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<document id="A0A64695B9613CF43ED3AC77043611FC" ID-CLB-Dataset="27316" ID-DOI="10.11646/zootaxa.4583.1.1" ID-GBIF-Dataset="027ed27d-5035-4646-93a6-c23b565f3d20" ID-ISSN="1175-5326" ID-Zenodo-Dep="2637603" ID-ZooBank="27F24995-359E-46F6-AB22-75568BACFDCF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555054242242" checkinUser="plazi" docAuthor="Zhang, - H." docDate="2019" docId="724987F6FF8D2413FF7EBEEEFEA63BAC" docLanguage="en" docName="zootaxa.4583.1.1.pdf" docOrigin="Zootaxa 4583 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Eulodrobia fenshami Zhang 2019, n. sp." docType="treatment" docVersion="16" lastPageNumber="15" masterDocId="8E70FF8EFF86241DFFE9BE79FFE73928" masterDocTitle="New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages" masterLastPageNumber="67" masterPageNumber="1" pageNumber="12" updateTime="1698720421338" updateUser="ExternalLinkService">
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<mods:title id="CB261AFD45E0FC5925AF4844B8881B89">New taxa of Tateidae (Caenogastropoda, Truncatelloidea) from springs associated with the Great Artesian Basin and Einasleigh Uplands, Queensland, with the description of two related taxa from eastern coastal drainages</mods:title>
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<mods:namePart id="D567F53AC770693025C17D2120075F78">Zhang, - H.</mods:namePart>
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<mods:title id="2A86564D74BCDD0D3B564511D2231B1D">Zootaxa</mods:title>
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<mods:part id="C765896248A76E42576503383BD4CA47">
<mods:date id="1F51A14C6D5E24A5D774E6FAF64051FD">2019</mods:date>
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<mods:number id="93D414E1EB16D91B2BBC3896B2C2DE0C">2019-04-10</mods:number>
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<mods:number id="751D55FD446AFBFF4E767AA6447AB6EE">4583</mods:number>
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<mods:classification id="FD2D964C5E10155014DAC94ABB140ACB">journal article</mods:classification>
<mods:identifier id="433861A5EAD4DDFFF81282D054A47BE4" type="CLB-Dataset">27316</mods:identifier>
<mods:identifier id="C9A0FE6973984310375E11222E94EEE7" type="DOI">10.11646/zootaxa.4583.1.1</mods:identifier>
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<treatment id="724987F6FF8D2413FF7EBEEEFEA63BAC" ID-DOI="http://doi.org/10.5281/zenodo.5582668" ID-GBIF-Taxon="156198445" ID-Zenodo-Dep="5582668" LSID="urn:lsid:plazi:treatment:724987F6FF8D2413FF7EBEEEFEA63BAC" httpUri="http://treatment.plazi.org/id/724987F6FF8D2413FF7EBEEEFEA63BAC" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="B2FA656BFF8D2416FF7EBEEEFE06399A" box="[151,481,151,178]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="FA5F36E0FF8D2416FF7EBEEEFE06399A" blockId="11.[151,481,151,178]" box="[151,481,151,178]" pageId="11" pageNumber="12">
<heading id="A117818CFF8D2416FF7EBEEEFE06399A" bold="true" box="[151,481,151,178]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="C894EAF2FF8D2416FF7EBEEEFE06399A" bold="true" box="[151,481,151,178]" pageId="11" pageNumber="12">
<taxonomicName id="3DE04D63FF8D2416FF7EBEEEFE703999" ID-CoL="7R9CW" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[151,407,151,177]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="fenshami" status="sp. nov.">
<emphasis id="C894EAF2FF8D2416FF7EBEEEFE703999" bold="true" box="[151,407,151,177]" italics="true" pageId="11" pageNumber="12">Eulodrobia fenshami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8D2416FE77BEE1FE06399A" box="[414,481,152,178]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF8D2416FF7EBEA7FE873B13" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="FA5F36E0FF8D2416FF7EBEA7FD2C384B" blockId="11.[151,1437,222,571]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF7EBEA7FE6739DF" bold="true" box="[151,384,222,247]" pageId="11" pageNumber="12">Material examined.</emphasis>
<materialsCitation id="4A883CBDFF8D2416FE6FBEA7FB763817" ID-GBIF-Occurrence="2235592656" ID-GBIF-Specimen="3070853407" collectingDate="2002-04-06" collectorName="Paroo R &amp; W. F. Ponder &amp; J. H. Waterhouse &amp; A. C. Miller" httpUri="https://www.gbif.org/occurrence/3070853407" latitude="-28.158333" location="Eulo township" longLatPrecision="20" longitude="145.03612" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="255B8842FF8D2416FE6FBEA7FE1439DF" box="[390,499,222,247]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
:
<collectingRegion id="3824F802FF8D2416FE14BEA7FD6339DF" box="[509,644,222,247]" country="Australia" name="Queensland" pageId="11" pageNumber="12">Queensland</collectingRegion>
, spring W side of
<collectorName id="57155336FF8D2416FCA6BEA6FC4B39DE" box="[847,940,223,246]" pageId="11" pageNumber="12">Paroo R</collectorName>
, near
<location id="FF3F603BFF8D2416FC07BEA7FB7339DF" LSID="urn:lsid:plazi:treatment:724987F6FF8D2413FF7EBEEEFEA63BAC:FF3F603BFF8D2416FC07BEA7FB7339DF" box="[1006,1172,222,247]" latitude="-28.158333" longLatPrecision="20" longitude="145.03612" name="Eulo township" pageId="11" pageNumber="12" stateProvince="Queensland">Eulo township</location>
,
<geoCoordinate id="9FD45027FF8D2416FB77BEA6FAD439DF" box="[1182,1331,223,247]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="-28.158333">28° 09' 30&quot; S</geoCoordinate>
,
<geoCoordinate id="9FD45027FF8D2416FAD6BEA7FF3F3832" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="145.03612">145° 02' 10&quot; E</geoCoordinate>
, small seeping mound amongst trees, snails in and on mud (some out of water), mostly under grasses in midsection of mound,
<collectorName id="57155336FF8D2416FE98BF5EFE183817" box="[369,511,294,319]" pageId="11" pageNumber="12">W.F. Ponder</collectorName>
,
<collectorName id="57155336FF8D2416FDE7BF5EFD2D3817" box="[526,714,294,319]" pageId="11" pageNumber="12">J.H. Waterhouse</collectorName>
&amp;
<collectorName id="57155336FF8D2416FD1ABF5EFC9D3817" box="[755,890,294,319]" pageId="11" pageNumber="12">A.C. Miller</collectorName>
,
<collectingDate id="9E1AE9C8FF8D2416FC60BF5FFBF53817" box="[905,1042,294,319]" pageId="11" pageNumber="12" value="2002-04-06">
<date id="8E5E1020FF8D2416FC60BF5FFBF53817" box="[905,1042,294,319]" pageId="11" pageNumber="12" value="2002-04-06">6 Apr 2002</date>
</collectingDate>
, C.479950
</materialsCitation>
.
<typeStatus id="255B8842FF8D2416FB76BF5EFAF73817" box="[1183,1296,295,319]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
: Same data, C.410721, 20+, C.431170, 20+;
<materialsCitation id="4A883CBDFF8D2416FDE8BF32FD20384B" ID-GBIF-Occurrence="2235592667" box="[513,711,330,355]" collectionCode="QM" pageId="11" pageNumber="12" specimenCode="MO85761, 5" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="9CF1AE25FF8D2416FDE8BF32FDC9384B" box="[513,558,331,355]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="12" type="Museum">QM</collectionCode>
<specimenCode id="AA469E9BFF8D2416FDDFBF33FD4B384B" box="[566,684,330,355]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="12" type="Museum">MO85761</specimenCode>
,
<specimenCode id="AA469E9BFF8D2416FD5EBF33FD20384B" box="[695,711,330,355]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="12" type="Museum">5</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="FA5F36E0FF8D2416FF2EBF17FE873B13" blockId="11.[151,1437,222,571]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EBF17FEE538AF" bold="true" box="[199,258,366,391]" pageId="11" pageNumber="12">Shell</emphasis>
(
<figureCitation id="62DB2A65FF8D2416FEFDBF17FE9738AF" box="[276,368,366,391]" captionStart="FIGURE 2" captionStartId="8.[151,250,1857,1879]" captionTargetBox="[185,1418,742,1819]" captionTargetId="figure@8.[150,1436,697,1836]" captionTargetPageId="8" captionText="FIGURE 2. Shells of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., holotype, spring at Town Common, Eulo (C.479950). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia cf. eulo (Ponder &amp; Clark, 1990), Tunga Spring (C.410643). D. Eulodrobia ovata n. sp., holotype, Yowah Springs, Bundoona Station (C.479944). E. Eulodrobia bundoona n. sp., holotype, Yowah Springs, Bundoona Station (C.479943). F. Eulodrobia carinata n. sp., holotype, Yowah Springs, Bundoona Station (C.479942). GI. Eulodrobia spirula n. sp., Yowah Springs, Bundoona Station. G. dorsal view of holotype (479945). H, I. apertural view of paratypes (C.400133). Scale bars 1 mm." httpUri="https://zenodo.org/record/2637607/files/figure.png" pageId="11" pageNumber="12">Fig. 2A</figureCitation>
,
<figureCitation id="62DB2A65FF8D2416FE96BF16FE4338AF" box="[383,420,367,391]" captionStart="FIGURE 3" captionStartId="9.[151,250,1476,1498]" captionTargetBox="[221,1370,184,1448]" captionTargetId="figure@9.[204,1382,169,1464]" captionTargetPageId="9" captionText="FIGURE 3. Protoconchs and protoconch microsculpture of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). D. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). E. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). F. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 100 µm (lowpowered images [left]), 10 µm (high-powered images [right])." httpUri="https://zenodo.org/record/2637609/files/figure.png" pageId="11" pageNumber="12">3A</figureCitation>
). Ovate, spire outline moderately convex, normally coiled, opaque. Length
<quantity id="3D189B05FF8D2416FAFBBF17FA7B38AE" box="[1298,1436,366,391]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="11" pageNumber="12" unit="mm" value="2.1" valueMax="2.5" valueMin="1.7">1.72.5 mm</quantity>
(mean
<quantity id="3D189B05FF8D2416FF0FBFEAFEA53883" box="[230,322,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="12" unit="mm" value="2.1">2.1 mm</quantity>
), width
<quantity id="3D189B05FF8D2416FE49BFEAFDCD3882" box="[416,554,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="11" pageNumber="12" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.82.3 mm</quantity>
(mean
<quantity id="3D189B05FF8D2416FD6BBFEAFD393883" box="[642,734,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="12" unit="mm" value="2.1">2.1 mm</quantity>
). Protoconch of 1.2 to 1.3 whorls, irregularly minutely pitted (
<figureCitation id="62DB2A65FF8D2416FF76BFCFFF1B38E7" box="[159,252,438,463]" captionStart="FIGURE 3" captionStartId="9.[151,250,1476,1498]" captionTargetBox="[221,1370,184,1448]" captionTargetId="figure@9.[204,1382,169,1464]" captionTargetPageId="9" captionText="FIGURE 3. Protoconchs and protoconch microsculpture of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). D. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). E. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). F. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 100 µm (lowpowered images [left]), 10 µm (high-powered images [right])." httpUri="https://zenodo.org/record/2637609/files/figure.png" pageId="11" pageNumber="12">Fig. 3A</figureCitation>
). Teleoconch whorls moderately convex, evenly rounded, total number 3.04.1 (mean 3.8). Umbilicus moderately wide. Sculpture of axial growth lines only. Aperture ovate, inner lip narrow, thick, slightly separated along whole length of parietal wall, outer lip medium. Periostracum moderately developed, yellow-brown or black or reddish-brown.
</paragraph>
</subSubSection>
<caption id="AE9F6668FF8D2416FF7EBC13FDB73B89" ID-Table-UUID="AE9F6668FF8D2416FF7EBC13FDB73B89" httpUri="http://table.plazi.org/id/AE9F6668FF8D2416FF7EBC13FDB73B89" pageId="11" pageNumber="12" targetBox="[159,1320,701,971]" targetIsTable="true" targetPageId="11">
<paragraph id="FA5F36E0FF8D2416FF7EBC13FDB73B89" blockId="11.[151,1436,617,673]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF7EBC13FEEA3BA9" bold="true" box="[151,269,618,641]" pageId="11" pageNumber="12">TABLE 2.</emphasis>
Shell measurements (in mm) of type material of
<taxonomicName id="3DE04D63FF8D2416FCD5BC10FBF93BA9" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[828,1054,617,641]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="fenshami" status="sp. nov.">
<emphasis id="C894EAF2FF8D2416FCD5BC10FBF93BA9" box="[828,1054,617,641]" italics="true" pageId="11" pageNumber="12">Eulodrobia fenshami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8D2416FBC0BC13FB8E3BA9" box="[1065,1129,618,641]" pageId="11" pageNumber="12" rank="species">
<emphasis id="C894EAF2FF8D2416FBC0BC13FB8E3BA9" bold="true" box="[1065,1129,618,641]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
(paratypes, C.410721). See Materials and Methods for abbreviations.
</paragraph>
</caption>
<paragraph id="FA5F36E0FF8D2416FE3FBCC4FACF3AE3" pageId="11" pageNumber="12">
<table id="88E0C440FF8DDBE2FF76BCC4FACF3AE3" box="[159,1320,701,971]" gridcols="7" gridrows="7" pageId="11" pageNumber="12">
<tr id="44D034A2FF8DDBE2FF76BCC4FACF3BFB" box="[159,1320,701,723]" gridrow="0" pageId="11" pageNumber="12" rowspan-0="1">
<th id="07015DDEFF8DDBE2FE3FBCC4FDE53BFB" box="[470,514,701,723]" gridcol="1" gridrow="0" pageId="11" pageNumber="12">SH</th>
<th id="07015DDEFF8DDBE2FD94BCC4FD4E3BFB" box="[637,681,701,723]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">SW</th>
<th id="07015DDEFF8DDBE2FCF8BCC4FCDA3BFB" box="[785,829,701,723]" gridcol="3" gridrow="0" pageId="11" pageNumber="12">AH</th>
<th id="07015DDEFF8DDBE2FC4DBCC4FC373BFB" box="[932,976,701,723]" gridcol="4" gridrow="0" pageId="11" pageNumber="12">AW</th>
<th id="07015DDEFF8DDBE2FBB9BCC4FB9B3BFB" box="[1104,1148,701,723]" gridcol="5" gridrow="0" pageId="11" pageNumber="12">BW</th>
<th id="07015DDEFF8DDBE2FB15BCC4FACF3BFB" box="[1276,1320,701,723]" gridcol="6" gridrow="0" pageId="11" pageNumber="12">TW</th>
</tr>
<tr id="44D034A2FF8DDBE2FF76BC9FFACF3BD4" box="[159,1320,742,764]" gridrow="1" pageId="11" pageNumber="12">
<th id="07015DDEFF8DDBE2FF76BC9FFF1A3BD4" box="[159,253,742,764]" gridcol="0" gridrow="1" pageId="11" pageNumber="12">Holotype</th>
<td id="07015DDEFF8DDBE2FE3FBC9FFDE53BD4" box="[470,514,742,764]" gridcol="1" gridrow="1" pageId="11" pageNumber="12">2.4</td>
<td id="07015DDEFF8DDBE2FD94BC9FFD4E3BD4" box="[637,681,742,764]" gridcol="2" gridrow="1" pageId="11" pageNumber="12">2.05</td>
<td id="07015DDEFF8DDBE2FCF8BC9FFCDA3BD4" box="[785,829,742,764]" gridcol="3" gridrow="1" pageId="11" pageNumber="12">1.18</td>
<td id="07015DDEFF8DDBE2FC4DBC9FFC373BD4" box="[932,976,742,764]" gridcol="4" gridrow="1" pageId="11" pageNumber="12">0.94</td>
<td id="07015DDEFF8DDBE2FBB9BC9FFB9B3BD4" box="[1104,1148,742,764]" gridcol="5" gridrow="1" pageId="11" pageNumber="12">1.85</td>
<td id="07015DDEFF8DDBE2FB15BC9FFACF3BD4" box="[1276,1320,742,764]" gridcol="6" gridrow="1" pageId="11" pageNumber="12">4</td>
</tr>
<tr id="44D034A2FF8DDBE2FF76BD69FACF3A0E" box="[159,1320,784,806]" gridrow="2" pageId="11" pageNumber="12">
<th id="07015DDEFF8DDBE2FF76BD69FACF3A0E" box="[159,1320,784,806]" colspan="7" colspanRight="6" gridcol="0" gridrow="2" pageId="11" pageNumber="12">Paratypes (20)</th>
</tr>
<tr id="44D034A2FF8DDBE2FF76BD40FACF3A67" box="[159,1320,825,847]" gridrow="3" pageId="11" pageNumber="12">
<th id="07015DDEFF8DDBE2FF76BD40FF1A3A67" box="[159,253,825,847]" gridcol="0" gridrow="3" pageId="11" pageNumber="12">Min</th>
<td id="07015DDEFF8DDBE2FE3FBD40FDE53A67" box="[470,514,825,847]" gridcol="1" gridrow="3" pageId="11" pageNumber="12">1.73</td>
<td id="07015DDEFF8DDBE2FD94BD40FD4E3A67" box="[637,681,825,847]" gridcol="2" gridrow="3" pageId="11" pageNumber="12">1.83</td>
<td id="07015DDEFF8DDBE2FCF8BD40FCDA3A67" box="[785,829,825,847]" gridcol="3" gridrow="3" pageId="11" pageNumber="12">0.95</td>
<td id="07015DDEFF8DDBE2FC4DBD40FC373A67" box="[932,976,825,847]" gridcol="4" gridrow="3" pageId="11" pageNumber="12">0.98</td>
<td id="07015DDEFF8DDBE2FBB9BD40FB9B3A67" box="[1104,1148,825,847]" gridcol="5" gridrow="3" pageId="11" pageNumber="12">1.48</td>
<td id="07015DDEFF8DDBE2FB15BD40FACF3A67" box="[1276,1320,825,847]" gridcol="6" gridrow="3" pageId="11" pageNumber="12">3.02</td>
</tr>
<tr id="44D034A2FF8DDBE2FF76BD1BFACF3A50" box="[159,1320,866,888]" gridrow="4" pageId="11" pageNumber="12">
<th id="07015DDEFF8DDBE2FF76BD1BFF1A3A50" box="[159,253,866,888]" gridcol="0" gridrow="4" pageId="11" pageNumber="12">Max</th>
<td id="07015DDEFF8DDBE2FE3FBD1BFDE53A50" box="[470,514,866,888]" gridcol="1" gridrow="4" pageId="11" pageNumber="12">2.46</td>
<td id="07015DDEFF8DDBE2FD94BD1BFD4E3A50" box="[637,681,866,888]" gridcol="2" gridrow="4" pageId="11" pageNumber="12">2.29</td>
<td id="07015DDEFF8DDBE2FCF8BD1BFCDA3A50" box="[785,829,866,888]" gridcol="3" gridrow="4" pageId="11" pageNumber="12">1.27</td>
<td id="07015DDEFF8DDBE2FC4DBD1BFC373A50" box="[932,976,866,888]" gridcol="4" gridrow="4" pageId="11" pageNumber="12">1.25</td>
<td id="07015DDEFF8DDBE2FBB9BD1BFB9B3A50" box="[1104,1148,866,888]" gridcol="5" gridrow="4" pageId="11" pageNumber="12">1.95</td>
<td id="07015DDEFF8DDBE2FB15BD1BFACF3A50" box="[1276,1320,866,888]" gridcol="6" gridrow="4" pageId="11" pageNumber="12">4.10</td>
</tr>
<tr id="44D034A2FF8DDBE2FF76BDF5FACF3A8A" box="[159,1320,908,930]" gridrow="5" pageId="11" pageNumber="12">
<th id="07015DDEFF8DDBE2FF76BDF5FF1A3A8A" box="[159,253,908,930]" gridcol="0" gridrow="5" pageId="11" pageNumber="12">Mean</th>
<td id="07015DDEFF8DDBE2FE3FBDF5FDE53A8A" box="[470,514,908,930]" gridcol="1" gridrow="5" pageId="11" pageNumber="12">2.11</td>
<td id="07015DDEFF8DDBE2FD94BDF5FD4E3A8A" box="[637,681,908,930]" gridcol="2" gridrow="5" pageId="11" pageNumber="12">2.07</td>
<td id="07015DDEFF8DDBE2FCF8BDF5FCDA3A8A" box="[785,829,908,930]" gridcol="3" gridrow="5" pageId="11" pageNumber="12">1.13</td>
<td id="07015DDEFF8DDBE2FC4DBDF5FC373A8A" box="[932,976,908,930]" gridcol="4" gridrow="5" pageId="11" pageNumber="12">1.11</td>
<td id="07015DDEFF8DDBE2FBB9BDF5FB9B3A8A" box="[1104,1148,908,930]" gridcol="5" gridrow="5" pageId="11" pageNumber="12">1.75</td>
<td id="07015DDEFF8DDBE2FB15BDF5FACF3A8A" box="[1276,1320,908,930]" gridcol="6" gridrow="5" pageId="11" pageNumber="12">3.80</td>
</tr>
<tr id="44D034A2FF8DDBE2FF76BDCCFACF3AE3" box="[159,1320,949,971]" gridrow="6" pageId="11" pageNumber="12">
<th id="07015DDEFF8DDBE2FF76BDCCFF1A3AE3" box="[159,253,949,971]" gridcol="0" gridrow="6" pageId="11" pageNumber="12">SD</th>
<td id="07015DDEFF8DDBE2FE3FBDCCFDE53AE3" box="[470,514,949,971]" gridcol="1" gridrow="6" pageId="11" pageNumber="12">0.18</td>
<td id="07015DDEFF8DDBE2FD94BDCCFD4E3AE3" box="[637,681,949,971]" gridcol="2" gridrow="6" pageId="11" pageNumber="12">0.13</td>
<td id="07015DDEFF8DDBE2FCF8BDCCFCDA3AE3" box="[785,829,949,971]" gridcol="3" gridrow="6" pageId="11" pageNumber="12">0.09</td>
<td id="07015DDEFF8DDBE2FC4DBDCCFC373AE3" box="[932,976,949,971]" gridcol="4" gridrow="6" pageId="11" pageNumber="12">0.07</td>
<td id="07015DDEFF8DDBE2FBB9BDCCFB9B3AE3" box="[1104,1148,949,971]" gridcol="5" gridrow="6" pageId="11" pageNumber="12">0.13</td>
<td id="07015DDEFF8DDBE2FB15BDCCFACF3AE3" box="[1276,1320,949,971]" gridcol="6" gridrow="6" pageId="11" pageNumber="12">0.30</td>
</tr>
</table>
</paragraph>
<subSubSection id="B2FA656BFF8D2416FF2EBA76FC213E4D" pageId="11" pageNumber="12" type="description">
<paragraph id="FA5F36E0FF8D2416FF2EBA76FDEB3D65" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EBA76FEA83D00" bold="true" box="[199,335,1039,1064]" pageId="11" pageNumber="12">Operculum</emphasis>
(
<figureCitation id="62DB2A65FF8D2416FEB6BA76FE3D3D00" box="[351,474,1039,1064]" captionStart="FIGURE 4" captionStartId="12.[151,250,1521,1543]" captionTargetBox="[196,1406,256,1477]" captionTargetId="figure@12.[151,1435,229,1500]" captionTargetPageId="12" captionText="FIGURE 4. Opercula of species of Eulodrobia n. gen., external and internal views of each species. A, B. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). C, D. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). E, F. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). G, H. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). I, J. Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131). K, L. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 500 µm." httpUri="https://zenodo.org/record/2637611/files/figure.png" pageId="11" pageNumber="12">Fig. 4A, B</figureCitation>
). Transparent, pale yellow or yellow-brown, slightly concave, nucleus acentric. Inner side lacking white smear, simple.
</paragraph>
<paragraph id="FA5F36E0FF8D2416FF2EBA2EFB3C3DBD" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EBA2EFDCE3D58" bold="true" box="[199,553,1111,1136]" pageId="11" pageNumber="12">Head-foot and external body.</emphasis>
Snout, tentacles, neck and opercular lobes pigmented, dorsal and lateral foot unpigmented, mantle roof weakly pigmented to black, visceral coil weakly to densely pigmented.
</paragraph>
<paragraph id="FA5F36E0FF8D2416FF2EBAE6FDCD3C45" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EBAE6FE9A3D90" bold="true" box="[199,381,1183,1208]" pageId="11" pageNumber="12">Mantle cavity.</emphasis>
Ctenidium well-developed, filaments 1719, broadly triangular, apex towards right. Osphradium narrowly oval, towards posterior end of ctenidium, length relative to gill 0.320.45. Hypobranchial gland thin (poorly developed). Rectum with U-shaped bend, faecal pellets orientated obliquely and longitudinally or orientated obliquely or sideways, anus behind mantle collar. Kidney extends for about quarter of length into mantle cavity roof. Renal gland transverse. Pericardium extends for about quarter of length into mantle cavity roof, abutting posterior end of ctenidium.
</paragraph>
<paragraph id="FA5F36E0FF8D2416FF2EBB0EFECD3CD4" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EBB0EFEF93CB8" bold="true" box="[199,286,1399,1424]" pageId="11" pageNumber="12">Radula</emphasis>
(
<figureCitation id="62DB2A65FF8D2416FED8BB0EFE543CB8" box="[305,435,1399,1424]" captionStart="FIGURE 5" captionStartId="13.[151,250,1812,1834]" captionTargetBox="[269,1322,207,1777]" captionTargetId="figure@13.[269,1322,193,1794]" captionTargetPageId="13" captionText="FIGURE 5. Radulae of Eulodrobia n. gen. A, B. Eulodrobia fenshami n. sp., paratype, spring at Town Common, Eulo (C.410721). A, half rows, B, detail of lateral tooth. CE. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779), C, half rows, D detail of lateral tooth, E, detail of central tooth. FH. Eulodrobia ovata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400130). F. Detail of central tooth. G. Detail of inner marginal tooth. H. Detail of outer lateral teeth. IK. Eulodrobia bundoona n. sp., paratype, Yowah Springs, Bundoona Stn (C.400132). I. Part of tooth row. J. Detail of lateral teeth. K. Detail of inner marginal tooth. L.Eulodrobia carinata n. sp., paratype, Yowah Springs, Bundoona Stn (C.400131), half rows. M. N. Eulodrobia spirula n. sp., paratype, Yowah Springs, Bundoona Stn (C.400133), half rows. Scale bars 10 µm." httpUri="https://zenodo.org/record/2637613/files/figure.png" pageId="11" pageNumber="12">Fig. 5A, B</figureCitation>
). Central teeth with cusp formula 24+1+23, basal cusps 2+2; median cusp narrow, sharp, about twice as long as adjacent cusps. Lateral teeth with cusp formula 23+1+3, main cusp pointed, slightly curved, less than twice as long as adjacent cusps. Inner marginal teeth with 1318 cusps. Outer marginal teeth with 2228 cusps.
</paragraph>
<paragraph id="FA5F36E0FF8D2416FF2EB87EFEC43FD0" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EB87EFDFE3F08" bold="true" box="[199,537,1543,1568]" pageId="11" pageNumber="12">Female reproductive system</emphasis>
(
<figureCitation id="62DB2A65FF8D2416FDC3B87EFD403F08" box="[554,679,1543,1568]" captionStart="FIGURE 6" captionStartId="15.[151,250,1512,1534]" captionTargetBox="[181,1411,211,1442]" captionTargetId="figure@15.[150,1436,193,1490]" captionTargetPageId="15" captionText="FIGURE 6. Female reproductive system of species of Eulodrobia n. gen., shown from left side. A, B. Eulodrobia fenshami n. sp. (C. 410721). C, F. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). D, E. Eulodrobia ovata n. sp. (C. 400130). G, H. Eulodrobia bundoona n. sp. (C.400132). I, L. Eulodrobia spirula n. sp. (C.400133). J, K. Eulodrobia carinata n. sp. (C.400131). Abbreviations: ag—albumen gland; bc—bursa copulatrix; cg—capsule gland; ga—opening of oviduct; pwposterior wall of mantle cavity; ro—renal oviduct; sr—seminal receptacle; v—vestibule; vc—ventral channel. Scale bars 500 µm." httpUri="https://zenodo.org/record/2637615/files/figure.png" pageId="11" pageNumber="12">Fig. 6A, B</figureCitation>
). Ovary weakly lobed. Renal oviduct makes wide, open loop over bursa copulatrix. Seminal receptacle oval, duct very short; opens to renal oviduct opposite anterior end of bursa; orientated sperm also in renal oviduct. Bursa copulatrix behind albumen gland, pyriform, about same length as albumen gland, bursal duct enters bursa mid anteriorly, bursal duct joins coiled oviduct in front of posterior mantle cavity wall. Albumen gland partly in mantle cavity. Capsule gland with two distinct glandular zones, medium thickness in cross section, markedly indented by rectum. Anterior vestibule small, opening anterior to capsule gland, short.
</paragraph>
<paragraph id="FA5F36E0FF8D2416FF2EB97DFC213E4D" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EB97DFE1B3E35" bold="true" box="[199,508,1796,1821]" pageId="11" pageNumber="12">Male reproductive system</emphasis>
(
<figureCitation id="62DB2A65FF8D2416FDE5B97DFD973E34" box="[524,624,1796,1821]" captionStart="FIGURE 7" captionStartId="16.[151,250,1324,1346]" captionTargetBox="[170,1416,747,1275]" captionTargetId="figure@16.[150,1436,735,1299]" captionTargetPageId="16" captionText="FIGURE 7. Prostate and vas deferens of species of Eulodrobia n. gen. Each species is shown with anterior at left and posterior at right, with line indicating position of posterior pallial wall. A. Eulodrobia fenshami n. sp. (C. 410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). C. Eulodrobia ovata n. sp. (C.400130). D. Eulodrobia bundoona n. sp. (C.400132). E, Eulodrobia carinata n. sp. (C. 400131). F. Eulodrobia spirula n. sp. (C.400133). Abbreviations: avd—anterior vas deferens; pw—position of posterior wall of pallial cavity; ve—posterior vas deferens. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637617/files/figure.png" pageId="11" pageNumber="12">Figs. 7A</figureCitation>
,
<figureCitation id="62DB2A65FF8D2416FD95B97DFD463E34" box="[636,673,1796,1820]" captionStart="FIGURE 8" captionStartId="17.[151,250,1042,1064]" captionTargetBox="[170,1435,206,1008]" captionTargetId="figure@17.[151,1435,195,1020]" captionTargetPageId="17" captionText="FIGURE 8. Penis and vas deferens of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp. (C. 410721). B. Eulodrobia eulo (Ponder &amp; Clark, 1990) (C. 410724). C. Eulodrobia ovata n. sp. (C. 400130). D. Eulodrobia bundoona n. sp. (C.400132). E, Eulodrobia carinata n. sp. (C. 400131). F. Eulodrobia spirula n. sp. (C.400133). Abbreviations: ngl—non-glandular lobe; pd—penial duct; vd—vas deferens. Scale bars 200 µm." httpUri="https://zenodo.org/record/2637619/files/figure.png" pageId="11" pageNumber="12">8A</figureCitation>
). Prostate gland less than half in mantle roof, bean-shaped, medium in cross section. Posterior and anterior pallial vas deferens slightly undulating. Penis towards middle of head, intermediate, distal end with non-glandular lobe, terminal papilla absent.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF8D2416FF2EB916FBBD3EA0" box="[199,1114,1903,1928]" pageId="11" pageNumber="12" type="etymology">
<paragraph id="FA5F36E0FF8D2416FF2EB916FBBD3EA0" blockId="11.[151,1437,1039,2037]" box="[199,1114,1903,1928]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EB916FEA93EA0" bold="true" box="[199,334,1903,1928]" pageId="11" pageNumber="12">Etymology.</emphasis>
Named for Dr. Rod Fensham, who originally discovered this species.
</paragraph>
</subSubSection>
<subSubSection id="B2FA656BFF8D2416FF2EB9EDFAB03EDC" pageId="11" pageNumber="12" type="distribution">
<paragraph id="FA5F36E0FF8D2416FF2EB9EDFAB03EDC" blockId="11.[151,1437,1039,2037]" pageId="11" pageNumber="12">
<emphasis id="C894EAF2FF8D2416FF2EB9EDFE123E85" bold="true" box="[199,501,1940,1965]" pageId="11" pageNumber="12">Distribution and habitat.</emphasis>
Known from a single small muddy spring on the edge of the western side of the Paroo River (Paroo River Springs in
<collectingCountry id="82F77670FF8D2416FDDEB9C1FC913EF9" box="[567,886,1976,2001]" name="Australia" pageId="11" pageNumber="12">
Commonwealth of
<bibRefCitation id="9E714B11FF8D2416FCE6B9C1FC913EF9" author="Commonwealth of Australia" box="[783,886,1976,2001]" pageId="11" pageNumber="12" refId="ref33702" refString="Commonwealth of Australia (2014) Ecological and hydrogeological survey of the Great Artesian Basin springs-Springsure, Eulo, Bourke and Bogan River Supergroups. Fol. 2. Hydrogeological profiles, Knowledge report, prepared by UniQuest for the Department of the Environment, Commonwealth of Australia, Canberra. Department of the Environment, Public Affairs, Canberra, xiii + 307 pp." type="book" year="2014">Australia</bibRefCitation>
</collectingCountry>
2014) in the town common near the township of Eulo, SW
<collectingRegion id="3824F802FF8D2416FEE2B9A5FE683EDD" box="[267,399,2012,2037]" country="Australia" name="Queensland" pageId="11" pageNumber="12">Queensland</collectingRegion>
(
<figureCitation id="62DB2A65FF8D2416FE77B9A5FE073EDC" box="[414,480,2012,2037]" captionStart="FIGURE 1" captionStartId="4.[151,250,1844,1866]" captionTargetBox="[151,1435,498,1802]" captionTargetId="figure@4.[151,1435,498,1802]" captionTargetPageId="4" captionText="FIGURE 1. Map of the spring supergroups associated with the Great Artesian Basin (GAB) in Queensland, showing the main non-town locations mentioned in the text (squares) and some of the towns in the area (ovals). The pale grey shading is the discharge part of the GAB, the dark grey the recharge part. The inset map of Australia shows the area illustrated. The approximate location of Conondale is indicated by a triangle and Nundle by a star. The spring supergroups are outlined with dotted lines and are as follows: A, Mulligan River; B, Springvale; C, Flinders River; D, Hughenden; E, Barcaldine (see inset at upper right for details); F, Springsure; G, Eulo." httpUri="https://zenodo.org/record/2637605/files/figure.png" pageId="11" pageNumber="12">Fig. 1</figureCitation>
). This small spring was vegetated with sedges and grass and shaded by trees.
</paragraph>
</subSubSection>
<caption id="AE9F6668FF8A2411FF7EBB88FEBE3F89" httpUri="https://zenodo.org/record/2637611/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1521,1543]" targetBox="[196,1406,256,1477]" targetPageId="12">
<paragraph id="FA5F36E0FF8A2411FF7EBB88FEBE3F89" blockId="12.[151,1437,1521,1697]" pageId="12" pageNumber="13">
<emphasis id="C894EAF2FF8A2411FF7EBB88FEF53F2F" bold="true" box="[151,274,1521,1543]" pageId="12" pageNumber="13">FIGURE 4.</emphasis>
Opercula of species of
<taxonomicName id="3DE04D63FF8A2411FDEBBB88FD923F2F" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[514,629,1521,1543]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="C894EAF2FF8A2411FDEBBB88FD923F2F" box="[514,629,1521,1543]" italics="true" pageId="12" pageNumber="13">Eulodrobia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8A2411FD92BB88FD253F2F" box="[635,706,1521,1543]" pageId="12" pageNumber="13" rank="genus">
<emphasis id="C894EAF2FF8A2411FD92BB88FD253F2F" bold="true" box="[635,706,1521,1543]" pageId="12" pageNumber="13">n. gen.</emphasis>
</taxonomicNameLabel>
, external and internal views of each species. A, B.
<taxonomicName id="3DE04D63FF8A2411FB21BB88FA7B3F2F" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[1224,1436,1521,1543]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fenshami" status="sp. nov.">
<emphasis id="C894EAF2FF8A2411FB21BB88FA7B3F2F" box="[1224,1436,1521,1543]" italics="true" pageId="12" pageNumber="13">Eulodrobia fenshami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8A2411FF7EB869FF373F0E" box="[151,208,1552,1574]" pageId="12" pageNumber="13" rank="species">
<emphasis id="C894EAF2FF8A2411FF7EB869FF373F0E" bold="true" box="[151,208,1552,1574]" pageId="12" pageNumber="13">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, spring at Town Common, Eulo (C.410721). C, D.
<taxonomicName id="3DE04D63FF8A2411FCA9B869FB383F0E" authority="(Ponder &amp; Clark, 1990)" authorityName="Ponder &amp; Clark" authorityYear="1990" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[832,1247,1552,1574]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8A2411FCA9B869FC013F0E" box="[832,998,1552,1574]" italics="true" pageId="12" pageNumber="13">Eulodrobia eulo</emphasis>
(Ponder &amp; Clark, 1990)
</taxonomicName>
, paratype, Massey Spring (C.156779). E, F.
<taxonomicName id="3DE04D63FF8A2411FE4DB856FDBE3F6D" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[420,601,1583,1605]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="ovata" status="sp. nov.">
<emphasis id="C894EAF2FF8A2411FE4DB856FDBE3F6D" box="[420,601,1583,1605]" italics="true" pageId="12" pageNumber="13">Eulodrobia ovata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8A2411FD8AB856FD473F6D" box="[611,672,1583,1605]" pageId="12" pageNumber="13" rank="species">
<emphasis id="C894EAF2FF8A2411FD8AB856FD473F6D" bold="true" box="[611,672,1583,1605]" pageId="12" pageNumber="13">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400130). G, H.
<taxonomicName id="3DE04D63FF8A2411FAC3B856FF1B3F4B" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="bundoona" status="sp. nov.">
<emphasis id="C894EAF2FF8A2411FAC3B856FF1B3F4B" italics="true" pageId="12" pageNumber="13">Eulodrobia bundoona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8A2411FEEFB834FEA33F4B" box="[262,324,1613,1635]" pageId="12" pageNumber="13" rank="species">
<emphasis id="C894EAF2FF8A2411FEEFB834FEA33F4B" bold="true" box="[262,324,1613,1635]" pageId="12" pageNumber="13">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400132). I, J.
<taxonomicName id="3DE04D63FF8A2411FC2BB834FB723F4B" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[962,1173,1613,1635]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="carinata" status="sp. nov.">
<emphasis id="C894EAF2FF8A2411FC2BB834FB723F4B" box="[962,1173,1613,1635]" italics="true" pageId="12" pageNumber="13">Eulodrobia carinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8A2411FB76B834FB3A3F4B" box="[1183,1245,1613,1635]" pageId="12" pageNumber="13" rank="species">
<emphasis id="C894EAF2FF8A2411FB76B834FB3A3F4B" bold="true" box="[1183,1245,1613,1635]" pageId="12" pageNumber="13">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400131). K, L.
<taxonomicName id="3DE04D63FF8A2411FDBFB815FCFC3FAA" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[598,795,1644,1666]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="spirula" status="sp. nov.">
<emphasis id="C894EAF2FF8A2411FDBFB815FCFC3FAA" box="[598,795,1644,1666]" italics="true" pageId="12" pageNumber="13">Eulodrobia spirula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8A2411FCCDB815FC873FAA" box="[804,864,1644,1666]" pageId="12" pageNumber="13" rank="species">
<emphasis id="C894EAF2FF8A2411FCCDB815FC873FAA" bold="true" box="[804,864,1644,1666]" pageId="12" pageNumber="13">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400133). Scale bars 500 µm.
</paragraph>
</caption>
<caption id="AE9F6668FF8B2410FF7EB96DFA743EDC" httpUri="https://zenodo.org/record/2637613/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1812,1834]" targetBox="[269,1322,207,1777]" targetPageId="13">
<paragraph id="FA5F36E0FF8B2410FF7EB96DFA743EDC" blockId="13.[151,1437,1812,2036]" pageId="13" pageNumber="14">
<emphasis id="C894EAF2FF8B2410FF7EB96DFEF03E02" bold="true" box="[151,279,1812,1834]" pageId="13" pageNumber="14">FIGURE 5.</emphasis>
Radulae of
<taxonomicName id="3DE04D63FF8B2410FE76B96DFDF53E02" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[415,530,1812,1834]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="C894EAF2FF8B2410FE76B96DFDF53E02" box="[415,530,1812,1834]" italics="true" pageId="13" pageNumber="14">Eulodrobia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8B2410FDF5B96DFD8F3E02" box="[540,616,1812,1834]" pageId="13" pageNumber="14" rank="genus">
<emphasis id="C894EAF2FF8B2410FDF5B96DFD8F3E02" bold="true" box="[540,616,1812,1834]" pageId="13" pageNumber="14">n. gen.</emphasis>
</taxonomicNameLabel>
A, B.
<taxonomicName id="3DE04D63FF8B2410FD51B96DFC763E02" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[696,913,1812,1834]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="fenshami" status="sp. nov.">
<emphasis id="C894EAF2FF8B2410FD51B96DFC763E02" box="[696,913,1812,1834]" italics="true" pageId="13" pageNumber="14">Eulodrobia fenshami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8B2410FC72B96DFC3F3E02" box="[923,984,1812,1834]" pageId="13" pageNumber="14" rank="species">
<emphasis id="C894EAF2FF8B2410FC72B96DFC3F3E02" bold="true" box="[923,984,1812,1834]" pageId="13" pageNumber="14">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, spring at Town Common, Eulo (C.410721). A, half rows, B, detail of lateral tooth. CE.
<taxonomicName id="3DE04D63FF8B2410FD02B94CFB763E63" authority="(Ponder &amp; Clark, 1990)" authorityName="Ponder &amp; Clark" authorityYear="1990" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[747,1169,1845,1867]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF8B2410FD02B94CFC763E63" box="[747,913,1845,1867]" italics="true" pageId="13" pageNumber="14">Eulodrobia eulo</emphasis>
(Ponder &amp; Clark, 1990)
</taxonomicName>
, paratype, Massey Spring (C.156779), C, half rows, D detail of lateral tooth, E, detail of central tooth. FH.
<taxonomicName id="3DE04D63FF8B2410FC04B92EFB783E45" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[1005,1183,1879,1901]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="ovata" status="sp. nov.">
<emphasis id="C894EAF2FF8B2410FC04B92EFB783E45" box="[1005,1183,1879,1901]" italics="true" pageId="13" pageNumber="14">Eulodrobia ovata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8B2410FB4EB92EFB053E45" box="[1191,1250,1879,1901]" pageId="13" pageNumber="14" rank="species">
<emphasis id="C894EAF2FF8B2410FB4EB92EFB053E45" bold="true" box="[1191,1250,1879,1901]" pageId="13" pageNumber="14">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400130). F. Detail of central tooth. G. Detail of inner marginal tooth. H. Detail of outer lateral teeth. IK.
<taxonomicName id="3DE04D63FF8B2410FF22B9E3FE4E3E98" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[203,425,1946,1968]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="bundoona" status="sp. nov.">
<emphasis id="C894EAF2FF8B2410FF22B9E3FE4E3E98" box="[203,425,1946,1968]" italics="true" pageId="13" pageNumber="14">Eulodrobia bundoona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8B2410FE59B9E3FE0E3E98" box="[432,489,1946,1968]" pageId="13" pageNumber="14" rank="species">
<emphasis id="C894EAF2FF8B2410FE59B9E3FE0E3E98" bold="true" box="[432,489,1946,1968]" pageId="13" pageNumber="14">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400132). I. Part of tooth row. J. Detail of lateral teeth. K. Detail of inner marginal tooth. L.
<taxonomicName id="3DE04D63FF8B2410FDBAB9C2FCC33EF9" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[595,804,1979,2001]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="carinata" status="sp. nov.">
<emphasis id="C894EAF2FF8B2410FDBAB9C2FCC33EF9" box="[595,804,1979,2001]" italics="true" pageId="13" pageNumber="14">Eulodrobia carinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8B2410FCC5B9C2FC803EF9" box="[812,871,1979,2001]" pageId="13" pageNumber="14" rank="species">
<emphasis id="C894EAF2FF8B2410FCC5B9C2FC803EF9" bold="true" box="[812,871,1979,2001]" pageId="13" pageNumber="14">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400131), half rows. M. N.
<taxonomicName id="3DE04D63FF8B2410FEADB9A7FDE23EDC" authority="Zhang, 2019" authorityName="Zhang" authorityYear="2019" box="[324,517,2014,2036]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="spirula" status="sp. nov.">
<emphasis id="C894EAF2FF8B2410FEADB9A7FDE23EDC" box="[324,517,2014,2036]" italics="true" pageId="13" pageNumber="14">Eulodrobia spirula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3A75789FF8B2410FDE2B9A7FDA33EDC" box="[523,580,2014,2036]" pageId="13" pageNumber="14" rank="species">
<emphasis id="C894EAF2FF8B2410FDE2B9A7FDA33EDC" bold="true" box="[523,580,2014,2036]" pageId="13" pageNumber="14">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype, Yowah Springs, Bundoona Stn (C.400133), half rows. Scale bars 10 µm.
</paragraph>
</caption>
<subSubSection id="B2FA656BFF882413FF2EBEEEFEA63BAC" pageId="14" pageNumber="15" type="discussion">
<paragraph id="FA5F36E0FF882413FF2EBEEEFBFC3B30" blockId="14.[151,1436,151,645]" pageId="14" pageNumber="15">
<emphasis id="C894EAF2FF882413FF2EBEEEFEDC3998" bold="true" box="[199,315,151,176]" pageId="14" pageNumber="15">Remarks.</emphasis>
This species is most similar to
<taxonomicName id="3DE04D63FF882413FD7EBEE0FD143998" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[663,755,151,176]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF882413FD7EBEE0FD143998" box="[663,755,151,176]" italics="true" pageId="14" pageNumber="15">Eu. eulo</emphasis>
</taxonomicName>
from which it differs in its slightly more ovate shell (see
<figureCitation id="62DB2A65FF882413FA99BEEEFF5E39FC" captionStart="FIGURE 2" captionStartId="8.[151,250,1857,1879]" captionTargetBox="[185,1418,742,1819]" captionTargetId="figure@8.[150,1436,697,1836]" captionTargetPageId="8" captionText="FIGURE 2. Shells of species of Eulodrobia n. gen. A. Eulodrobia fenshami n. sp., holotype, spring at Town Common, Eulo (C.479950). B. Eulodrobia eulo (Ponder &amp; Clark, 1990), paratype, Massey Spring (C.156779). C. Eulodrobia cf. eulo (Ponder &amp; Clark, 1990), Tunga Spring (C.410643). D. Eulodrobia ovata n. sp., holotype, Yowah Springs, Bundoona Station (C.479944). E. Eulodrobia bundoona n. sp., holotype, Yowah Springs, Bundoona Station (C.479943). F. Eulodrobia carinata n. sp., holotype, Yowah Springs, Bundoona Station (C.479942). GI. Eulodrobia spirula n. sp., Yowah Springs, Bundoona Station. G. dorsal view of holotype (479945). H, I. apertural view of paratypes (C.400133). Scale bars 1 mm." httpUri="https://zenodo.org/record/2637607/files/figure.png" pageId="14" pageNumber="15">Fig. 2A</figureCitation>
compared with 2B, C) and in the inner side of the operculum being simple. In addition, the female genital system differs in the posterior loop of the renal oviduct not extending to the posterior end of the bursa copulatrix and the bursal duct being markedly longer, joining the common duct some distance anterior to the seminal receptacle. The distal part of the penis has a double flange in
<taxonomicName id="3DE04D63FF882413FCAABF50FC473868" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[835,928,295,320]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF882413FCAABF50FC473868" box="[835,928,295,320]" italics="true" pageId="14" pageNumber="15">Eu. eulo</emphasis>
</taxonomicName>
but only a single flange in
<taxonomicName id="3DE04D63FF882413FB39BF50FA853868" authorityName="Zhang" authorityYear="2019" box="[1232,1378,295,320]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="fenshami">
<emphasis id="C894EAF2FF882413FB39BF50FA853868" box="[1232,1378,295,320]" italics="true" pageId="14" pageNumber="15">Eu. fenshami</emphasis>
</taxonomicName>
. The molecular results also show these species as distinct, with
<taxonomicName id="3DE04D63FF882413FCD1BF34FC2B384D" authorityName="Zhang" authorityYear="2019" box="[824,972,332,357]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="fenshami">
<emphasis id="C894EAF2FF882413FCD1BF34FC2B384D" box="[824,972,332,357]" italics="true" pageId="14" pageNumber="15">Eu. fenshami</emphasis>
</taxonomicName>
clustering with species from the Yowah Creek Springs on a tributary of the Paroo River system (see below), which are about
<quantity id="3D189B05FF882413FBA1BF16FB6838A0" box="[1096,1167,367,392]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="15" unit="km" value="35.0">35 km</quantity>
NW. The only spring in which
<taxonomicName id="3DE04D63FF882413FF0BBFECFE933885" authorityName="Zhang" authorityYear="2019" box="[226,372,404,429]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="fenshami">
<emphasis id="C894EAF2FF882413FF0BBFECFE933885" box="[226,372,404,429]" italics="true" pageId="14" pageNumber="15">Eu. fenshami</emphasis>
</taxonomicName>
is known to occur is on the main Paroo River, while the springs containing
<taxonomicName id="3DE04D63FF882413FB2CBFECFAC43885" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[1221,1315,404,429]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF882413FB2CBFECFAC43885" box="[1221,1315,404,429]" italics="true" pageId="14" pageNumber="15">Eu. eulo</emphasis>
</taxonomicName>
and
<taxonomicName id="3DE04D63FF882413FAB0BFECFF2038F8" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF882413FAB0BFECFA9E3884" box="[1369,1401,405,428]" italics="true" pageId="14" pageNumber="15">Eu</emphasis>
cf.
<emphasis id="C894EAF2FF882413FF7EBFCEFF2038F8" box="[151,199,439,464]" italics="true" pageId="14" pageNumber="15">eulo</emphasis>
</taxonomicName>
are on tributaries to the west and separated from them by about
<quantity id="3D189B05FF882413FC48BFCEFC0E38F8" box="[929,1001,439,464]" metricMagnitude="4" metricUnit="m" metricValue="5.2" pageId="14" pageNumber="15" unit="km" value="52.0">52 km</quantity>
and
<quantity id="3D189B05FF882413FBC8BFCEFB9938F8" box="[1057,1150,439,464]" metricMagnitude="4" metricUnit="m" metricValue="4.25" pageId="14" pageNumber="15" unit="km" value="42.5">42.5 km</quantity>
respectively. Snails were first noticed at this site by Dr R. Fensham. Regrettably, Dr Fensham has recently reported (pers. comm. to WFP in
<date id="8E5E1020FF882413FF7EBC79FEE03B30" box="[151,263,512,536]" pageId="14" pageNumber="15" value="2017-10">Oct. 2017</date>
) that in 2012, and confirmed in 2014, the spring appears to be extinct.
</paragraph>
<paragraph id="FA5F36E0FF882413FF2EBC5DFEA63BAC" blockId="14.[151,1436,151,645]" pageId="14" pageNumber="15">
A discriminant function analysis using the shell measurements of
<taxonomicName id="3DE04D63FF882413FC0ABC5CFBDE3B15" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[995,1081,548,573]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF882413FC0ABC5CFBDE3B15" box="[995,1081,548,573]" italics="true" pageId="14" pageNumber="15">E. eulo</emphasis>
</taxonomicName>
,
<taxonomicName id="3DE04D63FF882413FBA4BC5CFB2A3B15" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[1101,1229,548,573]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF882413FBA4BC5CFBB93B14" box="[1101,1118,549,572]" italics="true" pageId="14" pageNumber="15">E</emphasis>
. cf.
<emphasis id="C894EAF2FF882413FB74BC5DFB2A3B15" box="[1181,1229,548,573]" italics="true" pageId="14" pageNumber="15">eulo</emphasis>
</taxonomicName>
and
<taxonomicName id="3DE04D63FF882413FAF8BC5CFA7C3B15" authorityName="Zhang" authorityYear="2019" box="[1297,1435,548,573]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="fenshami">
<emphasis id="C894EAF2FF882413FAF8BC5CFA7C3B15" box="[1297,1435,548,573]" italics="true" pageId="14" pageNumber="15">E. fenshami</emphasis>
</taxonomicName>
successfully classified 95% of
<taxonomicName id="3DE04D63FF882413FE11BC30FDAE3B48" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[504,585,583,608]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF882413FE11BC30FDAE3B48" box="[504,585,583,608]" italics="true" pageId="14" pageNumber="15">E. eulo</emphasis>
</taxonomicName>
, 90% of
<taxonomicName id="3DE04D63FF882413FD5EBC30FCC93B48" baseAuthorityName="Ponder &amp; Clark" baseAuthorityYear="1990" box="[695,814,583,608]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="eulo">
<emphasis id="C894EAF2FF882413FD5EBC30FD2F3B48" box="[695,712,585,608]" italics="true" pageId="14" pageNumber="15">E</emphasis>
. cf.
<emphasis id="C894EAF2FF882413FD17BC3EFCC93B48" box="[766,814,583,608]" italics="true" pageId="14" pageNumber="15">eulo</emphasis>
</taxonomicName>
and 100% of
<taxonomicName id="3DE04D63FF882413FC3DBC30FBBD3B48" authorityName="Zhang" authorityYear="2019" box="[980,1114,583,608]" class="Gastropoda" family="Tateidae" genus="Eulodrobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="fenshami">
<emphasis id="C894EAF2FF882413FC3DBC30FBBD3B48" box="[980,1114,583,608]" italics="true" pageId="14" pageNumber="15">E. fenshami</emphasis>
</taxonomicName>
(Wilkss Lambda 0.096, p&lt;0.000) (
<figureCitation id="62DB2A65FF882413FF07BC15FED43BAC" box="[238,307,620,645]" captionStart="FIGURE 9" captionStartId="22.[151,250,1886,1908]" captionTargetBox="[426,1136,1132,1851]" captionTargetId="figure@22.[410,1181,1119,1868]" captionTargetPageId="22" captionText="FIGURE 9. Canonical scores plot of five shell measurements of the Eulodrobia eulo species-group: E. eulo (x), E. cf. eulo (Ǫ) and E. fenshami (+). Ellipse p: 0.6827." httpUri="https://zenodo.org/record/2637621/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>