treatments-xml/data/72/42/87/724287C1FF81FFCD3DE8FF1222A4BC93.xml

<document id="69C042A02D697B02A82A4E224E47A60E" ID-DOI="10.5281/zenodo.180479" ID-GBIF-Dataset="d1874f7a-7d67-4f0d-8e5e-16c18ececf86" ID-ISSN="1175-5326" ID-Zenodo-Dep="180479" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459808686165" checkinUser="plazi" docAuthor="Lindquist, Evert E. &amp; Moraza, Maria L." docDate="2008" docId="724287C1FF81FFCD3DE8FF1222A4BC93" docLanguage="en" docName="zt01685p037.pdf" docOrigin="Zootaxa 1685" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Spadiseius spathiphyllae Lindquist &amp; Moraza, 2008, new species" docType="treatment" docVersion="10" lastPageNumber="32" masterDocId="8E7BFFB9FF94FFD23D7FFF852731BA6E" masterDocTitle="A new genus of flower-dwelling melicharid mites (Acari: Mesostigmata: Ascoidea) phoretic on bats and insects in Costa Rica and Brazil" masterLastPageNumber="37" masterPageNumber="1" pageNumber="22" updateTime="1698220741931" updateUser="plazi">
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<mods:title id="B4D63B1BA3805068449C95DC831A30BB">A new genus of flower-dwelling melicharid mites (Acari: Mesostigmata: Ascoidea) phoretic on bats and insects in Costa Rica and Brazil</mods:title>
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<mods:namePart id="2676CEA9F638631FC8361327B771E06E">Lindquist, Evert E.</mods:namePart>
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<mods:namePart id="B417CE4129E9E34255E60FEDBD7FB494">Moraza, Maria L.</mods:namePart>
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<mods:date id="3DC89AC3FFF0D6CF44038F977236CBEF">2008</mods:date>
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<treatment id="724287C1FF81FFCD3DE8FF1222A4BC93" ID-DOI="http://doi.org/10.5281/zenodo.5620718" ID-GBIF-Taxon="119353314" ID-Zenodo-Dep="5620718" LSID="urn:lsid:plazi:treatment:724287C1FF81FFCD3DE8FF1222A4BC93" httpUri="http://treatment.plazi.org/id/724287C1FF81FFCD3DE8FF1222A4BC93" lastPageId="31" lastPageNumber="32" pageId="21" pageNumber="22">
<subSubSection id="B2F1655CFF81FFC73DE8FF12257ABABA" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="FA5436D7FF81FFC73DE8FF122566BADF" blockId="21.[151,599,151,212]" box="[151,599,151,177]" pageId="21" pageNumber="22">
<heading id="A11C81BBFF81FFC73DE8FF122566BADF" bold="true" box="[151,599,151,177]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="C89FEAC5FF81FFC73DE8FF122566BADF" bold="true" box="[151,599,151,177]" pageId="21" pageNumber="22">
<taxonomicName id="3DEB4D54FF81FFC73DE8FF1226F1BADF" box="[151,448,151,177]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF81FFC73DE8FF1226F1BADF" bold="true" box="[151,448,151,177]" italics="true" pageId="21" pageNumber="22">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF81FFC73CB8FF122566BADF" box="[455,599,151,177]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FA5436D7FF81FFC73DE8FF3F257ABABA" blockId="21.[151,599,151,212]" box="[151,587,186,212]" pageId="21" pageNumber="22">
(
<figureCitation id="62D02A52FF81FFC73DE0FF3F2621BABA" box="[159,272,186,212]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="21" pageNumber="22">Figs. 7–9</figureCitation>
, 
<figureCitation id="62D02A52FF81FFC73C63FF3F2659BABA" box="[284,360,186,212]" captionStart="FIGURES 35 – 37" captionStartId="22.[151,269,1758,1782]" captionTargetBox="[151,1434,598,1757]" captionTargetId="figure@22.[151,1436,594,1776]" captionTargetPageId="22" captionText="FIGURES 35 – 37. Spadiseius spathiphyllae n. sp. 35, larva, idiosoma, dorsal aspect; 36, protonymph, idiosoma, dorsal aspect; 37, larva, chelicera, adaxial view. Abbreviation: v m, ventral mucro." httpUri="https://zenodo.org/record/180490/files/figure.png" pageId="21" pageNumber="22">35–37</figureCitation>
, 
<figureCitation id="62D02A52FF81FFC73C0BFF3F2573BABA" box="[372,578,186,212]" captionStart-0="FIGURES 38 – 41" captionStart-1="FIGURES 42 – 45" captionStart-2="FIGURES 46 – 50" captionStart-3="FIGURES 51 – 55" captionStart-4="FIGURES 56 – 58" captionStart-5="FIGURES 59 – 64" captionStart-6="FIGURES 65 – 69" captionStartId-0="24.[151,269,1163,1187]" captionStartId-1="25.[151,269,1816,1840]" captionStartId-2="27.[151,269,1773,1797]" captionStartId-3="28.[151,269,1906,1930]" captionStartId-4="29.[151,269,1642,1666]" captionStartId-5="30.[151,269,1546,1570]" captionStartId-6="31.[151,269,1219,1243]" captionTargetBox-0="[168,1419,313,1131]" captionTargetBox-1="[154,1431,346,1815]" captionTargetBox-2="[186,1409,183,1772]" captionTargetBox-3="[206,1386,432,1878]" captionTargetBox-4="[155,1429,194,1600]" captionTargetBox-5="[152,1432,441,1545]" captionTargetBox-6="[164,1431,194,1163]" captionTargetId-0="figure@24.[148,1433,288,1233]" captionTargetId-1="figure@25.[151,1436,330,1936]" captionTargetId-2="figure@27.[186,1411,178,1937]" captionTargetId-3="figure@28.[206,1393,416,1899]" captionTargetId-4="figure@29.[151,1436,194,1637]" captionTargetId-5="figure@30.[151,1436,434,1616]" captionTargetId-6="figure@31.[151,1436,194,1283]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="27" captionTargetPageId-3="28" captionTargetPageId-4="29" captionTargetPageId-5="30" captionTargetPageId-6="31" captionText-0="FIGURES 38 – 41. Dorsal subapical femoral setae of legs. Figs. 38, 39, larval femora I – III: 38, Spadiseius calyptrogynae n. sp.; 39, Spadiseius spathiphyllae n. sp. Figs. 40, 41, protonymphal femora I – IV: 40, S. calyptrogynae n. sp.; 41, S." captionText-1="FIGURES 42 – 45. Figs. 42, 43, Spadiseius spathiphyllae n. sp., deutonymph: 42, idiosoma, dorsal aspect; 43, idiosoma, ventral aspect; supernumerary setae denoted by ‘ x’. Figs. 44, 45, dorsal subapical femoral setae legs I – IV, deutonymph: 44, Spadiseius calyptrogynae n. sp.; 45, Spadiseius spathiphyllae n. sp." captionText-2="FIGURES 46 – 50. Spadiseius spathiphyllae n. sp., adult female: 46, idiosoma, dorsal aspect, supernumerary setae denoted by ‘ x’; 47, idiosoma, ventral aspect; 48, spermathecae, detail of calyx; 49, position of spermatheca at base of leg IV;" captionText-3="FIGURES 51 – 55. Spadiseius spathiphyllae n. sp., adult male. Figs. 51 – 53, idiosomas, dorsal aspect: 51, “ long-haired ” morph; 52, intermediate morph; 53, “ short-haired ” morph, partial view. Fig. 54, detail of peritrematal shield; note variation in length of peritreme from Fig. 52. Fig. 55, chelicera and spermatodactyl, adaxial view." captionText-4="FIGURES 56 – 58. Spadiseius spathiphyllae n. sp., adult male. Figs. 56, 57, opisthogaster: 56, intermediate morph; 57, “ short-haired ” morph. Fig. 58, idiosoma, ventral aspect, “ long-haired ” morph." captionText-5="FIGURES 59 – 64. Spadiseius spathiphyllae n. sp., adult female: 59, subcapitulum, abbreviation: k, keel; 60, salivary stylus; 61, tarsus I, dorsal view, without pretarsal structures; 62, detail of dorsodistal sensillar complex of tarsus I; 63, enlarged detail of sigmoid seta “ s ”; 64, leg I, ventral view, excluding tarsus; note serrated apical sclerotized rims of segments." captionText-6="FIGURES 65 – 69. Figs. 65 – 67: Spadiseius spathiphyllae n. sp., adult male: 65, tarsus IV, dorsolateral view, without pretarsus; 66, 67, femora IV, dorsal view: 66, “ short-haired ” morph; 67, “ long-haired ” morph. Figs. 68, 69, Spadiseius calyptrogynae n. sp., adult male, femora IV, dorsal view: 68, “ wartless ” morph; 69, “ warted ” morph." httpUri-0="https://zenodo.org/record/180491/files/figure.png" httpUri-1="https://zenodo.org/record/180492/files/figure.png" httpUri-2="https://zenodo.org/record/180493/files/figure.png" httpUri-3="https://zenodo.org/record/180494/files/figure.png" httpUri-4="https://zenodo.org/record/180495/files/figure.png" httpUri-5="https://zenodo.org/record/180496/files/figure.png" httpUri-6="https://zenodo.org/record/180497/files/figure.png" pageId="21" pageNumber="22">39, 41–43, 45–67</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B2F1655CFF81FFC73DE8FE8F2673B93A" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="FA5436D7FF81FFC73DE8FE8F2673B93A" blockId="21.[151,1437,266,2012]" pageId="21" pageNumber="22">
<emphasis id="C89FEAC5FF81FFC73DE8FE8F26AEBB4A" bold="true" box="[151,415,266,292]" pageId="21" pageNumber="22">Diagnosis. All instars</emphasis>
: legs with apical sclerotized rim of femur, genu, tibia clearly serrated ventrally. 
<emphasis id="C89FEAC5FF81FFC73833FE8F22A2BB4A" bold="true" box="[1356,1427,266,292]" pageId="21" pageNumber="22">Adult</emphasis>
: peritremes extending anteriorly only to level of marginal setae 
<emphasis id="C89FEAC5FF81FFC73EE9FEB724E8BB25" box="[918,985,306,331]" italics="true" pageId="21" pageNumber="22">r2–r5</emphasis>
, those of female thick; hypostomatic setae 
<emphasis id="C89FEAC5FF81FFC73DA5FEDF2623BB1D" box="[218,274,346,371]" italics="true" pageId="21" pageNumber="22">hyp4</emphasis>
shorter than 
<emphasis id="C89FEAC5FF81FFC73CD0FEDF2501BB1D" box="[431,560,346,371]" italics="true" pageId="21" pageNumber="22">hyp1–hyp2</emphasis>
, but as long as 
<emphasis id="C89FEAC5FF81FFC73F94FEDF2415BB1D" box="[747,804,346,371]" italics="true" pageId="21" pageNumber="22">hyp3</emphasis>
. 
<emphasis id="C89FEAC5FF81FFC73E4BFEDF24E6BB1A" bold="true" box="[820,983,346,372]" pageId="21" pageNumber="22">Adult female</emphasis>
: marginal setae 
<emphasis id="C89FEAC5FF81FFC739E5FEDF23ECBB1D" box="[1178,1245,346,371]" italics="true" pageId="21" pageNumber="22">r2–r3</emphasis>
on edge of prodorsal shield, 
<emphasis id="C89FEAC5FF81FFC73C45FE072662BBF5" box="[314,339,386,411]" italics="true" pageId="21" pageNumber="22">r4</emphasis>
on soft cuticle; subcapitulum without pair of gland-like structures between insertions of setae 
<emphasis id="C89FEAC5FF81FFC73DE8FE2F27E1BBAD" box="[151,208,426,451]" italics="true" pageId="21" pageNumber="22">hyp1</emphasis>
, and with setae 
<emphasis id="C89FEAC5FF81FFC73CEEFE2F26FBBBAD" box="[401,458,426,451]" italics="true" pageId="21" pageNumber="22">hyp4</emphasis>
not swollen basally. 
<emphasis id="C89FEAC5FF81FFC73FBBFE2F247EBBAA" bold="true" box="[708,847,426,452]" pageId="21" pageNumber="22">Adult male</emphasis>
: dorsal shield with setae in central region moderately short, blade-like, not inserted on warts, and with a variable number of peripheral setae of 
<emphasis id="C89FEAC5FF81FFC73998FE572220BB85" box="[1255,1297,466,491]" italics="true" pageId="21" pageNumber="22">z–Z</emphasis>
, s–S and 
<emphasis id="C89FEAC5FF81FFC738FDFE572798B87D" italics="true" pageId="21" pageNumber="22">r– R</emphasis>
series forming a corona of slightly to clearly enlarged setae, either thickened and more or less capitate apically or attenuated; presternal area without platelets; peritrematal shield gland pore not enlarged at level between coxae II and III; spermatodactyl simple in form, sinuate, tapered apically; leg IV with a spur on basitarsus. 
<emphasis id="C89FEAC5FF81FFC73D94FDF726BFB8E2" bold="true" box="[235,398,626,652]" pageId="21" pageNumber="22">Deutonymph</emphasis>
: coxa IV seta subequally as long as setae on coxa III. 
<emphasis id="C89FEAC5FF81FFC7396AFDF722A5B8E2" bold="true" box="[1045,1428,626,652]" pageId="21" pageNumber="22">Deutonymph and protonymph</emphasis>
: legs I–IV with femoral seta 
<emphasis id="C89FEAC5FF81FFC73C95FD1F252EB8DD" box="[490,543,666,691]" italics="true" pageId="21" pageNumber="22">ad-1</emphasis>
thicker and different in spatulate form than adjacent tapered setae. 
<emphasis id="C89FEAC5FF81FFC73833FD1F27DEB8B2" bold="true" pageId="21" pageNumber="22">Protonymph</emphasis>
: pygidial shield without hump or tubercle laterad seta 
<emphasis id="C89FEAC5FF81FFC73E13FD4724B8B8B5" box="[876,905,706,731]" italics="true" pageId="21" pageNumber="22">S5</emphasis>
. 
<emphasis id="C89FEAC5FF81FFC73EE7FD4724D5B8B2" bold="true" box="[920,996,706,732]" pageId="21" pageNumber="22">Larva</emphasis>
: podonotal shield conspicuously rugose over much of surface; pygidial shield without puncta, with medial projection of anterior margin extending slightly beyond level of setal alveoli 
<emphasis id="C89FEAC5FF81FFC73F2FFC97255DB945" box="[592,620,786,811]" italics="true" pageId="21" pageNumber="22">J2</emphasis>
; legs I–III with femoral seta 
<emphasis id="C89FEAC5FF81FFC73EB5FC972331B945" box="[970,1024,786,811]" italics="true" pageId="21" pageNumber="22">ad-1</emphasis>
rod-like, thicker, less tapered than adjacent setae.
</paragraph>
</subSubSection>
<subSubSection id="B2F1655CFF81FFCC3DB9FCE72248BB14" lastPageId="30" lastPageNumber="31" pageId="21" pageNumber="22" type="description">
<paragraph id="FA5436D7FF81FFC73DB9FCE7239EBF62" blockId="21.[151,1437,266,2012]" pageId="21" pageNumber="22">
<emphasis id="C89FEAC5FF81FFC73DB9FCE72519B915" bold="true" box="[198,552,866,892]" pageId="21" pageNumber="22">Description. ADULT FEMALE</emphasis>
. Dorsal shield 420–485 long, 230–275 at its greatest width at level of setae 
<emphasis id="C89FEAC5FF81FFC73DE8FC0F2780B9CD" box="[151,177,906,931]" italics="true" pageId="21" pageNumber="22">r3</emphasis>
(n = 6); shield wrinkled on vertex and conspicuously reticulate over entire surface (
<figureCitation id="62D02A52FF81FFC739CCFC0F223FB9CA" box="[1203,1294,906,932]" captionStart="FIGURES 46 – 50" captionStartId="27.[151,269,1773,1797]" captionTargetBox="[186,1409,183,1772]" captionTargetId="figure@27.[186,1411,178,1937]" captionTargetPageId="27" captionText="FIGURES 46 – 50. Spadiseius spathiphyllae n. sp., adult female: 46, idiosoma, dorsal aspect, supernumerary setae denoted by ‘ x’; 47, idiosoma, ventral aspect; 48, spermathecae, detail of calyx; 49, position of spermatheca at base of leg IV;" httpUri="https://zenodo.org/record/180493/files/figure.png" pageId="21" pageNumber="22">Fig. 46</figureCitation>
); podonotal region with longitudinal distance from alveoli of 
<emphasis id="C89FEAC5FF81FFC73F9BFC3725CAB9A5" box="[740,763,946,971]" italics="true" pageId="21" pageNumber="22">j1</emphasis>
to 
<emphasis id="C89FEAC5FF81FFC73E5BFC37240BB9A5" box="[804,826,946,971]" italics="true" pageId="21" pageNumber="22">j5</emphasis>
subequal to that from 
<emphasis id="C89FEAC5FF81FFC73934FC372350B9A5" box="[1099,1121,946,971]" italics="true" pageId="21" pageNumber="22">j5</emphasis>
to 
<emphasis id="C89FEAC5FF81FFC739F4FC372390B9A5" box="[1163,1185,946,971]" italics="true" pageId="21" pageNumber="22">j6</emphasis>
. Peritrematal shields uniting with dorsal shield at level of setae 
<emphasis id="C89FEAC5FF81FFC73FFDFC5F25ADB99D" box="[642,668,986,1011]" italics="true" pageId="21" pageNumber="22">s1</emphasis>
; peritremes thick, extending to level of setae 
<emphasis id="C89FEAC5FF81FFC739D2FC5F23C1B99D" box="[1197,1264,986,1011]" italics="true" pageId="21" pageNumber="22">r2–r3</emphasis>
. Body dorsum with approximately 56–58 pairs of setae, 11 or 12 of which (
<emphasis id="C89FEAC5FF81FFC73E19FB872499BE75" box="[870,936,1026,1051]" italics="true" pageId="21" pageNumber="22">r4–r5</emphasis>
, 
<emphasis id="C89FEAC5FF81FFC73EC7FB87233FBE75" box="[952,1038,1026,1051]" italics="true" pageId="21" pageNumber="22">R1–R6,</emphasis>
and 3 or 4 
<emphasis id="C89FEAC5FF81FFC739E8FB8723FCBE75" box="[1175,1229,1026,1051]" italics="true" pageId="21" pageNumber="22">UR–</emphasis>
setae) inserted on soft cuticle; dorsal shield with 4 or 5 extra pairs of setae amidst 
<emphasis id="C89FEAC5FF81FFC73ED6FBAF2480BE2D" box="[937,945,1066,1091]" italics="true" pageId="21" pageNumber="22">j</emphasis>
, 
<emphasis id="C89FEAC5FF81FFC73EBDFBAF24FCBE2D" box="[962,973,1066,1091]" italics="true" pageId="21" pageNumber="22">z</emphasis>
and 
<emphasis id="C89FEAC5FF81FFC73973FBAF2326BE2D" box="[1036,1047,1066,1091]" italics="true" pageId="21" pageNumber="22">s</emphasis>
series on podonotal region, and sometimes an extra pair of setae between 
<emphasis id="C89FEAC5FF81FFC73FE4FBD62589BE02" box="[667,696,1107,1132]" italics="true" pageId="21" pageNumber="22">S1</emphasis>
and 
<emphasis id="C89FEAC5FF81FFC73F86FBD62426BE02" box="[761,791,1107,1132]" italics="true" pageId="21" pageNumber="22">Z1</emphasis>
and several paired or asymmetrically unpaired setae amidst 
<emphasis id="C89FEAC5FF81FFC73D92FBFE27CBBEFA" box="[237,250,1147,1172]" italics="true" pageId="21" pageNumber="22">J</emphasis>
series on opisthonotal region. Dorsal setae uniform in simple but slightly curved, scimitar-like form (most of them slightly swollen at base) and moderately short length (15–21), 
<emphasis id="C89FEAC5FF81FFC73952FB26237BBED2" box="[1069,1098,1187,1212]" italics="true" pageId="21" pageNumber="22">S5</emphasis>
(25–28) 
<emphasis id="C89FEAC5FF81FFC739C8FB2623E6BED2" box="[1207,1239,1187,1212]" italics="true" pageId="21" pageNumber="22">Z5</emphasis>
(28–30) slightly largest, 
<emphasis id="C89FEAC5FF81FFC73D8AFB4E2620BE8A" box="[245,273,1227,1252]" italics="true" pageId="21" pageNumber="22">J5</emphasis>
clearly smallest (10–12). Anterodorsal expansions of exopodal plates above legs I formed as lineated, wrinkly plates (not illustrated, difficult to discern) touching vertex of dorsal shield.
</paragraph>
<paragraph id="FA5436D7FF81FFC73DB9FA9F2698BDE2" blockId="21.[151,1437,266,2012]" pageId="21" pageNumber="22">
Tritosternum with length of trapezoidal base (15–18) slightly shorter than basal width (24–27); laciniae free for about 0.6–0.7 of total length (60–70 excluding base). Presternal area weakly sclerotized, lineate, insensibly consolidated with sternal shield. Sternal shield (
<figureCitation id="62D02A52FF81FFC73E22FAEF2487BFEA" box="[861,950,1386,1412]" captionStart="FIGURES 46 – 50" captionStartId="27.[151,269,1773,1797]" captionTargetBox="[186,1409,183,1772]" captionTargetId="figure@27.[186,1411,178,1937]" captionTargetPageId="27" captionText="FIGURES 46 – 50. Spadiseius spathiphyllae n. sp., adult female: 46, idiosoma, dorsal aspect, supernumerary setae denoted by ‘ x’; 47, idiosoma, ventral aspect; 48, spermathecae, detail of calyx; 49, position of spermatheca at base of leg IV;" httpUri="https://zenodo.org/record/180493/files/figure.png" pageId="21" pageNumber="22">Fig. 47</figureCitation>
) faintly reticulated anteriorly, becoming more weakly sclerotized and unornamented (or desclerotized) posterior to level of second poroids; posterior margin faint, slightly concave; setae 
<emphasis id="C89FEAC5FF81FFC73F36FA3E25AABFBA" box="[585,667,1467,1492]" italics="true" pageId="21" pageNumber="22">st1–st2</emphasis>
and first and second pairs of poroids inserted on sclerotized portion of shield, 
<emphasis id="C89FEAC5FF81FFC73C33FA66265FBF92" box="[332,366,1507,1532]" italics="true" pageId="21" pageNumber="22">st3</emphasis>
inserted on desclerotized portion; 
<emphasis id="C89FEAC5FF81FFC73E67FA66245DBF92" box="[792,876,1507,1532]" italics="true" pageId="21" pageNumber="22">st1–st3</emphasis>
slightly attenuated, similar in length (37–40), slightly longer than 
<emphasis id="C89FEAC5FF81FFC73CFAF98E2696BC4A" box="[389,423,1547,1572]" italics="true" pageId="21" pageNumber="22">st4</emphasis>
(33) and 
<emphasis id="C89FEAC5FF81FFC73F63F98E250FBC4A" box="[540,574,1547,1572]" italics="true" pageId="21" pageNumber="22">st5</emphasis>
(33) on soft cuticle. Endopodal strips faintly formed alongside coxae III and IV. Genital shield smooth or faintly lineated, its lateral margins nearly straight or slightly concave alongside genital setae. Anal shield weakly reticulate-lineate, about as long as wide (72–75) at level of anterior edge of anus; postanal seta subequally as long as para-anal setae (17). Ten pairs of opisthogastric setae (
<emphasis id="C89FEAC5FF81FFC7381FF9062677BCAA" italics="true" pageId="21" pageNumber="22">JV1– JV5, ZV1–ZV5</emphasis>
) plus 1 or 2 pairs of adjacent 
<emphasis id="C89FEAC5FF81FFC73FCBF92E25D8BCAA" box="[692,745,1707,1732]" italics="true" pageId="21" pageNumber="22">UR–</emphasis>
setae on soft cuticle around anal shield, all similar in size (17–25) except for slightly thicker 
<emphasis id="C89FEAC5FF81FFC73F4EF956256FBC82" box="[561,606,1747,1772]" italics="true" pageId="21" pageNumber="22">JV5</emphasis>
, 
<emphasis id="C89FEAC5FF81FFC73F12F95625EFBC82" box="[621,734,1747,1772]" italics="true" pageId="21" pageNumber="22">ZV4, ZV5</emphasis>
(22–25). Metapodal plates subdivided, larger piece slender, pointed anteriorly and posteriorly, about 23–32 long, 7–8 wide, smaller piece linear, difficult to discern. Spermathecal apparatus with thin, visibly dense, nearly straight conduit (section of major duct from embolus to apex of calyx, 30–35) leading from solenostome between bases of legs III and IV to a hyaline wrinkled sac (vesicle) (
<figureCitation id="62D02A52FF81FFC73C74F8F726ADBDE2" box="[267,412,1906,1932]" captionStart="FIGURES 46 – 50" captionStartId="27.[151,269,1773,1797]" captionTargetBox="[186,1409,183,1772]" captionTargetId="figure@27.[186,1411,178,1937]" captionTargetPageId="27" captionText="FIGURES 46 – 50. Spadiseius spathiphyllae n. sp., adult female: 46, idiosoma, dorsal aspect, supernumerary setae denoted by ‘ x’; 47, idiosoma, ventral aspect; 48, spermathecae, detail of calyx; 49, position of spermatheca at base of leg IV;" httpUri="https://zenodo.org/record/180493/files/figure.png" pageId="21" pageNumber="22">Figs. 48–49</figureCitation>
).
</paragraph>
<paragraph id="FA5436D7FF81FFC43DB9F81F2209B874" blockId="21.[151,1437,266,2012]" lastBlockId="22.[151,1436,152,538]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
Tectum with anterior margin smooth, sharply pointed medially (
<figureCitation id="62D02A52FF81FFC73EBAF81F233EBDDA" box="[965,1039,1946,1972]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="21" pageNumber="22">Fig. 7</figureCitation>
). Fixed chela with 2 or 3 minute teeth beside offset tooth subapically (
<figureCitation id="62D02A52FF81FFC73F25F8472585BDB2" box="[602,692,1986,2012]" captionStart="FIGURES 46 – 50" captionStartId="27.[151,269,1773,1797]" captionTargetBox="[186,1409,183,1772]" captionTargetId="figure@27.[186,1411,178,1937]" captionTargetPageId="27" captionText="FIGURES 46 – 50. Spadiseius spathiphyllae n. sp., adult female: 46, idiosoma, dorsal aspect, supernumerary setae denoted by ‘ x’; 47, idiosoma, ventral aspect; 48, spermathecae, detail of calyx; 49, position of spermatheca at base of leg IV;" httpUri="https://zenodo.org/record/180493/files/figure.png" pageId="21" pageNumber="22">Fig. 50</figureCitation>
); movable chela (length 30–35) usually weakly bidentate, the more apical tooth hardly discernible, sometimes absent; antiaxial face of movable chela with sclerotized process along its basal half (
<figureCitation id="62D02A52FF82FFC43CC3FF452532BAB4" box="[444,515,192,218]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
b); ventral mucro slender, attenuate, reaching mid-level of chela (
<figureCitation id="62D02A52FF82FFC4387FFF452263BAB4" box="[1280,1362,192,218]" captionStart="FIGURES 1 – 9" captionStartId="9.[151,269,1577,1601]" captionTargetBox="[152,1425,197,1576]" captionTargetId="figure@9.[151,1436,197,1643]" captionTargetPageId="9" captionText="FIGURES 1 – 9. Figs. 1 – 5, Spadiseius calyptrogynae n. sp., adult female: 1, subcapitulum; 2, labrum, ventral view; 3, salivary stylus; 4, tritosternum; 5, palpus, ventral view. Fig. 6, S. calyptrogynae n. sp., larva, chelicera, paraxial view. Figs. 7 – 9, Spadiseius spathiphyllae n. sp., adult female: 7, tectum; 8, chelicera, dorso-paraxial view; 9 a, chelicera, movable digit, dorsal view; 9 b, chelicera, movable digit, antiaxial-ventral view (Figs. 2 – 4, same scale)." httpUri="https://zenodo.org/record/180480/files/figure.png" pageId="22" pageNumber="23">Figs. 9</figureCitation>
a, 50). Corniculi strongly convergent, sometimes overlapping apically (
<figureCitation id="62D02A52FF82FFC43ED4FF6D2339BB6C" box="[939,1032,232,258]" captionStart="FIGURES 59 – 64" captionStartId="30.[151,269,1546,1570]" captionTargetBox="[152,1432,441,1545]" captionTargetId="figure@30.[151,1436,434,1616]" captionTargetPageId="30" captionText="FIGURES 59 – 64. Spadiseius spathiphyllae n. sp., adult female: 59, subcapitulum, abbreviation: k, keel; 60, salivary stylus; 61, tarsus I, dorsal view, without pretarsal structures; 62, detail of dorsodistal sensillar complex of tarsus I; 63, enlarged detail of sigmoid seta “ s ”; 64, leg I, ventral view, excluding tarsus; note serrated apical sclerotized rims of segments." httpUri="https://zenodo.org/record/180496/files/figure.png" pageId="22" pageNumber="23">Fig. 59</figureCitation>
); internal malae slender, slightly longer than corniculi, their lateral margins finely fringed. Deutosternum with 7 or 8 transverse rows of denticles, evenly multidenticulate (7–12 denticles), none distinctly widened. Subcapitulum lacking gland-like structures between bases of anterior subcapitular setae; setae finely tapered, medial pair longest; capitular pair not noticeably thickened basally, shorter (22) than anterior (25–28) and medial (35–37) pairs. Subcapitulum with serrated ridge extending on either side laterally from bases of capitular setae (
<figureCitation id="62D02A52FF82FFC439FFFE3523EBBBA4" box="[1152,1242,432,458]" captionStart="FIGURES 59 – 64" captionStartId="30.[151,269,1546,1570]" captionTargetBox="[152,1432,441,1545]" captionTargetId="figure@30.[151,1436,434,1616]" captionTargetPageId="30" captionText="FIGURES 59 – 64. Spadiseius spathiphyllae n. sp., adult female: 59, subcapitulum, abbreviation: k, keel; 60, salivary stylus; 61, tarsus I, dorsal view, without pretarsal structures; 62, detail of dorsodistal sensillar complex of tarsus I; 63, enlarged detail of sigmoid seta “ s ”; 64, leg I, ventral view, excluding tarsus; note serrated apical sclerotized rims of segments." httpUri="https://zenodo.org/record/180496/files/figure.png" pageId="22" pageNumber="23">Fig. 59</figureCitation>
). Palptrochanter setae slender, subequal in length (16–18); palpgenu with seta 
<emphasis id="C89FEAC5FF82FFC43E0BFE5D2492BB9F" box="[884,931,472,497]" italics="true" pageId="22" pageNumber="23">al-2</emphasis>
tapered, similar in form to 
<emphasis id="C89FEAC5FF82FFC43993FE5D222ABB9F" box="[1260,1307,472,497]" italics="true" pageId="22" pageNumber="23">
al- 
<quantity id="3D139B32FF82FFC43870FE5D222ABB9F" box="[1295,1307,472,497]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="22" pageNumber="23" unit="in" value="1.0">1</quantity>
</emphasis>
in distinction to spatulate 
<emphasis id="C89FEAC5FF82FFC43C26FD852641B877" box="[345,368,512,537]" italics="true" pageId="22" pageNumber="23">al</emphasis>
of palpfemur; apical sclerotized rim of palpfemur and palpgenu serrated ventrally.
</paragraph>
<caption id="AE94665FFF82FFC43DE8F95B24F0BD76" httpUri="https://zenodo.org/record/180490/files/figure.png" pageId="22" pageNumber="23" targetBox="[151,1434,598,1757]" targetPageId="22">
<paragraph id="FA5436D7FF82FFC43DE8F95B24F0BD76" blockId="22.[151,1436,1758,1817]" pageId="22" pageNumber="23">
<emphasis id="C89FEAC5FF82FFC43DE8F95B266FBC98" bold="true" box="[151,350,1758,1782]" pageId="22" pageNumber="23">FIGURES 35–37.</emphasis>
<taxonomicName id="3DEB4D54FF82FFC43C19F95A255FBC98" box="[358,622,1759,1782]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF82FFC43C19F95A255FBC98" box="[358,622,1759,1782]" italics="true" pageId="22" pageNumber="23">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF82FFC43F09F95B2582BC98" box="[630,691,1758,1782]" pageId="22" pageNumber="23" rank="species">
<emphasis id="C89FEAC5FF82FFC43F09F95B2582BC98" bold="true" box="[630,691,1758,1782]" pageId="22" pageNumber="23">n. sp.</emphasis>
</taxonomicNameLabel>
35, larva, idiosoma, dorsal aspect; 36, protonymph, idiosoma, dorsal aspect; 37, larva, chelicera, adaxial view. Abbreviation: 
<emphasis id="C89FEAC5FF82FFC43F92F8842426BD77" bold="true" box="[749,791,1793,1817]" pageId="22" pageNumber="23">v m</emphasis>
, ventral mucro.
</paragraph>
</caption>
<paragraph id="FA5436D7FF82FFC53DB9F8C9263DBB14" blockId="22.[151,1436,1868,2014]" lastBlockId="23.[151,1436,152,938]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
Legs IV the longest (415–440 excluding pretarsus) of legs, about as long as dorsal shield; other leg lengths: I 355–370, II 320–330, III 345–355. Tarsi II–IV with pair of apical setal processes (
<emphasis id="C89FEAC5FF82FFC439AFF8F12278BDE3" box="[1232,1353,1908,1933]" italics="true" pageId="22" pageNumber="23">ad-1, pd-1</emphasis>
) about one-third as long as length of pretarsi to base of claws. Leg II slightly thicker than legs III–IV, but thickness and form of its setae not differentiated from those of legs III–IV (as in 
<figureCitation id="62D02A52FF82FFC43E9FF8412341BDB0" box="[992,1136,1988,2014]" captionStart="FIGURES 30 – 34" captionStartId="19.[151,269,1464,1488]" captionTargetBox="[164,1434,194,1391]" captionTargetId="figure@19.[151,1436,194,1440]" captionTargetPageId="19" captionText="FIGURES 30 – 34. Spadiseius calyptrogynae n. sp., adult female. Figs. 30 – 33, dorsal views of legs excluding tarsi: 30, leg I; 31, leg II; 32, leg III; 33, leg IV; anterolateral (al) and posterolateral (pl) faces indicated by arrows. Fig. 34, tarsus IV, dorsal view." httpUri="https://zenodo.org/record/180489/files/figure.png" pageId="22" pageNumber="23">Figs. 31–33</figureCitation>
). Coxae I with many serrated ridges on ventral face (
<figureCitation id="62D02A52FF83FFC53C96FF1D2570BADC" box="[489,577,152,178]" captionStart="FIGURES 59 – 64" captionStartId="30.[151,269,1546,1570]" captionTargetBox="[152,1432,441,1545]" captionTargetId="figure@30.[151,1436,434,1616]" captionTargetPageId="30" captionText="FIGURES 59 – 64. Spadiseius spathiphyllae n. sp., adult female: 59, subcapitulum, abbreviation: k, keel; 60, salivary stylus; 61, tarsus I, dorsal view, without pretarsal structures; 62, detail of dorsodistal sensillar complex of tarsus I; 63, enlarged detail of sigmoid seta “ s ”; 64, leg I, ventral view, excluding tarsus; note serrated apical sclerotized rims of segments." httpUri="https://zenodo.org/record/180496/files/figure.png" pageId="23" pageNumber="24">Fig. 64</figureCitation>
), coxae II–IV with 2 or 3 lines on posteroventral faces, these serrated on II; seta on coxa IV about as long (25–34) as setae on coxa III. Apical sclerotized rim of femur, genu, tibia clearly serrated ventrally on legs I–IV (
<figureCitation id="62D02A52FF83FFC53F1DFF6D258BBB6C" box="[610,698,232,258]" captionStart="FIGURES 59 – 64" captionStartId="30.[151,269,1546,1570]" captionTargetBox="[152,1432,441,1545]" captionTargetId="figure@30.[151,1436,434,1616]" captionTargetPageId="30" captionText="FIGURES 59 – 64. Spadiseius spathiphyllae n. sp., adult female: 59, subcapitulum, abbreviation: k, keel; 60, salivary stylus; 61, tarsus I, dorsal view, without pretarsal structures; 62, detail of dorsodistal sensillar complex of tarsus I; 63, enlarged detail of sigmoid seta “ s ”; 64, leg I, ventral view, excluding tarsus; note serrated apical sclerotized rims of segments." httpUri="https://zenodo.org/record/180496/files/figure.png" pageId="23" pageNumber="24">Fig. 64</figureCitation>
). Femora I–IV with dorsal seta 
<emphasis id="C89FEAC5FF83FFC53953FF6D2350BB6F" box="[1068,1121,232,257]" italics="true" pageId="23" pageNumber="24">ad-1</emphasis>
thickened, untapered, rodlike or slightly oar-like in contrast with adjacent simple setae, slightly longer from I (14–17) to IV (22–25) (much as in 
<figureCitation id="62D02A52FF83FFC53C55FEBD268BBB3C" box="[298,442,312,338]" captionStart="FIGURES 30 – 34" captionStartId="19.[151,269,1464,1488]" captionTargetBox="[164,1434,194,1391]" captionTargetId="figure@19.[151,1436,194,1440]" captionTargetPageId="19" captionText="FIGURES 30 – 34. Spadiseius calyptrogynae n. sp., adult female. Figs. 30 – 33, dorsal views of legs excluding tarsi: 30, leg I; 31, leg II; 32, leg III; 33, leg IV; anterolateral (al) and posterolateral (pl) faces indicated by arrows. Fig. 34, tarsus IV, dorsal view." httpUri="https://zenodo.org/record/180489/files/figure.png" pageId="23" pageNumber="24">Figs. 30–33</figureCitation>
); legs without elongated macrosetae. Setation of genua and tibiae I–IV as presented for genus.
</paragraph>
<paragraph id="FA5436D7FF83FFC53DB9FE022601B9C4" blockId="23.[151,1436,152,938]" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC53DB9FE022645BBCF" bold="true" box="[198,372,391,417]" pageId="23" pageNumber="24">ADULT MALE.</emphasis>
Dorsal shield 380–475 long, 265–350 at its greatest width at level of setae 
<emphasis id="C89FEAC5FF83FFC53873FE0D2217BBCF" box="[1292,1318,392,417]" italics="true" pageId="23" pageNumber="24">r5</emphasis>
(n = 10); podonotal region with longitudinal distance from alveoli of 
<emphasis id="C89FEAC5FF83FFC53E27FE35245EBBA7" box="[856,879,432,457]" italics="true" pageId="23" pageNumber="24">j1</emphasis>
to 
<emphasis id="C89FEAC5FF83FFC53EEAFE35249DBBA7" box="[917,940,432,457]" italics="true" pageId="23" pageNumber="24">j5</emphasis>
subequal to that from 
<emphasis id="C89FEAC5FF83FFC539C9FE3523FDBBA7" box="[1206,1228,432,457]" italics="true" pageId="23" pageNumber="24">j5</emphasis>
to 
<emphasis id="C89FEAC5FF83FFC5398CFE352238BBA7" box="[1267,1289,432,457]" italics="true" pageId="23" pageNumber="24">j6</emphasis>
; central and posterior area of shield with uniformly blade-like setae of mostly similar length, including some hypertrichy, on a given specimen but of variably short (15–25) to moderate length (25–40) among different specimens, and with a more or less well developed corona of a variable number of enlarged (lengths from 30 to 160) and either attenuated or apically capitate or oar-like setae peripherally (
<figureCitation id="62D02A52FF83FFC53ECBFDD52374B804" box="[948,1093,592,618]" captionStart="FIGURES 51 – 55" captionStartId="28.[151,269,1906,1930]" captionTargetBox="[206,1386,432,1878]" captionTargetId="figure@28.[206,1393,416,1899]" captionTargetPageId="28" captionText="FIGURES 51 – 55. Spadiseius spathiphyllae n. sp., adult male. Figs. 51 – 53, idiosomas, dorsal aspect: 51, “ long-haired ” morph; 52, intermediate morph; 53, “ short-haired ” morph, partial view. Fig. 54, detail of peritrematal shield; note variation in length of peritreme from Fig. 52. Fig. 55, chelicera and spermatodactyl, adaxial view." httpUri="https://zenodo.org/record/180494/files/figure.png" pageId="23" pageNumber="24">Figs. 51–53</figureCitation>
); setae 
<emphasis id="C89FEAC5FF83FFC539E1FDD52384B807" box="[1182,1205,592,617]" italics="true" pageId="23" pageNumber="24">j1</emphasis>
variably blade-like (14–15) or untapered (22–30) or attenuated (50–57); whether elongated or not, 
<emphasis id="C89FEAC5FF83FFC53948FDFD2362B8FF" box="[1079,1107,632,657]" italics="true" pageId="23" pageNumber="24">S1</emphasis>
consistently shorter (1/3–2/ 3) than adjacent setae 
<emphasis id="C89FEAC5FF83FFC53CDBFD25268CB8D7" box="[420,445,672,697]" italics="true" pageId="23" pageNumber="24">s6</emphasis>
and 
<emphasis id="C89FEAC5FF83FFC53C84FD252529B8D7" box="[507,536,672,697]" italics="true" pageId="23" pageNumber="24">S2</emphasis>
(
<tableCitation id="B769036CFF83FFC53F55FD2525B6B8D4" box="[554,647,672,698]" captionStart="TABLE 2" captionStartId="23.[151,239,991,1015]" captionTargetBox="[159,1407,1048,1532]" captionTargetPageId="23" captionText="TABLE 2. Polymorphic male measurements (µm) - Spadiseius spathiphyllae n. sp." httpUri="http://table.plazi.org/id/AE94665FFF83FFC53DE8FC5A2323B999" pageId="23" pageNumber="24" tableUuid="AE94665FFF83FFC53DE8FC5A2323B999">Table 2</tableCitation>
). Dorsal shield entirely reticulated (not shown in figures), consistently without wart-like thickenings. Peritrematal shields uniting with dorsal shield at level between bases of legs II and III; peritrematal shield gland pore at that level not enlarged (
<figureCitation id="62D02A52FF83FFC53E9DFD75230BB964" box="[994,1082,752,778]" captionStart="FIGURES 51 – 55" captionStartId="28.[151,269,1906,1930]" captionTargetBox="[206,1386,432,1878]" captionTargetId="figure@28.[206,1393,416,1899]" captionTargetPageId="28" captionText="FIGURES 51 – 55. Spadiseius spathiphyllae n. sp., adult male. Figs. 51 – 53, idiosomas, dorsal aspect: 51, “ long-haired ” morph; 52, intermediate morph; 53, “ short-haired ” morph, partial view. Fig. 54, detail of peritrematal shield; note variation in length of peritreme from Fig. 52. Fig. 55, chelicera and spermatodactyl, adaxial view." httpUri="https://zenodo.org/record/180494/files/figure.png" pageId="23" pageNumber="24">Fig. 54</figureCitation>
); peritremes variable in thickness and length, extending to level of setae 
<emphasis id="C89FEAC5FF83FFC53FEAFC9D2581B95F" box="[661,688,792,817]" italics="true" pageId="23" pageNumber="24">r5</emphasis>
or nearly to 
<emphasis id="C89FEAC5FF83FFC53E3AFC9D246EB95F" box="[837,863,792,817]" italics="true" pageId="23" pageNumber="24">r2</emphasis>
. Dorsal shield complement of setae, poroids and gland-pores similar to that in female, except marginals 
<emphasis id="C89FEAC5FF83FFC53E67FCC52403B937" box="[792,818,832,857]" italics="true" pageId="23" pageNumber="24">r5</emphasis>
, 
<emphasis id="C89FEAC5FF83FFC53E3FFCC524A1B937" box="[832,912,832,857]" italics="true" pageId="23" pageNumber="24">R1–R2</emphasis>
, 
<emphasis id="C89FEAC5FF83FFC53EE1FCC524DFB937" box="[926,1006,832,857]" italics="true" pageId="23" pageNumber="24">R4–R6</emphasis>
, poroid 
<collectingCountry id="82FC7647FF83FFC53930FCC52341B937" box="[1103,1136,832,857]" name="Poland" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC53930FCC52341B937" box="[1103,1136,832,857]" italics="true" pageId="23" pageNumber="24">Rp</emphasis>
</collectingCountry>
and sometimes 1 or 2 
<emphasis id="C89FEAC5FF83FFC5380AFCC522ADB937" box="[1397,1436,832,857]" italics="true" pageId="23" pageNumber="24">UR</emphasis>
setae usually on weakly sclerotized expanded margins of shield, though variably on soft cuticle; 
<emphasis id="C89FEAC5FF83FFC53855FCED227BB9EF" box="[1322,1354,872,897]" italics="true" pageId="23" pageNumber="24">R3</emphasis>
sometimes absent.
</paragraph>
<caption id="AE94665FFF83FFC53DE8FC5A2323B999" ID-Table-UUID="AE94665FFF83FFC53DE8FC5A2323B999" box="[151,1042,991,1015]" httpUri="http://table.plazi.org/id/AE94665FFF83FFC53DE8FC5A2323B999" pageId="23" pageNumber="24" targetBox="[159,1407,1048,1532]" targetIsTable="true" targetPageId="23">
<paragraph id="FA5436D7FF83FFC53DE8FC5A2323B999" blockId="23.[151,1042,991,1015]" box="[151,1042,991,1015]" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC53DE8FC5A2639B999" bold="true" box="[151,264,991,1015]" pageId="23" pageNumber="24">TABLE 2.</emphasis>
Polymorphic male measurements (µm) - 
<taxonomicName id="3DEB4D54FF83FFC53FB9FC6524FEB999" box="[710,975,992,1015]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF83FFC53FB9FC6524FEB999" box="[710,975,992,1015]" italics="true" pageId="23" pageNumber="24">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF83FFC53EAAFC5A2323B999" box="[981,1042,991,1015]" pageId="23" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="FA5436D7FF83FFC53DE0FB9D2250BF92" pageId="23" pageNumber="24">
<table id="88EBC477FF83002D3DE0FB9D224EBF92" box="[159,1407,1048,1532]" gridcols="11" gridrows="8" pageId="23" pageNumber="24">
<tr id="44DB3495FF83002D3DE0FB9D224EBE3F" box="[159,1407,1048,1105]" gridrow="0" pageId="23" pageNumber="24">
<th id="070A5DE9FF83002D3DE0FB9D2636BE3F" box="[159,263,1048,1105]" gridcol="0" gridrow="0" pageId="23" pageNumber="24">Morph</th>
<th id="070A5DE9FF83002D3C4EFB9D2642BE3F" box="[305,371,1048,1105]" gridcol="1" gridrow="0" pageId="23" pageNumber="24">shield length</th>
<th id="070A5DE9FF83002D3CDBFB9D26D5BE3F" box="[420,484,1048,1105]" gridcol="2" gridrow="0" pageId="23" pageNumber="24">shield width</th>
<th id="070A5DE9FF83002D3F6DFB9D2592BE3F" box="[530,675,1048,1105]" gridcol="3" gridrow="0" pageId="23" pageNumber="24">leg IV length / ratio</th>
<th id="070A5DE9FF83002D3FA7FB9D2431BE3F" box="[728,768,1048,1105]" gridcol="4" gridrow="0" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC53FA7FB9D25C6BE41" box="[728,759,1048,1071]" italics="true" pageId="23" pageNumber="24">st5</emphasis>
</th>
<th id="070A5DE9FF83002D3E4FFB9D246BBE3F" box="[816,858,1048,1105]" gridcol="5" gridrow="0" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC53E4FFB9D246BBE41" box="[816,858,1048,1071]" italics="true" pageId="23" pageNumber="24">JV5</emphasis>
</th>
<th id="070A5DE9FF83002D3EF7FB9D2481BE3F" box="[904,944,1048,1105]" gridcol="6" gridrow="0" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC53EF7FB9D2491BE41" box="[904,928,1048,1071]" italics="true" pageId="23" pageNumber="24">s6</emphasis>
</th>
<th id="070A5DE9FF83002D3E9FFB9D24CABE3F" box="[992,1019,1048,1105]" gridcol="7" gridrow="0" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC53E9FFB9D24CBBE41" box="[992,1018,1048,1071]" italics="true" pageId="23" pageNumber="24">S1</emphasis>
</th>
<th id="070A5DE9FF83002D3954FB9D2365BE3F" box="[1067,1108,1048,1105]" gridcol="8" gridrow="0" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC53953FB9D2376BE41" box="[1068,1095,1048,1071]" italics="true" pageId="23" pageNumber="24">S2</emphasis>
</th>
<th id="070A5DE9FF83002D39FCFB9D239DBE3F" box="[1155,1196,1048,1105]" gridcol="9" gridrow="0" pageId="23" pageNumber="24">
<emphasis id="C89FEAC5FF83FFC539FBFB9D23AFBE41" box="[1156,1182,1048,1071]" italics="true" pageId="23" pageNumber="24">S5</emphasis>
</th>
<th id="070A5DE9FF83002D39A3FB9D224EBE3F" box="[1244,1407,1048,1105]" gridcol="10" gridrow="0" pageId="23" pageNumber="24">
IVFe 
<emphasis id="C89FEAC5FF83FFC5386DFB9D2216BE41" box="[1298,1319,1048,1071]" italics="true" pageId="23" pageNumber="24">-v</emphasis>
IVFe- 
<emphasis id="C89FEAC5FF83FFC53840FBBF2241BE3F" box="[1343,1392,1082,1105]" italics="true" pageId="23" pageNumber="24">ad-1</emphasis>
</th>
</tr>
<tr id="44DB3495FF83002D3DE0FBE0224EBE12" box="[159,1407,1125,1148]" gridrow="1" pageId="23" pageNumber="24">
<th id="070A5DE9FF83002D3DE0FBE02636BE12" box="[159,263,1125,1148]" gridcol="0" gridrow="1" pageId="23" pageNumber="24">short-hair</th>
<td id="070A5DE9FF83002D3C4EFBE02642BE12" box="[305,371,1125,1148]" gridcol="1" gridrow="1" pageId="23" pageNumber="24">395</td>
<td id="070A5DE9FF83002D3CDBFBE026D5BE12" box="[420,484,1125,1148]" gridcol="2" gridrow="1" pageId="23" pageNumber="24">275</td>
<td id="070A5DE9FF83002D3F6DFBE02592BE12" box="[530,675,1125,1148]" gridcol="3" gridrow="1" pageId="23" pageNumber="24">425 1.08</td>
<td id="070A5DE9FF83002D3FA7FBE02431BE12" box="[728,768,1125,1148]" gridcol="4" gridrow="1" pageId="23" pageNumber="24">65</td>
<td id="070A5DE9FF83002D3E4FFBE0246BBE12" box="[816,858,1125,1148]" gridcol="5" gridrow="1" pageId="23" pageNumber="24">30</td>
<td id="070A5DE9FF83002D3EF7FBE02481BE12" box="[904,944,1125,1148]" gridcol="6" gridrow="1" pageId="23" pageNumber="24">47</td>
<td id="070A5DE9FF83002D3E9FFBE024CABE12" box="[992,1019,1125,1148]" gridcol="7" gridrow="1" pageId="23" pageNumber="24">25</td>
<td id="070A5DE9FF83002D3954FBE02365BE12" box="[1067,1108,1125,1148]" gridcol="8" gridrow="1" pageId="23" pageNumber="24">47</td>
<td id="070A5DE9FF83002D39FCFBE0239DBE12" box="[1155,1196,1125,1148]" gridcol="9" gridrow="1" pageId="23" pageNumber="24">65</td>
<td id="070A5DE9FF83002D39A3FBE0224EBE12" box="[1244,1407,1125,1148]" gridcol="10" gridrow="1" pageId="23" pageNumber="24">42 42</td>
</tr>
<tr id="44DB3495FF83002D3DE0FB15224EBEBF" box="[159,1407,1168,1233]" gridrow="2" pageId="23" pageNumber="24">
<th id="070A5DE9FF83002D3DE0FB152636BEBF" box="[159,263,1168,1233]" gridcol="0" gridrow="2" pageId="23" pageNumber="24">short-hair short-hair</th>
<td id="070A5DE9FF83002D3C4EFB152642BEBF" box="[305,371,1168,1233]" gridcol="1" gridrow="2" pageId="23" pageNumber="24">382 405</td>
<td id="070A5DE9FF83002D3CDBFB1526D5BEBF" box="[420,484,1168,1233]" gridcol="2" gridrow="2" pageId="23" pageNumber="24">265 287</td>
<td id="070A5DE9FF83002D3F6DFB152592BEBF" box="[530,675,1168,1233]" gridcol="3" gridrow="2" pageId="23" pageNumber="24">412 1.08 440 1.09</td>
<td id="070A5DE9FF83002D3FA7FB152431BEBF" box="[728,768,1168,1233]" gridcol="4" gridrow="2" pageId="23" pageNumber="24">53 55</td>
<td id="070A5DE9FF83002D3E4FFB15246BBEBF" box="[816,858,1168,1233]" gridcol="5" gridrow="2" pageId="23" pageNumber="24">33 22</td>
<td id="070A5DE9FF83002D3EF7FB152481BEBF" box="[904,944,1168,1233]" gridcol="6" gridrow="2" pageId="23" pageNumber="24">45 35</td>
<td id="070A5DE9FF83002D3E9FFB1524CABEBF" box="[992,1019,1168,1233]" gridcol="7" gridrow="2" pageId="23" pageNumber="24">20 21</td>
<td id="070A5DE9FF83002D3954FB152365BEBF" box="[1067,1108,1168,1233]" gridcol="8" gridrow="2" pageId="23" pageNumber="24">42 27</td>
<td id="070A5DE9FF83002D39FCFB15239DBEBF" box="[1155,1196,1168,1233]" gridcol="9" gridrow="2" pageId="23" pageNumber="24">60 40</td>
<td id="070A5DE9FF83002D39A3FB15224EBEBF" box="[1244,1407,1168,1233]" gridcol="10" gridrow="2" pageId="23" pageNumber="24">45 40 32 32</td>
</tr>
<tr id="44DB3495FF83002D3DE0FB60224EBE92" box="[159,1407,1253,1276]" gridrow="3" pageId="23" pageNumber="24">
<th id="070A5DE9FF83002D3DE0FB602636BE92" box="[159,263,1253,1276]" gridcol="0" gridrow="3" pageId="23" pageNumber="24">short-hair</th>
<td id="070A5DE9FF83002D3C4EFB602642BE92" box="[305,371,1253,1276]" gridcol="1" gridrow="3" pageId="23" pageNumber="24">392</td>
<td id="070A5DE9FF83002D3CDBFB6026D5BE92" box="[420,484,1253,1276]" gridcol="2" gridrow="3" pageId="23" pageNumber="24">262</td>
<td id="070A5DE9FF83002D3F6DFB602592BE92" box="[530,675,1253,1276]" gridcol="3" gridrow="3" pageId="23" pageNumber="24">432 1.10</td>
<td id="070A5DE9FF83002D3FA7FB602431BE92" box="[728,768,1253,1276]" gridcol="4" gridrow="3" pageId="23" pageNumber="24">55</td>
<td id="070A5DE9FF83002D3E4FFB60246BBE92" box="[816,858,1253,1276]" gridcol="5" gridrow="3" pageId="23" pageNumber="24">25</td>
<td id="070A5DE9FF83002D3EF7FB602481BE92" box="[904,944,1253,1276]" gridcol="6" gridrow="3" pageId="23" pageNumber="24">30</td>
<td id="070A5DE9FF83002D3E9FFB6024CABE92" box="[992,1019,1253,1276]" gridcol="7" gridrow="3" pageId="23" pageNumber="24">20</td>
<td id="070A5DE9FF83002D3954FB602365BE92" box="[1067,1108,1253,1276]" gridcol="8" gridrow="3" pageId="23" pageNumber="24">30</td>
<td id="070A5DE9FF83002D39FCFB60239DBE92" box="[1155,1196,1253,1276]" gridcol="9" gridrow="3" pageId="23" pageNumber="24">32</td>
<td id="070A5DE9FF83002D39A3FB60224EBE92" box="[1244,1407,1253,1276]" gridcol="10" gridrow="3" pageId="23" pageNumber="24">35 30</td>
</tr>
<tr id="44DB3495FF83002D3DE0FA95224EBF3F" box="[159,1407,1296,1361]" gridrow="4" pageId="23" pageNumber="24">
<th id="070A5DE9FF83002D3DE0FA952636BF3F" box="[159,263,1296,1361]" gridcol="0" gridrow="4" pageId="23" pageNumber="24">long-hair long-hair</th>
<td id="070A5DE9FF83002D3C4EFA952642BF3F" box="[305,371,1296,1361]" gridcol="1" gridrow="4" pageId="23" pageNumber="24">430 442</td>
<td id="070A5DE9FF83002D3CDBFA9526D5BF3F" box="[420,484,1296,1361]" gridcol="2" gridrow="4" pageId="23" pageNumber="24">312 325</td>
<td id="070A5DE9FF83002D3F6DFA952592BF3F" box="[530,675,1296,1361]" gridcol="3" gridrow="4" pageId="23" pageNumber="24">555 1.29 575 1.30</td>
<td id="070A5DE9FF83002D3FA7FA952431BF3F" box="[728,768,1296,1361]" gridcol="4" gridrow="4" pageId="23" pageNumber="24">110 105</td>
<td id="070A5DE9FF83002D3E4FFA95246BBF3F" box="[816,858,1296,1361]" gridcol="5" gridrow="4" pageId="23" pageNumber="24">95 105</td>
<td id="070A5DE9FF83002D3EF7FA952481BF3F" box="[904,944,1296,1361]" gridcol="6" gridrow="4" pageId="23" pageNumber="24">117 137</td>
<td id="070A5DE9FF83002D3E9FFA9524CABF3F" box="[992,1019,1296,1361]" gridcol="7" gridrow="4" pageId="23" pageNumber="24">45 45</td>
<td id="070A5DE9FF83002D3954FA952365BF3F" box="[1067,1108,1296,1361]" gridcol="8" gridrow="4" pageId="23" pageNumber="24">125 137</td>
<td id="070A5DE9FF83002D39FCFA95239DBF3F" box="[1155,1196,1296,1361]" gridcol="9" gridrow="4" pageId="23" pageNumber="24">125 137</td>
<td id="070A5DE9FF83002D39A3FA95224EBF3F" box="[1244,1407,1296,1361]" gridcol="10" gridrow="4" pageId="23" pageNumber="24">67 82 87 125</td>
</tr>
<tr id="44DB3495FF83002D3DE0FAE0224EBF12" box="[159,1407,1381,1404]" gridrow="5" pageId="23" pageNumber="24">
<th id="070A5DE9FF83002D3DE0FAE02636BF12" box="[159,263,1381,1404]" gridcol="0" gridrow="5" pageId="23" pageNumber="24">long-hair</th>
<td id="070A5DE9FF83002D3C4EFAE02642BF12" box="[305,371,1381,1404]" gridcol="1" gridrow="5" pageId="23" pageNumber="24">430</td>
<td id="070A5DE9FF83002D3CDBFAE026D5BF12" box="[420,484,1381,1404]" gridcol="2" gridrow="5" pageId="23" pageNumber="24">305</td>
<td id="070A5DE9FF83002D3F6DFAE02592BF12" box="[530,675,1381,1404]" gridcol="3" gridrow="5" pageId="23" pageNumber="24">582 1.35</td>
<td id="070A5DE9FF83002D3FA7FAE02431BF12" box="[728,768,1381,1404]" gridcol="4" gridrow="5" pageId="23" pageNumber="24">110</td>
<td id="070A5DE9FF83002D3E4FFAE0246BBF12" box="[816,858,1381,1404]" gridcol="5" gridrow="5" pageId="23" pageNumber="24">95</td>
<td id="070A5DE9FF83002D3EF7FAE02481BF12" box="[904,944,1381,1404]" gridcol="6" gridrow="5" pageId="23" pageNumber="24">150</td>
<td id="070A5DE9FF83002D3E9FFAE024CABF12" box="[992,1019,1381,1404]" gridcol="7" gridrow="5" pageId="23" pageNumber="24">65</td>
<td id="070A5DE9FF83002D3954FAE02365BF12" box="[1067,1108,1381,1404]" gridcol="8" gridrow="5" pageId="23" pageNumber="24">135</td>
<td id="070A5DE9FF83002D39FCFAE0239DBF12" box="[1155,1196,1381,1404]" gridcol="9" gridrow="5" pageId="23" pageNumber="24">125</td>
<td id="070A5DE9FF83002D39A3FAE0224EBF12" box="[1244,1407,1381,1404]" gridcol="10" gridrow="5" pageId="23" pageNumber="24">75 112</td>
</tr>
<tr id="44DB3495FF83002D3DE0FA15224EBFBF" box="[159,1407,1424,1489]" gridrow="6" pageId="23" pageNumber="24">
<th id="070A5DE9FF83002D3DE0FA152636BFBF" box="[159,263,1424,1489]" gridcol="0" gridrow="6" pageId="23" pageNumber="24">long-hair intermed.</th>
<td id="070A5DE9FF83002D3C4EFA152642BFBF" box="[305,371,1424,1489]" gridcol="1" gridrow="6" pageId="23" pageNumber="24">455 412</td>
<td id="070A5DE9FF83002D3CDBFA1526D5BFBF" box="[420,484,1424,1489]" gridcol="2" gridrow="6" pageId="23" pageNumber="24">315 300</td>
<td id="070A5DE9FF83002D3F6DFA152592BFBF" box="[530,675,1424,1489]" gridcol="3" gridrow="6" pageId="23" pageNumber="24">587 129 525 1.27</td>
<td id="070A5DE9FF83002D3FA7FA152431BFBF" box="[728,768,1424,1489]" gridcol="4" gridrow="6" pageId="23" pageNumber="24">125 100</td>
<td id="070A5DE9FF83002D3E4FFA15246BBFBF" box="[816,858,1424,1489]" gridcol="5" gridrow="6" pageId="23" pageNumber="24">105 70</td>
<td id="070A5DE9FF83002D3EF7FA152481BFBF" box="[904,944,1424,1489]" gridcol="6" gridrow="6" pageId="23" pageNumber="24">155 105</td>
<td id="070A5DE9FF83002D3E9FFA1524CABFBF" box="[992,1019,1424,1489]" gridcol="7" gridrow="6" pageId="23" pageNumber="24">87 40</td>
<td id="070A5DE9FF83002D3954FA152365BFBF" box="[1067,1108,1424,1489]" gridcol="8" gridrow="6" pageId="23" pageNumber="24">135 105</td>
<td id="070A5DE9FF83002D39FCFA15239DBFBF" box="[1155,1196,1424,1489]" gridcol="9" gridrow="6" pageId="23" pageNumber="24">150 112</td>
<td id="070A5DE9FF83002D39A3FA15224EBFBF" box="[1244,1407,1424,1489]" gridcol="10" gridrow="6" pageId="23" pageNumber="24">77 115 67 82</td>
</tr>
<tr id="44DB3495FF83002D3DE0FA60224EBF92" box="[159,1407,1509,1532]" gridrow="7" pageId="23" pageNumber="24">
<th id="070A5DE9FF83002D3DE0FA602636BF92" box="[159,263,1509,1532]" gridcol="0" gridrow="7" pageId="23" pageNumber="24">intermed.</th>
<td id="070A5DE9FF83002D3C4EFA602642BF92" box="[305,371,1509,1532]" gridcol="1" gridrow="7" pageId="23" pageNumber="24">432</td>
<td id="070A5DE9FF83002D3CDBFA6026D5BF92" box="[420,484,1509,1532]" gridcol="2" gridrow="7" pageId="23" pageNumber="24">290</td>
<td id="070A5DE9FF83002D3F6DFA602592BF92" box="[530,675,1509,1532]" gridcol="3" gridrow="7" pageId="23" pageNumber="24">500 1.16</td>
<td id="070A5DE9FF83002D3FA7FA602431BF92" box="[728,768,1509,1532]" gridcol="4" gridrow="7" pageId="23" pageNumber="24">82</td>
<td id="070A5DE9FF83002D3E4FFA60246BBF92" box="[816,858,1509,1532]" gridcol="5" gridrow="7" pageId="23" pageNumber="24">57</td>
<td id="070A5DE9FF83002D3EF7FA602481BF92" box="[904,944,1509,1532]" gridcol="6" gridrow="7" pageId="23" pageNumber="24">82</td>
<td id="070A5DE9FF83002D3E9FFA6024CABF92" box="[992,1019,1509,1532]" gridcol="7" gridrow="7" pageId="23" pageNumber="24">42</td>
<td id="070A5DE9FF83002D3954FA602365BF92" box="[1067,1108,1509,1532]" gridcol="8" gridrow="7" pageId="23" pageNumber="24">80</td>
<td id="070A5DE9FF83002D39FCFA60239DBF92" box="[1155,1196,1509,1532]" gridcol="9" gridrow="7" pageId="23" pageNumber="24">90</td>
<td id="070A5DE9FF83002D39A3FA60224EBF92" box="[1244,1407,1509,1532]" gridcol="10" gridrow="7" pageId="23" pageNumber="24">75 60</td>
</tr>
</table>
</paragraph>
<paragraph id="FA5436D7FF83FFCA3DB9F9BD2696BB6C" blockId="23.[151,1436,1592,2018]" lastBlockId="24.[151,1436,152,258]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Tritosternum much as in female, laciniae free for about 0.7 of total length (50–55 excluding base). Presternal area much as in female, without platelets (
<figureCitation id="62D02A52FF83FFC53F9BF9E5240EBC14" box="[740,831,1632,1658]" captionStart="FIGURES 56 – 58" captionStartId="29.[151,269,1642,1666]" captionTargetBox="[155,1429,194,1600]" captionTargetId="figure@29.[151,1436,194,1637]" captionTargetPageId="29" captionText="FIGURES 56 – 58. Spadiseius spathiphyllae n. sp., adult male. Figs. 56, 57, opisthogaster: 56, intermediate morph; 57, “ short-haired ” morph. Fig. 58, idiosoma, ventral aspect, “ long-haired ” morph." httpUri="https://zenodo.org/record/180495/files/figure.png" pageId="23" pageNumber="24">Fig. 58</figureCitation>
). Sternogenital shield smooth or ornamented with arched lines behind setae 
<emphasis id="C89FEAC5FF83FFC53CB5F90D26DDBCCF" box="[458,492,1672,1697]" italics="true" pageId="23" pageNumber="24">st5</emphasis>
; of 4 or 5 pairs of setae on shield, 
<emphasis id="C89FEAC5FF83FFC53EEEF90D2485BCCF" box="[913,948,1672,1697]" italics="true" pageId="23" pageNumber="24">st4</emphasis>
(35–55) longer than 
<emphasis id="C89FEAC5FF83FFC539CEF90D2232BCCF" box="[1201,1283,1672,1697]" italics="true" pageId="23" pageNumber="24">st1–st3</emphasis>
(20–30), 
<emphasis id="C89FEAC5FF83FFC53805F90D22ADBCCF" box="[1402,1436,1672,1697]" italics="true" pageId="23" pageNumber="24">st5</emphasis>
variably much longer (50–110) and often inserted near paragenital poroids (these hard to discern) on soft cuticle closely beside constricted posterolateral corners of shield; posterior margin of shield strongly delineated, nearly truncated, often bilobed (
<figureCitation id="62D02A52FF83FFC53F6AF8852596BD74" box="[533,679,1792,1818]" captionStart="FIGURES 56 – 58" captionStartId="29.[151,269,1642,1666]" captionTargetBox="[155,1429,194,1600]" captionTargetId="figure@29.[151,1436,194,1637]" captionTargetPageId="29" captionText="FIGURES 56 – 58. Spadiseius spathiphyllae n. sp., adult male. Figs. 56, 57, opisthogaster: 56, intermediate morph; 57, “ short-haired ” morph. Fig. 58, idiosoma, ventral aspect, “ long-haired ” morph." httpUri="https://zenodo.org/record/180495/files/figure.png" pageId="23" pageNumber="24">Figs. 56–58</figureCitation>
). Ventrianal shield unornamented, with weakly delineated anterior and often strongly sloping anterolateral margins; shield length (155–160) about 0.75 its greatest width (195–220) anteriorly where metapodal plates incorporated; anal region slightly more sclerotized, sometimes appearing delineated like an anal plate; ventrianal shield with circumanal setae and usually 7 pairs of opisthogastric setae (
<emphasis id="C89FEAC5FF83FFC53C45F8252695BDD7" box="[314,420,1952,1977]" italics="true" pageId="23" pageNumber="24">JV1–JV4</emphasis>
, 
<emphasis id="C89FEAC5FF83FFC53CCBF8252515BDD7" box="[436,548,1952,1977]" italics="true" pageId="23" pageNumber="24">ZV1–ZV3</emphasis>
), these setae similar in length as in female (15–25) or variably much more attenuated (40–90), 
<emphasis id="C89FEAC5FF83FFC53CF9F84D2682BD8F" box="[390,435,1992,2017]" italics="true" pageId="23" pageNumber="24">JV3</emphasis>
often shorter than others; postanal seta (18–25) slightly longer than para-anal setae (15–20); 
<emphasis id="C89FEAC5FF8CFFCA3C79FF1D2605BADF" box="[262,308,152,177]" italics="true" pageId="24" pageNumber="25">JV5</emphasis>
and 
<emphasis id="C89FEAC5FF8CFFCA3C0CFF1D2695BADF" box="[371,420,152,177]" italics="true" pageId="24" pageNumber="25">ZV4</emphasis>
variably short (15–30) to attenuated (105–115) not clavate distally, inserted on soft cuticle, 
<emphasis id="C89FEAC5FF8CFFCA3D8BFF452615BAB7" box="[244,292,192,217]" italics="true" pageId="24" pageNumber="25">ZV5</emphasis>
absent. Form of peritrematal and exopodal shields posteriorly as in female, except exopodal rims wider behind coxae IV.
</paragraph>
<caption id="AE94665FFF8CFFCA3DE8FB0E22ADBEAB" httpUri="https://zenodo.org/record/180491/files/figure.png" pageId="24" pageNumber="25" targetBox="[168,1419,313,1131]" targetPageId="24">
<paragraph id="FA5436D7FF8CFFCA3DE8FB0E22ADBEAB" blockId="24.[151,1436,1163,1221]" pageId="24" pageNumber="25">
<emphasis id="C89FEAC5FF8CFFCA3DE8FB0E266CBECD" bold="true" box="[151,349,1163,1187]" pageId="24" pageNumber="25">FIGURES 38–41.</emphasis>
Dorsal subapical femoral setae of legs. Figs. 38, 39, larval femora I–III: 38, 
<taxonomicName id="3DEB4D54FF8CFFCA39F3FB0E22ADBECC" box="[1164,1436,1163,1186]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
<emphasis id="C89FEAC5FF8CFFCA39F3FB0E22ADBECC" box="[1164,1436,1163,1186]" italics="true" pageId="24" pageNumber="25">Spadiseius calyptrogynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8CFFCA3DE8FB2827E5BEAB" box="[151,212,1197,1221]" pageId="24" pageNumber="25" rank="species">
<emphasis id="C89FEAC5FF8CFFCA3DE8FB2827E5BEAB" bold="true" box="[151,212,1197,1221]" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
; 39, 
<taxonomicName id="3DEB4D54FF8CFFCA3C71FB2B2529BEAB" box="[270,536,1198,1221]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF8CFFCA3C71FB2B2529BEAB" box="[270,536,1198,1221]" italics="true" pageId="24" pageNumber="25">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8CFFCA3F5EFB28256EBEAB" box="[545,607,1197,1221]" pageId="24" pageNumber="25" rank="species">
<emphasis id="C89FEAC5FF8CFFCA3F5EFB28256EBEAB" bold="true" box="[545,607,1197,1221]" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
Figs. 40, 41, protonymphal femora I–IV: 40, 
<taxonomicName id="3DEB4D54FF8CFFCA392BFB2B2239BEAB" box="[1108,1288,1198,1221]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
<emphasis id="C89FEAC5FF8CFFCA392BFB2B2239BEAB" box="[1108,1288,1198,1221]" italics="true" pageId="24" pageNumber="25">S. calyptrogynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8CFFCA386FFB28227FBEAB" box="[1296,1358,1197,1221]" pageId="24" pageNumber="25" rank="species">
<emphasis id="C89FEAC5FF8CFFCA386FFB28227FBEAB" bold="true" box="[1296,1358,1197,1221]" pageId="24" pageNumber="25">n. sp.</emphasis>
</taxonomicNameLabel>
; 41, S.
</paragraph>
</caption>
<paragraph id="FA5436D7FF8CFFCA3DB9FAC623EEBCF3" blockId="24.[151,1436,1347,2013]" pageId="24" pageNumber="25">
Tectum with narrower base and more acutely pointed apex than in female. Cheliceral shaft narrowing dorsally at base of fixed digit (
<figureCitation id="62D02A52FF8CFFCA3CA9FAEE251CBFEB" box="[470,557,1387,1413]" captionStart="FIGURES 51 – 55" captionStartId="28.[151,269,1906,1930]" captionTargetBox="[206,1386,432,1878]" captionTargetId="figure@28.[206,1393,416,1899]" captionTargetPageId="28" captionText="FIGURES 51 – 55. Spadiseius spathiphyllae n. sp., adult male. Figs. 51 – 53, idiosomas, dorsal aspect: 51, “ long-haired ” morph; 52, intermediate morph; 53, “ short-haired ” morph, partial view. Fig. 54, detail of peritrematal shield; note variation in length of peritreme from Fig. 52. Fig. 55, chelicera and spermatodactyl, adaxial view." httpUri="https://zenodo.org/record/180494/files/figure.png" pageId="24" pageNumber="25">Fig. 55</figureCitation>
); fixed digit with a small offset tooth followed by 1–2 minute teeth at level of hyaline pilus dentilis, and with a large pointed tooth basally; spermatodactyl simple in form, extending sinuously antero-ventrally about 30 beyond apex of chela (
<figureCitation id="62D02A52FF8CFFCA3E5EFA3E2449BFBB" box="[801,888,1467,1493]" captionStart="FIGURES 51 – 55" captionStartId="28.[151,269,1906,1930]" captionTargetBox="[206,1386,432,1878]" captionTargetId="figure@28.[206,1393,416,1899]" captionTargetPageId="28" captionText="FIGURES 51 – 55. Spadiseius spathiphyllae n. sp., adult male. Figs. 51 – 53, idiosomas, dorsal aspect: 51, “ long-haired ” morph; 52, intermediate morph; 53, “ short-haired ” morph, partial view. Fig. 54, detail of peritrematal shield; note variation in length of peritreme from Fig. 52. Fig. 55, chelicera and spermatodactyl, adaxial view." httpUri="https://zenodo.org/record/180494/files/figure.png" pageId="24" pageNumber="25">Fig. 55</figureCitation>
). Corniculi more widely spaced basally than in female, their apices not overlapping; internal malae larger that in female, with fringed lateral margins and bifid apices (as in 
<figureCitation id="62D02A52FF8CFFCA3C12F98E26F2BC4B" box="[365,451,1547,1573]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,269,1250,1274]" captionTargetBox="[151,1435,354,1192]" captionTargetId="figure@18.[151,1436,354,1270]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Spadiseius calyptrogynae n. sp., adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation: k, keel." httpUri="https://zenodo.org/record/180488/files/figure.png" pageId="24" pageNumber="25">Fig. 26</figureCitation>
). Subcapitulum with rows of deutosternal denticles as in female. Subcapitulum with form and relative lengths of setae as in female, and with serrated ridge on either side of capitular setae and ventrally serrated apical sclerotized rim of palpfemur and palpgenu as in female. Palptrochanter setae subequal in length (15–20); palptibia with ventral seta 
<emphasis id="C89FEAC5FF8CFFCA3F90F906243ABCF2" box="[751,779,1667,1692]" italics="true" pageId="24" pageNumber="25">av</emphasis>
thicker, more spine-like than in female.
</paragraph>
<paragraph id="FA5436D7FF8CFFCB3DB9F92E24E2BB44" blockId="24.[151,1436,1347,2013]" lastBlockId="25.[151,1437,152,298]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
Relative leg lengths, excluding pretarsi, similar to those of female, I (350–420), II (295–385), III (345– 485), IV (395–620). Coxa IV seta at least as long (32–42) as coxa III setae (28–30). Coxal ornamention and serration of apically sclerotized rims of segments of legs I–IV as in female, but size and form of many setae variably polymorphic on femora, genua, tibiae and tarsi. Tarsi II–IV with pair of apical setal processes (
<emphasis id="C89FEAC5FF8CFFCA3820F8A627FDBD0A" italics="true" pageId="24" pageNumber="25">ad-1, pd-1</emphasis>
) as in female. Legs II–IV sometimes curved and slightly thicker relative to leg I than in female; tarsus II with setae 
<emphasis id="C89FEAC5FF8CFFCA3C6BF8F62679BDE2" box="[276,328,1907,1932]" italics="true" pageId="24" pageNumber="25">av-1</emphasis>
, 
<emphasis id="C89FEAC5FF8CFFCA3C28F8F626BABDE2" box="[343,395,1907,1932]" italics="true" pageId="24" pageNumber="25">av-2</emphasis>
modified to short, stout spines, and 
<emphasis id="C89FEAC5FF8CFFCA3E4AF8F6245BBDE2" box="[821,874,1907,1932]" italics="true" pageId="24" pageNumber="25">av-3</emphasis>
(
<emphasis id="C89FEAC5FF8CFFCA3E04F8F624ACBDE2" box="[891,925,1907,1932]" italics="true" pageId="24" pageNumber="25">mv</emphasis>
) variably short setiform or spine-like (as in 
<figureCitation id="62D02A52FF8CFFCA3DE8F81E27DEBDDB" box="[151,239,1947,1973]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,269,1250,1274]" captionTargetBox="[151,1435,354,1192]" captionTargetId="figure@18.[151,1436,354,1270]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Spadiseius calyptrogynae n. sp., adult male: 26, subcapitulum; 27, chelicera and spermatodactyl, paraxial view; 28, tarsus II, anterolateral view; 29, tarsus IV, dorsal view, without pretarsus. Abbreviation: k, keel." httpUri="https://zenodo.org/record/180488/files/figure.png" pageId="24" pageNumber="25">Fig. 28</figureCitation>
); basitarsus of legs IV and sometimes of III with conical spur posteroventrally beside seta 
<emphasis id="C89FEAC5FF8CFFCA3853F81E226ABDDA" box="[1324,1371,1947,1972]" italics="true" pageId="24" pageNumber="25">pl-4</emphasis>
(
<figureCitation id="62D02A52FF8CFFCA3813F81E2789BDB3" captionStart="FIGURES 65 – 69" captionStartId="31.[151,269,1219,1243]" captionTargetBox="[164,1431,194,1163]" captionTargetId="figure@31.[151,1436,194,1283]" captionTargetPageId="31" captionText="FIGURES 65 – 69. Figs. 65 – 67: Spadiseius spathiphyllae n. sp., adult male: 65, tarsus IV, dorsolateral view, without pretarsus; 66, 67, femora IV, dorsal view: 66, “ short-haired ” morph; 67, “ long-haired ” morph. Figs. 68, 69, Spadiseius calyptrogynae n. sp., adult male, femora IV, dorsal view: 68, “ wartless ” morph; 69, “ warted ” morph." httpUri="https://zenodo.org/record/180497/files/figure.png" pageId="24" pageNumber="25">Fig. 65</figureCitation>
). Femur I with dorsal seta 
<emphasis id="C89FEAC5FF8CFFCA3C86F846251EBDB2" box="[505,559,1987,2012]" italics="true" pageId="24" pageNumber="25">ad-1</emphasis>
apically capitate or oar-like and variably longer (20–100) than in female; 
<emphasis id="C89FEAC5FF8DFFCB3DE8FF1D27FCBADF" box="[151,205,152,177]" italics="true" pageId="25" pageNumber="26">ad-1</emphasis>
on femora II–IV variably oar-like or attenuated, or apically capitate, and increasingly longer from II (20– 75) to IV (30–125). Legs I–IV with some dorsal and lateral setae variably oar-like or attenuated on femur of legs I–IV, genua II–IV and tibiae III–IV, and legs III–IV with some ventral and anterolateral setae variably attenuated (40 up to 75) on femur, genu, tibia and tarsus (
<figureCitation id="62D02A52FF8DFFCB3E48FE9524F7BB44" box="[823,966,272,298]" captionStart="FIGURES 65 – 69" captionStartId="31.[151,269,1219,1243]" captionTargetBox="[164,1431,194,1163]" captionTargetId="figure@31.[151,1436,194,1283]" captionTargetPageId="31" captionText="FIGURES 65 – 69. Figs. 65 – 67: Spadiseius spathiphyllae n. sp., adult male: 65, tarsus IV, dorsolateral view, without pretarsus; 66, 67, femora IV, dorsal view: 66, “ short-haired ” morph; 67, “ long-haired ” morph. Figs. 68, 69, Spadiseius calyptrogynae n. sp., adult male, femora IV, dorsal view: 68, “ wartless ” morph; 69, “ warted ” morph." httpUri="https://zenodo.org/record/180497/files/figure.png" pageId="25" pageNumber="26">Figs. 66, 67</figureCitation>
).
</paragraph>
<caption id="AE94665FFF8DFFCB3DE8F89D24A2BD18" httpUri="https://zenodo.org/record/180492/files/figure.png" pageId="25" pageNumber="26" targetBox="[154,1431,346,1815]" targetPageId="25">
<paragraph id="FA5436D7FF8DFFCB3DE8F89D24A2BD18" blockId="25.[151,1437,1816,1910]" pageId="25" pageNumber="26">
<emphasis id="C89FEAC5FF8DFFCB3DE8F89D266CBD5E" bold="true" box="[151,349,1816,1840]" pageId="25" pageNumber="26">FIGURES 42–45.</emphasis>
Figs. 42, 43, 
<taxonomicName id="3DEB4D54FF8DFFCB3C90F89C25C9BD5E" box="[495,760,1817,1840]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF8DFFCB3C90F89C25C9BD5E" box="[495,760,1817,1840]" italics="true" pageId="25" pageNumber="26">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8DFFCB3F80F89D240ABD5E" box="[767,827,1816,1840]" pageId="25" pageNumber="26" rank="species">
<emphasis id="C89FEAC5FF8DFFCB3F80F89D240ABD5E" bold="true" box="[767,827,1816,1840]" pageId="25" pageNumber="26">n. sp.</emphasis>
</taxonomicNameLabel>
, deutonymph: 42, idiosoma, dorsal aspect; 43, idiosoma, ventral aspect; supernumerary setae denoted by ‘x’. Figs. 44, 45, dorsal subapical femoral setae legs I–IV, deutonymph: 44, 
<taxonomicName id="3DEB4D54FF8DFFCB3DC0F8DB26FEBD1B" box="[191,463,1886,1909]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
<emphasis id="C89FEAC5FF8DFFCB3DC0F8DB26FEBD1B" box="[191,463,1886,1909]" italics="true" pageId="25" pageNumber="26">Spadiseius calyptrogynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8DFFCB3CAAF8DB2520BD18" box="[469,529,1886,1910]" pageId="25" pageNumber="26" rank="species">
<emphasis id="C89FEAC5FF8DFFCB3CAAF8DB2520BD18" bold="true" box="[469,529,1886,1910]" pageId="25" pageNumber="26">n. sp.</emphasis>
</taxonomicNameLabel>
; 45, 
<taxonomicName id="3DEB4D54FF8DFFCB3F37F8DB2461BD1B" box="[584,848,1886,1909]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF8DFFCB3F37F8DB2461BD1B" box="[584,848,1886,1909]" italics="true" pageId="25" pageNumber="26">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8DFFCB3E28F8DB24A2BD18" box="[855,915,1886,1910]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="FA5436D7FF8DFFC83DB9F82D26F8BBA4" blockId="25.[151,1436,1960,2027]" lastBlockId="26.[151,1437,152,2018]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="C89FEAC5FF8DFFCB3DB9F82D264CBDAC" bold="true" box="[198,381,1960,1986]" pageId="25" pageNumber="26">DEUTONYMPH.</emphasis>
Dorsal shield 365–400 long, 200–270 wide at level of lateral incisions by setae 
<emphasis id="C89FEAC5FF8DFFCB385BF82C220CBDAC" box="[1316,1341,1961,1986]" italics="true" pageId="25" pageNumber="26">s6</emphasis>
(n = 5); shield lightly evenly reticulate over nearly entire surface (
<figureCitation id="62D02A52FF8DFFCB3E2CF854249CBD85" box="[851,941,2001,2027]" captionStart="FIGURES 42 – 45" captionStartId="25.[151,269,1816,1840]" captionTargetBox="[154,1431,346,1815]" captionTargetId="figure@25.[151,1436,330,1936]" captionTargetPageId="25" captionText="FIGURES 42 – 45. Figs. 42, 43, Spadiseius spathiphyllae n. sp., deutonymph: 42, idiosoma, dorsal aspect; 43, idiosoma, ventral aspect; supernumerary setae denoted by ‘ x’. Figs. 44, 45, dorsal subapical femoral setae legs I – IV, deutonymph: 44, Spadiseius calyptrogynae n. sp.; 45, Spadiseius spathiphyllae n. sp." httpUri="https://zenodo.org/record/180492/files/figure.png" pageId="25" pageNumber="26">Fig. 42</figureCitation>
); podonotal region with longitudinal distance from alveoli of 
<emphasis id="C89FEAC5FF8EFFC83CEDFF1D2698BADF" box="[402,425,152,177]" italics="true" pageId="26" pageNumber="27">j1</emphasis>
to 
<emphasis id="C89FEAC5FF8EFFC83CB1FF1D26D4BADF" box="[462,485,152,177]" italics="true" pageId="26" pageNumber="27">j5</emphasis>
subequal to that from 
<emphasis id="C89FEAC5FF8EFFC83F92FF1D2435BADF" box="[749,772,152,177]" italics="true" pageId="26" pageNumber="27">j5</emphasis>
to 
<emphasis id="C89FEAC5FF8EFFC83E56FF1D240EBADF" box="[809,831,152,177]" italics="true" pageId="26" pageNumber="27">j6</emphasis>
. Dorsal shield with approximately 35 to 40 pairs of setae: about 17–18 pairs (
<emphasis id="C89FEAC5FF8EFFC83CBDFF452591BAB7" box="[450,672,192,217]" italics="true" pageId="26" pageNumber="27">j1–j6, z2–z6, s3–s6</emphasis>
, and 2 to 4 pairs of extra setae) on podonotal region, and 18 pairs (
<emphasis id="C89FEAC5FF8EFFC83DDEFF6D26AEBB6F" box="[161,415,232,257]" italics="true" pageId="26" pageNumber="27">J1–J5, Z1–Z5, S1–S5</emphasis>
, and a few paired or asymmetrically unpaired setae amidst 
<emphasis id="C89FEAC5FF8EFFC8390AFF6D23B3BB6F" box="[1141,1154,232,257]" italics="true" pageId="26" pageNumber="27">J</emphasis>
series) on opisthonotal region. Dorsal shield setae mostly alike in simple form (larger ones slightly swollen at base), mostly very short (5–7), similar in length to clunals 
<emphasis id="C89FEAC5FF8EFFC83F1BFEBD254EBB3F" box="[612,639,312,337]" italics="true" pageId="26" pageNumber="27">J5</emphasis>
; 
<emphasis id="C89FEAC5FF8EFFC83FF0FEBD2414BB3F" box="[655,805,312,337]" italics="true" pageId="26" pageNumber="27">j1, s6, S2–S5</emphasis>
slightly longer (9–12), 
<emphasis id="C89FEAC5FF8EFFC83945FEBD2369BB3F" box="[1082,1112,312,337]" italics="true" pageId="26" pageNumber="27">S4</emphasis>
, 
<emphasis id="C89FEAC5FF8EFFC8391AFEBD23B3BB3F" box="[1125,1154,312,337]" italics="true" pageId="26" pageNumber="27">S5</emphasis>
and 
<emphasis id="C89FEAC5FF8EFFC839C4FEBD23EABB3F" box="[1211,1243,312,337]" italics="true" pageId="26" pageNumber="27">Z5</emphasis>
barbed, 
<emphasis id="C89FEAC5FF8EFFC83840FEBD226FBB3F" box="[1343,1374,312,337]" italics="true" pageId="26" pageNumber="27">Z5</emphasis>
largest (16–18), tapered or untapered. Dorsal setae on soft cuticle mostly 12–15 long, 
<emphasis id="C89FEAC5FF8EFFC83911FEE523BEBB17" box="[1134,1167,352,377]" italics="true" pageId="26" pageNumber="27">R1</emphasis>
longest (19–20). Peritremes extending anteriorly to level of setae 
<emphasis id="C89FEAC5FF8EFFC83FBBFE0D25EFBBCF" box="[708,734,392,417]" italics="true" pageId="26" pageNumber="27">s1</emphasis>
; peritrematal shields extending further anteriorly but not uniting with dorsal shield.
</paragraph>
<paragraph id="FA5436D7FF8EFFC83DB9FE5D249AB8D4" blockId="26.[151,1437,152,2018]" pageId="26" pageNumber="27">
Tritosternum similar in form to that in adult female. Sternal shield (
<figureCitation id="62D02A52FF8EFFC83968FE5D2343BB9C" box="[1047,1138,472,498]" captionStart="FIGURES 42 – 45" captionStartId="25.[151,269,1816,1840]" captionTargetBox="[154,1431,346,1815]" captionTargetId="figure@25.[151,1436,330,1936]" captionTargetPageId="25" captionText="FIGURES 42 – 45. Figs. 42, 43, Spadiseius spathiphyllae n. sp., deutonymph: 42, idiosoma, dorsal aspect; 43, idiosoma, ventral aspect; supernumerary setae denoted by ‘ x’. Figs. 44, 45, dorsal subapical femoral setae legs I – IV, deutonymph: 44, Spadiseius calyptrogynae n. sp.; 45, Spadiseius spathiphyllae n. sp." httpUri="https://zenodo.org/record/180492/files/figure.png" pageId="26" pageNumber="27">Fig. 43</figureCitation>
) smooth, with narrowly rounded posterior margin; setae 
<emphasis id="C89FEAC5FF8EFFC83F68FD852508B877" box="[535,569,512,537]" italics="true" pageId="26" pageNumber="27">st4</emphasis>
(10–13) clearly shorter than 
<emphasis id="C89FEAC5FF8EFFC83EEBFD8524D7B877" box="[916,998,512,537]" italics="true" pageId="26" pageNumber="27">st1–st3</emphasis>
(23–25). Anal shield unornamented, length (63–68) nearly equal to greatest width (65–75) at level of anterior edge of anus; postanal seta (13) slightly shorter than para-anal setae (15). Opisthogaster with 9 or 10 pairs of ventral setae (
<emphasis id="C89FEAC5FF8EFFC8399DFDD5227DB807" box="[1250,1356,592,617]" italics="true" pageId="26" pageNumber="27">JV1–JV5</emphasis>
, 
<emphasis id="C89FEAC5FF8EFFC83823FDD527F9B8FF" italics="true" pageId="26" pageNumber="27">ZV1– ZV4</emphasis>
) plus 1 or 2 adjacent 
<emphasis id="C89FEAC5FF8EFFC83CB3FDFD2533B8FF" box="[460,514,632,657]" italics="true" pageId="26" pageNumber="27">UR–</emphasis>
setae (
<emphasis id="C89FEAC5FF8EFFC83F28FDFD25B9B8FF" box="[599,648,632,657]" italics="true" pageId="26" pageNumber="27">ZV5</emphasis>
sometimes absent), most similarly short (5–10) but 
<emphasis id="C89FEAC5FF8EFFC8398BFDFD226FB8FF" box="[1268,1374,632,657]" italics="true" pageId="26" pageNumber="27">JV1–JV2</emphasis>
(15– 18), 
<emphasis id="C89FEAC5FF8EFFC83DB3FD2527CBB8D7" box="[204,250,672,697]" italics="true" pageId="26" pageNumber="27">JV5</emphasis>
(17–20). Metapodal plates faint, oval (15 length, 8 width).
</paragraph>
<paragraph id="FA5436D7FF8EFFC83DB9FD4D2604B95C" blockId="26.[151,1437,152,2018]" pageId="26" pageNumber="27">Tectum, chelicerae, most ventral gnathosomatic structures and palpi, including serrated apical sclerotized rim of palpfemur and palpgenu, similar to those in adult female, except corniculi less convergent, not overlapping apically.</paragraph>
<paragraph id="FA5436D7FF8EFFC83DB9FCC527F4BE24" blockId="26.[151,1437,152,2018]" pageId="26" pageNumber="27">
Relative leg lengths, excluding pretarsi, similar to those of female, I (315–325), II (275–285), III (295– 300), IV (350–357). Coxal ornamentation, serration of apically sclerotized rims of segments, and form of legs I–IV and their setae as in adult female, except leg II no thicker than legs III–IV; tarsus I with pedestal bearing pretarsus less defined than on protonymph; tarsi II–IV tapered apically beyond insertions of 
<emphasis id="C89FEAC5FF8EFFC839A7FC3D227FB9BF" box="[1240,1358,952,977]" italics="true" pageId="26" pageNumber="27">av-1, pv-1</emphasis>
. Coxa IV seta as long as coxa III setae (18–20). Dorsal seta 
<emphasis id="C89FEAC5FF8EFFC83E7DFC652409B997" box="[770,824,992,1017]" italics="true" pageId="26" pageNumber="27">ad-1</emphasis>
thicker, rod- or oar-like on femora I–III, much as on femur IV, in contrast with adjacent tapered setae, progressively slightly longer from I (12) to IV (18–19) (
<figureCitation id="62D02A52FF8EFFC83813FB8D2789BE24" captionStart="FIGURES 42 – 45" captionStartId="25.[151,269,1816,1840]" captionTargetBox="[154,1431,346,1815]" captionTargetId="figure@25.[151,1436,330,1936]" captionTargetPageId="25" captionText="FIGURES 42 – 45. Figs. 42, 43, Spadiseius spathiphyllae n. sp., deutonymph: 42, idiosoma, dorsal aspect; 43, idiosoma, ventral aspect; supernumerary setae denoted by ‘ x’. Figs. 44, 45, dorsal subapical femoral setae legs I – IV, deutonymph: 44, Spadiseius calyptrogynae n. sp.; 45, Spadiseius spathiphyllae n. sp." httpUri="https://zenodo.org/record/180492/files/figure.png" pageId="26" pageNumber="27">Fig. 45</figureCitation>
).
</paragraph>
<paragraph id="FA5436D7FF8EFFC83DB9FBD22336BFE4" blockId="26.[151,1437,152,2018]" pageId="26" pageNumber="27">
<emphasis id="C89FEAC5FF8EFFC83DB9FBD2264DBE1F" bold="true" box="[198,380,1111,1137]" pageId="26" pageNumber="27">PROTONYMPH.</emphasis>
Idiosoma 310–340 long, 210–255 wide at level of setae 
<emphasis id="C89FEAC5FF8EFFC83951FBDD2376BE1F" box="[1070,1095,1112,1137]" italics="true" pageId="26" pageNumber="27">r5</emphasis>
(n = 5), with lightly sclerotized, fully reticulated podonotal and pygidial shields (
<figureCitation id="62D02A52FF8EFFC83E5BFB05244FBEF4" box="[804,894,1152,1178]" captionStart="FIGURES 35 – 37" captionStartId="22.[151,269,1758,1782]" captionTargetBox="[151,1434,598,1757]" captionTargetId="figure@22.[151,1436,594,1776]" captionTargetPageId="22" captionText="FIGURES 35 – 37. Spadiseius spathiphyllae n. sp. 35, larva, idiosoma, dorsal aspect; 36, protonymph, idiosoma, dorsal aspect; 37, larva, chelicera, adaxial view. Abbreviation: v m, ventral mucro." httpUri="https://zenodo.org/record/180490/files/figure.png" pageId="26" pageNumber="27">Fig. 36</figureCitation>
); podonotal shield 200–202 long, 175–188 at widest level near posterior margin, with longitudinal distance from alveoli of 
<emphasis id="C89FEAC5FF8EFFC8395BFB2D230ABEAF" box="[1060,1083,1192,1217]" italics="true" pageId="26" pageNumber="27">j1</emphasis>
to 
<emphasis id="C89FEAC5FF8EFFC8391EFB2D2349BEAF" box="[1121,1144,1192,1217]" italics="true" pageId="26" pageNumber="27">j5</emphasis>
about 0.8 that from 
<emphasis id="C89FEAC5FF8EFFC83818FB2D224CBEAF" box="[1383,1405,1192,1217]" italics="true" pageId="26" pageNumber="27">j5</emphasis>
to 
<emphasis id="C89FEAC5FF8EFFC83DE8FB55279FBE87" box="[151,174,1232,1257]" italics="true" pageId="26" pageNumber="27">j6</emphasis>
; pygidial shield 82–87 along midlength, 140–145 at widest level near seta 
<emphasis id="C89FEAC5FF8EFFC83965FB552307BE87" box="[1050,1078,1232,1257]" italics="true" pageId="26" pageNumber="27">S4</emphasis>
, with anterior margin strongly protruding anteriorly slightly beyond level of bases of setae 
<emphasis id="C89FEAC5FF8EFFC83E20FB7D244ABF7F" box="[863,891,1272,1297]" italics="true" pageId="26" pageNumber="27">J2</emphasis>
, and without protuberance near seta 
<emphasis id="C89FEAC5FF8EFFC8384FFB7D227DBF7F" box="[1328,1356,1272,1297]" italics="true" pageId="26" pageNumber="27">S5</emphasis>
. Body dorsum with setae on mid-surfaces simple, very short, similar in length to clunals 
<emphasis id="C89FEAC5FF8EFFC83914FAA523B6BF57" box="[1131,1159,1312,1337]" italics="true" pageId="26" pageNumber="27">J5</emphasis>
(3–5), lateral and marginal setae slightly longer (10–13), 
<emphasis id="C89FEAC5FF8EFFC83F42FACD25B7BF0F" box="[573,646,1352,1377]" italics="true" pageId="26" pageNumber="27">S4–S5</emphasis>
(11–14), 
<emphasis id="C89FEAC5FF8EFFC83F84FACD242BBF0F" box="[763,794,1352,1377]" italics="true" pageId="26" pageNumber="27">Z5</emphasis>
slightly longest (15), 
<emphasis id="C89FEAC5FF8EFFC8395EFACD230FBF0F" box="[1057,1086,1352,1377]" italics="true" pageId="26" pageNumber="27">S4</emphasis>
, 
<emphasis id="C89FEAC5FF8EFFC83931FACD235ABF0F" box="[1102,1131,1352,1377]" italics="true" pageId="26" pageNumber="27">S5</emphasis>
, 
<emphasis id="C89FEAC5FF8EFFC83904FACD23AABF0F" box="[1147,1179,1352,1377]" italics="true" pageId="26" pageNumber="27">Z5</emphasis>
slightly barbed. Peritremes extending anteriorly nearly to level of anterior margins of coxae III.
</paragraph>
<paragraph id="FA5436D7FF8EFFC83DB9FA1D2648BC14" blockId="26.[151,1437,152,2018]" pageId="26" pageNumber="27">
Tritosternum similar in form to that in deutonymph and adult female, but base longer (15) relative to laciniae (40–45), which free for 0.7–0.8 of length. Sternal shield (as in 
<figureCitation id="62D02A52FF8EFFC83E9EFA452304BFB4" box="[993,1077,1472,1498]" captionStart="FIGURES 13 – 15" captionStartId="11.[151,269,1529,1553]" captionTargetBox="[156,1435,492,1528]" captionTargetId="figure@11.[151,1436,394,1627]" captionTargetPageId="11" captionText="FIGURES 13 – 15. Spadiseius calyptrogynae n. sp., protonymph: 13, idiosoma, dorsal aspect; 14, idiosoma, ventral aspect; 15, tectum. Setae added to larval complement denoted without parentheses." httpUri="https://zenodo.org/record/180482/files/figure.png" pageId="26" pageNumber="27">Fig. 14</figureCitation>
) smooth; setae 
<emphasis id="C89FEAC5FF8EFFC83990FA452223BFB7" box="[1263,1298,1472,1497]" italics="true" pageId="26" pageNumber="27">st5</emphasis>
inserted on soft cuticle much shorter (7) than 
<emphasis id="C89FEAC5FF8EFFC83F5FFA6D2545BC6F" box="[544,628,1512,1537]" italics="true" pageId="26" pageNumber="27">st1–st3</emphasis>
(20–22); paragenital poroids absent. Anal shield unornamented, similar in form and structures to those on deutonymph (length 50–60, width 53–60), postanal seta (12) nearly as long as para-anal setae (11–14). Of opisthogastric setae on soft cuticle, 
<emphasis id="C89FEAC5FF8EFFC8396BF9BD23B7BC3F" box="[1044,1158,1592,1617]" italics="true" pageId="26" pageNumber="27">JV2, ZV2</emphasis>
short (5–7), 
<emphasis id="C89FEAC5FF8EFFC83852F9BD22ADBC3F" box="[1325,1436,1592,1617]" italics="true" pageId="26" pageNumber="27">JV1, JV5</emphasis>
slightly larger (12).
</paragraph>
<paragraph id="FA5436D7FF8EFFC83DB9F90D24B9BC9C" blockId="26.[151,1437,152,2018]" pageId="26" pageNumber="27">Tectum more broadly irregularly triangular and bluntly pointed than in deutonymph; other gnathosomatic structures similar to those in deutonymph, except apical sclerotized rims of palpfemur and palpgenu less serrated and palpi with normal protonymphal complement of setae.</paragraph>
<paragraph id="FA5436D7FF8EFFC83DB9F88523C8BD8C" blockId="26.[151,1437,152,2018]" pageId="26" pageNumber="27">
Leg lengths, excluding pretarsi, I (245–275), II (225–245), III (245–260), IV (260–280); pretarsi II–IV with apical setalike processes about one-fourth as long as length of pretarsi to base of claws. Form of legs and their setae, and patterns of serration on coxal faces and apical sclerotized rims of leg segments generally as in deutonymph, except tarsus I apically with more clearly defined pedestal (5) bearing pretarsus. Coxa IV seta as long as coxa III setae (15–16). Femora I–IV with dorsal seta 
<emphasis id="C89FEAC5FF8EFFC83E1CF82524A8BDD7" box="[867,921,1952,1977]" italics="true" pageId="26" pageNumber="27">ad-1</emphasis>
thicker and rod- or oar-like in contrast with adjacent tapered setae, and progressively slightly longer from I (10–11) to IV (13–15) (
<figureCitation id="62D02A52FF8EFFC839EBF84D23DABD8C" box="[1172,1259,1992,2018]" captionStart="FIGURES 38 – 41" captionStartId="24.[151,269,1163,1187]" captionTargetBox="[168,1419,313,1131]" captionTargetId="figure@24.[148,1433,288,1233]" captionTargetPageId="24" captionText="FIGURES 38 – 41. Dorsal subapical femoral setae of legs. Figs. 38, 39, larval femora I – III: 38, Spadiseius calyptrogynae n. sp.; 39, Spadiseius spathiphyllae n. sp. Figs. 40, 41, protonymphal femora I – IV: 40, S. calyptrogynae n. sp.; 41, S." httpUri="https://zenodo.org/record/180491/files/figure.png" pageId="26" pageNumber="27">Fig. 41</figureCitation>
).
</paragraph>
<caption id="AE94665FFF8FFFC93DE8F9682789BD24" httpUri="https://zenodo.org/record/180493/files/figure.png" pageId="27" pageNumber="28" targetBox="[186,1409,183,1772]" targetPageId="27">
<paragraph id="FA5436D7FF8FFFC93DE8F9682789BD24" blockId="27.[151,1436,1773,1866]" pageId="27" pageNumber="28">
<emphasis id="C89FEAC5FF8FFFC93DE8F9682652BD6B" bold="true" box="[151,355,1773,1797]" pageId="27" pageNumber="28">FIGURES 46–50.</emphasis>
<taxonomicName id="3DEB4D54FF8FFFC93C11F96B254DBD6B" box="[366,636,1774,1797]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF8FFFC93C11F96B254DBD6B" box="[366,636,1774,1797]" italics="true" pageId="27" pageNumber="28">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<emphasis id="C89FEAC5FF8FFFC93FF8F96825F9BD6B" bold="true" box="[647,712,1773,1797]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="D3AC57BEFF8FFFC93FF8F96825F9BD6B" box="[647,712,1773,1797]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult female: 46, idiosoma, dorsal aspect, supernumerary setae denoted by ‘x’; 47, idiosoma, ventral aspect; 48, spermathecae, detail of calyx; 49, position of spermatheca at base of leg IV;
</paragraph>
</caption>
<paragraph id="FA5436D7FF8FFFCE3DB9F8F823ABBB14" blockId="27.[151,1436,1917,2023]" lastBlockId="28.[151,1436,152,378]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="C89FEAC5FF8FFFC93DB9F8F82615BDF9" bold="true" box="[198,292,1917,1943]" pageId="27" pageNumber="28">LARVA.</emphasis>
Idiosoma 260–280 long, 175–180 wide at level of setae 
<emphasis id="C89FEAC5FF8FFFC93EBEF8FB24EABDF9" box="[961,987,1918,1943]" italics="true" pageId="27" pageNumber="28">s4</emphasis>
(n = 2); podonotal shield rugose (
<figureCitation id="62D02A52FF8FFFC93812F8F82786BDD1" captionStart="FIGURES 35 – 37" captionStartId="22.[151,269,1758,1782]" captionTargetBox="[151,1434,598,1757]" captionTargetId="figure@22.[151,1436,594,1776]" captionTargetPageId="22" captionText="FIGURES 35 – 37. Spadiseius spathiphyllae n. sp. 35, larva, idiosoma, dorsal aspect; 36, protonymph, idiosoma, dorsal aspect; 37, larva, chelicera, adaxial view. Abbreviation: v m, ventral mucro." httpUri="https://zenodo.org/record/180490/files/figure.png" pageId="27" pageNumber="28">Fig. 35</figureCitation>
), 190–205 long, 148–155 at widest level near posterior margin, with longitudinal distance from alveoli of 
<emphasis id="C89FEAC5FF8FFFC93DE8F84B279FBD89" box="[151,174,1998,2023]" italics="true" pageId="27" pageNumber="28">j1</emphasis>
to 
<emphasis id="C89FEAC5FF8FFFC93DABF84B27DABD89" box="[212,235,1998,2023]" italics="true" pageId="27" pageNumber="28">j5</emphasis>
0.7–0.8 that from 
<emphasis id="C89FEAC5FF8FFFC93CB8F84B26ECBD89" box="[455,477,1998,2023]" italics="true" pageId="27" pageNumber="28">j5</emphasis>
to 
<emphasis id="C89FEAC5FF8FFFC93F7BF84B252BBD89" box="[516,538,1998,2023]" italics="true" pageId="27" pageNumber="28">j6</emphasis>
, and with lateral margins slightly concave between levels of setae 
<emphasis id="C89FEAC5FF8FFFC93851F84B2279BD89" box="[1326,1352,1998,2023]" italics="true" pageId="27" pageNumber="28">z2</emphasis>
and 
<emphasis id="C89FEAC5FF8FFFC938FDF84B22AABD89" box="[1410,1435,1998,2023]" italics="true" pageId="27" pageNumber="28">z4</emphasis>
but not behind 
<emphasis id="C89FEAC5FF88FFCE3C3AFF1D266EBADF" box="[325,351,152,177]" italics="true" pageId="28" pageNumber="29">s4</emphasis>
; pygidial shield with few lines, without puncta, about 65 along midlength in dorsal view (but further length of ca 15 folded under ventrally), 138–145 at widest level near anterior margin, with anterior margin projecting medially to level of setal alveoli 
<emphasis id="C89FEAC5FF88FFCE3F94FF6D2436BB6F" box="[747,775,232,257]" italics="true" pageId="28" pageNumber="29">J2</emphasis>
and deeply invaginated on either side of midline at level of setal alveoli 
<emphasis id="C89FEAC5FF88FFCE3C35FE952654BB47" box="[330,357,272,297]" italics="true" pageId="28" pageNumber="29">J3</emphasis>
(
<figureCitation id="62D02A52FF88FFCE3C0AFE9526FDBB44" box="[373,460,272,298]" captionStart="FIGURES 35 – 37" captionStartId="22.[151,269,1758,1782]" captionTargetBox="[151,1434,598,1757]" captionTargetId="figure@22.[151,1436,594,1776]" captionTargetPageId="22" captionText="FIGURES 35 – 37. Spadiseius spathiphyllae n. sp. 35, larva, idiosoma, dorsal aspect; 36, protonymph, idiosoma, dorsal aspect; 37, larva, chelicera, adaxial view. Abbreviation: v m, ventral mucro." httpUri="https://zenodo.org/record/180490/files/figure.png" pageId="28" pageNumber="29">Fig. 35</figureCitation>
), posterior margin also invaginated on either side of midline at level of setal alveoli 
<emphasis id="C89FEAC5FF88FFCE3DE8FEBD2783BB3F" box="[151,178,312,337]" italics="true" pageId="28" pageNumber="29">J5</emphasis>
. Dorsal body setae similarly short (8–10), simple on podonotal shield, 
<emphasis id="C89FEAC5FF88FFCE397FFEBD232BBB3F" box="[1024,1050,312,337]" italics="true" pageId="28" pageNumber="29">s6</emphasis>
, 
<emphasis id="C89FEAC5FF88FFCE3955FEBD2376BB3F" box="[1066,1095,312,337]" italics="true" pageId="28" pageNumber="29">S3</emphasis>
, 
<emphasis id="C89FEAC5FF88FFCE3928FEBD2344BB3F" box="[1111,1141,312,337]" italics="true" pageId="28" pageNumber="29">S4</emphasis>
slightly shorter (6–8) on soft cuticle, 
<emphasis id="C89FEAC5FF88FFCE3C58FEE52645BB17" box="[295,372,352,377]" italics="true" pageId="28" pageNumber="29">Z3–Z4</emphasis>
on pygidial shield much larger (25–30) and thicker, capitate distally.
</paragraph>
<caption id="AE94665FFF88FFCE3DE8F8F7235DBDB8" httpUri="https://zenodo.org/record/180494/files/figure.png" pageId="28" pageNumber="29" targetBox="[206,1386,432,1878]" targetPageId="28">
<paragraph id="FA5436D7FF88FFCE3DE8F8F7235DBDB8" blockId="28.[151,1436,1906,2006]" pageId="28" pageNumber="29">
<emphasis id="C89FEAC5FF88FFCE3DE8F8F7266FBDE4" bold="true" box="[151,350,1906,1930]" pageId="28" pageNumber="29">FIGURES 51–55.</emphasis>
<taxonomicName id="3DEB4D54FF88FFCE3C1AF8F6255EBDE4" box="[357,623,1907,1930]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF88FFCE3C1AF8F6255EBDE4" box="[357,623,1907,1930]" italics="true" pageId="28" pageNumber="29">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF88FFCE3F09F8F72582BDE4" box="[630,691,1906,1930]" pageId="28" pageNumber="29" rank="species">
<emphasis id="C89FEAC5FF88FFCE3F09F8F72582BDE4" bold="true" box="[630,691,1906,1930]" pageId="28" pageNumber="29">n. sp.</emphasis>
</taxonomicNameLabel>
, adult male. Figs. 51–53, idiosomas, dorsal aspect: 51, “long-haired” morph; 52, intermediate morph; 53, “short-haired” morph, partial view. Fig. 54, detail of peritrematal shield; note variation in length of peritreme from Fig. 52. Fig.55, chelicera and spermatodactyl, adaxial view.
</paragraph>
</caption>
<caption id="AE94665FFF89FFCF3DE8F9EF24E6BCCA" httpUri="https://zenodo.org/record/180495/files/figure.png" pageId="29" pageNumber="30" targetBox="[155,1429,194,1600]" targetPageId="29">
<paragraph id="FA5436D7FF89FFCF3DE8F9EF24E6BCCA" blockId="29.[151,1436,1642,1700]" pageId="29" pageNumber="30">
<emphasis id="C89FEAC5FF89FFCF3DE8F9EF266EBCEC" bold="true" box="[151,351,1642,1666]" pageId="29" pageNumber="30">FIGURES 56–58.</emphasis>
<taxonomicName id="3DEB4D54FF89FFCF3C19F9EE2541BCEC" box="[358,624,1643,1666]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF89FFCF3C19F9EE2541BCEC" box="[358,624,1643,1666]" italics="true" pageId="29" pageNumber="30">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF89FFCF3F07F9EF2584BCEC" box="[632,693,1642,1666]" pageId="29" pageNumber="30" rank="species">
<emphasis id="C89FEAC5FF89FFCF3F07F9EF2584BCEC" bold="true" box="[632,693,1642,1666]" pageId="29" pageNumber="30">n. sp.</emphasis>
</taxonomicNameLabel>
, adult male. Figs. 56, 57, opisthogaster: 56, intermediate morph; 57, “short-haired” morph. Fig. 58, idiosoma, ventral aspect, “long-haired” morph.
</paragraph>
</caption>
<paragraph id="FA5436D7FF89FFCF3DB9F95D2508BDFC" blockId="29.[151,1437,1752,2018]" pageId="29" pageNumber="30">
Tritosternum similar in form to that in protonymph, with laciniae free for about two-thirds of length (29– 37, excluding base). Sternal shield smooth (as in 
<figureCitation id="62D02A52FF89FFCF3F9DF8852406BD74" box="[738,823,1792,1818]" captionStart="FIGURES 10 – 12" captionStartId="10.[151,269,1581,1605]" captionTargetBox="[168,1435,385,1580]" captionTargetId="figure@10.[151,1436,354,1612]" captionTargetPageId="10" captionText="FIGURES 10 – 12. Spadiseius calyptrogynae n. sp., larva: 10, idiosoma, dorsal aspect; 11, idiosoma, ventral aspect; 12, tectum, variations." httpUri="https://zenodo.org/record/180481/files/figure.png" pageId="29" pageNumber="30">Fig. 11</figureCitation>
), with setae 
<emphasis id="C89FEAC5FF89FFCF3EB0F8852313BD77" box="[975,1058,1792,1817]" italics="true" pageId="29" pageNumber="30">st1–st3</emphasis>
similar in length (20–22). Anal shield smooth, slightly wider (48–50) than long (45–47); para-anal setae (19–22) longer than postanal seta (12). Three or four pairs of short opisthogastric setae plus 
<emphasis id="C89FEAC5FF89FFCF3E42F8D4246BBD04" box="[829,858,1873,1898]" italics="true" pageId="29" pageNumber="30">S5</emphasis>
and 
<emphasis id="C89FEAC5FF89FFCF3EECF8D42483BD04" box="[915,946,1873,1898]" italics="true" pageId="29" pageNumber="30">Z5</emphasis>
on soft cuticle, 
<emphasis id="C89FEAC5FF89FFCF3913F8D423ABBD04" box="[1132,1178,1873,1898]" italics="true" pageId="29" pageNumber="30">JV5</emphasis>
minute (3), 
<emphasis id="C89FEAC5FF89FFCF3855F8D42267BD04" box="[1322,1366,1873,1898]" italics="true" pageId="29" pageNumber="30">JV2</emphasis>
longest (10–12), 
<emphasis id="C89FEAC5FF89FFCF3C55F8FC266BBDFC" box="[298,346,1913,1938]" italics="true" pageId="29" pageNumber="30">ZV2</emphasis>
sometimes absent.
</paragraph>
<paragraph id="FA5436D7FF89FFCC3DB9F8252666BADC" blockId="29.[151,1437,1752,2018]" lastBlockId="30.[151,1437,152,378]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
Tectum with anterior margin bluntly unevenly triangular. Fixed digit of chelicera with dentition and pilus dentilis lobe as in protonymph; movable chela 17–18 long, edentate or with a minute tooth, and with ventral mucro (
<figureCitation id="62D02A52FF8AFFCC3D8DFF1D2678BADC" box="[242,329,152,178]" captionStart="FIGURES 35 – 37" captionStartId="22.[151,269,1758,1782]" captionTargetBox="[151,1434,598,1757]" captionTargetId="figure@22.[151,1436,594,1776]" captionTargetPageId="22" captionText="FIGURES 35 – 37. Spadiseius spathiphyllae n. sp. 35, larva, idiosoma, dorsal aspect; 36, protonymph, idiosoma, dorsal aspect; 37, larva, chelicera, adaxial view. Abbreviation: v m, ventral mucro." httpUri="https://zenodo.org/record/180490/files/figure.png" pageId="30" pageNumber="31">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="FA5436D7FF8AFFCC3DB9FF452248BB14" blockId="30.[151,1437,152,378]" pageId="30" pageNumber="31">
Leg lengths, excluding pretarsi: I (220–235), II (195–205), III (220–225); pretarsi II–III with apical setalike processes about one-third as long as length of pretarsi to base of claws. Form of legs and their setae, and patterns of serration on apical sclerotized rims ventrally on femora, genua, tibiae generally as in protonymph. Tarsus I apically with well defined pedestal (6) bearing pretarsus. Femora I–III with dorsal seta 
<emphasis id="C89FEAC5FF8AFFCC398FFEBD2217BB3F" box="[1264,1318,312,337]" italics="true" pageId="30" pageNumber="31">ad-1</emphasis>
similar in size (ca 15), thicker and untapered, rod- or oar-like, apically in contrast to adjacent femoral setae (
<figureCitation id="62D02A52FF8AFFCC386BFEE5225DBB14" box="[1300,1388,352,378]" captionStart="FIGURES 38 – 41" captionStartId="24.[151,269,1163,1187]" captionTargetBox="[168,1419,313,1131]" captionTargetId="figure@24.[148,1433,288,1233]" captionTargetPageId="24" captionText="FIGURES 38 – 41. Dorsal subapical femoral setae of legs. Figs. 38, 39, larval femora I – III: 38, Spadiseius calyptrogynae n. sp.; 39, Spadiseius spathiphyllae n. sp. Figs. 40, 41, protonymphal femora I – IV: 40, S. calyptrogynae n. sp.; 41, S." httpUri="https://zenodo.org/record/180491/files/figure.png" pageId="30" pageNumber="31">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="AE94665FFF8AFFCC3DE8F98F27EFBCE7" httpUri="https://zenodo.org/record/180496/files/figure.png" pageId="30" pageNumber="31" targetBox="[152,1432,441,1545]" targetPageId="30">
<paragraph id="FA5436D7FF8AFFCC3DE8F98F27EFBCE7" blockId="30.[151,1435,1546,1673]" pageId="30" pageNumber="31">
<emphasis id="C89FEAC5FF8AFFCC3DE8F98F2651BC4C" bold="true" box="[151,352,1546,1570]" pageId="30" pageNumber="31">FIGURES 59–64.</emphasis>
<taxonomicName id="3DEB4D54FF8AFFCC3C16F98F2545BC4F" box="[361,628,1546,1569]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF8AFFCC3C16F98F2545BC4F" box="[361,628,1546,1569]" italics="true" pageId="30" pageNumber="31">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8AFFCC3F03F98F258ABC4C" box="[636,699,1546,1570]" pageId="30" pageNumber="31" rank="species">
<emphasis id="C89FEAC5FF8AFFCC3F03F98F258ABC4C" bold="true" box="[636,699,1546,1570]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
, adult female: 59, subcapitulum, abbreviation: 
<emphasis id="C89FEAC5FF8AFFCC39BDF98F23E0BC4C" bold="true" box="[1218,1233,1546,1570]" pageId="30" pageNumber="31">k</emphasis>
, keel; 60, salivary stylus; 61, tarsus I, dorsal view, without pretarsal structures; 62, detail of dorsodistal sensillar complex of tarsus I; 63, enlarged detail of sigmoid seta “s”; 64, leg I, ventral view, excluding tarsus; note serrated apical sclerotized rims of segments.
</paragraph>
</caption>
<subSubSection id="B2F1655CFF8AFFCC3DBAF93823C5BD81" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="FA5436D7FF8AFFCC3DBAF93823C5BD81" blockId="30.[151,1437,1725,2031]" pageId="30" pageNumber="31">
<emphasis id="C89FEAC5FF8AFFCC3DBAF9382649BCB9" bold="true" box="[197,376,1725,1751]" pageId="30" pageNumber="31">
<typeStatus id="25508875FF8AFFCC3DBAF9382631BCB9" box="[197,256,1725,1751]" pageId="30" pageNumber="31">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="82FC7647FF8AFFCC3CFDF938251EBCB9" box="[386,559,1725,1751]" name="Costa Rica" pageId="30" pageNumber="31">COSTA RICA</collectingCountry>
, Heredia Province, La Selva Biological Station, (
<geoCoordinate id="9FDF5010FF8AFFCC3913F93823CEBCB9" box="[1132,1279,1725,1751]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="15" value="10.433611">10° 26' 1&quot; N</geoCoordinate>
, 
<geoCoordinate id="9FDF5010FF8AFFCC3874F93822A5BCB9" box="[1291,1428,1725,1751]" direction="west" orientation="longitude" pageId="30" pageNumber="31" precision="15" value="-84.01723">84° 1' 2&quot; W</geoCoordinate>
) elevation 
<quantity id="3D139B32FF8AFFCC3C76F96026B2BC91" box="[265,387,1765,1791]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="30" pageNumber="31" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50–150 m</quantity>
: 
<typeStatus id="25508875FF8AFFCC3CF2F960253CBC91" box="[397,525,1765,1791]" pageId="30" pageNumber="31" type="holotype">HOLOTYPE</typeStatus>
: adult female, Research Swamp, ex inflorescences of 
<taxonomicName id="3DEB4D54FF8AFFCC39FAF96023D9BC91" box="[1157,1256,1765,1791]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
, 
<taxonomicName id="3DEB4D54FF8AFFCC398CF960267EBD48" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="friedrichsthallii">
<emphasis id="C89FEAC5FF8AFFCC398CF960267EBD48" italics="true" pageId="30" pageNumber="31">Spathiphyllum friedrichsthallii</emphasis>
</taxonomicName>
, 
<date id="8E551017FF8AFFCC3C21F8882503BD49" box="[350,562,1805,1831]" pageId="30" pageNumber="31" value="1994-02-10">10 February 1994</date>
, coll. S. Cunningham &amp; E.E. Lindquist. 
<typeStatus id="25508875FF8AFFCC396DF88823AABD49" box="[1042,1179,1805,1831]" pageId="30" pageNumber="31" type="paratype">PARATYPES</typeStatus>
: 
<specimenCount id="ECEDFD5EFF8AFFCC39D3F888223CBD49" box="[1196,1293,1805,1831]" pageId="30" pageNumber="31" type="adult">18 adult</specimenCount>
females, 
<specimenCount id="ECEDFD5EFF8AFFCC3800F88827E3BD21" pageId="30" pageNumber="31" type="adult">15 adult</specimenCount>
males, 8 deutonymphs, 12 protonymphs, 
<specimenCount id="ECEDFD5EFF8AFFCC3FBBF8B02414BD21" box="[708,805,1845,1871]" pageId="30" pageNumber="31" type="larva">8 larvae</specimenCount>
, with same data as 
<typeStatus id="25508875FF8AFFCC3971F8B02349BD21" box="[1038,1144,1845,1871]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
; 
<specimenCount id="ECEDFD5EFF8AFFCC39FAF8B023E9BD21" box="[1157,1240,1845,1871]" pageId="30" pageNumber="31" type="adult">3 adult</specimenCount>
females, 
<specimenCount id="ECEDFD5EFF8AFFCC3834F8B022ACBD21" box="[1355,1437,1845,1871]" pageId="30" pageNumber="31" type="adult">1 adult</specimenCount>
male, 1 deutonymph, Research Swamp, ex “flowers of Cantarrona aroid” (sic, Cantarrana Swamp, 
<taxonomicName id="3DEB4D54FF8AFFCC38F9F8D8267EBDF0" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="friedrichsthallii">
<emphasis id="C89FEAC5FF8AFFCC38F9F8D8267EBDF0" italics="true" pageId="30" pageNumber="31">S. friedrichsthallii</emphasis>
</taxonomicName>
), 
<date id="8E551017FF8AFFCC3C18F800253FBDF1" box="[359,526,1925,1951]" pageId="30" pageNumber="31" value="1991-04-11">11 April 1991</date>
, coll. S. Cunningham; 
<specimenCount id="ECEDFD5EFF8AFFCC3E64F800245DBDF1" box="[795,876,1925,1951]" pageId="30" pageNumber="31" type="adult">7 adult</specimenCount>
females, Research Swamp, ex bee, 
<taxonomicName id="3DEB4D54FF8AFFCC386DF800266CBDA9" authority="Vachal" authorityName="Vachal" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="silvestriana">
<emphasis id="C89FEAC5FF8AFFCC386DF8002635BDA8" italics="true" pageId="30" pageNumber="31">Trigona silvestriana</emphasis>
Vachal
</taxonomicName>
(on inflorescences of 
<taxonomicName id="3DEB4D54FF8AFFCC3F15F8282470BDA8" box="[618,833,1965,1990]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="friedrichsthallii">
<emphasis id="C89FEAC5FF8AFFCC3F15F8282470BDA8" box="[618,833,1965,1990]" italics="true" pageId="30" pageNumber="31">S. friedrichsthallii</emphasis>
</taxonomicName>
), 
<date id="8E551017FF8AFFCC3E23F82824DFBDA9" box="[860,1006,1965,1991]" pageId="30" pageNumber="31" value="1925-02">25 February</date>
, 
<date id="8E551017FF8AFFCC3E84F8282208BDA9" box="[1019,1337,1965,1991]" pageId="30" pageNumber="31" value="1994-03-08" valueMax="1994-04-04" valueMin="1994-03-08">8 March and 4 April 1994</date>
, coll. S. Cunningham; 
<specimenCount id="ECEDFD5EFF8AFFCC3C41F85026BCBD81" box="[318,397,2005,2031]" pageId="30" pageNumber="31" type="adult">1 adult</specimenCount>
male, “Puesto 
<quantity id="3D139B32FF8AFFCC3F40F8502599BD81" box="[575,680,2005,2031]" metricMagnitude="-2" metricUnit="kg" metricValue="1.5322" pageId="30" pageNumber="31" unit="g" value="15.322">15, 322g</quantity>
, 
<quantity id="3D139B32FF8AFFCC3FCAF85025D9BD81" box="[693,744,2005,2031]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="m" value="10.0">10m</quantity>
(
<geoCoordinate id="9FDF5010FF8AFFCC3F86F85024BFBD81" box="[761,910,2005,2031]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="15" value="10.433333">10° 26' 0&quot; N</geoCoordinate>
, 
<geoCoordinate id="9FDF5010FF8AFFCC3EE5F8502319BD81" box="[922,1064,2005,2031]" direction="west" orientation="longitude" pageId="30" pageNumber="31" precision="15" value="-84.01667">84° 1' 0&quot; W</geoCoordinate>
)”, 
<date id="8E551017FF8AFFCC3937F85023C1BD81" box="[1096,1264,2005,2031]" pageId="30" pageNumber="31" value="1993-03-03">3 March 1993</date>
.
</paragraph>
</subSubSection>
<caption id="AE94665FFF8BFFCD3DE8FB4623FABF71" httpUri="https://zenodo.org/record/180497/files/figure.png" pageId="31" pageNumber="32" targetBox="[164,1431,194,1163]" targetPageId="31">
<paragraph id="FA5436D7FF8BFFCD3DE8FB4623FABF71" blockId="31.[151,1436,1219,1312]" pageId="31" pageNumber="32">
<emphasis id="C89FEAC5FF8BFFCD3DE8FB46266CBEB5" bold="true" box="[151,349,1219,1243]" pageId="31" pageNumber="32">FIGURES 65–69.</emphasis>
Figs. 65–67: 
<taxonomicName id="3DEB4D54FF8BFFCD3C90FB4625C6BEB4" box="[495,759,1219,1242]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="spathiphyllae" status="sp. nov.">
<emphasis id="C89FEAC5FF8BFFCD3C90FB4625C6BEB4" box="[495,759,1219,1242]" italics="true" pageId="31" pageNumber="32">Spadiseius spathiphyllae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8BFFCD3F82FB462408BEB5" box="[765,825,1219,1243]" pageId="31" pageNumber="32" rank="species">
<emphasis id="C89FEAC5FF8BFFCD3F82FB462408BEB5" bold="true" box="[765,825,1219,1243]" pageId="31" pageNumber="32">n. sp.</emphasis>
</taxonomicNameLabel>
, adult male: 65, tarsus IV, dorsolateral view, without pretarsus; 66, 67, femora IV, dorsal view: 66, “short-haired” morph; 67, “long-haired” morph. Figs. 68, 69, 
<taxonomicName id="3DEB4D54FF8BFFCD3855FB63261EBF4E" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="calyptrogynae" status="sp. nov.">
<emphasis id="C89FEAC5FF8BFFCD3855FB63261EBF4E" italics="true" pageId="31" pageNumber="32">Spadiseius calyptrogynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3AC57BEFF8BFFCD3C4AFA8D2640BF4E" box="[309,369,1288,1312]" pageId="31" pageNumber="32" rank="species">
<emphasis id="C89FEAC5FF8BFFCD3C4AFA8D2640BF4E" bold="true" box="[309,369,1288,1312]" pageId="31" pageNumber="32">n. sp.</emphasis>
</taxonomicNameLabel>
, adult male, femora IV, dorsal view: 68, “wartless” morph; 69, “warted” morph.
</paragraph>
</caption>
<subSubSection id="B2F1655CFF8BFFCD3DB9FAD624C9BFFB" pageId="31" pageNumber="32" type="etymology">
<paragraph id="FA5436D7FF8BFFCD3DB9FAD624C9BFFB" blockId="31.[151,1436,1363,1789]" pageId="31" pageNumber="32">
<emphasis id="C89FEAC5FF8BFFCD3DB9FAD62662BF03" bold="true" box="[198,339,1363,1389]" pageId="31" pageNumber="32">Etymology.</emphasis>
The specific epithet, 
<taxonomicName id="3DEB4D54FF8BFFCD3F14FAD62438BF02" box="[619,777,1363,1388]" class="Arachnida" family="Ascidae" genus="Spadiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="spathiphyllae">
<emphasis id="C89FEAC5FF8BFFCD3F14FAD62438BF02" box="[619,777,1363,1388]" italics="true" pageId="31" pageNumber="32">spathiphyllae</emphasis>
</taxonomicName>
, is based on the name of the genus of aroid plants (
<taxonomicName id="3DEB4D54FF8BFFCD3DE0FAFE2632BFFB" box="[159,259,1403,1429]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
) in whose inflorescences these mites undergo their life histories.
</paragraph>
</subSubSection>
<subSubSection id="B2F1655CFF8BFFCD3DB9FA2622A4BC93" pageId="31" pageNumber="32" type="discussion">
<paragraph id="FA5436D7FF8BFFCD3DB9FA26265BBC33" blockId="31.[151,1436,1363,1789]" pageId="31" pageNumber="32">
<emphasis id="C89FEAC5FF8BFFCD3DB9FA262671BFD3" bold="true" box="[198,320,1443,1469]" pageId="31" pageNumber="32">Remarks.</emphasis>
<taxonomicName id="3DEB4D54FF8BFFCD3C35FA262587BFD2" box="[330,694,1443,1468]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="friedrichsthallii">
<emphasis id="C89FEAC5FF8BFFCD3C35FA262587BFD2" box="[330,694,1443,1468]" italics="true" pageId="31" pageNumber="32">Spathiphyllum friedrichsthallii</emphasis>
</taxonomicName>
is widespread in tropical lowland rainforest along the Caribbean coast, from southern 
<collectingCountry id="82FC7647FF8BFFCD3CAEFA4E251ABF8B" box="[465,555,1483,1509]" name="Mexico" pageId="31" pageNumber="32">Mexico</collectingCountry>
to 
<collectingCountry id="82FC7647FF8BFFCD3F2EFA4E25F4BF8B" box="[593,709,1483,1509]" name="Colombia" pageId="31" pageNumber="32">Colombia</collectingCountry>
(Graysum 2003a). A sample of one fully developed inflorescence (spadix) of 
<taxonomicName id="3DEB4D54FF8BFFCD3C17FA76250CBC62" box="[360,573,1523,1548]" class="Liliopsida" family="Araceae" genus="Spathiphyllum" kingdom="Plantae" order="Alismatales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="friedrichsthallii">
<emphasis id="C89FEAC5FF8BFFCD3C17FA76250CBC62" box="[360,573,1523,1548]" italics="true" pageId="31" pageNumber="32">S. friedrichsthallii</emphasis>
</taxonomicName>
taken at the La Selva Biological Station, 10 Februrary 1994, by one of us (EEL) contained more than 300 mites, not accounting for eggs, including an abundance of all developmental instars and adults.
</paragraph>
<paragraph id="FA5436D7FF8BFFCD3DB9F9EE22A4BC93" blockId="31.[151,1436,1363,1789]" pageId="31" pageNumber="32">
In males of this species, the presence of a basitarsal spur on leg IV may correlate with the strongly sclerotized, often bilobate ridge on the posterior edge of the sternogenital shield. The two structures may be in apposition in a grasping function while mating with females. We have observed similarly apposable structures among some fungus-inhabiting species of 
<taxonomicName id="3DEB4D54FF8BFFCD3FFBF9662431BC92" box="[644,768,1763,1788]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="C89FEAC5FF8BFFCD3FFBF9662431BC92" box="[644,768,1763,1788]" italics="true" pageId="31" pageNumber="32">Lasioseius</emphasis>
</taxonomicName>
, e.g., 
<taxonomicName id="3DEB4D54FF8BFFCD3E38F9662304BC93" authority="Koch" authorityName="Koch" box="[839,1077,1763,1789]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="muricatus">
<emphasis id="C89FEAC5FF8BFFCD3E38F96624EABC92" box="[839,987,1763,1788]" italics="true" pageId="31" pageNumber="32">L. muricatus</emphasis>
(Koch)
</taxonomicName>
and 
<taxonomicName id="3DEB4D54FF8BFFCD3912F96622A0BC93" authority="Berlese" authorityName="Berlese" box="[1133,1425,1763,1789]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tuberculiger">
<emphasis id="C89FEAC5FF8BFFCD3912F966222DBC92" box="[1133,1308,1763,1788]" italics="true" pageId="31" pageNumber="32">L. tuberculiger</emphasis>
(Berlese)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>