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<document id="E3C359B2815B9E14F14DE335ED72D47D" ID="10.11646/zootaxa.4236.2.1SLASH10281" ID-CLB-Dataset="36468" ID-DOI="10.11646/zootaxa.4236.2.1" ID-GBIF-Dataset="ec0dbb00-2a4a-41c9-9e60-185f972be9bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="321720" ID-ZooBank="BC5E331F-045C-47FF-BA0E-042C69DE3F80" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487837593550" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2017" docId="724387B7D26F505EBF85292081F1F9B2" docLanguage="en" docName="zootaxa.4236.2.1SLASH10281.pdf" docOrigin="Zootaxa 4236 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus reflexus subsp. rugatus Häckel, 2017, n. ssp." docType="treatment" docVersion="9" lastPageNumber="225" masterDocId="8E7AFFCFD2775046BF122D3A8244FFFC" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 2. Revision of the Craspedophorus leprieuri and C. regalis groups (Coleoptera: Carabidae)" masterLastPageNumber="243" masterPageNumber="201" pageNumber="225" updateTime="1698437222970" updateUser="ExternalLinkService">
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<mods:title id="B78930F9CCDE377801A5A311204AEC31">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 2. Revision of the Craspedophorus leprieuri and C. regalis groups (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="9742A961464F4BB815363814708D69CD">Häckel, Martin</mods:namePart>
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<mods:typeOfResource id="043C58EBFDE08325AB9EFBEAC1C1A732">text</mods:typeOfResource>
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<mods:date id="848CF5A9672D86D54185AD3153D4C95E">2017</mods:date>
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<mods:identifier id="38175554CFE9E3A0BE5C69A182F3A559" type="CLB-Dataset">36468</mods:identifier>
<mods:identifier id="2D0D99C51F26AD08B9A41B96E4F7D581" type="DOI">10.11646/zootaxa.4236.2.1</mods:identifier>
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<treatment id="724387B7D26F505EBF85292081F1F9B2" ID-DOI="http://doi.org/10.5281/zenodo.5668738" ID-GBIF-Taxon="127709712" ID-Zenodo-Dep="5668738" LSID="urn:lsid:plazi:treatment:724387B7D26F505EBF85292081F1F9B2" httpUri="http://treatment.plazi.org/id/724387B7D26F505EBF85292081F1F9B2" lastPageNumber="225" pageId="24" pageNumber="225">
<subSubSection id="B2F0652AD26F505EBF85292083C6FBAA" pageId="24" pageNumber="225" type="nomenclature">
<paragraph id="FA5536A1D26F505EBF852920803BFBC9" blockId="24.[151,639,1050,1110]" box="[151,639,1050,1077]" pageId="24" pageNumber="225">
<heading id="A11D81CDD26F505EBF852920803BFBC9" bold="true" box="[151,639,1050,1077]" fontSize="11" level="1" pageId="24" pageNumber="225" reason="1">
<emphasis id="C89EEAB3D26F505EBF852920803BFBC9" bold="true" box="[151,639,1050,1077]" pageId="24" pageNumber="225">
<taxonomicName id="3DEA4D22D26F505EBF852920806FFBC8" ID-CoL="7GPNV" box="[151,555,1050,1076]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subSpecies" species="reflexus" status="subsp. nov." subSpecies="rugatus">
<emphasis id="C89EEAB3D26F505EBF852920806FFBC8" bold="true" box="[151,555,1050,1076]" italics="true" pageId="24" pageNumber="225">Craspedophorus reflexus rugatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AD57C8D26F505EBD202921803BFBC9" box="[562,639,1051,1077]" pageId="24" pageNumber="225" rank="subSpecies">n. ssp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FA5536A1D26F505EBF85290783C6FBAA" blockId="24.[151,639,1050,1110]" box="[151,386,1085,1110]" pageId="24" pageNumber="225">(Plate 9, Figs 6768)</paragraph>
</subSubSection>
<subSubSection id="B2F0652AD26F505EBF8529BF801EFA8A" pageId="24" pageNumber="225" type="materials_examined">
<paragraph id="FA5536A1D26F505EBF8529BF801EFA8A" blockId="24.[151,1437,1157,1614]" pageId="24" pageNumber="225">
<materialsCitation id="4A823CFCD26F505EBF8529BF8026FB3E" ID-GBIF-Occurrence="1437069683" country="Cameroon" county="Bafoussam" elevation="740" latitude="5.4166665" location="Mandoripa" municipality="Foumban" pageId="24" pageNumber="225" specimenCount="1" specimenCount-male="1" stateProvince="West" typeStatus="holotype">
<emphasis id="C89EEAB3D26F505EBF8529BF830DFB62" bold="true" box="[151,329,1157,1182]" pageId="24" pageNumber="225">
<typeStatus id="25518803D26F505EBF8529BF8296FB62" box="[151,210,1157,1182]" pageId="24" pageNumber="225">Type</typeStatus>
material.
</emphasis>
<typeStatus id="25518803D26F505EBE4729BF83F9FB62" box="[341,445,1157,1182]" pageId="24" pageNumber="225" type="holotype">Holotype</typeStatus>
(
<specimenCount id="ECECFD28D26F505EBEC029BC83ACFB62" box="[466,488,1158,1182]" pageId="24" pageNumber="225" type="male"></specimenCount>
) “
<collectingCountry id="82FD7631D26F505EBD1529BC8030FB62" box="[519,628,1158,1182]" name="Cameroon" pageId="24" pageNumber="225">Kamerun</collectingCountry>
, Prov. Nord-Quest [=
<collectingRegion id="382EF843D26F505EBC6229BC81EAFB61" box="[880,942,1158,1181]" country="Cameroon" name="West" pageId="24" pageNumber="225">West</collectingRegion>
] / Str. zw.
<collectingCounty id="13344E2DD26F505EBB2429BF86F7FB62" box="[1078,1203,1157,1182]" pageId="24" pageNumber="225">Bafoussam</collectingCounty>
u.
<collectingMunicipality id="1A31ACDBD26F505EBBC229BF877AFB62" box="[1232,1342,1157,1182]" pageId="24" pageNumber="225">Foumban</collectingMunicipality>
/ Umg. Koutaba,
<location id="FF35607AD26F505EBE14299383C0FB3E" LSID="urn:lsid:plazi:treatment:724387B7D26F505EBF85292081F1F9B2:FF35607AD26F505EBE14299383C0FB3E" box="[262,388,1193,1218]" country="Cameroon" county="Bafoussam" latitude="5.4166665" municipality="Foumban" name="Mandoripa" pageId="24" pageNumber="225" stateProvince="West">Mandoripa</location>
,
<quantity id="3D129B44D26F505EBE8329908395FB3E" box="[401,465,1194,1218]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="24" pageNumber="225" unit="m" value="740.0">
<elevation id="71C7D192D26F505EBE8329908395FB3E" box="[401,465,1194,1218]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="24" pageNumber="225" unit="m" value="740.0">740m</elevation>
</quantity>
/
<geoCoordinate id="9FDE5066D26F505EBEF92990801DFB3E" box="[491,601,1193,1218]" direction="north" orientation="latitude" pageId="24" pageNumber="225" precision="925" value="5.4166665">N 05°25'</geoCoordinate>
1
</materialsCitation>
.
<materialsCitation id="4A823CFCD26F505EBD7D299080A1FB1A" ID-GBIF-Occurrence="1437069689" collectingDate="2013-08" collectorName="A. Puchner" country="Cameroon" county="Bafoussam" elevation="740" latitude="5.4166665" location="Mandoripa" longitude="-11.0" municipality="Foumban" pageId="24" pageNumber="225" specimenCount="2" specimenCount-male="2" stateProvince="West" typeStatus="paratype">
<geoCoordinate id="9FDE5066D26F505EBD7D299080A2FB3E" box="[623,742,1194,1218]" direction="west" orientation="longitude" pageId="24" pageNumber="225" precision="925" value="-11.0">O 011°00'</geoCoordinate>
7. / leg.
<collectorName id="571F5377D26F505EBC5629908180FB3E" box="[836,964,1193,1218]" pageId="24" pageNumber="225">A. Puchner</collectorName>
<date id="8E541061D26F505EBCDF29908615FB3E" box="[973,1105,1194,1218]" bridgedPair="/" pageId="24" pageNumber="225" value="2013-08">
<collectingDate id="9E10E989D26F505EBCDF29908615FB3E" box="[973,1105,1194,1218]" bridgedPair="/" pageId="24" pageNumber="225" value="2013-08">VIII / 2013</collectingDate>
</date>
” (cAP, Plate 9, Fig. 67 left).
<typeStatus id="25518803D26F505EBF8529F4834CFB1A" box="[151,264,1230,1254]" pageId="24" pageNumber="225" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="ECECFD28D26F505EBE0729F4837FFB1A" box="[277,315,1230,1254]" pageId="24" pageNumber="225" type="male">2♂</specimenCount>
, same data as holotype (cAP, cMH)
</materialsCitation>
,
<materialsCitation id="4A823CFCD26F505EBDE629F481F3FAF6" ID-GBIF-Occurrence="1437069694" collectingDate="2013-06" collectorName="J. M. Dountio" country="Cameroon" county="Bafoussam" elevation="1210" latitude="5.4166665" location="Mandoripa" municipality="Foumban" pageId="24" pageNumber="225" specimenCount="1" specimenCount-female="1" stateProvince="West" typeStatus="paratype">
<specimenCount id="ECECFD28D26F505EBDE629F48153FB1A" box="[756,791,1230,1254]" pageId="24" pageNumber="225" type="female">1♀</specimenCount>
, same data as holotype except: “
<quantity id="3D129B44D26F505EBB8929F486AFFB1A" box="[1179,1259,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="24" pageNumber="225" unit="m" value="1210.0">
<elevation id="71C7D192D26F505EBB8929F486AFFB1A" box="[1179,1259,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="24" pageNumber="225" unit="m" value="1210.0">1210m</elevation>
</quantity>
/ N 05°095' O 010°464' / leg.
<collectorName id="571F5377D26F505EBE5729C883A7FAF6" box="[325,483,1265,1290]" pageId="24" pageNumber="225">J. M. Dountio</collectorName>
<date id="8E541061D26F505EBEF929C8801FFAF6" box="[491,603,1266,1290]" bridgedPair="/" pageId="24" pageNumber="225" value="2013-06">
<collectingDate id="9E10E989D26F505EBEF929C8801FFAF6" box="[491,603,1266,1290]" bridgedPair="/" pageId="24" pageNumber="225" value="2013-06">VI / 2013</collectingDate>
</date>
” (cAP, cMH, Plate 9, Fig. 68)
</materialsCitation>
;
<materialsCitation id="4A823CFCD26F505EBCD629C88156FAD2" ID-GBIF-Occurrence="1437069699" country="Cameroon" elevation="500" latitude="10.15" location="Yagoua" municipality="Maroua" pageId="24" pageNumber="225" specimenCount="1" specimenCount-female="1" stateProvince="Nord" typeStatus="paratype">
<specimenCount id="ECECFD28D26F505EBCD629C881ADFAF6" box="[964,1001,1266,1290]" pageId="24" pageNumber="225" type="female">1♀</specimenCount>
: “
<collectingCountry id="82FD7631D26F505EBB1329C8862AFAF6" box="[1025,1134,1266,1290]" name="Cameroon" pageId="24" pageNumber="225">Kamerun</collectingCountry>
, Prov. Extrême
<collectingRegion id="382EF843D26F505EBA3729CB871AFAF6" box="[1317,1374,1265,1290]" country="Cameroon" name="North" pageId="24" pageNumber="225">Nord</collectingRegion>
/ Str. zw.
<collectingMunicipality id="1A31ACDBD26F505EBFD5282C835AFAD1" box="[199,286,1302,1325]" pageId="24" pageNumber="225">Maroua</collectingMunicipality>
u.
<location id="FF35607AD26F505EBE28282C83D6FAD2" LSID="urn:lsid:plazi:treatment:724387B7D26F505EBF85292081F1F9B2:FF35607AD26F505EBE28282C83D6FAD2" box="[314,402,1302,1326]" country="Cameroon" latitude="10.15" municipality="Maroua" name="Yagoua" pageId="24" pageNumber="225" stateProvince="Nord">Yagoua</location>
/ Umg. Lara,
<quantity id="3D129B44D26F505EBD2F282F8039FAD2" box="[573,637,1301,1326]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="225" unit="m" value="500.0">
<elevation id="71C7D192D26F505EBD2F282F8039FAD2" box="[573,637,1301,1326]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="225" unit="m" value="500.0">500m</elevation>
</quantity>
/
<geoCoordinate id="9FDE5066D26F505EBD88282C8141FAD2" box="[666,773,1301,1326]" direction="north" orientation="latitude" pageId="24" pageNumber="225" precision="925" value="10.15">N 10°09'</geoCoordinate>
5
</materialsCitation>
. O 014°30'0. / leg. J. M. Dountio
<date id="8E541061D26F505EBBBA282C8743FAD2" box="[1192,1287,1302,1326]" pageId="24" pageNumber="225" value="2013-10">X. 2013</date>
” (cAP);
<materialsCitation id="4A823CFCD26F505EBA63282C867DFAAE" ID-GBIF-Occurrence="1437069704" country="Cameroon" elevation="680" latitude="3.5" location="Mbalmayo" municipality="Region" pageId="24" pageNumber="225" specimenCount="1" specimenCount-male="1" stateProvince="Centre" typeStatus="paratype">
<specimenCount id="ECECFD28D26F505EBA63282C87D3FAD2" box="[1393,1431,1302,1326]" pageId="24" pageNumber="225" type="male">1♂</specimenCount>
: “
<collectingCountry id="82FD7631D26F505EBFB32800834AFAAE" box="[161,270,1338,1362]" name="Cameroon" pageId="24" pageNumber="225">Kamerun</collectingCountry>
, Prov.
<collectingRegion id="382EF843D26F505EBE4E280083E3FAAE" box="[348,423,1338,1362]" country="Cameroon" name="Centre" pageId="24" pageNumber="225">Centre</collectingRegion>
/
<collectingMunicipality id="1A31ACDBD26F505EBEAF2803804AFAAE" box="[445,526,1337,1362]" pageId="24" pageNumber="225">Region</collectingMunicipality>
um
<location id="FF35607AD26F505EBD53280380FEFAAE" LSID="urn:lsid:plazi:treatment:724387B7D26F505EBF85292081F1F9B2:FF35607AD26F505EBD53280380FEFAAE" box="[577,698,1337,1362]" country="Cameroon" latitude="3.5" municipality="Region" name="Mbalmayo" pageId="24" pageNumber="225" stateProvince="Centre">Mbalmayo</location>
/ Umg. Obout,
<quantity id="3D129B44D26F505EBC7F280381E9FAAE" box="[877,941,1337,1362]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="24" pageNumber="225" unit="m" value="680.0">
<elevation id="71C7D192D26F505EBC7F280381E9FAAE" box="[877,941,1337,1362]" metricMagnitude="2" metricUnit="m" metricValue="6.8" pageId="24" pageNumber="225" unit="m" value="680.0">680m</elevation>
</quantity>
/
<geoCoordinate id="9FDE5066D26F505EBCD628008668FAAE" box="[964,1068,1338,1362]" direction="north" orientation="latitude" pageId="24" pageNumber="225" precision="925" value="3.5">N 03°30'</geoCoordinate>
9
</materialsCitation>
. O 011°43'4. / leg. J. M. Mbida
<date id="8E541061D26F505EBF8528648342FA8A" box="[151,262,1374,1398]" pageId="24" pageNumber="225" value="2013-12">XII. 2013</date>
” (cAP, Plate 9, Fig. 67 right).
</paragraph>
</subSubSection>
<subSubSection id="B2F0652AD26F505EBFD528BB860DF9D6" pageId="24" pageNumber="225" type="description">
<paragraph id="FA5536A1D26F505EBFD528BB8113FA65" blockId="24.[151,1437,1157,1614]" box="[199,855,1409,1434]" pageId="24" pageNumber="225">
<emphasis id="C89EEAB3D26F505EBFD528BB83A0FA66" bold="true" box="[199,484,1409,1434]" pageId="24" pageNumber="225">
Description of
<typeStatus id="25518803D26F505EBE6428BB839BFA66" box="[374,479,1409,1434]" pageId="24" pageNumber="225" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="3D129B44D26F505EBD5028BB80ECFA65" box="[578,680,1409,1434]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="24" pageNumber="225" unit="mm" value="24.5">24.5 mm</quantity>
, width
<quantity id="3D129B44D26F505EBDE828B88117FA65" box="[762,851,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="24" pageNumber="225" unit="mm" value="9.8">9.8 mm</quantity>
.
</paragraph>
<paragraph id="FA5536A1D26F505EBFD5289F860DF9D6" blockId="24.[151,1437,1157,1614]" pageId="24" pageNumber="225">
Head and pronotum as in variants of other subspecies of
<taxonomicName id="3DEA4D22D26F505EBC2F289C81F0FA41" box="[829,948,1445,1469]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="C89EEAB3D26F505EBC2F289C81F0FA41" box="[829,948,1445,1469]" italics="true" pageId="24" pageNumber="225">C. reflexus</emphasis>
</taxonomicName>
, pronotum somewhat smaller than in HT of
<taxonomicName id="3DEA4D22D26F505EBF8528F08377FA1D" box="[151,307,1481,1505]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subSpecies" species="reflexus" subSpecies="reflexus">
<emphasis id="C89EEAB3D26F505EBF8528F08377FA1D" box="[151,307,1481,1505]" italics="true" pageId="24" pageNumber="225">C. r. reflexus</emphasis>
</taxonomicName>
. Statue smaller and broader, namely elytra less elongate than in
<taxonomicName id="3DEA4D22D26F505EBB5F28F086B1FA1D" box="[1101,1269,1481,1505]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subSpecies" species="reflexus" subSpecies="crampeli">
<emphasis id="C89EEAB3D26F505EBB5F28F086B1FA1D" box="[1101,1269,1481,1505]" italics="true" pageId="24" pageNumber="225">C. r. crampeli</emphasis>
</taxonomicName>
. Differs from populations of
<taxonomicName id="3DEA4D22D26F505EBE2D28D48388F9F9" box="[319,460,1517,1541]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subSpecies" species="reflexus" subSpecies="reflexus">
<emphasis id="C89EEAB3D26F505EBE2D28D48388F9F9" box="[319,460,1517,1541]" italics="true" pageId="24" pageNumber="225">C. l. reflexus</emphasis>
</taxonomicName>
and
<taxonomicName id="3DEA4D22D26F505EBD1628D480DAF9F9" box="[516,670,1517,1541]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="225" phylum="Arthropoda" rank="subSpecies" species="reflexus" subSpecies="crampeli">
<emphasis id="C89EEAB3D26F505EBD1628D480DAF9F9" box="[516,670,1517,1541]" italics="true" pageId="24" pageNumber="225">C. r. crampeli</emphasis>
</taxonomicName>
by coarser elytral sculpture. Strial punctures larger and deeper, their margins reach centers of intervals, in some instances interrupt or obliterate intervals.
</paragraph>
</subSubSection>
<subSubSection id="B2F0652AD26F505EBFD52B0F81F1F9B2" box="[199,949,1589,1614]" pageId="24" pageNumber="225" type="distribution">
<paragraph id="FA5536A1D26F505EBFD52B0F81F1F9B2" blockId="24.[151,1437,1157,1614]" box="[199,949,1589,1614]" pageId="24" pageNumber="225">
<emphasis id="C89EEAB3D26F505EBFD52B0F8324F9B2" bold="true" box="[199,352,1589,1614]" pageId="24" pageNumber="225">Distribution.</emphasis>
<collectingCountry id="82FD7631D26F505EBE752B0C83A4F9B2" box="[359,480,1590,1614]" name="Cameroon" pageId="24" pageNumber="225">Cameroon</collectingCountry>
:
<collectingRegion id="382EF843D26F505EBEF92B0F8004F9B2" box="[491,576,1589,1614]" country="Zambia" name="Central" pageId="24" pageNumber="225">Central</collectingRegion>
, Extreme
<collectingRegion id="382EF843D26F505EBDA32B0F80B1F9B2" box="[689,757,1589,1614]" country="Cameroon" name="North" pageId="24" pageNumber="225">North</collectingRegion>
,
<collectingRegion id="382EF843D26F505EBC122B0C8173F9B1" box="[768,823,1590,1613]" country="Cameroon" name="West" pageId="24" pageNumber="225">West</collectingRegion>
Provinces.
</paragraph>
</subSubSection>
</treatment>
</document>