90EFC893EA4" box="[808,878,1911,1932]" italics="true" pageId="29" pageNumber="30">rugosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9B2400FC9DB90CFC4A3EA3" box="[884,941,1909,1931]" pageId="29" pageNumber="30" rank="species">
<emphasis id="C894EAF2FF9B2400FC9DB90CFC4A3EA3" bold="true" box="[884,941,1909,1931]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
, internal view (C.457753). I, J.
<taxonomicName id="3DE04D63FF9B2400FB06B90CFE963E85" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="subSpecies" species="corrugata" status="subsp. nov." subGenus="Barcaldinia" subSpecies="umbilicata">
<emphasis id="C894EAF2FF9B2400FB06B90CFA883EA3" box="[1263,1391,1909,1931]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9B2400FA97B90EFA733EA4" box="[1406,1428,1911,1932]" italics="true" pageId="29" pageNumber="30">B.</emphasis>
)
<emphasis id="C894EAF2FF9B2400FF7EB9E1FE963E85" box="[151,369,1943,1965]" italics="true" pageId="29" pageNumber="30">corrugata umbilicata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9B2400FE92B9EEFE3A3E85" box="[379,477,1943,1965]" pageId="29" pageNumber="30" rank="subSpecies">
<emphasis id="C894EAF2FF9B2400FE92B9EEFE3A3E85" bold="true" box="[379,477,1943,1965]" pageId="29" pageNumber="30">n. subsp.</emphasis>
</taxonomicNameLabel>
I. External view. J. Internal view (C.415845). K, L.
<taxonomicName id="3DE04D63FF9B2400FBF9B9EEFB773E85" box="[1040,1168,1943,1965]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="C894EAF2FF9B2400FBF9B9EEFB773E85" box="[1040,1168,1943,1965]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
</taxonomicName>
?
<emphasis id="C894EAF2FF9B2400FB4DB9EEFAD43E85" box="[1188,1331,1943,1965]" italics="true" pageId="29" pageNumber="30">conjuboyensis</emphasis>
<emphasis id="C894EAF2FF9B2400FAD4B9EEFA9D3E85" bold="true" box="[1341,1402,1943,1965]" pageId="29" pageNumber="30">n. sp.</emphasis>
K. External view. L. Internal view (C.445416). M, N.
<taxonomicName id="3DE04D63FF9B2400FD43B9C1FC013EE6" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[682,998,1976,1999]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="chillagoensis" status="sp. nov." subGenus="Barcaldinia">
<emphasis id="C894EAF2FF9B2400FD43B9C1FCCD3EE6" box="[682,810,1976,1998]" italics="true" pageId="29" pageNumber="30">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF9B2400FCD3B9C3FCB63EE7" box="[826,849,1978,1999]" italics="true" pageId="29" pageNumber="30">B.</emphasis>
)
<emphasis id="C894EAF2FF9B2400FC88B9C1FC013EE6" box="[865,998,1976,1998]" italics="true" pageId="29" pageNumber="30">chillagoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF9B2400FC06B9C1FBCD3EE6" box="[1007,1066,1976,1998]" pageId="29" pageNumber="30" rank="species">
<emphasis id="C894EAF2FF9B2400FC06B9C1FBCD3EE6" bold="true" box="[1007,1066,1976,1998]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
M. External view. N. Internal view (C.438557). Scale bar 200 µm.
</paragraph>
</caption>
<caption id="AE9F6668FF982403FF7EBEE1FC9F39E7" ID-Table-UUID="AE9F6668FF982403FF7EBEE1FC9F39E7" httpUri="http://table.plazi.org/id/AE9F6668FF982403FF7EBEE1FC9F39E7" pageId="30" pageNumber="31" targetBox="[159,1315,234,504]" targetIsTable="true" targetPageId="30">
<paragraph id="FA5F36E0FF982403FF7EBEE1FC9F39E7" blockId="30.[151,1437,151,207]" pageId="30" pageNumber="31">
<emphasis id="C894EAF2FF982403FF7EBEE1FEEB3987" bold="true" box="[151,268,152,175]" pageId="30" pageNumber="31">TABLE 9.</emphasis>
Shell measurements (in mm) of type material of
<taxonomicName id="3DE04D63FF982403FCCEBEEEFAD33987" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[807,1332,151,175]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" status="subsp. nov." subGenus="Barcaldinia" subSpecies="umbilicata">
<emphasis id="C894EAF2FF982403FCCEBEEEFC493987" box="[807,942,151,175]" italics="true" pageId="30" pageNumber="31">Edgbastonia</emphasis>
(
<emphasis id="C894EAF2FF982403FC56BEEEFBDA3987" box="[959,1085,151,175]" italics="true" pageId="30" pageNumber="31">Barcaldinia</emphasis>
)
<emphasis id="C894EAF2FF982403FBA6BEE1FAD33987" box="[1103,1332,151,175]" italics="true" pageId="30" pageNumber="31">corrugata umbilicata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF982403FAD5BEE1FA7A3987" box="[1340,1437,152,175]" pageId="30" pageNumber="31" rank="subSpecies">n. subsp.</taxonomicNameLabel>
(paratypes, C.415845). See Materials and Methods for abbreviations.
</paragraph>
</caption>
<paragraph id="FA5F36E0FF982403FE1BBE93FAC438D0" pageId="30" pageNumber="31">
<table id="88E0C440FF98DBE2FF76BE93FAC438D0" box="[159,1315,234,504]" gridcols="7" gridrows="7" pageId="30" pageNumber="31">
<tr id="44D034A2FF98DBE2FF76BE93FAC43828" box="[159,1315,234,256]" gridrow="0" pageId="30" pageNumber="31" rowspan-0="1">
<th id="07015DDEFF98DBE2FE1BBE93FDF93828" box="[498,542,234,256]" gridcol="1" gridrow="0" pageId="30" pageNumber="31">SH</th>
<th id="07015DDEFF98DBE2FD8ABE93FD683828" box="[611,655,234,256]" gridcol="2" gridrow="0" pageId="30" pageNumber="31">SW</th>
<th id="07015DDEFF98DBE2FCE8BE93FCCA3828" box="[769,813,234,256]" gridcol="3" gridrow="0" pageId="30" pageNumber="31">AH</th>
<th id="07015DDEFF98DBE2FC71BE93FC233828" box="[920,964,234,256]" gridcol="4" gridrow="0" pageId="30" pageNumber="31">AW</th>
<th id="07015DDEFF98DBE2FBAEBE93FB943828" box="[1095,1139,234,256]" gridcol="5" gridrow="0" pageId="30" pageNumber="31">BW</th>
<th id="07015DDEFF98DBE2FB1EBE93FAC43828" box="[1271,1315,234,256]" gridcol="6" gridrow="0" pageId="30" pageNumber="31">TW</th>
</tr>
<tr id="44D034A2FF98DBE2FF76BF6DFAC43802" box="[159,1315,276,298]" gridrow="1" pageId="30" pageNumber="31">
<th id="07015DDEFF98DBE2FF76BF6DFF1A3802" box="[159,253,276,298]" gridcol="0" gridrow="1" pageId="30" pageNumber="31">Holotype</th>
<td id="07015DDEFF98DBE2FE1BBF6DFDF93802" box="[498,542,276,298]" gridcol="1" gridrow="1" pageId="30" pageNumber="31">3.75</td>
<td id="07015DDEFF98DBE2FD8ABF6DFD683802" box="[611,655,276,298]" gridcol="2" gridrow="1" pageId="30" pageNumber="31">2.79</td>
<td id="07015DDEFF98DBE2FCE8BF6DFCCA3802" box="[769,813,276,298]" gridcol="3" gridrow="1" pageId="30" pageNumber="31">1.95</td>
<td id="07015DDEFF98DBE2FC71BF6DFC233802" box="[920,964,276,298]" gridcol="4" gridrow="1" pageId="30" pageNumber="31">1.36</td>
<td id="07015DDEFF98DBE2FBAEBF6DFB943802" box="[1095,1139,276,298]" gridcol="5" gridrow="1" pageId="30" pageNumber="31">3.14</td>
<td id="07015DDEFF98DBE2FB1EBF6DFAC43802" box="[1271,1315,276,298]" gridcol="6" gridrow="1" pageId="30" pageNumber="31">4</td>
</tr>
<tr id="44D034A2FF98DBE2FF76BF44FAC4387B" box="[159,1315,317,339]" gridrow="2" pageId="30" pageNumber="31">
<th id="07015DDEFF98DBE2FF76BF44FAC4387B" box="[159,1315,317,339]" colspan="7" colspanRight="6" gridcol="0" gridrow="2" pageId="30" pageNumber="31">Paratypes (40)</th>
</tr>
<tr id="44D034A2FF98DBE2FF76BF1FFAC43854" box="[159,1315,358,380]" gridrow="3" pageId="30" pageNumber="31">
<th id="07015DDEFF98DBE2FF76BF1FFF1A3854" box="[159,253,358,380]" gridcol="0" gridrow="3" pageId="30" pageNumber="31">Min</th>
<td id="07015DDEFF98DBE2FE1BBF1FFDF93854" box="[498,542,358,380]" gridcol="1" gridrow="3" pageId="30" pageNumber="31">2.18</td>
<td id="07015DDEFF98DBE2FD8ABF1FFD683854" box="[611,655,358,380]" gridcol="2" gridrow="3" pageId="30" pageNumber="31">1.78</td>
<td id="07015DDEFF98DBE2FCE8BF1FFCCA3854" box="[769,813,358,380]" gridcol="3" gridrow="3" pageId="30" pageNumber="31">1.23</td>
<td id="07015DDEFF98DBE2FC71BF1FFC233854" box="[920,964,358,380]" gridcol="4" gridrow="3" pageId="30" pageNumber="31">1.05</td>
<td id="07015DDEFF98DBE2FBAEBF1FFB943854" box="[1095,1139,358,380]" gridcol="5" gridrow="3" pageId="30" pageNumber="31">1.92</td>
<td id="07015DDEFF98DBE2FB1EBF1FFAC43854" box="[1271,1315,358,380]" gridcol="6" gridrow="3" pageId="30" pageNumber="31">3.20</td>
</tr>
<tr id="44D034A2FF98DBE2FF76BFE9FAC4388E" box="[159,1315,400,422]" gridrow="4" pageId="30" pageNumber="31">
<th id="07015DDEFF98DBE2FF76BFE9FF1A388E" box="[159,253,400,422]" gridcol="0" gridrow="4" pageId="30" pageNumber="31">Max</th>
<td id="07015DDEFF98DBE2FE1BBFE9FDF9388E" box="[498,542,400,422]" gridcol="1" gridrow="4" pageId="30" pageNumber="31">3.51</td>
<td id="07015DDEFF98DBE2FD8ABFE9FD68388E" box="[611,655,400,422]" gridcol="2" gridrow="4" pageId="30" pageNumber="31">3.06</td>
<td id="07015DDEFF98DBE2FCE8BFE9FCCA388E" box="[769,813,400,422]" gridcol="3" gridrow="4" pageId="30" pageNumber="31">2.12</td>
<td id="07015DDEFF98DBE2FC71BFE9FC23388E" box="[920,964,400,422]" gridcol="4" gridrow="4" pageId="30" pageNumber="31">1.66</td>
<td id="07015DDEFF98DBE2FBAEBFE9FB94388E" box="[1095,1139,400,422]" gridcol="5" gridrow="4" pageId="30" pageNumber="31">3.07</td>
<td id="07015DDEFF98DBE2FB1EBFE9FAC4388E" box="[1271,1315,400,422]" gridcol="6" gridrow="4" pageId="30" pageNumber="31">4.38</td>
</tr>
<tr id="44D034A2FF98DBE2FF76BFC0FAC438E7" box="[159,1315,441,463]" gridrow="5" pageId="30" pageNumber="31">
<th id="07015DDEFF98DBE2FF76BFC0FF1A38E7" box="[159,253,441,463]" gridcol="0" gridrow="5" pageId="30" pageNumber="31">Mean</th>
<td id="07015DDEFF98DBE2FE1BBFC0FDF938E7" box="[498,542,441,463]" gridcol="1" gridrow="5" pageId="30" pageNumber="31">2.46</td>
<td id="07015DDEFF98DBE2FD8ABFC0FD6838E7" box="[611,655,441,463]" gridcol="2" gridrow="5" pageId="30" pageNumber="31">2.10</td>
<td id="07015DDEFF98DBE2FCE8BFC0FCCA38E7" box="[769,813,441,463]" gridcol="3" gridrow="5" pageId="30" pageNumber="31">1.46</td>
<td id="07015DDEFF98DBE2FC71BFC0FC2338E7" box="[920,964,441,463]" gridcol="4" gridrow="5" pageId="30" pageNumber="31">1.21</td>
<td id="07015DDEFF98DBE2FBAEBFC0FB9438E7" box="[1095,1139,441,463]" gridcol="5" gridrow="5" pageId="30" pageNumber="31">2.20</td>
<td id="07015DDEFF98DBE2FB1EBFC0FAC438E7" box="[1271,1315,441,463]" gridcol="6" gridrow="5" pageId="30" pageNumber="31">3.87</td>
</tr>
<tr id="44D034A2FF98DBE2FF76BF9BFAC438D0" box="[159,1315,482,504]" gridrow="6" pageId="30" pageNumber="31">
<th id="07015DDEFF98DBE2FF76BF9BFF1A38D0" box="[159,253,482,504]" gridcol="0" gridrow="6" pageId="30" pageNumber="31">SD</th>
<td id="07015DDEFF98DBE2FE1BBF9BFDF938D0" box="[498,542,482,504]" gridcol="1" gridrow="6" pageId="30" pageNumber="31">0.23</td>
<td id="07015DDEFF98DBE2FD8ABF9BFD6838D0" box="[611,655,482,504]" gridcol="2" gridrow="6" pageId="30" pageNumber="31">0.21</td>
<td id="07015DDEFF98DBE2FCE8BF9BFCCA38D0" box="[769,813,482,504]" gridcol="3" gridrow="6" pageId="30" pageNumber="31">0.14</td>
<td id="07015DDEFF98DBE2FC71BF9BFC2338D0" box="[920,964,482,504]" gridcol="4" gridrow="6" pageId="30" pageNumber="31">0.11</td>
<td id="07015DDEFF98DBE2FBAEBF9BFB9438D0" box="[1095,1139,482,504]" gridcol="5" gridrow="6" pageId="30" pageNumber="31">0.20</td>
<td id="07015DDEFF98DBE2FB1EBF9BFAC438D0" box="[1271,1315,482,504]" gridcol="6" gridrow="6" pageId="30" pageNumber="31">0.23</td>
</tr>
</table>
</paragraph>
<paragraph id="FA5F36E0FF982403FF2EBC18FD7A3BCD" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">
<emphasis id="C894EAF2FF982403FF2EBC18FEF93B52" bold="true" box="[199,286,609,634]" pageId="30" pageNumber="31">Radula</emphasis>
(
<figureCitation id="62DB2A65FF982403FEC4BC18FE523B52" box="[301,437,609,634]" captionStart="FIGURE 19" captionStartId="36.[151,250,1281,1303]" captionTargetBox="[168,1402,212,1252]" captionTargetId="figure@36.[151,1435,193,1280]" captionTargetPageId="36" captionText="FIGURE 19. Radulae of species of Edgbastonia (Barcaldonia) n. subgen. A, B. Edgbastonia (B.) rugosa n. sp. A. Half row. B. Lateral teeth (C.457753). C, D. Edgbastonia (B.) corrugata umbilicata n. subsp. C. Half row. D. Lateral and marginal teeth (C.415845). E, F. Edgbastonia? conjuboyensis n. sp. E. Half row. F. Marginal teeth (C.445416). G, H, I. Edgbastonia (B.) chillagoensis n. sp. G. Half row. H. Lateral and marginal teeth. I. Outer marginal teeth (C.438557). Scale bars 10 µm." httpUri="https://zenodo.org/record/2637641/files/figure.png" pageId="30" pageNumber="31">Fig. 19C, D</figureCitation>
). Central teeth with cusp formula 3–4+3–5, basal cusps 1+1+1; median cusp sharp, about twice as long as adjacent cusps. Lateral teeth with cusp formula 3–4+1+3–4; main cusp sharp, about twice as long as adjacent cusp on outer side, slightly longer than adjacent cusp on inner side. Inner marginal teeth with 15–19 cusps. Outer marginal teeth with 12–15 cusps.
</paragraph>
<paragraph id="FA5F36E0FF982403FF2EBC88FCD43AC9" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">
<emphasis id="C894EAF2FF982403FF2EBC88FDF03A22" bold="true" box="[199,535,753,778]" pageId="30" pageNumber="31">Female reproductive system</emphasis>
(
<figureCitation id="62DB2A65FF982403FDCFBC88FD4E3A22" box="[550,681,753,778]" captionStart="FIGURE 16" captionStartId="33.[151,250,1801,1823]" captionTargetBox="[150,1389,213,1728]" captionTargetId="figure@33.[150,1436,193,1779]" captionTargetPageId="33" captionText="FIGURE 16. Female reproductive system of species of Edgbastonia (Barcaldonia) n. subgen., shown from left side. A, B. Edgbastonia (B.) hufferensis n. sp. (C.423238). C, F. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990) (C.457755). D, E. Edgbastonia (B.) routhensis n. sp. (C.438551). G, H, Edgbastonia (B.) pagoda n. sp. (C.457756). I, L. Edgbastonia (B.) rugosa n. sp. (C.457753). J, K. Edgbastonia (B.) corrugata umbilicata n. subsp., (C.415845). M, N. Edgbastonia? conjuboyensis n. sp. (C.445416). O, P. Edgbastonia (B.) chillagoensis n. sp. (C.438557). Abbreviations: ag—albumen gland; bc—bursa copulatrix; cg—capsule gland; o—opening of oviduct; pw—posterior wall of mantle cavity; ro—renal oviduct; sr—seminal receptacle; vc—ventral channel. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637635/files/figure.png" pageId="30" pageNumber="31">Fig. 16J, K</figureCitation>
). Ovary weakly to strongly lobed. Renal oviduct with rather small, U-shaped arch. Seminal receptacle anterior to bursa, oval, duct intermediate; sometimes orientated sperm in renal oviduct and/or bursal duct. Bursa copulatrix overlapping albumen gland on left, round or triangular, shorter than albumen gland, bursal duct enters bursa antero-ventrally, bursal duct joins coiled oviduct little behind posterior mantle cavity wall. Albumen gland partly in mantle cavity. Capsule gland with two distinct glandular zones, thick in cross section, markedly indented by rectum. Anterior vestibule large, opening anterior to capsule gland, short, cowl and/or gutter associated with oviduct opening present.
</paragraph>
<paragraph id="FA5F36E0FF982403FF2EBD94FB363D65" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">
<emphasis id="C894EAF2FF982403FF2EBD94FE1D3D2E" bold="true" box="[199,506,1005,1030]" pageId="30" pageNumber="31">Male reproductive system</emphasis>
(
<figureCitation id="62DB2A65FF982403FDE0BD94FD943D2E" box="[521,627,1005,1030]" captionStart="FIGURE 17" captionStartId="34.[151,250,1813,1835]" captionTargetBox="[183,1412,762,1756]" captionTargetId="figure@34.[151,1436,741,1775]" captionTargetPageId="34" captionText="FIGURE 17. Prostate and vas deferens of species of Edgbastonia (Barcaldonia) n. subgen. A. Edgbastonia (B.) hufferensis n. sp. (C.423238). B. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990) (C.457755). C. Edgbastonia (B.) routhensis n. sp. (C.438551). D. Edgbastonia (B.) pagoda n. sp. (C.457756). E. Edgbastonia (B.) rugosa n. sp. (C.457753). F. Edgbastonia (B.) corrugata umbilicata n. subsp. (C.415845). G. Edgbastonia? conjuboyensis n. sp. (C. 445416). H. Edgbastonia (B.) chillagoensis n. sp. (C.438557). Abbreviations: avd—anterior vas deferens; pg—prostate gland; pvd—posterior vas deferens; pw—position of posterior wall of pallial cavity. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637637/files/figure.png" pageId="30" pageNumber="31">Figs. 17F</figureCitation>
,
<figureCitation id="62DB2A65FF982403FD97BD97FD553D2E" box="[638,690,1006,1030]" captionStart="FIGURE 18" captionStartId="35.[151,250,1854,1876]" captionTargetBox="[200,1347,213,1826]" captionTargetId="figure@35.[185,1406,193,1841]" captionTargetPageId="35" captionText="FIGURE 18. Penis and vas deferens of species of Edgbastonia (Barcaldonia) n. subgen. A. Edgbastonia (B.) hufferensis n. sp. (C.423238). B, C. Edgbastonia (B.) cf. pallida (Ponder &amp; Clark, 1990) (C.457755). D, Edgbastonia (B.) routhensis n. sp. (C.438551). E. Edgbastonia (B.) pagoda n. sp. (C.457756). F. Edgbastonia (B.) rugosa n. sp. (C.457753). G. Edgbastonia (B.) corrugata umbilicata n. subsp. H, I. Edgbastonia? conjuboyensis n. sp. (C.445416). J, K. Edgbastonia (B.) chilagoensis n. sp. (C.438557). Abbreviations: avd—anterior vas deferens; ngl—non-glandular lobe; p—papilla; pd—penial duct. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637639/files/figure.png" pageId="30" pageNumber="31">18G</figureCitation>
). Prostate gland less than half in mantle roof, bean-shaped, thick in cross section. Posterior pallial vas deferens coiled, anteriorly slightly undulating. Penis towards middle of head, intermediate, distal end tapering, terminal papilla absent, large non-glandular distal lobe present.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF982403FF2EBA20FF1B3DBE" pageId="30" pageNumber="31" type="etymology">
<paragraph id="FA5F36E0FF982403FF2EBA20FF1B3DBE" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">
<emphasis id="C894EAF2FF982403FF2EBA20FEA93D5A" bold="true" box="[199,334,1113,1138]" pageId="30" pageNumber="31">Etymology.</emphasis>
Umbilicus—Latin, navel. A term used in gastropods for a depression or perforation in the base of the shell.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF982403FF2EBAD8FF373C2A" pageId="30" pageNumber="31" type="distribution">
<paragraph id="FA5F36E0FF982403FF2EBAD8FF373C2A" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">
<emphasis id="C894EAF2FF982403FF2EBAD8FE1E3D92" bold="true" box="[199,505,1185,1210]" pageId="30" pageNumber="31">Distribution and habitat.</emphasis>
Known only from a large, shallow spring-fed pool on Myross Station (
<figureCitation id="62DB2A65FF982403FAACBAD8FA6C3D92" box="[1349,1419,1185,1210]" captionStart="FIGURE 1" captionStartId="4.[151,250,1844,1866]" captionTargetBox="[151,1435,498,1802]" captionTargetId="figure@4.[151,1435,498,1802]" captionTargetPageId="4" captionText="FIGURE 1. Map of the spring supergroups associated with the Great Artesian Basin (GAB) in Queensland, showing the main non-town locations mentioned in the text (squares) and some of the towns in the area (ovals). The pale grey shading is the discharge part of the GAB, the dark grey the recharge part. The inset map of Australia shows the area illustrated. The approximate location of Conondale is indicated by a triangle and Nundle by a star. The spring supergroups are outlined with dotted lines and are as follows: A, Mulligan River; B, Springvale; C, Flinders River; D, Hughenden; E, Barcaldine (see inset at upper right for details); F, Springsure; G, Eulo." httpUri="https://zenodo.org/record/2637605/files/figure.png" pageId="30" pageNumber="31">Fig. 1</figureCitation>
). Abundant, living in fine silt on the bottom of the pool. It was not found in vegetated areas around the edge of the pool.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF982403FF2EBB74FEEC3EFA" pageId="30" pageNumber="31" type="discussion">
<paragraph id="FA5F36E0FF982403FF2EBB74FCAB3CF1" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">
<emphasis id="C894EAF2FF982403FF2EBB74FEDC3C0E" bold="true" box="[199,315,1293,1318]" pageId="30" pageNumber="31">Remarks.</emphasis>
This subspecies is very similar to
<taxonomicName id="3DE04D63FF982403FD31BB74FBCE3C0D" authority="corrugata" authorityName="corrugata (Ponder &amp; Clark" authorityYear="1990" box="[728,1065,1293,1318]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="38" phylum="Mollusca" rank="subSpecies" species="corrugata" subGenus="Barcaldinia" subSpecies="corrugata">
<emphasis id="C894EAF2FF982403FD31BB74FD193C0E" box="[728,766,1293,1318]" italics="true" pageId="30" pageNumber="31">Ed.</emphasis>
(
<emphasis id="C894EAF2FF982403FCFBBB77FCC43C0D" box="[786,803,1294,1317]" italics="true" pageId="30" pageNumber="31">B</emphasis>
.)
<emphasis id="C894EAF2FF982403FCD7BB77FBCE3C0D" box="[830,1065,1294,1317]" italics="true" pageId="30" pageNumber="31">corrugata corrugata</emphasis>
</taxonomicName>
but differs in having less welldeveloped axial sculpture, and the inner lip is in contact with the parietal wall or only slightly separated from it, instead of being widely separated. The female genital system is very similar to that of
<taxonomicName id="3DE04D63FF982403FBBEBB2CFA7B3C45" authority="corrugata" authorityName="corrugata (Ponder &amp; Clark" authorityYear="1990" box="[1111,1436,1365,1390]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="38" phylum="Mollusca" rank="subSpecies" species="corrugata" subGenus="Barcaldinia" subSpecies="corrugata">
<emphasis id="C894EAF2FF982403FBBEBB2CFB993C46" box="[1111,1150,1365,1390]" italics="true" pageId="30" pageNumber="31">Ed.</emphasis>
(
<emphasis id="C894EAF2FF982403FB64BB2FFB793C45" box="[1165,1182,1366,1389]" italics="true" pageId="30" pageNumber="31">B</emphasis>
.)
<emphasis id="C894EAF2FF982403FB5CBB2FFA7B3C45" box="[1205,1436,1366,1389]" italics="true" pageId="30" pageNumber="31">corrugata corrugata</emphasis>
</taxonomicName>
but differs in having a smaller, shorter bursa copulatrix with a narrower duct. The central teeth of the radula in the new subspecies are unusual in having only a single pair of basal cusps developed (cf.,
<bibRefCitation id="9E714B11FF982403FB70BBE4FA7F3C9E" author="Ponder, W. F. &amp; Clark, G. A." box="[1177,1432,1437,1462]" pageId="30" pageNumber="31" pagination="301 - 363" refId="ref35763" refString="Ponder, W. F. &amp; Clark, G. A. (1990) A radiation of hydrobiid snails in threatened artesian springs in Western Queensland. Records of the Australian Museum, 42, 301 - 363. https: // doi. org / 10.3853 / j. 0067 - 1975.42.1990.119" type="journal article" year="1990">Ponder &amp; Clark 1990</bibRefCitation>
, fig.8D, E), although a rudimentary second cusp is rarely seen.
</paragraph>
<paragraph id="FA5F36E0FF982403FF2EBB9FFBE23FD2" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">
A discriminant function analysis using five shell measurements of the four large globular species,
<taxonomicName id="3DE04D63FF982403FA9FBB9CFE673F09" authority="corrugata" authorityName="corrugata (Ponder &amp; Clark" authorityYear="1990" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" subSpecies="corrugata">
<emphasis id="C894EAF2FF982403FA9FBB9CFE673F09" italics="true" pageId="30" pageNumber="31">Ed. corrugata corrugata</emphasis>
</taxonomicName>
,
<taxonomicName id="3DE04D63FF982403FE78B870FD553F0A" authorityName="Zhang" authorityYear="2019" box="[401,690,1545,1570]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" subSpecies="umbilicata">
<emphasis id="C894EAF2FF982403FE78B870FD553F0A" box="[401,690,1545,1570]" italics="true" pageId="30" pageNumber="31">Ed. corrugata umbilicata</emphasis>
</taxonomicName>
,
<taxonomicName id="3DE04D63FF982403FD2BB870FC7A3F0A" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[706,925,1545,1570]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="edgbastonensis">
<emphasis id="C894EAF2FF982403FD2BB870FC7A3F0A" box="[706,925,1545,1570]" italics="true" pageId="30" pageNumber="31">Ed. edgbastonensis</emphasis>
</taxonomicName>
and
<taxonomicName id="3DE04D63FF982403FC30B870FB863F09" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[985,1121,1545,1570]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="coreena">
<emphasis id="C894EAF2FF982403FC30B870FB863F09" box="[985,1121,1545,1570]" italics="true" pageId="30" pageNumber="31">Ed. coreena</emphasis>
</taxonomicName>
, gave the following results (Wilks’s Lambda 0.236 p&lt;0.000); overall 78% were correctly classified with
<taxonomicName id="3DE04D63FF982403FC16B854FB613F6D" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[1023,1158,1581,1606]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="coreena">
<emphasis id="C894EAF2FF982403FC16B854FB613F6D" box="[1023,1158,1581,1606]" italics="true" pageId="30" pageNumber="31">Ed. coreena</emphasis>
</taxonomicName>
being the least correctly placed (42%) with specimens distributed between
<taxonomicName id="3DE04D63FF982403FD0EB828FBE33F41" authority="corrugata" authorityName="corrugata (Ponder &amp; Clark" authorityYear="1990" box="[743,1028,1617,1642]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" subSpecies="corrugata">
<emphasis id="C894EAF2FF982403FD0EB828FBE33F41" box="[743,1028,1617,1642]" italics="true" pageId="30" pageNumber="31">Ed. corrugata corrugata</emphasis>
</taxonomicName>
and
<taxonomicName id="3DE04D63FF982403FBADB828FAC73F42" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[1092,1312,1617,1642]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="edgbastonensis">
<emphasis id="C894EAF2FF982403FBADB828FAC73F42" box="[1092,1312,1617,1642]" italics="true" pageId="30" pageNumber="31">Ed. edgbastonensis</emphasis>
</taxonomicName>
while
<taxonomicName id="3DE04D63FF982403FA9FB828FE623FA5" authority="corrugata" authorityName="corrugata (Ponder &amp; Clark" authorityYear="1990" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" subSpecies="corrugata">
<emphasis id="C894EAF2FF982403FA9FB828FE623FA5" italics="true" pageId="30" pageNumber="31">Ed. corrugata corrugata</emphasis>
</taxonomicName>
was 77% correctly classified with some specimens being incorrectly classified as
<taxonomicName id="3DE04D63FF982403FA9FB80CFEA53F9A" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="edgbastonensis">
<emphasis id="C894EAF2FF982403FA9FB80CFEA53F9A" italics="true" pageId="30" pageNumber="31">Ed. edgbastonensis</emphasis>
</taxonomicName>
and
<taxonomicName id="3DE04D63FF982403FE93B8E0FD7F3F9A" authorityName="Zhang" authorityYear="2019" box="[378,664,1689,1714]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" subSpecies="umbilicata">
<emphasis id="C894EAF2FF982403FE93B8E0FD7F3F9A" box="[378,664,1689,1714]" italics="true" pageId="30" pageNumber="31">Ed. corrugata umbilicata</emphasis>
</taxonomicName>
.
<taxonomicName id="3DE04D63FF982403FD4FB8E0FC983F9A" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[678,895,1689,1714]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="edgbastonensis">
<emphasis id="C894EAF2FF982403FD4FB8E0FC983F9A" box="[678,895,1689,1714]" italics="true" pageId="30" pageNumber="31">Ed. edgbastonensis</emphasis>
</taxonomicName>
was only 64% correctly classified with most of those going to
<emphasis id="C894EAF2FF982403FED4B8C4FC9B3FFE" box="[317,892,1725,1750]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="3DE04D63FF982403FED4B8C4FDB23FFD" authority="corrugata" authorityName="corrugata (Ponder &amp; Clark" authorityYear="1990" box="[317,597,1725,1750]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" subSpecies="corrugata">Ed. corrugata corrugata</taxonomicName>
.
<taxonomicName id="3DE04D63FF982403FDB6B8C4FC9B3FFE" authorityName="Zhang" authorityYear="2019" box="[607,892,1725,1750]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" subSpecies="umbilicata">Ed. corrugata umbilicata</taxonomicName>
</emphasis>
was the most successfully classified (98%) with only one specimen being misclassified as
<taxonomicName id="3DE04D63FF982403FD85B898FC663FD1" authority="corrugata" authorityName="corrugata (Ponder &amp; Clark" authorityYear="1990" box="[620,897,1761,1786]" class="Gastropoda" family="Hydrobiidae" genus="Edgbastonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="subSpecies" species="corrugata" subSpecies="corrugata">
<emphasis id="C894EAF2FF982403FD85B898FC663FD1" box="[620,897,1761,1786]" italics="true" pageId="30" pageNumber="31">Ed. corrugata corrugata</emphasis>
</taxonomicName>
(
<figureCitation id="62DB2A65FF982403FC66B898FC1E3FD2" box="[911,1017,1761,1786]" captionStart="FIGURE 20" captionStartId="37.[151,250,1474,1496]" captionTargetBox="[203,1358,231,1417]" captionTargetId="figure@37.[151,1435,193,1453]" captionTargetPageId="37" captionText="FIGURE 20. Canonical scores plot of five shell measurements of species of Edgbastonia (Barcaldonia) n. subgen. A. Members of the Ed. (B.) edgbastonensis-corrugata species-group: Ed. (B.) corrugata corrugata (x), Ed. (B.) coreena (Ǫ), Ed. (B.) edgbastonensis (+), and Ed. (B.) corrugata umbilicata (D). B. Members of the Ed. (B.) colmani-pallida species-group: Ed. (B.) colmani (x), Ed. (B.) pallida (Ǫ) and Ed. (B.) cf. pallida (+). C. Members of the Ed. (B.) jesswiseae- zeidlerorum speciesgroup: Ed. (B.) jesswiseae (x), Ed. (B.) acuminata (Ǫ), Ed. (B.) carinata (+), Ed. (B.) zeidlerorum (s) and Ed. (B.) rugosa (D). Ellipse p: 0.6827." httpUri="https://zenodo.org/record/2637643/files/figure.png" pageId="30" pageNumber="31">Fig. 20A</figureCitation>
).
</paragraph>
<paragraph id="FA5F36E0FF982403FF2EB97CFE8D3E4E" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">The new subspecies is unusual in that the egg capsules are laid on the shell, a habit not observed in other taxa, including the typical subspecies, and this is presumably due to the lack of hard substrate and vegetation in the habitat it occupies.</paragraph>
<paragraph id="FA5F36E0FF982403FF2EB908FEEC3EFA" blockId="30.[151,1437,609,2002]" pageId="30" pageNumber="31">
Various water birds such as spoonbills (
<taxonomicName id="3DE04D63FF982403FD70B908FD1F3EA2" authorityName="Linnaeus" authorityYear="1758" box="[665,760,1905,1930]" class="Aves" family="Threskiornithidae" genus="Platalea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelecaniformes" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="C894EAF2FF982403FD70B908FD1F3EA2" box="[665,760,1905,1930]" italics="true" pageId="30" pageNumber="31">Platalea</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="3DE04D63FF982403FCDDB908FC1A3EA2" authorityName="Poche" authorityYear="1904" box="[820,1021,1905,1930]" class="Aves" family="Threskiornithidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pelecaniformes" pageId="30" pageNumber="31" phylum="Chordata" rank="family">Threskiornithidae</taxonomicName>
) and several species of ducks were common in the pool and were feeding in the mud, suggesting that this rather abundant snail is probably utilised as a food item.
</paragraph>
</subSubSection>
</treatment>
</document>
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