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<document id="849BE81BF0DC6B428844360FF213869A" ID-CLB-Dataset="53971" ID-DOI="10.11646/zootaxa.5222.2.7" ID-GBIF-Dataset="7e0388db-28c3-477b-a1a6-161c6db00984" ID-ISSN="1175-5326" ID-Zenodo-Dep="7456761" ID-ZooBank="53D96A2E-27CF-482E-8661-ABD55A28BE59" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671439025026" checkinUser="plazi" docAuthor="Sundukov, Yurii N. &amp; Makarov, Kirill V." docDate="2022" docId="72349F6E9E76FFFBFF61FC25FCA7CBB4" docLanguage="en" docName="zootaxa.5222.2.7.pdf" docOrigin="Zootaxa 5222 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Agostenus) quadrisulcatus" docType="treatment" docVersion="7" lastPageNumber="198" masterDocId="8E0DE7169E73FFF3FFF6FFEFFFA3C87D" masterDocTitle="Notes on the systematics and nomenclature of some taxa of the genus Chlaenius Bonelli, 1810 (Coleoptera, Carabidae) from the Far East" masterLastPageNumber="200" masterPageNumber="190" pageNumber="195" updateTime="1699413662263" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D8F02D68735C6DABDF5529538AE20B6E">Notes on the systematics and nomenclature of some taxa of the genus Chlaenius Bonelli, 1810 (Coleoptera, Carabidae) from the Far East</mods:title>
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<mods:namePart id="13EB1A5E46CC48D42390804F3E23866A">Sundukov, Yurii N.</mods:namePart>
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<mods:affiliation id="9A318DA5A0AE94866E5DD370197EE8B7">Federal Scientific Center of East Asia Terrestrial Biodiversity, Zoology Department, Laboratory of Entomology, pr. 100 - letiya Vladivostoka 159, 690022 Vladivostok, Russia. yun-sundukov @ mail. ru; https: // orcid. org / 0000 - 0003 - 3312 - 4029</mods:affiliation>
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<mods:namePart id="EFEA98422C99474A3950B059C486CE36">Makarov, Kirill V.</mods:namePart>
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<mods:affiliation id="2D49D95DD76CEBBBB837610B99E06ACF">Moscow State Pedagogical University, Institute of Biology &amp; Chemistry, Zoology &amp; Ecology Department, Kibalchicha str. 6, bld. 3, Moscow 129164, Russia. https: // orcid. org / 0000 - 0002 - 9184 - 7869</mods:affiliation>
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<treatment id="72349F6E9E76FFFBFF61FC25FCA7CBB4" ID-DOI="http://doi.org/10.5281/zenodo.7469991" ID-GBIF-Taxon="207891982" ID-Zenodo-Dep="7469991" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4" httpUri="http://treatment.plazi.org/id/72349F6E9E76FFFBFF61FC25FCA7CBB4" lastPageId="8" lastPageNumber="198" pageId="5" pageNumber="195">
<subSubSection id="B2877DF39E76FFF6FF61FC25FC89CB99" box="[151,810,970,997]" pageId="5" pageNumber="195" type="nomenclature">
<paragraph id="FA222E789E76FFF6FF61FC25FC89CB99" blockId="5.[151,810,970,1033]" box="[151,810,970,997]" pageId="5" pageNumber="195">
<heading id="A16A99149E76FFF6FF61FC25FC89CB99" bold="true" box="[151,810,970,997]" fontSize="11" level="1" pageId="5" pageNumber="195" reason="1">
<taxonomicName id="3D9D55FB9E76FFF6FF61FC25FC89CB99" ID-CoL="69VTY" authority="(Paykull, 1790)" baseAuthorityName="Paykull" baseAuthorityYear="1790" box="[151,810,970,997]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="quadrisulcatus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E76FFF6FF61FC25FC89CB99" bold="true" box="[151,810,970,997]" pageId="5" pageNumber="195">
<emphasis id="C8E9F26A9E76FFF6FF61FC25FEB0CB98" bold="true" box="[151,275,970,997]" italics="true" pageId="5" pageNumber="195">Chlaenius</emphasis>
(
<emphasis id="C8E9F26A9E76FFF6FED2FC24FDC5CB98" bold="true" box="[292,614,970,997]" italics="true" pageId="5" pageNumber="195">Agostenus) quadrisulcatus</emphasis>
(
<bibRefCitation id="9E0C53899E76FFF6FD83FC25FC81CB99" author="Paykull, G. von" box="[629,802,970,996]" pageId="5" pageNumber="195" refId="ref9070" refString="Paykull, G. von (1790) Monographia caraborum Sueciae. J. F. Edman, Upsaliae, [5] + 138 pp. https: // doi. org / 10.5962 / bhl. title. 146996" type="book" year="1790">Paykull, 1790</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="B2877DF39E76FFF6FF61FC00FED8CC74" box="[151,379,1007,1033]" pageId="5" pageNumber="195" type="description">
<paragraph id="FA222E789E76FFF6FF61FC00FED8CC74" blockId="5.[151,810,970,1033]" box="[151,379,1007,1033]" pageId="5" pageNumber="195">
(
<figureCitation id="62A632FD9E76FFF6FF69FC00FE82CC74" box="[159,289,1007,1033]" captionStart="FIGURES 1318" captionStartId="6.[151,264,1880,1904]" captionTargetBox="[305,1285,676,1862]" captionTargetId="figure-277@6.[304,1286,675,1865]" captionTargetPageId="6" captionText="FIGURES 1318. Far Eastern species of the subgenus Agostenus Fischer von Waldheim, 1829, dorsal view. 13, Chlaenius alutaceus Gеbler, 1830 (Lake Khanka, Primorsky Territory, Russia); 1415, Chlaenius sulcicollis (Paykul, 1798) (14, Belarus; 15, Golubiny Utes, Primorsky Territory, Russia); 1618, Chlaenius quadrisulcatus (Paykul, 1790) (16, Kedrovaya Pad Reserve, Primorsky Territory, Russia; 17, Blagoveshchensk Distr., Amurskaya Oblast, Russia; 18, Kunashir Island, Kuril Islands, Russia)." figureDoi="http://doi.org/10.5281/zenodo.7456767" httpUri="https://zenodo.org/record/7456767/files/figure.png" pageId="5" pageNumber="195">Figs 1618</figureCitation>
,
<figureCitation id="62A632FD9E76FFF6FEDAFC00FED0CC74" box="[300,371,1007,1033]" captionStart="FIGURES 1927" captionStartId="7.[151,264,1785,1809]" captionTargetBox="[161,1433,424,1751]" captionTargetId="figure-143@7.[151,1435,417,1762]" captionTargetPageId="7" captionText="FIGURES 1927. Aedeagi of the subgenus Agostenus Fischervon Waldheim, 1829, apical lamella (dorsal view) and median lobe (lateral view). 1920, Chlaenius alutaceus Gеbler, 1830 (19, Moldova; 20, Buryatia, Russia); 2124, Chlaenius quadrisulcatus (Paykul, 1790) (21, Belarus; 22, Amurskaya Oblast, Russia; 23, Narva River, Khasansky Distr., Primorsky Territory, Russia; 24, Kunashir Island, Kurile Islands, Russia); 2527, Chlaenius sulcicollis (Paykul, 1798) (25, Belovezhskaya Pushcha, Belarus; 26, Vladimir.[skaya] Vol.[olst], Belarus; 27, Golubinyi Utes, Primorsky Territory, Russia)." figureDoi="http://doi.org/10.5281/zenodo.7456769" httpUri="https://zenodo.org/record/7456769/files/figure.png" pageId="5" pageNumber="195">2124</figureCitation>
)
</paragraph>
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<subSubSection id="B2877DF39E76FFF6FF61FBC5FB20CCE3" pageId="5" pageNumber="195" type="reference_group">
<paragraph id="FA222E789E76FFF6FF61FBC5FB09CC3F" blockId="5.[151,1194,1066,1182]" box="[151,1194,1066,1090]" pageId="5" pageNumber="195">
<treatmentCitationGroup id="DA8D09569E76FFF6FF61FBC5FB09CC3F" box="[151,1194,1066,1090]" pageId="5" pageNumber="195">
<treatmentCitation id="7B3C08699E76FFF6FF61FBC5FD2ECC3F" author="Paykull, G. von" box="[151,653,1066,1090]" page="110" pageId="5" pageNumber="195" year="1790">
<taxonomicName id="3D9D55FB9E76FFF6FF61FBC5FD2ECC3F" ID-CoL="QYLT" authority="Paykull, 1790: 109 - 110" authorityName="Paykull" authorityPageNumber="109 - 110" authorityYear="1790" box="[151,653,1066,1090]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="quadrisulcatus">
<emphasis id="C8E9F26A9E76FFF6FF61FBC5FE32CC3F" box="[151,401,1066,1090]" italics="true" pageId="5" pageNumber="195">Carabus Quadrisulcatus</emphasis>
<bibRefCitation id="9E0C53899E76FFF6FE61FBC5FD2ECC3F" author="Paykull, G. von" box="[407,653,1066,1090]" pageId="5" pageNumber="195" refId="ref9070" refString="Paykull, G. von (1790) Monographia caraborum Sueciae. J. F. Edman, Upsaliae, [5] + 138 pp. https: // doi. org / 10.5962 / bhl. title. 146996" type="book" year="1790">Paykull, 1790: 109110</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<materialsCitation id="4AF524259E76FFF6FD61FBC5FB04CC3F" ID-GBIF-Occurrence="4000632378" box="[663,1191,1066,1090]" country="Sweden" location="Island" pageId="5" pageNumber="195" specimenCount="1" stateProvince="Gotland">
<typeStatus id="252690DA9E76FFF6FD61FBC5FD60CC3F" box="[663,707,1066,1090]" pageId="5" pageNumber="195">type</typeStatus>
locality: “Gottlandia”,
<collectingRegion id="3859E09A9E76FFF6FC46FBC5FBA1CC3F" box="[944,1026,1066,1090]" country="Sweden" name="Gotland" pageId="5" pageNumber="195">Gotland</collectingRegion>
<location id="FF4278A39E76FFF6FBFEFBC5FBEBCC3F" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FBFEFBC5FBEBCC3F" box="[1032,1096,1066,1090]" country="Sweden" name="Island" pageId="5" pageNumber="195" stateProvince="Gotland">Island</location>
,
<collectingCountry id="828A6EE89E76FFF6FBA5FBC5FB04CC3F" box="[1107,1191,1066,1090]" name="Sweden" pageId="5" pageNumber="195">Sweden</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA222E789E76FFF6FF61FBA7FBB7CC1C" blockId="5.[151,1194,1066,1182]" box="[151,1044,1096,1121]" pageId="5" pageNumber="195">
<treatmentCitationGroup id="DA8D09569E76FFF6FF61FBA7FBB7CC1C" box="[151,1044,1096,1121]" pageId="5" pageNumber="195">
<treatmentCitation id="7B3C08699E76FFF6FF61FBA7FD84CC1C" author="Weber, F." box="[151,551,1096,1121]" page="43" pageId="5" pageNumber="195" year="1801">
<taxonomicName id="3D9D55FB9E76FFF6FF61FBA7FD84CC1C" ID-CoL="54DB9" authority="Weber, 1801: 42 - 43" authorityName="Weber" authorityPageNumber="42 - 43" authorityYear="1801" box="[151,551,1096,1121]" class="Insecta" family="Carabidae" genus="Tachypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="caelatus">
<emphasis id="C8E9F26A9E76FFF6FF61FBA7FEF3CC1D" box="[151,336,1096,1120]" italics="true" pageId="5" pageNumber="195">Tachypus caelatus</emphasis>
<bibRefCitation id="9E0C53899E76FFF6FEA1FBA7FD84CC1C" author="Weber, F." box="[343,551,1096,1121]" pageId="5" pageNumber="195" pagination="1" refId="ref9620" refString="Weber, F. (1801) Observationes entomologicae, continentes novorum quae condidit generum characteres, et nuper detectarum specierum descriptiones. Impensis Bibliopolii Academici Novi, Kiliae, xii + 116 + [1] pp. https: // doi. org / 10.5962 / bhl. title. 8639" type="journal article" year="1801">Weber, 1801: 4243</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<materialsCitation id="4AF524259E76FFF6FDC7FBA6FBB3CC1C" ID-GBIF-Occurrence="4000632301" box="[561,1040,1096,1121]" country="Germany" location="Holstein" pageId="5" pageNumber="195" specimenCount="1">
<typeStatus id="252690DA9E76FFF6FDC7FBA6FDFECC1C" box="[561,605,1097,1121]" pageId="5" pageNumber="195">type</typeStatus>
locality: “In Holfatia”,
<location id="FF4278A39E76FFF6FCB8FBA7FC05CC1D" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FCB8FBA7FC05CC1D" box="[846,934,1096,1120]" country="Germany" name="Holstein" pageId="5" pageNumber="195">Holstein</location>
,
<collectingCountry id="828A6EE89E76FFF6FC46FBA6FBB3CC1C" box="[944,1040,1097,1121]" name="Germany" pageId="5" pageNumber="195">Germany</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA222E789E76FFF6FF61FB88FC44CC02" blockId="5.[151,1194,1066,1182]" box="[151,999,1127,1151]" pageId="5" pageNumber="195">
<taxonomicName id="3D9D55FB9E76FFF6FF61FB88FD29CC02" ID-CoL="5XVCZ" authority="Ganglbauer, 1891" authorityName="Ganglbauer" authorityYear="1891" box="[151,650,1127,1151]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="gebleri" subGenus="Agostenus">
<emphasis id="C8E9F26A9E76FFF6FF61FB88FF5DCC02" box="[151,254,1127,1151]" italics="true" pageId="5" pageNumber="195">Chlaenius</emphasis>
(
<emphasis id="C8E9F26A9E76FFF6FEFBFB88FE6FCC02" box="[269,460,1127,1151]" italics="true" pageId="5" pageNumber="195">Agostenus) gebleri</emphasis>
<bibRefCitation id="9E0C53899E76FFF6FE24FB88FD29CC02" author="Ganglbauer, L." box="[466,650,1127,1151]" pageId="5" pageNumber="195" refId="ref7652" refString="Ganglbauer, L. (1891) Die Kafer von Mitteleuropa. Die Kafer der Osterreichisch-Ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. Erster Band. Familienreihe Caraboidea. C. Gerold's Sohn, Wien, iii + 557 pp. https: // doi. org / 10.5962 / bhl. title. 8764" type="book" year="1891">Ganglbauer, 1891</bibRefCitation>
</taxonomicName>
: records from
<collectingCountry id="828A6EE89E76FFF6FCEAFB87FCF5CC02" box="[796,854,1128,1151]" name="Japan" pageId="5" pageNumber="195">Japan</collectingCountry>
and Sakhalin.
</paragraph>
<paragraph id="FA222E789E76FFF6FF61FB69FB20CCE3" blockId="5.[151,1194,1066,1182]" box="[151,1155,1158,1182]" pageId="5" pageNumber="195">
<taxonomicName id="3D9D55FB9E76FFF6FF61FB69FD31CCE3" ID-CoL="69W5Y" authority="(Paykull, 1798)" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[151,658,1158,1182]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="sulcicollis" subGenus="Agostenus">
<emphasis id="C8E9F26A9E76FFF6FF61FB69FF5DCCE3" box="[151,254,1158,1182]" italics="true" pageId="5" pageNumber="195">Chlaenius</emphasis>
(
<emphasis id="C8E9F26A9E76FFF6FEFBFB69FE48CCE3" box="[269,491,1158,1182]" italics="true" pageId="5" pageNumber="195">Agostenus) sulcicollis</emphasis>
(Paykull, 1798)
</taxonomicName>
: records from
<collectingCountry id="828A6EE89E76FFF6FCD2FB69FCFDCCE0" box="[804,862,1158,1181]" name="Japan" pageId="5" pageNumber="195">Japan</collectingCountry>
by
<bibRefCitation id="9E0C53899E76FFF6FC75FB69FBDCCCE3" author="Yoshimatsu, S. - I. &amp; Ito, N. &amp; Nakatani, Y. &amp; Yoshitake, H." box="[899,1151,1158,1182]" pageId="5" pageNumber="195" pagination="15 - 191" refId="ref9751" refString="Yoshimatsu, S. - I., Ito, N., Nakatani, Y. &amp; Yoshitake, H. (2018) A list of ground beetles (Insecta: Coleoptera: Caraboidea) in Dr. Kazuo Tanaka's collection preserved in the Insect Museum of Institute for Agro-Environmental Sciences, Naro. Bulletin of National Institute for Agro-Environmental Sciences, 39, 15 - 191." type="journal article" year="2018">
Yoshimatsu
<emphasis id="C8E9F26A9E76FFF6FBF6FB68FB90CCE3" box="[1024,1075,1158,1182]" italics="true" pageId="5" pageNumber="195">et al.</emphasis>
(2018)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2877DF39E76FFF5FF61FB2DFC71C93C" lastPageId="6" lastPageNumber="196" pageId="5" pageNumber="195" type="materials_examined">
<paragraph id="FA222E789E76FFF6FF61FB2DFBA1CE38" blockId="5.[151,1437,1218,2039]" pageId="5" pageNumber="195">
<emphasis id="C8E9F26A9E76FFF6FF61FB2DFE24CCA1" bold="true" box="[151,391,1218,1244]" pageId="5" pageNumber="195">Material examined.</emphasis>
<emphasis id="C8E9F26A9E76FFF6FE62FB2CFD86CCA0" box="[404,549,1219,1245]" italics="true" pageId="5" pageNumber="195">Kuril Islands</emphasis>
:
<materialsCitation id="4AF524259E76FFF6FDCEFB2CFD74CD7C" ID-GBIF-Occurrence="4000632363" collectingDate="2016-05-28" collectionCode="FEB" collectorName="Yu. Sundukov" county="Belozerskaya" location="Kunashir Island" municipality="Rikord" pageId="5" pageNumber="195" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location id="FF4278A39E76FFF6FDCEFB2CFD4ACCA0" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FDCEFB2CFD4ACCA0" box="[568,745,1219,1245]" county="Belozerskaya" municipality="Rikord" name="Kunashir Island" pageId="5" pageNumber="195">Kunashir Island</location>
, swamp in the interfluve of
<collectingMunicipality id="1A46B4029E76FFF6FBC3FB2CFBDCCCA0" box="[1077,1151,1219,1245]" pageId="5" pageNumber="195">Rikord</collectingMunicipality>
and
<collectingCounty id="134356F49E76FFF6FB48FB2CFAEFCCA0" box="[1214,1356,1219,1245]" pageId="5" pageNumber="195">Belozerskaya</collectingCounty>
rivers,
<date id="8E2308B89E76FFF6FF61FB08FEA5CD7C" box="[151,262,1255,1281]" pageId="5" pageNumber="195" value="2016-05-28">
<collectingDate id="9E67F1509E76FFF6FF61FB08FEA5CD7C" box="[151,262,1255,1281]" pageId="5" pageNumber="195" value="2016-05-28">28.V.2016</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FEF9FB08FE06CD7C" box="[271,421,1255,1281]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FE2BFB08FD84CD7C" box="[477,551,1255,1281]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FDC6FB08FD2FCD7C" box="[560,652,1255,1281]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FD6CFB08FD73CD7C" box="[666,720,1255,1281]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4AF524259E76FFF6FD17FB08FCBACD58" ID-GBIF-Occurrence="4000632366" collectingDate="1998-06-17" collectingDateMax="1998-06-22" collectingDateMin="1998-06-17" collectionCode="FEB" collectorName="Yu. Sundukov" county="Lake Blagodatnoe" location="Primorsky Territory" municipality="Sikhote-Alinsky Reserve" pageId="5" pageNumber="195" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<emphasis id="C8E9F26A9E76FFF6FD17FB08FC10CD7D" box="[737,947,1255,1281]" italics="true" pageId="5" pageNumber="195">
<location id="FF4278A39E76FFF6FD17FB08FC10CD7D" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FD17FB08FC10CD7D" box="[737,947,1255,1281]" county="Lake Blagodatnoe" municipality="Sikhote-Alinsky Reserve" name="Primorsky Territory" pageId="5" pageNumber="195">Primorsky Territory</location>
</emphasis>
:
<collectingMunicipality id="1A46B4029E76FFF6FC36FB08FB68CD7C" box="[960,1227,1255,1281]" pageId="5" pageNumber="195">Sikhote-Alinsky Reserve</collectingMunicipality>
,
<collectingCounty id="134356F49E76FFF6FB22FB08FA3BCD7C" box="[1236,1432,1255,1281]" pageId="5" pageNumber="195">Lake Blagodatnoe</collectingCounty>
,
<date id="8E2308B89E76FFF6FF61FAE4FE99CD58" box="[151,314,1291,1317]" pageId="5" pageNumber="195" value="1998-06-17" valueMax="1998-06-22" valueMin="1998-06-17">
<collectingDate id="9E67F1509E76FFF6FF61FAE4FE99CD58" box="[151,314,1291,1317]" pageId="5" pageNumber="195" value="1998-06-17" valueMax="1998-06-22" valueMin="1998-06-17">1722.VI.1998</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FEB5FAE4FE79CD58" box="[323,474,1291,1317]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FDE5FAE4FDC4CD58" box="[531,615,1291,1317]" pageId="5" pageNumber="195" type="male">2 males</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FD84FAE4FD6ECD58" box="[626,717,1291,1317]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FD2AFAE4FCB1CD58" box="[732,786,1291,1317]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FCD2FAE4FF7ECD34" ID-GBIF-Occurrence="4000632391" collectingDate="1999-07-08" collectingDateMax="1999-07-12" collectingDateMin="1999-07-08" collectionCode="FEB" collectorName="Yu. Sundukov" county="Lake Blagodatnoe" location="Primorsky Territory" municipality="Sikhote-Alinsky Reserve" pageId="5" pageNumber="195" specimenCount="10" specimenCount-female="6" specimenCount-male="4">
ibid.,
<date id="8E2308B89E76FFF6FC96FAE4FC5DCD58" box="[864,1022,1291,1317]" pageId="5" pageNumber="195" value="1999-07-08" valueMax="1999-07-12" valueMin="1999-07-08">
<collectingDate id="9E67F1509E76FFF6FC96FAE4FC5DCD58" box="[864,1022,1291,1317]" pageId="5" pageNumber="195" value="1999-07-08" valueMax="1999-07-12" valueMin="1999-07-08">812.VII.1999</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FBF1FAE4FB3DCD58" box="[1031,1182,1291,1317]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FB21FAE4FA88CD58" box="[1239,1323,1291,1317]" pageId="5" pageNumber="195" type="male">4 males</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FAC3FAE4FA3FCD58" box="[1333,1436,1291,1317]" pageId="5" pageNumber="195" type="female">6 females</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FF69FAC0FF76CD34" box="[159,213,1327,1353]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FF11FAC0FA89CD34" ID-GBIF-Occurrence="4000632393" box="[231,1322,1327,1353]" collectingDate="2001-08-12" collectionCode="FEB" collectorName="V. Kuznetsov" location="Ilistaya River" pageId="5" pageNumber="195" specimenCount="1" specimenCount-female="1">
hunting base
<locationDeviation id="EB90375A9E76FFF6FE85FAC0FD20CD34" box="[371,643,1327,1353]" location="Chernigovka" pageId="5" pageNumber="195">
<quantity id="3D65839D9E76FFF6FE85FAC0FE15CD34" box="[371,438,1327,1353]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="195" unit="km" value="10.0">10 km</quantity>
W of Chernigovka
</locationDeviation>
,
<location id="FF4278A39E76FFF6FD7BFAC0FCBFCD34" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FD7BFAC0FCBFCD34" box="[653,796,1327,1353]" name="Ilistaya River" pageId="5" pageNumber="195">Ilistaya River</location>
,
<date id="8E2308B89E76FFF6FCD0FAC0FC11CD34" box="[806,946,1327,1353]" pageId="5" pageNumber="195" value="2001-08-12">
<collectingDate id="9E67F1509E76FFF6FCD0FAC0FC11CD34" box="[806,946,1327,1353]" pageId="5" pageNumber="195" value="2001-08-12">12.VIII.2001</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FC4AFAC0FBEACD34" box="[956,1097,1327,1353]" pageId="5" pageNumber="195">V. Kuznetsov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FB75FAC0FB7DCD34" box="[1155,1246,1327,1353]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FB1BFAC0FA80CD34" box="[1261,1315,1327,1353]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FAC2FAC0FB96CD10" ID-GBIF-Occurrence="4000632306" collectingDate="1995-06-09" collectingDateMax="1995-06-12" collectingDateMin="1995-06-09" collectionCode="FEB, MPU" collectorName="Yu. Sundukov" location="Lazovsky Reserve" municipality="Petrova Bay" pageId="5" pageNumber="195" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<location id="FF4278A39E76FFF6FAC2FAC0FF4CCD10" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FAC2FAC0FF4CCD10" municipality="Petrova Bay" name="Lazovsky Reserve" pageId="5" pageNumber="195">Lazovsky Reserve</location>
,
<collectingMunicipality id="1A46B4029E76FFF6FF0EFABCFEDACD10" box="[248,377,1363,1389]" pageId="5" pageNumber="195">Petrova Bay</collectingMunicipality>
,
<date id="8E2308B89E76FFF6FE75FABDFDBBCD10" box="[387,536,1362,1389]" pageId="5" pageNumber="195" value="1995-06-09" valueMax="1995-06-12" valueMin="1995-06-09">
<collectingDate id="9E67F1509E76FFF6FE75FABDFDBBCD10" box="[387,536,1362,1389]" pageId="5" pageNumber="195" value="1995-06-09" valueMax="1995-06-12" valueMin="1995-06-09">912.VI.1995</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FDD7FABCFD15CD10" box="[545,694,1363,1389]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FD18FABCFCE2CD10" box="[750,833,1363,1389]" pageId="5" pageNumber="195" type="male">3 males</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FCBCFABCFC06CD10" box="[842,933,1363,1389]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FC44FABCFC45CD10" box="[946,998,1363,1389]" pageId="5" pageNumber="195">FEB</collectionCode>
,
<collectionCode id="9C8CB6BD9E76FFF6FC06FABCFB8CCD10" box="[1008,1071,1363,1389]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FBC9FABCFECECDEC" ID-GBIF-Occurrence="4000632308" collectingDate="1991-06-20" collectingDateMax="1991-06-23" collectingDateMin="1991-06-20" collectionCode="MPU" collectorName="D. Obydov" location="Lazovsky Reserve" municipality="Petrova Bay" pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1">
ibid.,
<date id="8E2308B89E76FFF6FB8CFABCFABECD10" box="[1146,1309,1363,1389]" pageId="5" pageNumber="195" value="1991-06-20" valueMax="1991-06-23" valueMin="1991-06-20">
<collectingDate id="9E67F1509E76FFF6FB8CFABCFABECD10" box="[1146,1309,1363,1389]" pageId="5" pageNumber="195" value="1991-06-20" valueMax="1991-06-23" valueMin="1991-06-20">2023.VI.1991</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FAD0FABCFA3ECD10" box="[1318,1437,1363,1389]" pageId="5" pageNumber="195">D. Obydov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FF3AFA98FEB5CDEC" box="[204,278,1399,1425]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FED3FA98FEC6CDEC" box="[293,357,1399,1425]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FE8FFA98FA3BCDEC" ID-GBIF-Occurrence="4000632341" box="[377,1432,1398,1425]" collectingDate="2004-06-13" collectingDateMax="2004-06-15" collectingDateMin="2004-06-13" collectionCode="FEB" collectorName="Yu. Sundukov &amp; L. Sundukova" location="Lake Zarya" municipality="Petrova Bay" pageId="5" pageNumber="195" specimenCount="14" specimenCount-female="6" specimenCount-male="8">
ibid.,
<location id="FF4278A39E76FFF6FE40FA98FD97CDEC" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FE40FA98FD97CDEC" box="[438,564,1399,1425]" name="Lake Zarya" pageId="5" pageNumber="195">Lake Zarya</location>
,
<date id="8E2308B89E76FFF6FDB6FA99FD41CDEC" box="[576,738,1398,1425]" pageId="5" pageNumber="195" value="2004-06-13" valueMax="2004-06-15" valueMin="2004-06-13">
<collectingDate id="9E67F1509E76FFF6FDB6FA99FD41CDEC" box="[576,738,1398,1425]" pageId="5" pageNumber="195" value="2004-06-13" valueMax="2004-06-15" valueMin="2004-06-13">1315.VI.2004</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FD18FA98FC25CDEC" box="[750,902,1399,1425]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
&amp;
<collectorName id="57684BAE9E76FFF6FC5AFA98FBE0CDEC" box="[940,1091,1399,1425]" pageId="5" pageNumber="195">L. Sundukova</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FB76FA98FB76CDEC" box="[1152,1237,1399,1425]" pageId="5" pageNumber="195" type="male">8 males</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FB14FA98FAEACDEC" box="[1250,1353,1399,1425]" pageId="5" pageNumber="195" type="female">6 females</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FAACFA98FA33CDEC" box="[1370,1424,1399,1425]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FF61FA74FB1ACDC8" ID-GBIF-Occurrence="4000632416" box="[151,1209,1434,1461]" collectingDate="2007-08-20" collectingDateMax="2004-06-15" collectingDateMin="2004-06-13" collectionCode="MPU" collectorName="K. Makarov &amp; A. Zaitsev" latitude="43.031387" location="Lake Zarya" longLatPrecision="19" longitude="134.14555" municipality="Petrova Bay" pageId="5" pageNumber="195" specimenCount="1" specimenCount-female="1">
ibid.,
<geoCoordinate id="9FA948BF9E76FFF6FF21FA75FEF9CDC8" box="[215,346,1434,1461]" degrees="43" direction="north" minutes="01" orientation="latitude" pageId="5" pageNumber="195" precision="15" seconds="53" value="43.031387">43°0153”N</geoCoordinate>
<geoCoordinate id="9FA948BF9E76FFF6FE92FA74FE51CDC8" box="[356,498,1435,1461]" degrees="134" direction="east" minutes="08" orientation="longitude" pageId="5" pageNumber="195" precision="15" seconds="44" value="134.14555">134°0844”E</geoCoordinate>
,
<date id="8E2308B89E76FFF6FDF4FA74FD2DCDC8" box="[514,654,1435,1461]" pageId="5" pageNumber="195" value="2007-08-20">
<collectingDate id="9E67F1509E76FFF6FDF4FA74FD2DCDC8" box="[514,654,1435,1461]" pageId="5" pageNumber="195" value="2007-08-20">20.VIII.2007</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FD6BFA74FC82CDC8" box="[669,801,1435,1461]" pageId="5" pageNumber="195">K. Makarov</collectorName>
&amp;
<collectorName id="57684BAE9E76FFF6FCBCFA74FC1ECDC8" box="[842,957,1435,1461]" pageId="5" pageNumber="195">A. Zaitsev</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FC09FA74FBFCCDC8" box="[1023,1119,1435,1461]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FB87FA74FB12CDC8" box="[1137,1201,1435,1461]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FB31FA74FCCDCDA4" ID-GBIF-Occurrence="4000632307" collectingDate="2019-06-03" collectionCode="FEB" collectorName="Yu. Sundukov" location="Khasansky District" municipality="Melkovodnaya Bay" pageId="5" pageNumber="195" specimenCount="2" specimenCount-male="2">
<location id="FF4278A39E76FFF6FB31FA74FA3ACDC8" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FB31FA74FA3ACDC8" box="[1223,1433,1435,1461]" municipality="Melkovodnaya Bay" name="Khasansky District" pageId="5" pageNumber="195">Khasansky District</location>
,
<collectingMunicipality id="1A46B4029E76FFF6FF61FA50FEC8CDA4" box="[151,363,1471,1497]" pageId="5" pageNumber="195">Melkovodnaya Bay</collectingMunicipality>
,
<date id="8E2308B89E76FFF6FE8FFA50FE45CDA4" box="[377,486,1471,1497]" pageId="5" pageNumber="195" value="2019-06-03">
<collectingDate id="9E67F1509E76FFF6FE8FFA50FE45CDA4" box="[377,486,1471,1497]" pageId="5" pageNumber="195" value="2019-06-03">3.VI.2019</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FE05FA50FD2FCDA4" box="[499,652,1471,1497]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FD3DFA50FC83CDA4" box="[715,800,1471,1497]" pageId="5" pageNumber="195" type="male">2 males</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FCC7FA50FCC4CDA4" box="[817,871,1471,1497]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FC8AFA50FA3BCDA4" ID-GBIF-Occurrence="4000632335" box="[892,1432,1471,1497]" collectingDate="2019-06-03" collectionCode="MPU" collectorName="K. Makarov" location="Khasansky District" municipality="Melkovodnaya Bay" pageId="5" pageNumber="195" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
ibid.,
<collectorName id="57684BAE9E76FFF6FC4CFA50FB9ECDA4" box="[954,1085,1471,1497]" pageId="5" pageNumber="195">K. Makarov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FB8BFA50FB70CDA4" box="[1149,1235,1471,1497]" pageId="5" pageNumber="195" type="male">2 males</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FB17FA50FA9CCDA4" box="[1249,1343,1471,1497]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FAA6FA50FA33CDA4" box="[1360,1424,1471,1497]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FF61FA0CFB1FCD80" ID-GBIF-Occurrence="4000632310" box="[151,1212,1507,1533]" collectingDate="2021-05-24" collectionCode="FEB, MPU" collectorName="Yu. Sundukov" location="Narva River" municipality="Siniy Utes" pageId="5" pageNumber="195" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
ibid.,
<location id="FF4278A39E76FFF6FF23FA0CFEFACD80" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FF23FA0CFEFACD80" box="[213,345,1507,1533]" municipality="Siniy Utes" name="Narva River" pageId="5" pageNumber="195">Narva River</location>
near
<collectingMunicipality id="1A46B4029E76FFF6FE61FA0CFDA9CD80" box="[407,522,1507,1533]" pageId="5" pageNumber="195">Siniy Utes</collectingMunicipality>
,
<date id="8E2308B89E76FFF6FDE0FA0CFD26CD80" box="[534,645,1507,1533]" pageId="5" pageNumber="195" value="2021-05-24">
<collectingDate id="9E67F1509E76FFF6FDE0FA0CFD26CD80" box="[534,645,1507,1533]" pageId="5" pageNumber="195" value="2021-05-24">24.V.2021</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FD66FA0CFC8BCD80" box="[656,808,1507,1533]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FC93FA0CFC18CD80" box="[869,955,1507,1533]" pageId="5" pageNumber="195" type="male">3 males</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FC31FA0CFB86CD80" box="[967,1061,1507,1533]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FBC3FA0CFBCACD80" box="[1077,1129,1507,1533]" pageId="5" pageNumber="195">FEB</collectionCode>
,
<collectionCode id="9C8CB6BD9E76FFF6FB80FA0CFB16CD80" box="[1142,1205,1507,1533]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4AF524259E76FFF6FB3EFA0CFA9BCE5C" ID-GBIF-Occurrence="4000632380" collectingDate="2003-07-27" collectingDateMax="2003-07-31" collectingDateMin="2003-07-27" collectionCode="ZISP" collectorName="B. Kataev" county="Blagoveshchensk Distr." location="Amur region" municipality="Amur Prov." pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1">
<emphasis id="C8E9F26A9E76FFF6FB3EFA0CFAECCD81" box="[1224,1359,1507,1533]" italics="true" pageId="5" pageNumber="195">
<location id="FF4278A39E76FFF6FB3EFA0CFAECCD81" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FB3EFA0CFAECCD81" box="[1224,1359,1507,1533]" county="Blagoveshchensk Distr." municipality="Amur Prov." name="Amur region" pageId="5" pageNumber="195">Amur region</location>
</emphasis>
:
<collectingMunicipality id="1A46B4029E76FFF6FAABFA0CFF72CE5C" pageId="5" pageNumber="195">Amur Prov.</collectingMunicipality>
,
<collectingCounty id="134356F49E76FFF6FF2AF9E8FE7ACE5C" box="[220,473,1543,1569]" pageId="5" pageNumber="195">Blagoveshchensk Distr.</collectingCounty>
, ca.
<locationDeviation id="EB90375A9E76FFF6FDFFF9E9FD49CE5C" box="[521,746,1542,1569]" location="Natalino" pageId="5" pageNumber="195">
<quantity id="3D65839D9E76FFF6FDFFF9E9FDEECE5C" box="[521,589,1542,1569]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="195" unit="km" value="15.0">15 km</quantity>
N of Natalino
</locationDeviation>
, forest,
<date id="8E2308B89E76FFF6FCC9F9E8FC49CE5C" box="[831,1002,1543,1569]" pageId="5" pageNumber="195" value="2003-07-27" valueMax="2003-07-31" valueMin="2003-07-27">
<collectingDate id="9E67F1509E76FFF6FCC9F9E8FC49CE5C" box="[831,1002,1543,1569]" pageId="5" pageNumber="195" value="2003-07-27" valueMax="2003-07-31" valueMin="2003-07-27">2731.VII.2003</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FC00F9E8FBC2CE5C" box="[1014,1121,1543,1569]" pageId="5" pageNumber="195">B. Kataev</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FB6AF9E8FB46CE5C" box="[1180,1253,1543,1569]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FB02F9E8FA93CE5C" box="[1268,1328,1543,1569]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="195">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4AF524259E76FFF6FAB4F9E8FC5DCE38" ID-GBIF-Occurrence="4000632314" collectingDate="1999-06-05" collectionCode="MPU" collectorName="R. Minets" country="Belarus" location="Belovezhskaya Pushcha" municipality="Dikoe Swamp" pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1">
<emphasis id="C8E9F26A9E76FFF6FAB4F9E8FA37CE5C" box="[1346,1428,1543,1569]" italics="true" pageId="5" pageNumber="195">
<collectingCountry id="828A6EE89E76FFF6FAB4F9E8FA37CE5C" box="[1346,1428,1543,1569]" name="Belarus" pageId="5" pageNumber="195">Belarus</collectingCountry>
</emphasis>
:
<location id="FF4278A39E76FFF6FF61F9C4FE3ACE38" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FF61F9C4FE3ACE38" box="[151,409,1579,1605]" country="Belarus" municipality="Dikoe Swamp" name="Belovezhskaya Pushcha" pageId="5" pageNumber="195">Belovezhskaya Pushcha</location>
,
<collectingMunicipality id="1A46B4029E76FFF6FE55F9C4FD9ECE38" box="[419,573,1579,1605]" pageId="5" pageNumber="195">Dikoe Swamp</collectingMunicipality>
,
<date id="8E2308B89E76FFF6FDB1F9C5FD17CE38" box="[583,692,1578,1605]" pageId="5" pageNumber="195" value="1999-06-05">
<collectingDate id="9E67F1509E76FFF6FDB1F9C5FD17CE38" box="[583,692,1578,1605]" pageId="5" pageNumber="195" value="1999-06-05">5.VI.1999</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FD49F9C4FC8BCE38" box="[703,808,1579,1605]" pageId="5" pageNumber="195">R. Minets</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FC97F9C4FC0BCE38" box="[865,936,1579,1605]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FC40F9C4FC55CE38" box="[950,1014,1579,1605]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="FA222E789E76FFF6FF31F9A1FBA3CF3C" blockId="5.[151,1437,1218,2039]" pageId="5" pageNumber="195">
<emphasis id="C8E9F26A9E76FFF6FF31F9A1FE7BCE15" bold="true" box="[199,472,1614,1640]" pageId="5" pageNumber="195">Comparative material.</emphasis>
<taxonomicName id="3D9D55FB9E76FFF6FE12F9A0FC4BCE14" authority="(Paykull, 1798)" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[484,1000,1615,1641]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="sulcicollis" subGenus="Agostenus">
<emphasis id="C8E9F26A9E76FFF6FE12F9A0FDEACE14" box="[484,585,1615,1641]" italics="true" pageId="5" pageNumber="195">Chlaenius</emphasis>
(
<emphasis id="C8E9F26A9E76FFF6FDABF9A0FD67CE14" box="[605,708,1615,1641]" italics="true" pageId="5" pageNumber="195">Agostenus</emphasis>
)
<emphasis id="C8E9F26A9E76FFF6FD21F9A0FC9ACE14" box="[727,825,1615,1641]" italics="true" pageId="5" pageNumber="195">sulcicollis</emphasis>
(Paykull, 1798)
</taxonomicName>
:
<materialsCitation id="4AF524259E76FFF6FC00F9A0FD2DCEF0" ID-GBIF-Occurrence="4000632396" collectingDate="1999-06-05" collectionCode="MPU" collectorName="R. Minets" country="Belarus" location="Dikoe Swamp" municipality="Belovezhskaya Pushcha" pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1">
<emphasis id="C8E9F26A9E76FFF6FC00F9A0FBE0CE14" box="[1014,1091,1615,1641]" italics="true" pageId="5" pageNumber="195">
<collectingCountry id="828A6EE89E76FFF6FC00F9A0FBE0CE14" box="[1014,1091,1615,1641]" name="Belarus" pageId="5" pageNumber="195">Belarus</collectingCountry>
</emphasis>
:
<collectingMunicipality id="1A46B4029E76FFF6FBA0F9A0FAECCE14" box="[1110,1359,1615,1641]" pageId="5" pageNumber="195">Belovezhskaya Pushcha</collectingMunicipality>
,
<location id="FF4278A39E76FFF6FAABF9A0FF45CEF0" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FAABF9A0FF45CEF0" country="Belarus" municipality="Belovezhskaya Pushcha" name="Dikoe Swamp" pageId="5" pageNumber="195">Dikoe Swamp</location>
,
<date id="8E2308B89E76FFF6FF06F99DFEF5CEF0" box="[240,342,1650,1677]" pageId="5" pageNumber="195" value="1999-06-05">
<collectingDate id="9E67F1509E76FFF6FF06F99DFEF5CEF0" box="[240,342,1650,1677]" pageId="5" pageNumber="195" value="1999-06-05">5.VI.1999</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FE97F99CFE60CEF0" box="[353,451,1651,1677]" pageId="5" pageNumber="195">R. Minets</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FE0CF99CFD9DCEF0" box="[506,574,1651,1677]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FDBDF99CFD24CEF0" box="[587,647,1651,1677]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FD6FF99CFA3ACEF0" ID-GBIF-Occurrence="4000632370" box="[665,1433,1651,1677]" collectingDate="1904-03-13" collectionCode="ZISP" collectorName="Olsufiev" country="Belarus" location="Vladimirskaya" municipality="Markovichi" pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1">
<collectingMunicipality id="1A46B4029E76FFF6FD6FF99CFCB3CEF0" box="[665,784,1651,1677]" pageId="5" pageNumber="195">Markovichi</collectingMunicipality>
,
<location id="FF4278A39E76FFF6FCEFF99CFC0FCEF0" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FCEFF99CFC0FCEF0" box="[793,940,1651,1677]" country="Belarus" municipality="Markovichi" name="Vladimirskaya" pageId="5" pageNumber="195">Vladimirskaya</location>
volost,
<date id="8E2308B89E76FFF6FBF4F99CFBD1CEF0" box="[1026,1138,1651,1677]" pageId="5" pageNumber="195" value="1904-03-13">
<collectingDate id="9E67F1509E76FFF6FBF4F99CFBD1CEF0" box="[1026,1138,1651,1677]" pageId="5" pageNumber="195" value="1904-03-13">13.III.1904</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FB8AF99CFB72CEF0" box="[1148,1233,1651,1677]" pageId="5" pageNumber="195">Olsufiev</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FAF1F99CFAE8CEF0" box="[1287,1355,1651,1677]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FAAFF99CFA31CEF0" box="[1369,1426,1651,1677]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="195">ZISP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4AF524259E76FFF6FF61F978FBC4CECC" ID-GBIF-Occurrence="4000632365" box="[151,1127,1687,1713]" collectingDate="1983-06-07" collectionCode="ZISP" collectorName="V. Starikov" country="Russia" location="Guberl'" pageId="5" pageNumber="195" specimenCount="1" specimenCount-female="1" stateProvince="Orenburg">
<emphasis id="C8E9F26A9E76FFF6FF61F978FF7ACECD" box="[151,217,1687,1712]" italics="true" pageId="5" pageNumber="195">
<collectingCountry id="828A6EE89E76FFF6FF61F978FF7ACECD" box="[151,217,1687,1712]" name="Russia" pageId="5" pageNumber="195">Russia</collectingCountry>
</emphasis>
:
<collectingRegion id="3859E09A9E76FFF6FF19F978FE34CECC" box="[239,407,1687,1713]" country="Russia" name="Orenburg" pageId="5" pageNumber="195">Orenburg region</collectingRegion>
, near
<location id="FF4278A39E76FFF6FE23F978FD87CECC" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FE23F978FD87CECC" box="[469,548,1687,1713]" country="Russia" name="Guberl'" pageId="5" pageNumber="195" stateProvince="Orenburg">Guberl</location>
, lake, reeds,
<date id="8E2308B89E76FFF6FD51F978FCADCECC" box="[679,782,1687,1713]" pageId="5" pageNumber="195" value="1983-06-07">
<collectingDate id="9E67F1509E76FFF6FD51F978FCADCECC" box="[679,782,1687,1713]" pageId="5" pageNumber="195" value="1983-06-07">7.VI.1983</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FCEFF978FC2ACECC" box="[793,905,1687,1713]" pageId="5" pageNumber="195">V. Starikov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FC34F978FBB9CECC" box="[962,1050,1687,1713]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FBDEF978FBC3CECC" box="[1064,1120,1687,1713]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="195">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FB85F978FCBFCEA8" ID-GBIF-Occurrence="4000632359" collectingDate="1972-07-30" collectionCode="ZISP" collectorName="Lafer" country="Russia" county="South" location="Khasan" municipality="Territory" pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy">
<collectingCounty id="134356F49E76FFF6FB85F978FB0ECECC" box="[1139,1197,1687,1713]" pageId="5" pageNumber="195">South</collectingCounty>
of
<collectingRegion id="3859E09A9E76FFF6FB27F978FA9ACECC" box="[1233,1337,1687,1713]" country="Russia" name="Primorskiy" pageId="5" pageNumber="195">Primorsky</collectingRegion>
<collectingMunicipality id="1A46B4029E76FFF6FAB6F978FA3BCECC" box="[1344,1432,1687,1713]" pageId="5" pageNumber="195">Territory</collectingMunicipality>
,
<location id="FF4278A39E76FFF6FF61F954FF47CEA8" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FF61F954FF47CEA8" box="[151,228,1723,1749]" country="Russia" county="South" municipality="Territory" name="Khasan" pageId="5" pageNumber="195" stateProvince="Primorskiy">Khasan</location>
,
<location id="FF4278A39E76FFF6FF06F954FE24CEA8" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FF06F954FE24CEA8" box="[240,391,1723,1749]" country="Russia" county="South" municipality="Territory" name="Golubiny Utes" pageId="5" pageNumber="195" stateProvince="Primorskiy">Golubiny Utes</location>
,
<date id="8E2308B89E76FFF6FE65F954FDADCEA8" box="[403,526,1723,1749]" pageId="5" pageNumber="195" value="1972-07-30">
<collectingDate id="9E67F1509E76FFF6FE65F954FDADCEA8" box="[403,526,1723,1749]" pageId="5" pageNumber="195" value="1972-07-30">30.VII.1972</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FDECF954FDF3CEA8" box="[538,592,1723,1749]" pageId="5" pageNumber="195">Lafer</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FD7FF954FD6DCEA8" box="[649,718,1723,1749]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FD2BF954FCB6CEA8" box="[733,789,1723,1749]" country="Russia" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="195">ZISP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FCDFF954FCBBCE84" ID-GBIF-Occurrence="4000632357" collectingDate="2001-08-12" collectionCode="FEB" collectorName="V. Kuznetsov" country="Russia" county="South" location="Chernigovka" municipality="Territory" pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy">
<collectingCounty id="134356F49E76FFF6FCDFF954FCC0CEA8" box="[809,867,1723,1749]" pageId="5" pageNumber="195">South</collectingCounty>
of
<collectingRegion id="3859E09A9E76FFF6FC7EF954FC53CEA8" box="[904,1008,1723,1749]" country="Russia" name="Primorskiy" pageId="5" pageNumber="195">Primorsky</collectingRegion>
<collectingMunicipality id="1A46B4029E76FFF6FC01F954FBEDCEA8" box="[1015,1102,1723,1749]" pageId="5" pageNumber="195">Territory</collectingMunicipality>
, hunting base
<quantity id="3D65839D9E76FFF6FB14F954FA87CEA8" box="[1250,1316,1723,1749]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="195" unit="km" value="10.0">10 km</quantity>
to the west of
<location id="FF4278A39E76FFF6FF45F930FE95CE84" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FF45F930FE95CE84" box="[179,310,1759,1785]" country="Russia" county="South" municipality="Territory" name="Chernigovka" pageId="5" pageNumber="195" stateProvince="Primorskiy">Chernigovka</location>
,
<date id="8E2308B89E76FFF6FEB6F930FE60CE84" box="[320,451,1759,1785]" pageId="5" pageNumber="195" value="2001-08-12">
<collectingDate id="9E67F1509E76FFF6FEB6F930FE60CE84" box="[320,451,1759,1785]" pageId="5" pageNumber="195" value="2001-08-12">12.VIII.2001</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FE38F930FDF7CE84" box="[462,596,1759,1785]" pageId="5" pageNumber="195">V. Kuznetsov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FD7AF930FD73CE84" box="[652,720,1759,1785]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FD28F930FCB2CE84" box="[734,785,1759,1785]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FCD5F930FC86CF60" ID-GBIF-Occurrence="4000632321" collectingDate="2004-07-26" collectionCode="FEB" collectorName="Yu. Sundukov &amp; L. Sundukova" country="Russia" county="South" location="Yakovlevka" municipality="Territory" pageId="5" pageNumber="195" specimenCount="1" specimenCount-female="1" stateProvince="Primorskiy">
<collectingCounty id="134356F49E76FFF6FCD5F930FCFECE84" box="[803,861,1759,1785]" pageId="5" pageNumber="195">South</collectingCounty>
of
<collectingRegion id="3859E09A9E76FFF6FC76F930FC4BCE84" box="[896,1000,1759,1785]" country="Russia" name="Primorskiy" pageId="5" pageNumber="195">Primorsky</collectingRegion>
<collectingMunicipality id="1A46B4029E76FFF6FC18F930FBE6CE84" box="[1006,1093,1759,1785]" pageId="5" pageNumber="195">Territory</collectingMunicipality>
,
<location id="FF4278A39E76FFF6FBB8F930FB64CE84" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FBB8F930FB64CE84" box="[1102,1223,1759,1785]" country="Russia" county="South" municipality="Territory" name="Yakovlevka" pageId="5" pageNumber="195" stateProvince="Primorskiy">Yakovlevka</location>
, sedge-moss swamp,
<date id="8E2308B89E76FFF6FF61F8ECFEB1CF60" box="[151,274,1795,1821]" pageId="5" pageNumber="195" value="2004-07-26">
<collectingDate id="9E67F1509E76FFF6FF61F8ECFEB1CF60" box="[151,274,1795,1821]" pageId="5" pageNumber="195" value="2004-07-26">26.VII.2004</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FEEDF8ECFE0BCF60" box="[283,424,1795,1821]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
&amp;
<collectorName id="57684BAE9E76FFF6FE3EF8ECFDF7CF60" box="[456,596,1795,1821]" pageId="5" pageNumber="195">L. Sundukova</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FD7EF8ECFD7CCF60" box="[648,735,1795,1821]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FD1DF8ECFCBDCF60" box="[747,798,1795,1821]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FCD9F8ECFC5ECF3C" ID-GBIF-Occurrence="4000632395" collectingDate="2022-05-24" collectionCode="FEB" collectorName="Yu. Sundukov" country="Russia" county="South" location="Khasansky distr." municipality="Territory" pageId="5" pageNumber="195" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Primorskiy">
<collectingCounty id="134356F49E76FFF6FCD9F8ECFCCACF60" box="[815,873,1795,1821]" pageId="5" pageNumber="195">South</collectingCounty>
of
<collectingRegion id="3859E09A9E76FFF6FC7FF8ECFC52CF60" box="[905,1009,1795,1821]" country="Russia" name="Primorskiy" pageId="5" pageNumber="195">Primorsky</collectingRegion>
<collectingMunicipality id="1A46B4029E76FFF6FC00F8ECFBEECF60" box="[1014,1101,1795,1821]" pageId="5" pageNumber="195">Territory</collectingMunicipality>
,
<location id="FF4278A39E76FFF6FBA0F8ECFB58CF60" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FBA0F8ECFB58CF60" box="[1110,1275,1795,1821]" country="Russia" county="South" municipality="Territory" name="Khasansky distr." pageId="5" pageNumber="195" stateProvince="Primorskiy">Khasansky distr.</location>
,
<location id="FF4278A39E76FFF6FAF5F8ECFA3BCF60" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FAF5F8ECFA3BCF60" box="[1283,1432,1795,1821]" country="Russia" county="South" municipality="Territory" name="Golubiny Utes" pageId="5" pageNumber="195" stateProvince="Primorskiy">Golubiny Utes</location>
,
<quantity id="3D65839D9E76FFF6FF61F8C9FF55CF3C" box="[151,246,1830,1857]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="5" pageNumber="195" unit="km" value="2.25" valueMax="2.5" valueMin="2.0">22.5 km</quantity>
on the way to
<location id="FF4278A39E76FFF6FE70F8C8FE70CF3C" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FE70F8C8FE70CF3C" box="[390,467,1831,1857]" country="Russia" county="South" municipality="Territory" name="Khasan" pageId="5" pageNumber="195" stateProvince="Primorskiy">Khasan</location>
,
<date id="8E2308B89E76FFF6FE2BF8C8FDE6CF3C" box="[477,581,1831,1857]" pageId="5" pageNumber="195" value="2022-05-24">
<collectingDate id="9E67F1509E76FFF6FE2BF8C8FDE6CF3C" box="[477,581,1831,1857]" pageId="5" pageNumber="195" value="2022-05-24">24.V.2022</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FDBBF8C8FD78CF3C" box="[589,731,1831,1857]" pageId="5" pageNumber="195">Yu. Sundukov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FCE6F8C8FCF6CF3C" box="[784,853,1831,1857]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FCA8F8C8FC17CF3C" box="[862,948,1831,1857]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FC37F8C8FC56CF3C" box="[961,1013,1831,1857]" pageId="5" pageNumber="195">FEB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="FA222E789E76FFF5FF31F8A4FC71C93C" blockId="5.[151,1437,1218,2039]" lastBlockId="6.[151,1437,150,645]" lastPageId="6" lastPageNumber="196" pageId="5" pageNumber="195">
<taxonomicName id="3D9D55FB9E76FFF6FF31F8A4FD00CF18" authority="Gebler, 1830" authorityName="Gebler" authorityYear="1830" box="[199,675,1867,1893]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="alutaceus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E76FFF6FF31F8A4FE8FCF18" box="[199,300,1867,1893]" italics="true" pageId="5" pageNumber="195">Chlaenius</emphasis>
(
<emphasis id="C8E9F26A9E76FFF6FECBF8A4FDB0CF18" box="[317,531,1867,1893]" italics="true" pageId="5" pageNumber="195">Agostenus) alutaceus</emphasis>
<materialsCitation id="4AF524259E76FFF6FDEDF8A4FD00CF18" ID-GBIF-Occurrence="4000632368" box="[539,675,1867,1893]" collectingDate="1982-07-22" collectionCode="MPU" collectorName="A. Matalin" country="Moldova" county="Kagulsky Distr." location="Prut River" municipality="Roshu" pageId="5" pageNumber="195" specimenCount="5" specimenCount-female="2" specimenCount-male="3">Gebler, 1830</materialsCitation>
</taxonomicName>
:
<emphasis id="C8E9F26A9E76FFF6FD58F8A4FCABCF18" box="[686,776,1867,1893]" italics="true" pageId="5" pageNumber="195">
<collectingCountry id="828A6EE89E76FFF6FD58F8A4FCABCF18" box="[686,776,1867,1893]" name="Moldova" pageId="5" pageNumber="195">Moldova</collectingCountry>
</emphasis>
:
<collectingCounty id="134356F49E76FFF6FCE1F8A4FC1BCF18" box="[791,952,1867,1893]" pageId="5" pageNumber="195">Kagulsky Distr.</collectingCounty>
, near
<collectingMunicipality id="1A46B4029E76FFF6FC01F8A4FB9ACF18" box="[1015,1081,1867,1893]" pageId="5" pageNumber="195">Roshu</collectingMunicipality>
, floodplain of
<location id="FF4278A39E76FFF6FB27F8A4FA98CF18" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FB27F8A4FA98CF18" box="[1233,1339,1867,1893]" country="Moldova" county="Kagulsky Distr." municipality="Roshu" name="Prut River" pageId="5" pageNumber="195">Prut River</location>
, on light,
<date id="8E2308B89E76FFF6FF61F880FEB0CFF4" box="[151,275,1903,1929]" pageId="5" pageNumber="195" value="1982-07-22">
<collectingDate id="9E67F1509E76FFF6FF61F880FEB0CFF4" box="[151,275,1903,1929]" pageId="5" pageNumber="195" value="1982-07-22">22.VII.1982</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FEEDF880FE25CFF4" box="[283,390,1903,1929]" pageId="5" pageNumber="195">A. Matalin</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FE4DF880FDA9CFF4" box="[443,522,1903,1929]" pageId="5" pageNumber="195" type="male">3 males</specimenCount>
,
<specimenCount id="EC9BE5F19E76FFF6FDE2F880FDD0CFF4" box="[532,627,1903,1929]" pageId="5" pageNumber="195" type="female">2 females</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FD77F880FD1ECFF4" box="[641,701,1903,1929]" pageId="5" pageNumber="195">MPU</collectionCode>
);
<materialsCitation id="4AF524259E76FFF6FD38F880FB67CFF4" ID-GBIF-Occurrence="4000632332" box="[718,1220,1903,1929]" collectingDate="1981-07-07" collectionCode="MPU" collectorName="A. Matalin" country="Moldova" county="Kagulsky Distr." location="Prut River" municipality="Roshu" pageId="5" pageNumber="195" specimenCount="1" specimenCount-female="1">
ibid.,
<date id="8E2308B89E76FFF6FCF3F880FCD0CFF4" box="[773,883,1903,1929]" pageId="5" pageNumber="195" value="1981-07-07">
<collectingDate id="9E67F1509E76FFF6FCF3F880FCD0CFF4" box="[773,883,1903,1929]" pageId="5" pageNumber="195" value="1981-07-07">7.VII.1981</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FC8AF880FC44CFF4" box="[892,999,1903,1929]" pageId="5" pageNumber="195">A. Matalin</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FBEBF880FBD0CFF4" box="[1053,1139,1903,1929]" pageId="5" pageNumber="195" type="female">1 female</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FB76F880FB1FCFF4" box="[1152,1212,1903,1929]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4AF524259E76FFF6FB3BF880FB98CFD0" ID-GBIF-Occurrence="4000632303" collectingDate="1987-07-24" collectionCode="MPU" collectorName="V. Grachev" country="Russia" location="Novomikhaylovka" municipality="Kochenevskiy Distr." pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Novosibirsk">
<emphasis id="C8E9F26A9E76FFF6FB3BF880FAACCFF5" box="[1229,1295,1903,1928]" italics="true" pageId="5" pageNumber="195">
<collectingCountry id="828A6EE89E76FFF6FB3BF880FAACCFF5" box="[1229,1295,1903,1928]" name="Russia" pageId="5" pageNumber="195">Russia</collectingCountry>
</emphasis>
:
<collectingRegion id="3859E09A9E76FFF6FAD7F880FF42CFD0" country="Russia" name="Novosibirsk" pageId="5" pageNumber="195">Novosibirsk Region</collectingRegion>
,
<collectingMunicipality id="1A46B4029E76FFF6FF1CF87CFE14CFD0" box="[234,439,1939,1965]" pageId="5" pageNumber="195">Kochenevskiy Distr.</collectingMunicipality>
,
<location id="FF4278A39E76FFF6FE48F87CFDD8CFD0" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FE48F87CFDD8CFD0" box="[446,635,1939,1965]" country="Russia" municipality="Kochenevskiy Distr." name="Novomikhaylovka" pageId="5" pageNumber="195" stateProvince="Novosibirsk">Novomikhaylovka</location>
,
<date id="8E2308B89E76FFF6FD72F87CFD5CCFD0" box="[644,767,1939,1965]" pageId="5" pageNumber="195" value="1987-07-24">
<collectingDate id="9E67F1509E76FFF6FD72F87CFD5CCFD0" box="[644,767,1939,1965]" pageId="5" pageNumber="195" value="1987-07-24">24.VII.1987</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FCFEF87CFCD5CFD0" box="[776,886,1939,1965]" pageId="5" pageNumber="195">V. Grachev</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FC5FF87CFC4FCFD0" box="[937,1004,1939,1965]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FC0EF87CFB97CFD0" box="[1016,1076,1939,1965]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FBB2F87CFBDDCFAC" ID-GBIF-Occurrence="4000632386" collectingDate="1999-07-01" collectingDateMax="1999-07-07" collectingDateMin="1999-07-01" collectionCode="MPU" collectorName="I. Melnik" country="Russia" latitude="50.64167" location="Lake Beloe" longLatPrecision="18" longitude="105.73833" municipality="Dzhidinsky Distr." pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Buryatiya">
<collectingRegion id="3859E09A9E76FFF6FBB2F87CFB0BCFD0" box="[1092,1192,1939,1965]" country="Russia" name="Buryatiya" pageId="5" pageNumber="195">Buryatiya</collectingRegion>
,
<collectingMunicipality id="1A46B4029E76FFF6FB47F87CFAC2CFD0" box="[1201,1377,1939,1965]" pageId="5" pageNumber="195">Dzhidinsky Distr.</collectingMunicipality>
,
<locationDeviation id="EB90375A9E76FFF6FA9EF87CFE93CFAC" location="Belogorsk" pageId="5" pageNumber="195">
<quantity id="3D65839D9E76FFF6FA9EF87CFA3FCFD0" box="[1384,1436,1939,1965]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="195" unit="km" value="2.0">2 km</quantity>
E of Belogorsk
</locationDeviation>
,
<location id="FF4278A39E76FFF6FECDF858FE13CFAC" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FECDF858FE13CFAC" box="[315,432,1975,2001]" country="Russia" latitude="50.64167" longLatPrecision="18" longitude="105.73833" municipality="Dzhidinsky Distr." name="Lake Beloe" pageId="5" pageNumber="195" stateProvince="Buryatiya">Lake Beloe</location>
,
<geoCoordinate id="9FA948BF9E76FFF6FE4CF857FD92CFAE" box="[442,561,1976,2003]" degrees="50" direction="north" minutes="38" orientation="latitude" pageId="5" pageNumber="195" precision="15" seconds="30" value="50.64167">50°3830″N</geoCoordinate>
<geoCoordinate id="9FA948BF9E76FFF6FDC1F857FD19CFAE" box="[567,698,1976,2003]" degrees="105" direction="east" minutes="44" orientation="longitude" pageId="5" pageNumber="195" precision="15" seconds="18" value="105.73833">105°4418″E</geoCoordinate>
,
<date id="8E2308B89E76FFF6FD32F858FCE8CFAC" box="[708,843,1975,2001]" pageId="5" pageNumber="195" value="1999-07-01" valueMax="1999-07-07" valueMin="1999-07-01">
<collectingDate id="9E67F1509E76FFF6FD32F858FCE8CFAC" box="[708,843,1975,2001]" pageId="5" pageNumber="195" value="1999-07-01" valueMax="1999-07-07" valueMin="1999-07-01">17.VII.1999</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FCA0F858FC11CFAC" box="[854,946,1975,2001]" pageId="5" pageNumber="195">I. Melnik</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FC1FF858FB8ECFAC" box="[1001,1069,1975,2001]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FBCCF858FBD5CFAC" box="[1082,1142,1975,2001]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF6FB7EF858FBB1CF88" ID-GBIF-Occurrence="4000632312" collectingDate="2000-06-16" collectingDateMax="2000-06-18" collectingDateMin="2000-06-16" collectionCode="MPU" collectorName="I. Melnik" country="Russia" latitude="50.008335" location="Duroy" longLatPrecision="18" longitude="118.92694" pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Chita Region">
<collectingRegion id="3859E09A9E76FFF6FB7EF858FAAFCFAC" box="[1160,1292,1975,2001]" inferredBySuffix="true" pageId="5" pageNumber="195">Chita Region</collectingRegion>
[Zabaykalsky Krai], near
<location id="FF4278A39E76FFF6FEF3F834FEE5CF88" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FEF3F834FEE5CF88" box="[261,326,2011,2037]" country="Russia" latitude="50.008335" longLatPrecision="18" longitude="118.92694" name="Duroy" pageId="5" pageNumber="195" stateProvince="Chita Region">Duroy</location>
,
<geoCoordinate id="9FA948BF9E76FFF6FEBBF833FE66CF8A" box="[333,453,2012,2039]" degrees="50" direction="north" minutes="00" orientation="latitude" pageId="5" pageNumber="195" precision="15" seconds="30" value="50.008335">50°0030″N</geoCoordinate>
<geoCoordinate id="9FA948BF9E76FFF6FE3FF833FDE9CF8A" box="[457,586,2012,2039]" degrees="118" direction="east" minutes="55" orientation="longitude" pageId="5" pageNumber="195" precision="15" seconds="37" value="118.92694">118°5537″E</geoCoordinate>
,
<date id="8E2308B89E76FFF6FDA5F834FD4FCF88" box="[595,748,2011,2037]" pageId="5" pageNumber="195" value="2000-06-16" valueMax="2000-06-18" valueMin="2000-06-16">
<collectingDate id="9E67F1509E76FFF6FDA5F834FD4FCF88" box="[595,748,2011,2037]" pageId="5" pageNumber="195" value="2000-06-16" valueMax="2000-06-18" valueMin="2000-06-16">1618.VI.2000</collectingDate>
</date>
,
<collectorName id="57684BAE9E76FFF6FD02F834FCEDCF88" box="[756,846,2011,2037]" pageId="5" pageNumber="195">I. Melnik</collectorName>
leg.,
<specimenCount id="EC9BE5F19E76FFF6FC77F834FC60CF88" box="[897,963,2011,2037]" pageId="5" pageNumber="195" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E76FFF6FC39F834FBA8CF88" box="[975,1035,2011,2037]" pageId="5" pageNumber="195">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E76FFF5FBEDF834FD67C8CC" ID-GBIF-Occurrence="4000632318" collectingDate="1993-07-14" collectingDateMax="1993-08-02" collectingDateMin="1993-07-14" collectionCode="MPU" collectorName="P. Udovichenko" country="Russia" lastPageId="6" lastPageNumber="196" location="Bira" municipality="Autonomous Region" pageId="5" pageNumber="195" specimenCount="1" specimenCount-male="1" stateProvince="Yevrey">
<collectingRegion id="3859E09A9E76FFF6FBEDF834FBFCCF88" box="[1051,1119,2011,2037]" country="Russia" name="Yevrey" pageId="5" pageNumber="195">Jewish</collectingRegion>
<collectingRegion id="3859E09A9E76FFF6FB94F834FA90CF88" box="[1122,1331,2011,2037]" inferredBySuffix="true" pageId="5" pageNumber="195">
<collectingMunicipality id="1A46B4029E76FFF6FB94F834FA90CF88" box="[1122,1331,2011,2037]" pageId="5" pageNumber="195">Autonomous Region</collectingMunicipality>
</collectingRegion>
, near
<location id="FF4278A39E76FFF6FA9DF834FA3ACF88" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E76FFF6FA9DF834FA3ACF88" box="[1387,1433,2011,2037]" country="Russia" municipality="Autonomous Region" name="Bira" pageId="5" pageNumber="195" stateProvince="Yevrey">Bira</location>
,
<date id="8E2308B89E75FFF5FF61FF78FEFFC8CC" box="[151,348,151,177]" pageId="6" pageNumber="196" value="1993-07-14" valueMax="1993-08-02" valueMin="1993-07-14">
<collectingDate id="9E67F1509E75FFF5FF61FF78FEFFC8CC" box="[151,348,151,177]" pageId="6" pageNumber="196" value="1993-07-14" valueMax="1993-08-02" valueMin="1993-07-14">14.VII2.VIII.1993</collectingDate>
</date>
,
<collectorName id="57684BAE9E75FFF5FE92FF78FDA3C8CC" box="[356,512,151,177]" pageId="6" pageNumber="196">P.Udovichenko</collectorName>
leg.,
<specimenCount id="EC9BE5F19E75FFF5FDC5FF78FDD6C8CC" box="[563,629,151,177]" pageId="6" pageNumber="196" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E75FFF5FD76FF78FD1FC8CC" box="[640,700,151,177]" pageId="6" pageNumber="196">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E75FFF5FD3AFF78FE51C8A8" ID-GBIF-Occurrence="4000632319" collectingDate="1998-07-15" collectionCode="MPU" collectorName="V. Patrikeev" country="Russia" county="South" location="Lake Khanka" municipality="Territory" pageId="6" pageNumber="196" specimenCount="3" specimenCount-male="3" stateProvince="Primorskiy">
<collectingCounty id="134356F49E75FFF5FD3AFF78FCA5C8CC" box="[716,774,151,177]" pageId="6" pageNumber="196">South</collectingCounty>
of
<collectingRegion id="3859E09A9E75FFF5FCD2FF78FC2FC8CC" box="[804,908,151,177]" country="Russia" name="Primorskiy" pageId="6" pageNumber="196">Primorsky</collectingRegion>
<collectingMunicipality id="1A46B4029E75FFF5FC66FF78FC44C8CC" box="[912,999,151,177]" pageId="6" pageNumber="196">Territory</collectingMunicipality>
, near
<location id="FF4278A39E75FFF5FBE8FF78FB06C8CC" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E75FFF5FBE8FF78FB06C8CC" box="[1054,1189,151,177]" country="Russia" county="South" municipality="Territory" name="Lake Khanka" pageId="6" pageNumber="196" stateProvince="Primorskiy">Lake Khanka</location>
,
<location id="FF4278A39E75FFF5FB5BFF78FAB0C8CC" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E75FFF5FB5BFF78FAB0C8CC" box="[1197,1299,151,177]" country="Russia" county="South" municipality="Territory" name="Turiy Rog" pageId="6" pageNumber="196" stateProvince="Primorskiy">Turiy Rog</location>
,
<date id="8E2308B89E75FFF5FAEAFF79FA3BC8CC" box="[1308,1432,150,177]" pageId="6" pageNumber="196" value="1998-07-15">
<collectingDate id="9E67F1509E75FFF5FAEAFF79FA3BC8CC" box="[1308,1432,150,177]" pageId="6" pageNumber="196" value="1998-07-15">15.VII.1998</collectingDate>
</date>
,
<collectorName id="57684BAE9E75FFF5FF61FF54FEB1C8A8" box="[151,274,187,213]" pageId="6" pageNumber="196">V. Patrikeev</collectorName>
leg.,
<specimenCount id="EC9BE5F19E75FFF5FEB8FF54FE3DC8A8" box="[334,414,187,213]" pageId="6" pageNumber="196" type="male">3 males</specimenCount>
(
<collectionCode id="9C8CB6BD9E75FFF5FE59FF54FE48C8A8" box="[431,491,187,213]" pageId="6" pageNumber="196">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E75FFF5FE09FF54FEC1C884" ID-GBIF-Occurrence="4000632381" collectingDate="1991-07-20" collectionCode="MPU" collectorName="A. Sochivko" country="Russia" county="Territory" location="Shkotovo" municipality="southern Sikhote-Alin Mts" pageId="6" pageNumber="196" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy">
<collectingRegion id="3859E09A9E75FFF5FE09FF54FDC4C8A8" box="[511,615,187,213]" country="Russia" name="Primorskiy" pageId="6" pageNumber="196">Primorsky</collectingRegion>
<collectingCounty id="134356F49E75FFF5FD86FF54FD64C8A8" box="[624,711,187,213]" pageId="6" pageNumber="196">Territory</collectingCounty>
,
<collectingMunicipality id="1A46B4029E75FFF5FD22FF54FC47C8A8" box="[724,996,187,213]" pageId="6" pageNumber="196">southern Sikhote-Alin Mts</collectingMunicipality>
, near
<location id="FF4278A39E75FFF5FBD3FF54FB24C8A8" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E75FFF5FBD3FF54FB24C8A8" box="[1061,1159,187,213]" country="Russia" county="Territory" municipality="southern Sikhote-Alin Mts" name="Shkotovo" pageId="6" pageNumber="196" stateProvince="Primorskiy">Shkotovo</location>
,
<date id="8E2308B89E75FFF5FB62FF54FAACC8A8" box="[1172,1295,187,213]" pageId="6" pageNumber="196" value="1991-07-20">
<collectingDate id="9E67F1509E75FFF5FB62FF54FAACC8A8" box="[1172,1295,187,213]" pageId="6" pageNumber="196" value="1991-07-20">20.VII.1991</collectingDate>
</date>
,
<collectorName id="57684BAE9E75FFF5FAEDFF54FA3FC8A8" box="[1307,1436,187,213]" pageId="6" pageNumber="196">A. Sochivko</collectorName>
leg.,
<specimenCount id="EC9BE5F19E75FFF5FF3FFF30FEACC884" box="[201,271,223,249]" pageId="6" pageNumber="196" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E75FFF5FEE8FF30FEF9C884" box="[286,346,223,249]" pageId="6" pageNumber="196">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E75FFF5FE98FF30FE8EC960" ID-GBIF-Occurrence="4000632333" collectingDate="1998-08-12" collectingDateMax="1998-08-15" collectingDateMin="1998-08-12" collectionCode="MPU" collectorName="B. Zhuravlev" country="Russia" county="South" location="Pogranichnyi Distr." municipality="Territory" pageId="6" pageNumber="196" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy">
<collectingCounty id="134356F49E75FFF5FE98FF30FE0BC884" box="[366,424,223,249]" pageId="6" pageNumber="196">South</collectingCounty>
of
<collectingRegion id="3859E09A9E75FFF5FE38FF30FD95C884" box="[462,566,223,249]" country="Russia" name="Primorskiy" pageId="6" pageNumber="196">Primorsky</collectingRegion>
<collectingMunicipality id="1A46B4029E75FFF5FDC8FF30FD36C884" box="[574,661,223,249]" pageId="6" pageNumber="196">Territory</collectingMunicipality>
,
<location id="FF4278A39E75FFF5FD57FF30FCC5C884" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E75FFF5FD57FF30FCC5C884" box="[673,870,223,249]" country="Russia" county="South" municipality="Territory" name="Pogranichnyi Distr." pageId="6" pageNumber="196" stateProvince="Primorskiy">Pogranichnyi Distr.</location>
,
<location id="FF4278A39E75FFF5FC87FF30FB81C884" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E75FFF5FC87FF30FB81C884" box="[881,1058,223,249]" country="Russia" county="South" municipality="Territory" name="Barabash-Levada" pageId="6" pageNumber="196" stateProvince="Primorskiy">Barabash-Levada</location>
,
<date id="8E2308B89E75FFF5FBD8FF31FB75C884" box="[1070,1238,222,249]" pageId="6" pageNumber="196" value="1998-08-12" valueMax="1998-08-15" valueMin="1998-08-12">
<collectingDate id="9E67F1509E75FFF5FBD8FF31FB75C884" box="[1070,1238,222,249]" pageId="6" pageNumber="196" value="1998-08-12" valueMax="1998-08-15" valueMin="1998-08-12">1215.VIII.1998</collectingDate>
</date>
,
<collectorName id="57684BAE9E75FFF5FB15FF30FACAC884" box="[1251,1385,223,249]" pageId="6" pageNumber="196">B. Zhuravlev</collectorName>
leg.,
<specimenCount id="EC9BE5F19E75FFF5FF61FEECFF7FC960" box="[151,220,259,285]" pageId="6" pageNumber="196" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E75FFF5FF1CFEECFE85C960" box="[234,294,259,285]" pageId="6" pageNumber="196">MPU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AF524259E75FFF5FECEFEECFAB9C960" ID-GBIF-Occurrence="4000632325" box="[312,1306,258,285]" collectingDate="1992-07-05" collectingDateMax="1992-07-25" collectingDateMin="1992-07-05" collectionCode="MPU" collectorName="A. Generalov" country="Russia" county="South" location="Pogranichnyi" municipality="Territory" pageId="6" pageNumber="196" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy">
<collectingCounty id="134356F49E75FFF5FECEFEECFED1C960" box="[312,370,259,285]" pageId="6" pageNumber="196">South</collectingCounty>
of
<collectingRegion id="3859E09A9E75FFF5FE63FEECFE5EC960" box="[405,509,259,285]" country="Russia" name="Primorskiy" pageId="6" pageNumber="196">Primorsky</collectingRegion>
<collectingMunicipality id="1A46B4029E75FFF5FDF2FEECFDF8C960" box="[516,603,259,285]" pageId="6" pageNumber="196">Territory</collectingMunicipality>
, near
<location id="FF4278A39E75FFF5FD61FEECFCBDC960" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E75FFF5FD61FEECFCBDC960" box="[663,798,259,285]" country="Russia" county="South" municipality="Territory" name="Pogranichnyi" pageId="6" pageNumber="196" stateProvince="Primorskiy">Pogranichnyi</location>
,
<date id="8E2308B89E75FFF5FCDEFEEDFC1FC960" box="[808,956,258,285]" pageId="6" pageNumber="196" value="1992-07-05" valueMax="1992-07-25" valueMin="1992-07-05">
<collectingDate id="9E67F1509E75FFF5FCDEFEEDFC1FC960" box="[808,956,258,285]" pageId="6" pageNumber="196" value="1992-07-05" valueMax="1992-07-25" valueMin="1992-07-05">525.VII.1992</collectingDate>
</date>
,
<collectorName id="57684BAE9E75FFF5FC30FEECFBEEC960" box="[966,1101,259,285]" pageId="6" pageNumber="196">A. Generalov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E75FFF5FB72FEECFB6AC960" box="[1156,1225,259,285]" pageId="6" pageNumber="196" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E75FFF5FB21FEECFAB0C960" box="[1239,1299,259,285]" pageId="6" pageNumber="196">MPU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4AF524259E75FFF5FAD3FEECFC6DC93C" ID-GBIF-Occurrence="4000632369" collectingDate="1984-05-29" collectionCode="MPU" collectorName="A. Zamesov" country="Uzbekistan" location="Takhtakupyr" municipality="Karakalpakstan" pageId="6" pageNumber="196" specimenCount="1" specimenCount-male="1">
<collectingCountry id="828A6EE89E75FFF5FAD3FEECFA39C960" box="[1317,1434,259,285]" name="Uzbekistan" pageId="6" pageNumber="196">Uzbekistan</collectingCountry>
:
<collectingMunicipality id="1A46B4029E75FFF5FF61FEC8FE97C93C" box="[151,308,295,321]" pageId="6" pageNumber="196">Karakalpakstan</collectingMunicipality>
,
<location id="FF4278A39E75FFF5FECAFEC8FE18C93C" LSID="urn:lsid:plazi:treatment:72349F6E9E76FFFBFF61FC25FCA7CBB4:FF4278A39E75FFF5FECAFEC8FE18C93C" box="[316,443,295,321]" country="Uzbekistan" municipality="Karakalpakstan" name="Takhtakupyr" pageId="6" pageNumber="196">Takhtakupyr</location>
, on light,
<date id="8E2308B89E75FFF5FDEDFEC8FD21C93C" box="[539,642,295,321]" pageId="6" pageNumber="196" value="1984-05-29">
<collectingDate id="9E67F1509E75FFF5FDEDFEC8FD21C93C" box="[539,642,295,321]" pageId="6" pageNumber="196" value="1984-05-29">29.V.1984</collectingDate>
</date>
,
<collectorName id="57684BAE9E75FFF5FD7CFEC8FCA6C93C" box="[650,773,295,321]" pageId="6" pageNumber="196">A. Zamesov</collectorName>
leg.,
<specimenCount id="EC9BE5F19E75FFF5FCCCFEC8FCDEC93C" box="[826,893,295,321]" pageId="6" pageNumber="196" type="male">1 male</specimenCount>
(
<collectionCode id="9C8CB6BD9E75FFF5FC7FFEC8FC64C93C" box="[905,967,295,321]" pageId="6" pageNumber="196">MPU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2877DF39E75FFF5FF31FEA5FB88C988" pageId="6" pageNumber="196" type="distribution">
<paragraph id="FA222E789E75FFF5FF31FEA5FB88C988" blockId="6.[151,1437,150,645]" pageId="6" pageNumber="196">
<emphasis id="C8E9F26A9E75FFF5FF31FEA5FEFEC919" bold="true" box="[199,349,330,356]" pageId="6" pageNumber="196">Distribution.</emphasis>
<collectingCountry id="828A6EE89E75FFF5FE98FEA4FE18C918" box="[366,443,331,357]" name="Russia" pageId="6" pageNumber="196">Russia</collectingCountry>
: Southern Kuriles: Kunashir (new record),
<collectingRegion id="3859E09A9E75FFF5FC38FEA4FB93C918" box="[974,1072,331,357]" country="Russia" name="Sakhalin" pageId="6" pageNumber="196">Sakhalin</collectingRegion>
,
<collectingRegion id="3859E09A9E75FFF5FBB3FEA4FB14C918" box="[1093,1207,331,357]" country="Russia" name="Primorskiy" pageId="6" pageNumber="196">Primorsky</collectingRegion>
Territory, southern
<collectingRegion id="3859E09A9E75FFF5FF61FE80FEB8C9F4" box="[151,283,367,393]" country="Russia" name="Khabarovsk" pageId="6" pageNumber="196">Khabarovsk</collectingRegion>
Territory,
<collectingRegion id="3859E09A9E75FFF5FE63FE80FE43C9F4" box="[405,480,367,393]" country="Russia" name="Yevrey" pageId="6" pageNumber="196">Jewish</collectingRegion>
Autonomous Region,
<collectingRegion id="3859E09A9E75FFF5FD28FE80FCD8C9F4" box="[734,891,367,393]" country="Russia" name="Amur" pageId="6" pageNumber="196">Amur Region</collectingRegion>
; southern Yakutia, forest zone of Siberia, Urals, north and center of the European part.—
<collectingCountry id="828A6EE89E75FFF5FDBBFE7CFD2DC9D0" box="[589,654,403,429]" name="Japan" pageId="6" pageNumber="196">Japan</collectingCountry>
:
<collectingRegion id="3859E09A9E75FFF5FD61FE7CFCA5C9D0" box="[663,774,403,429]" country="Japan" name="Hokkaido" pageId="6" pageNumber="196">Hokkaido</collectingRegion>
, northern Honshu,
<collectingCountry id="828A6EE89E75FFF5FC22FE7CFBF4C9D0" box="[980,1111,403,429]" name="Kazakhstan" pageId="6" pageNumber="196">Kazakhstan</collectingCountry>
, Central and Northern Europe (
<bibRefCitation id="9E0C53899E75FFF5FF68FE58FEBDC9AC" author="Lafer, G. Sh." box="[158,286,439,465]" pageId="6" pageNumber="196" pagination="71 - 222" refId="ref8582" refString="Lafer, G. Sh. (1989) 4. Semeystvo Carabidae - Zhuzhelitsy. In: Lehr, P. A. (Ed.), Opredelitel nasekomykh Dalnego Vostoka SSSR. Tom 3. Zhestkokrylye, ili zhuki. Part 1. Nauka, Leningrad, pp. 71 - 222. [in Russian]" type="book chapter" year="1989">Lafer 1989</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FEDDFE58FD94C9AC" author="Kimoto, S. &amp; Yasuda, N." box="[299,567,438,465]" pageId="6" pageNumber="196" refId="ref8048" refString="Kimoto, S. &amp; Yasuda, N. (1995) The ground beetles of Hokkaido. Ecological and biological survey. Tokai University Press, Tokyo, 315 pp. [in Japanese]" type="book" year="1995">Kimoto &amp; Yasuda 1995</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FDB5FE58FD58C9AC" author="Averensky, A. I." box="[579,763,439,465]" pageId="6" pageNumber="196" refId="ref7538" refString="Averensky, A. I. (1999) The catalog of beetles of Yakutia. Yakutsk Scientific Center SD RAS, Yakutsk, 78 pp. [in Russian]" type="book" year="1999">Averensky 1999</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FCF1FE58FC55C9AC" author="Yoshitake, H. &amp; Kurihara, T. &amp; Yoshimatsu, S. - I. &amp; Nakatani, Y. &amp; Yasuda, K." box="[775,1014,439,465]" pageId="6" pageNumber="196" pagination="1 - 327" refId="ref9821" refString="Yoshitake, H., Kurihara, T., Yoshimatsu, S. - I., Nakatani, Y. &amp; Yasuda, K. (2011) A list of carabid specimens (Insecta: Coleoptera) collected by the late Dr. Akinobu Habu preserved in the Insect Museum of the National Institute for Agro-Environmental Sciences. Bulletin of National Institute for Agro-Environmental Sciences, 28, 1 - 327." type="journal article" year="2011">
Yoshitake
<emphasis id="C8E9F26A9E75FFF5FC8AFE58FC17C9AC" box="[892,948,439,465]" italics="true" pageId="6" pageNumber="196">et al.</emphasis>
2011
</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FBF4FE58FABCC9AC" author="Vertyankin, A. V. &amp; Lafer, G. Sh." box="[1026,1311,439,465]" pageId="6" pageNumber="196" pagination="433 - 436" refId="ref9508" refString="Vertyankin, A. V. &amp; Lafer, G. Sh. (2012) New records of ground beetles (Coleoptera, Carabidae) from Sakhalin and Moneron Islands of the Russian Far East. Euroasian Entomological Journal, 11 (5), 433 - 436. [in Russian]" type="journal article" year="2012">Vertyankin &amp; Lafer 2012</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FADBFE58FF72C988" author="Sundukov, Yu. N." pageId="6" pageNumber="196" refId="ref9272" refString="Sundukov, Yu. N. (2013) An annotated catalog of the ground beetles (Coleoptera: Caraboidea) of Sikhote-Alin. Dalnauka, Vladivostok, 271 pp. [in Russian]" type="book" year="2013">Sundukov 2013</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FF2DFE34FE23C988" author="Akopyan, E. K." box="[219,384,475,501]" pageId="6" pageNumber="196" pagination="1669 - 1674" refId="ref7428" refString="Akopyan, E. K. (2014) Ground beetles of the Volga River valley. Izvestiya Samarskogo nauchnogo tsentra RAS, 16 (5), 1669 - 1674. [in Russian] https: // doi. org / 10.0000 / cyberleninka. ru / arcitle / n / zhuzhelitsy-doliny-reki-volya" type="journal article" year="2014">Akopyan 2014</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FE7DFE34FD32C988" author="Khobrakova, L. T. &amp; Shilenkov, V. G. &amp; Dudko, R. Yu." box="[395,657,475,501]" pageId="6" pageNumber="196" refId="ref8326" refString="Khobrakova, L. T., Shilenkov, V. G. &amp; Dudko, R. Yu. (2014) The ground beetles (Coleoptera, Carabidae) of Buryatia. Buryat Scientific Center SB RAS Press, Ulan-Ude, 380 pp. [in Russian]" type="book" year="2014">
Khobrakova
<emphasis id="C8E9F26A9E75FFF5FDECFE34FDF3C988" box="[538,592,475,501]" italics="true" pageId="6" pageNumber="196">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FD6AFE34FC96C988" author="Budilov, P. V." box="[668,821,475,501]" pageId="6" pageNumber="196" pagination="136 - 145" refId="ref7569" refString="Budilov, P. V. (2017) Ground beetles (Coleoptera, Carabidae) in the small-leaved forest of a restricted area of the &quot; Bastak &quot; Nature Reserve, Jewish Autonomous Region. A. I. Kurentsov's Annual Memorial Meetings, 28, 136 - 145. [in Russian]" type="journal article" year="2017">Budilov 2017</bibRefCitation>
;
<bibRefCitation id="9E0C53899E75FFF5FCB6FE34FBBCC988" author="Kirschenhofer, E." box="[832,1055,475,501]" pageId="6" pageNumber="196" pagination="482 - 497" refId="ref8266" refString="Kirschenhofer, E. (2017) Tribe Chlaeniini Brulle, 1834. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalog of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 482 - 497." type="book chapter" year="2017">Kirschenhofer 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2877DF39E75FFFBFF31FE10FCA7CBB4" lastPageId="8" lastPageNumber="198" pageId="6" pageNumber="196" type="discussion">
<paragraph id="FA222E789E75FFF5FF31FE10FC65CA64" blockId="6.[151,1437,150,645]" box="[199,966,511,537]" pageId="6" pageNumber="196">
<emphasis id="C8E9F26A9E75FFF5FF31FE10FE9ACA64" bold="true" box="[199,313,511,537]" pageId="6" pageNumber="196">Remarks.</emphasis>
This species is firstly recorded from the Kuril Archipelago.
</paragraph>
<paragraph id="FA222E789E75FFF5FF31FDCCFC31CAF8" blockId="6.[151,1437,150,645]" pageId="6" pageNumber="196">
Finding
<taxonomicName id="3D9D55FB9E75FFF5FEDFFDCCFDA3CA40" authorityName="Paykull" authorityYear="1790" box="[297,512,547,573]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="species" species="quadrisulcatus">
<emphasis id="C8E9F26A9E75FFF5FEDFFDCCFDA3CA40" box="[297,512,547,573]" italics="true" pageId="6" pageNumber="196">Ch. quadrisulcatus</emphasis>
</taxonomicName>
in the extreme south of Kunashir, in the immediate vicinity of
<collectingRegion id="3859E09A9E75FFF5FB0EFDCCFAC4CA40" box="[1272,1383,547,573]" country="Japan" name="Hokkaido" pageId="6" pageNumber="196">Hokkaido</collectingRegion>
, has prompted the authors to clarify the confusing situation concerning the distribution of
<taxonomicName id="3D9D55FB9E75FFF5FBBDFDA8FB19CA1C" authorityName="Bonelli" authorityYear="1810" box="[1099,1210,583,609]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="C8E9F26A9E75FFF5FBBDFDA8FB19CA1C" box="[1099,1210,583,609]" italics="true" pageId="6" pageNumber="196">Chlaenius</emphasis>
</taxonomicName>
in the Far East, viz.
<emphasis id="C8E9F26A9E75FFF5FF61FD84FDB3CAF8" box="[151,528,619,645]" italics="true" pageId="6" pageNumber="196">
<taxonomicName id="3D9D55FB9E75FFF5FF61FD84FEC9CAF8" authorityName="Paykull" authorityYear="1790" box="[151,362,619,645]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="species" species="quadrisulcatus">Ch. quadrisulcatus</taxonomicName>
,
<taxonomicName id="3D9D55FB9E75FFF5FE83FD84FDB3CAF8" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[373,528,619,645]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="species" species="sulcicollis">Ch. sulcicollis</taxonomicName>
</emphasis>
and
<taxonomicName id="3D9D55FB9E75FFF5FDB0FD84FC2DCAF8" authority="Ganglbauer, 1891" authorityName="Ganglbauer" authorityYear="1891" box="[582,910,619,645]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E75FFF5FDB0FD84FD62CAF8" box="[582,705,619,645]" italics="true" pageId="6" pageNumber="196">Ch. gebleri</emphasis>
<bibRefCitation id="9E0C53899E75FFF5FD31FD84FC2DCAF8" author="Ganglbauer, L." box="[711,910,619,645]" pageId="6" pageNumber="196" refId="ref7652" refString="Ganglbauer, L. (1891) Die Kafer von Mitteleuropa. Die Kafer der Osterreichisch-Ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. Erster Band. Familienreihe Caraboidea. C. Gerold's Sohn, Wien, iii + 557 pp. https: // doi. org / 10.5962 / bhl. title. 8764" type="book" year="1891">Ganglbauer, 1891</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="AEE27EF09E75FFF5FF61F8B7FA3ECFA1" ID-DOI="http://doi.org/10.5281/zenodo.7456767" ID-Zenodo-Dep="7456767" httpUri="https://zenodo.org/record/7456767/files/figure.png" pageId="6" pageNumber="196" startId="6.[151,264,1880,1904]" targetBox="[305,1285,676,1862]" targetPageId="6">
<paragraph id="FA222E789E75FFF5FF61F8B7FA3ECFA1" blockId="6.[151,1437,1880,2013]" pageId="6" pageNumber="196">
<emphasis id="C8E9F26A9E75FFF5FF61F8B7FEF9CF0D" bold="true" box="[151,346,1880,1904]" pageId="6" pageNumber="196">FIGURES 1318.</emphasis>
Far Eastern species of the subgenus
<taxonomicName id="3D9D55FB9E75FFF5FD13F8B6FBDFCF0C" authority="Fischer von Waldheim, 1829" authorityName="Fischer von Waldheim" authorityYear="1829" box="[741,1148,1880,1905]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="subGenus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E75FFF5FD13F8B6FCE4CF0D" box="[741,839,1881,1904]" italics="true" pageId="6" pageNumber="196">Agostenus</emphasis>
Fischer von Waldheim, 1829
</taxonomicName>
, dorsal view. 13,
<taxonomicName id="3D9D55FB9E75FFF5FACDF8B7FED9CFE8" authority="Gibler, 1830" authorityName="Gibler" authorityYear="1830" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="species" species="alutaceus">
<emphasis id="C8E9F26A9E75FFF5FACDF8B7FF51CFE9" italics="true" pageId="6" pageNumber="196">Chlaenius alutaceus</emphasis>
Gеbler, 1830
</taxonomicName>
(Lake Khanka, Primorsky Territory, Russia); 1415,
<taxonomicName id="3D9D55FB9E75FFF5FC7DF893FB4CCFE8" authority="(Paykul, 1798)" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[907,1263,1916,1941]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="C8E9F26A9E75FFF5FC7DF893FBF6CFE9" box="[907,1109,1916,1940]" italics="true" pageId="6" pageNumber="196">Chlaenius sulcicollis</emphasis>
(Paykul, 1798)
</taxonomicName>
(14, Belarus; 15, Golubiny Utes, Primorsky Territory, Russia); 1618,
<taxonomicName id="3D9D55FB9E75FFF5FD4BF84FFBFFCFC4" authority="(Paykul, 1790)" baseAuthorityName="Paykull" baseAuthorityYear="1790" box="[701,1116,1952,1977]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="species" species="quadrisulcatus">
<emphasis id="C8E9F26A9E75FFF5FD4BF84FFC1ACFC5" box="[701,953,1952,1976]" italics="true" pageId="6" pageNumber="196">Chlaenius quadrisulcatus</emphasis>
(Paykul, 1790)
</taxonomicName>
(16, Kedrovaya Pad Reserve, Primorsky Territory, Russia; 17, Blagoveshchensk Distr., Amurskaya Oblast, Russia; 18, Kunashir Island, Kuril Islands, Russia).
</paragraph>
</caption>
<paragraph id="FA222E789E74FFF4FF31FF78FB3EC9F4" blockId="7.[151,1437,151,393]" pageId="7" pageNumber="197">
According to some Japanese authors, only one species of the subgenus
<taxonomicName id="3D9D55FB9E74FFF4FC26FF78FA27C8CC" authority="Fischer von Waldheim, 1829" authorityName="Fischer von Waldheim" authorityYear="1829" box="[976,1412,151,177]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="subGenus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E74FFF4FC26FF78FBE0C8CC" box="[976,1091,151,177]" italics="true" pageId="7" pageNumber="197">Agostenus</emphasis>
Fischer von Waldheim, 1829
</taxonomicName>
is known to populate
<collectingCountry id="828A6EE89E74FFF4FE9FFF54FE0AC8A8" box="[361,425,187,213]" name="Japan" pageId="7" pageNumber="197">Japan</collectingCountry>
(
<collectingRegion id="3859E09A9E74FFF4FE43FF54FD86C8A8" box="[437,549,187,213]" country="Japan" name="Hokkaido" pageId="7" pageNumber="197">Hokkaido</collectingRegion>
and northern Honshu): either
<taxonomicName id="3D9D55FB9E74FFF4FC98FF54FC49C8A8" authorityName="Ganglbauer" authorityYear="1891" box="[878,1002,187,213]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E74FFF4FC98FF54FC49C8A8" box="[878,1002,187,213]" italics="true" pageId="7" pageNumber="197">Ch. gebleri</emphasis>
</taxonomicName>
(
<bibRefCitation id="9E0C53899E74FFF4FC01FF54FB02C8A8" author="Kasahara, S." box="[1015,1185,186,213]" pageId="7" pageNumber="197" pagination="155 - 159" refId="ref7944" refString="Kasahara, S. (1985) Carabidae (Callistinae). In: Ueno, S. I. &amp; Kurosawa, Y. (Eds.), The Coleoptera of Japan in Color. Vol. II. Hoikusha Publishing Co., Osaka, pp. 155 - 159." type="book chapter" year="1985">Kasahara 1985</bibRefCitation>
;
<bibRefCitation id="9E0C53899E74FFF4FB5DFF54FA84C8A8" author="Habu, A." box="[1195,1319,187,213]" pageId="7" pageNumber="197" pagination="1 - 36" refId="ref7771" refString="Habu, A. (1987) Classification of the Callistini of Japan (Coleoptera, Carabidae). The Entomological Review of Japan, 42 (1), 1 - 36." type="journal article" year="1987">Habu 1987</bibRefCitation>
;
<bibRefCitation id="9E0C53899E74FFF4FAC6FF54FEFBC884" author="Yasuda, N. &amp; Matsumoto, H." pageId="7" pageNumber="197" pagination="1 - 93" refId="ref9716" refString="Yasuda, N. &amp; Matsumoto, H. (1993) Ground beetles from Hokkaido, Northern Japan. Bulletin of the Sounkyo Museum of Natural History, 13, 1 - 93." type="journal article" year="1993">Yasuda &amp; Matsumoto 1993</bibRefCitation>
;
<bibRefCitation id="9E0C53899E74FFF4FE95FF30FDCEC884" author="Kimoto, S. &amp; Yasuda, N." box="[355,621,222,249]" pageId="7" pageNumber="197" refId="ref8048" refString="Kimoto, S. &amp; Yasuda, N. (1995) The ground beetles of Hokkaido. Ecological and biological survey. Tokai University Press, Tokyo, 315 pp. [in Japanese]" type="book" year="1995">Kimoto &amp; Yasuda 1995</bibRefCitation>
; etc.) or
<taxonomicName id="3D9D55FB9E74FFF4FD3BFF30FB2BC884" authority="(Yoshimatsu et al. 2018)" baseAuthorityName="Yoshimatsu" baseAuthorityYear="2018" box="[717,1160,223,249]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="C8E9F26A9E74FFF4FD3BFF30FCCEC884" box="[717,877,223,249]" italics="true" pageId="7" pageNumber="197">Ch. sulcicollis</emphasis>
(
<bibRefCitation id="9E0C53899E74FFF4FC8CFF30FB22C884" author="Yoshimatsu, S. - I. &amp; Ito, N. &amp; Nakatani, Y. &amp; Yoshitake, H." box="[890,1153,223,249]" pageId="7" pageNumber="197" pagination="15 - 191" refId="ref9751" refString="Yoshimatsu, S. - I., Ito, N., Nakatani, Y. &amp; Yoshitake, H. (2018) A list of ground beetles (Insecta: Coleoptera: Caraboidea) in Dr. Kazuo Tanaka's collection preserved in the Insect Museum of Institute for Agro-Environmental Sciences, Naro. Bulletin of National Institute for Agro-Environmental Sciences, 39, 15 - 191." type="journal article" year="2018">
Yoshimatsu
<emphasis id="C8E9F26A9E74FFF4FBF0FF30FB9DC884" box="[1030,1086,223,249]" italics="true" pageId="7" pageNumber="197">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
. According to a different point of view, there are no representatives of this subgenus in
<collectingCountry id="828A6EE89E74FFF4FCBBFEECFC2EC960" box="[845,909,259,285]" name="Japan" pageId="7" pageNumber="197">Japan</collectingCountry>
at all, if
<taxonomicName id="3D9D55FB9E74FFF4FC07FEECFBCCC960" authorityName="Ganglbauer" authorityYear="1891" box="[1009,1135,259,285]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E74FFF4FC07FEECFBCCC960" box="[1009,1135,259,285]" italics="true" pageId="7" pageNumber="197">Ch. gebleri</emphasis>
</taxonomicName>
is regarded as a subjective junior synonym of
<taxonomicName id="3D9D55FB9E74FFF4FE86FEC8FDB1C93C" box="[368,530,295,321]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="C8E9F26A9E74FFF4FE86FEC8FDB1C93C" box="[368,530,295,321]" italics="true" pageId="7" pageNumber="197">Ch. sulcicollis</emphasis>
</taxonomicName>
(
<bibRefCitation id="9E0C53899E74FFF4FDD7FEC8FCA9C93C" author="Kryzhanovskij, O. L." box="[545,778,295,321]" pageId="7" pageNumber="197" pagination="8 - 17" refId="ref8438" refString="Kryzhanovskij, O. L. (1976) Obzor zhuzhelits triby Callistini (Coleoptera, Carabidae) Dalnego Vostoka. Proceedings of the Institute of Soil Biology, New Series, 43 (146), 8 - 17. [in Russian]" type="journal article" year="1976">Kryzhanovskij 1976</bibRefCitation>
;
<bibRefCitation id="9E0C53899E74FFF4FCE1FEC8FC34C93C" author="Lafer, G. Sh." box="[791,919,295,321]" pageId="7" pageNumber="197" pagination="71 - 222" refId="ref8582" refString="Lafer, G. Sh. (1989) 4. Semeystvo Carabidae - Zhuzhelitsy. In: Lehr, P. A. (Ed.), Opredelitel nasekomykh Dalnego Vostoka SSSR. Tom 3. Zhestkokrylye, ili zhuki. Part 1. Nauka, Leningrad, pp. 71 - 222. [in Russian]" type="book chapter" year="1989">Lafer 1989</bibRefCitation>
;
<bibRefCitation id="9E0C53899E74FFF4FC55FEC8FB24C93C" author="Kirschenhofer, E." box="[931,1159,295,321]" pageId="7" pageNumber="197" pagination="1834" refId="ref8115" refString="Kirschenhofer, E. (2003) Tribe Chlaeniini Brulle, 1834. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalog of Palearctic Coleoptera. Vol. 1. Archostemmata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 347 - 356." type="book chapter" year="2003">Kirschenhofer 2003</bibRefCitation>
,
<bibRefCitation id="9E0C53899E74FFF4FB65FEC8FB6DC93C" author="Kirschenhofer, E." box="[1171,1230,295,321]" pageId="7" pageNumber="197" pagination="482 - 497" refId="ref8266" refString="Kirschenhofer, E. (2017) Tribe Chlaeniini Brulle, 1834. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalog of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 482 - 497." type="book chapter" year="2017">2017</bibRefCitation>
;
<bibRefCitation id="9E0C53899E74FFF4FB2CFEC8FA32C93C" author="Sundukov, Yu. N." box="[1242,1425,295,321]" pageId="7" pageNumber="197" refId="ref9272" refString="Sundukov, Yu. N. (2013) An annotated catalog of the ground beetles (Coleoptera: Caraboidea) of Sikhote-Alin. Dalnauka, Vladivostok, 271 pp. [in Russian]" type="book" year="2013">Sundukov 2013</bibRefCitation>
). Finally, because the species status
<taxonomicName id="3D9D55FB9E74FFF4FDD4FEA4FD01C918" authorityName="Ganglbauer" authorityYear="1891" box="[546,674,331,357]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E74FFF4FDD4FEA4FD01C918" box="[546,674,331,357]" italics="true" pageId="7" pageNumber="197">Ch. gebleri</emphasis>
</taxonomicName>
has been resurrected, this species is widely distributed in southern Siberia, the southern Russian Far East, and the Japanese Islands (
<bibRefCitation id="9E0C53899E74FFF4FC86FE80FB32C9F4" author="Vertyankin, A. V. &amp; Lafer, G. Sh." box="[880,1169,367,393]" pageId="7" pageNumber="197" pagination="433 - 436" refId="ref9508" refString="Vertyankin, A. V. &amp; Lafer, G. Sh. (2012) New records of ground beetles (Coleoptera, Carabidae) from Sakhalin and Moneron Islands of the Russian Far East. Euroasian Entomological Journal, 11 (5), 433 - 436. [in Russian]" type="journal article" year="2012">Vertyankin &amp; Lafer 2012</bibRefCitation>
).
</paragraph>
<caption id="AEE27EF09E74FFF4FF61F916FBAFCFDC" ID-DOI="http://doi.org/10.5281/zenodo.7456769" ID-Zenodo-Dep="7456769" httpUri="https://zenodo.org/record/7456769/files/figure.png" pageId="7" pageNumber="197" startId="7.[151,264,1785,1809]" targetBox="[161,1433,424,1751]" targetPageId="7">
<paragraph id="FA222E789E74FFF4FF61F916FBAFCFDC" blockId="7.[151,1437,1785,1953]" pageId="7" pageNumber="197">
<emphasis id="C8E9F26A9E74FFF4FF61F916FEF1CF6C" bold="true" box="[151,338,1785,1809]" pageId="7" pageNumber="197">FIGURES 1927.</emphasis>
Aedeagi of the subgenus
<taxonomicName id="3D9D55FB9E74FFF4FDBBF916FC7ACF6C" authority="Fischer von Waldheim, 1829" authorityName="Fischer von Waldheim" authorityYear="1829" box="[589,985,1785,1809]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="subGenus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E74FFF4FDBBF916FD16CF6D" box="[589,693,1785,1808]" italics="true" pageId="7" pageNumber="197">Agostenus</emphasis>
Fischer von Waldheim, 1829
</taxonomicName>
, apical lamella (dorsal view) and median lobe (lateral view). 1920,
<taxonomicName id="3D9D55FB9E74FFF4FE81F8F2FD6DCF48" authority="Gibler, 1830" authorityName="Gibler" authorityYear="1830" box="[375,718,1821,1845]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="alutaceus">
<emphasis id="C8E9F26A9E74FFF4FE81F8F2FDE6CF48" box="[375,581,1821,1845]" italics="true" pageId="7" pageNumber="197">Chlaenius alutaceus</emphasis>
Gеbler, 1830
</taxonomicName>
(19, Moldova; 20, Buryatia, Russia); 2124,
<taxonomicName id="3D9D55FB9E74FFF4FB6FF8F2FE8ECF24" authority="(Paykul, 1790)" baseAuthorityName="Paykull" baseAuthorityYear="1790" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="quadrisulcatus">
<emphasis id="C8E9F26A9E74FFF4FB6FF8F2FA3ECF48" box="[1177,1437,1821,1845]" italics="true" pageId="7" pageNumber="197">Chlaenius quadrisulcatus</emphasis>
(Paykul, 1790)
</taxonomicName>
(21, Belarus; 22, Amurskaya Oblast, Russia; 23, Narva River, Khasansky Distr., Primorsky Territory, Russia; 24, Kunashir Island, Kurile Islands, Russia); 2527,
<taxonomicName id="3D9D55FB9E74FFF4FD77F88AFC53CF00" authority="(Paykul, 1798)" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[641,1008,1893,1917]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="C8E9F26A9E74FFF4FD77F88AFCF6CF00" box="[641,853,1893,1917]" italics="true" pageId="7" pageNumber="197">Chlaenius sulcicollis</emphasis>
(Paykul, 1798)
</taxonomicName>
(25, Belovezhskaya Pushcha, Belarus; 26, Vladimir.[skaya] Vol.[olst], Belarus; 27, Golubinyi Utes, Primorsky Territory, Russia).
</paragraph>
</caption>
<paragraph id="FA222E789E7BFFFBFF31FF78FD60C9AC" blockId="8.[151,1437,151,969]" pageId="8" pageNumber="198">
<taxonomicName id="3D9D55FB9E7BFFFBFF31FF78FE2EC8CC" authorityName="Ganglbauer" authorityYear="1891" box="[199,397,151,177]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E7BFFFBFF31FF78FE2EC8CC" box="[199,397,151,177]" italics="true" pageId="8" pageNumber="198">Chlaenius gebleri</emphasis>
</taxonomicName>
has been described as
<taxonomicName id="3D9D55FB9E7BFFFBFD74FF78FC57C8CC" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[642,1012,151,177]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="sulcicollis" subGenus="Agostenus">
<emphasis id="C8E9F26A9E7BFFFBFD74FF78FD57C8CC" box="[642,756,151,177]" italics="true" pageId="8" pageNumber="198">Chlaenius</emphasis>
(
<emphasis id="C8E9F26A9E7BFFFBFCF7FF78FCD7C8CC" box="[769,884,151,177]" italics="true" pageId="8" pageNumber="198">Agostenus</emphasis>
)
<emphasis id="C8E9F26A9E7BFFFBFC74FF78FC57C8CC" box="[898,1012,151,177]" italics="true" pageId="8" pageNumber="198">sulcicollis</emphasis>
</taxonomicName>
var.
<taxonomicName id="3D9D55FB9E7BFFFBFBD1FF78FBDEC8CC" authorityName="Ganglbauer" authorityYear="1891" box="[1063,1149,151,177]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E7BFFFBFBD1FF78FBDEC8CC" box="[1063,1149,151,177]" italics="true" pageId="8" pageNumber="198">Gebleri</emphasis>
</taxonomicName>
from Siberia (
<bibRefCitation id="9E0C53899E7BFFFBFAEEFF78FF70C8A8" author="Ganglbauer, L." pageId="8" pageNumber="198" refId="ref7652" refString="Ganglbauer, L. (1891) Die Kafer von Mitteleuropa. Die Kafer der Osterreichisch-Ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. Erster Band. Familienreihe Caraboidea. C. Gerold's Sohn, Wien, iii + 557 pp. https: // doi. org / 10.5962 / bhl. title. 8764" type="book" year="1891">Ganglbauer 1891</bibRefCitation>
). When describing it, L. Ganglbauer well distinguished
<taxonomicName id="3D9D55FB9E7BFFFBFCBFFF54FC49C8A8" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[841,1002,187,213]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="C8E9F26A9E7BFFFBFCBFFF54FC49C8A8" box="[841,1002,187,213]" italics="true" pageId="8" pageNumber="198">Ch. sulcicollis</emphasis>
</taxonomicName>
(
<figureCitation id="62A632FD9E7BFFFBFC0FFF54FBDAC8A8" box="[1017,1145,186,213]" captionStart="FIGURES 1318" captionStartId="6.[151,264,1880,1904]" captionTargetBox="[305,1285,676,1862]" captionTargetId="figure-277@6.[304,1286,675,1865]" captionTargetPageId="6" captionText="FIGURES 1318. Far Eastern species of the subgenus Agostenus Fischer von Waldheim, 1829, dorsal view. 13, Chlaenius alutaceus Gеbler, 1830 (Lake Khanka, Primorsky Territory, Russia); 1415, Chlaenius sulcicollis (Paykul, 1798) (14, Belarus; 15, Golubiny Utes, Primorsky Territory, Russia); 1618, Chlaenius quadrisulcatus (Paykul, 1790) (16, Kedrovaya Pad Reserve, Primorsky Territory, Russia; 17, Blagoveshchensk Distr., Amurskaya Oblast, Russia; 18, Kunashir Island, Kuril Islands, Russia)." figureDoi="http://doi.org/10.5281/zenodo.7456767" httpUri="https://zenodo.org/record/7456767/files/figure.png" pageId="8" pageNumber="198">Figs 1415</figureCitation>
) from
<taxonomicName id="3D9D55FB9E7BFFFBFB31FF54FA3FC8A8" baseAuthorityName="Paykull" baseAuthorityYear="1790" box="[1223,1436,187,213]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="quadrisulcatus">
<emphasis id="C8E9F26A9E7BFFFBFB31FF54FA3FC8A8" box="[1223,1436,187,213]" italics="true" pageId="8" pageNumber="198">Ch. quadrisulcatus</emphasis>
</taxonomicName>
(
<figureCitation id="62A632FD9E7BFFFBFF69FF30FE86C884" box="[159,293,223,249]" captionStart="FIGURES 1318" captionStartId="6.[151,264,1880,1904]" captionTargetBox="[305,1285,676,1862]" captionTargetId="figure-277@6.[304,1286,675,1865]" captionTargetPageId="6" captionText="FIGURES 1318. Far Eastern species of the subgenus Agostenus Fischer von Waldheim, 1829, dorsal view. 13, Chlaenius alutaceus Gеbler, 1830 (Lake Khanka, Primorsky Territory, Russia); 1415, Chlaenius sulcicollis (Paykul, 1798) (14, Belarus; 15, Golubiny Utes, Primorsky Territory, Russia); 1618, Chlaenius quadrisulcatus (Paykul, 1790) (16, Kedrovaya Pad Reserve, Primorsky Territory, Russia; 17, Blagoveshchensk Distr., Amurskaya Oblast, Russia; 18, Kunashir Island, Kuril Islands, Russia)." figureDoi="http://doi.org/10.5281/zenodo.7456767" httpUri="https://zenodo.org/record/7456767/files/figure.png" pageId="8" pageNumber="198">Figs 1618</figureCitation>
), clearly indicating that the latter species had an elevated, smooth and shiny elytral suture (in the original, “... ist die Naht der Flügeldecken vorn erhobt, glatt und glänzend”) (
<bibRefCitation id="9E0C53899E7BFFFBFBF0FEECFAAAC960" author="Ganglbauer, L." box="[1030,1289,259,285]" pageId="8" pageNumber="198" refId="ref7652" refString="Ganglbauer, L. (1891) Die Kafer von Mitteleuropa. Die Kafer der Osterreichisch-Ungarischen Monarchie, Deutschlands, der Schweiz, sowie des franzosischen und italienischen Alpengebietes. Erster Band. Familienreihe Caraboidea. C. Gerold's Sohn, Wien, iii + 557 pp. https: // doi. org / 10.5962 / bhl. title. 8764" type="book" year="1891">Ganglbauer 1891: 391</bibRefCitation>
). Thus, there is no doubt that Ganglbauer described the form as
<emphasis id="C8E9F26A9E7BFFFBFD14FEC8FC28C93C" box="[738,907,295,321]" italics="true" pageId="8" pageNumber="198">
<taxonomicName id="3D9D55FB9E7BFFFBFD14FEC8FC24C93C" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[738,903,295,321]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="sulcicollis">Ch. sulcicollis</taxonomicName>
,
</emphasis>
reminiscent of
<emphasis id="C8E9F26A9E7BFFFBFBB7FEC8FA83C93C" box="[1089,1312,295,321]" italics="true" pageId="8" pageNumber="198">
<taxonomicName id="3D9D55FB9E7BFFFBFBB7FEC8FABFC93C" baseAuthorityName="Paykull" baseAuthorityYear="1790" box="[1089,1308,295,321]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="quadrisulcatus">Ch. quadrisulcatus</taxonomicName>
.
</emphasis>
It remains unclear who and when assigned to
<taxonomicName id="3D9D55FB9E7BFFFBFDDBFEA4FD0EC918" authorityName="Ganglbauer" authorityYear="1891" box="[557,685,331,357]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E7BFFFBFDDBFEA4FD0EC918" box="[557,685,331,357]" italics="true" pageId="8" pageNumber="198">Ch. gebleri</emphasis>
</taxonomicName>
specimens of
<taxonomicName id="3D9D55FB9E7BFFFBFCA0FEA4FC6AC918" authorityName="Fischer von Waldheim" authorityYear="1829" box="[854,969,331,357]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="subGenus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E7BFFFBFCA0FEA4FC6AC918" box="[854,969,331,357]" italics="true" pageId="8" pageNumber="198">Agostenus</emphasis>
</taxonomicName>
from
<collectingCountry id="828A6EE89E7BFFFBFBE4FEA4FBF6C918" box="[1042,1109,331,357]" name="Japan" pageId="8" pageNumber="198">Japan</collectingCountry>
. A study of descriptions and illustrations (
<bibRefCitation id="9E0C53899E7BFFFBFEDCFE80FDAEC9F4" author="Kasahara, S." box="[298,525,366,393]" pageId="8" pageNumber="198" pagination="155 - 159" refId="ref7944" refString="Kasahara, S. (1985) Carabidae (Callistinae). In: Ueno, S. I. &amp; Kurosawa, Y. (Eds.), The Coleoptera of Japan in Color. Vol. II. Hoikusha Publishing Co., Osaka, pp. 155 - 159." type="book chapter" year="1985">Kasahara 1985: 159</bibRefCitation>
, pl. 29, fig. 19;
<bibRefCitation id="9E0C53899E7BFFFBFD48FE80FCFBC9F4" author="Habu, A." box="[702,856,367,393]" pageId="8" pageNumber="198" pagination="1 - 36" refId="ref7771" refString="Habu, A. (1987) Classification of the Callistini of Japan (Coleoptera, Carabidae). The Entomological Review of Japan, 42 (1), 1 - 36." type="journal article" year="1987">Habu 1987: 4</bibRefCitation>
, 810, fig. 16;
<bibRefCitation id="9E0C53899E7BFFFBFBF6FE80FB4CC9F4" author="Yoshitake, H. &amp; Kurihara, T. &amp; Yoshimatsu, S. - I. &amp; Nakatani, Y. &amp; Yasuda, K." box="[1024,1263,367,393]" pageId="8" pageNumber="198" pagination="1 - 327" refId="ref9821" refString="Yoshitake, H., Kurihara, T., Yoshimatsu, S. - I., Nakatani, Y. &amp; Yasuda, K. (2011) A list of carabid specimens (Insecta: Coleoptera) collected by the late Dr. Akinobu Habu preserved in the Insect Museum of the National Institute for Agro-Environmental Sciences. Bulletin of National Institute for Agro-Environmental Sciences, 28, 1 - 327." type="journal article" year="2011">
Yoshitake
<emphasis id="C8E9F26A9E7BFFFBFB80FE80FB0DC9F4" box="[1142,1198,367,393]" italics="true" pageId="8" pageNumber="198">et al.</emphasis>
2011
</bibRefCitation>
: fig. 33) allows us to reveal that, contrary to the original description, over the last half-century the name
<taxonomicName id="3D9D55FB9E7BFFFBFB86FE7CFB4EC9D0" authorityName="Ganglbauer" authorityYear="1891" box="[1136,1261,403,429]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E7BFFFBFB86FE7CFB4EC9D0" box="[1136,1261,403,429]" italics="true" pageId="8" pageNumber="198">Ch. gebleri</emphasis>
</taxonomicName>
from
<collectingCountry id="828A6EE89E7BFFFBFAC7FE7CFAD2C9D0" box="[1329,1393,403,429]" name="Japan" pageId="8" pageNumber="198">Japan</collectingCountry>
has concealed
<emphasis id="C8E9F26A9E7BFFFBFEF8FE58FE49C9AC" box="[270,490,439,465]" italics="true" pageId="8" pageNumber="198">
<taxonomicName id="3D9D55FB9E7BFFFBFEF8FE58FE45C9AC" baseAuthorityName="Paykull" baseAuthorityYear="1790" box="[270,486,439,465]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="quadrisulcatus">Ch. quadrisulcatus</taxonomicName>
,
</emphasis>
not
<taxonomicName id="3D9D55FB9E7BFFFBFDEAFE58FD1EC9AC" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[540,701,439,465]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="sulcicollis">
<emphasis id="C8E9F26A9E7BFFFBFDEAFE58FD1EC9AC" box="[540,701,439,465]" italics="true" pageId="8" pageNumber="198">Ch. sulcicollis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FA222E789E7BFFFBFF31FE34FD18CAD5" blockId="8.[151,1437,151,969]" pageId="8" pageNumber="198">
The study of all available material of
<taxonomicName id="3D9D55FB9E7BFFFBFD8CFE34FD4EC988" authorityName="Fischer von Waldheim" authorityYear="1829" box="[634,749,475,501]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="subGenus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E7BFFFBFD8CFE34FD4EC988" box="[634,749,475,501]" italics="true" pageId="8" pageNumber="198">Agostenus</emphasis>
</taxonomicName>
shows that three species range from Europe to the Far East of
<collectingCountry id="828A6EE89E7BFFFBFF41FE10FEA6CA64" box="[183,261,511,537]" name="Russia" pageId="8" pageNumber="198">Russia</collectingCountry>
, with clear differences both in external characters (
<figureCitation id="62A632FD9E7BFFFBFCBEFE10FC6BCA64" box="[840,968,511,537]" captionStart="FIGURES 1318" captionStartId="6.[151,264,1880,1904]" captionTargetBox="[305,1285,676,1862]" captionTargetId="figure-277@6.[304,1286,675,1865]" captionTargetPageId="6" captionText="FIGURES 1318. Far Eastern species of the subgenus Agostenus Fischer von Waldheim, 1829, dorsal view. 13, Chlaenius alutaceus Gеbler, 1830 (Lake Khanka, Primorsky Territory, Russia); 1415, Chlaenius sulcicollis (Paykul, 1798) (14, Belarus; 15, Golubiny Utes, Primorsky Territory, Russia); 1618, Chlaenius quadrisulcatus (Paykul, 1790) (16, Kedrovaya Pad Reserve, Primorsky Territory, Russia; 17, Blagoveshchensk Distr., Amurskaya Oblast, Russia; 18, Kunashir Island, Kuril Islands, Russia)." figureDoi="http://doi.org/10.5281/zenodo.7456767" httpUri="https://zenodo.org/record/7456767/files/figure.png" pageId="8" pageNumber="198">Figs 1318</figureCitation>
) and in the structure of the aedeagus and endophallus (
<figureCitation id="62A632FD9E7BFFFBFEDBFDCCFE0CCA40" box="[301,431,547,573]" captionStart="FIGURES 1927" captionStartId="7.[151,264,1785,1809]" captionTargetBox="[161,1433,424,1751]" captionTargetId="figure-143@7.[151,1435,417,1762]" captionTargetPageId="7" captionText="FIGURES 1927. Aedeagi of the subgenus Agostenus Fischervon Waldheim, 1829, apical lamella (dorsal view) and median lobe (lateral view). 1920, Chlaenius alutaceus Gеbler, 1830 (19, Moldova; 20, Buryatia, Russia); 2124, Chlaenius quadrisulcatus (Paykul, 1790) (21, Belarus; 22, Amurskaya Oblast, Russia; 23, Narva River, Khasansky Distr., Primorsky Territory, Russia; 24, Kunashir Island, Kurile Islands, Russia); 2527, Chlaenius sulcicollis (Paykul, 1798) (25, Belovezhskaya Pushcha, Belarus; 26, Vladimir.[skaya] Vol.[olst], Belarus; 27, Golubinyi Utes, Primorsky Territory, Russia)." figureDoi="http://doi.org/10.5281/zenodo.7456769" httpUri="https://zenodo.org/record/7456769/files/figure.png" pageId="8" pageNumber="198">Figs 1927</figureCitation>
). At the same time, the variability of the shape of the lamella and inner sac is insignificant, whereas the differences between these species are stable and persist throughout the entire distribution range. Thus, there is no reason to believe that any species of
<taxonomicName id="3D9D55FB9E7BFFFBFD23FD84FCEBCAF8" authorityName="Fischer von Waldheim" authorityYear="1829" box="[725,840,619,645]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="subGenus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E7BFFFBFD23FD84FCEBCAF8" box="[725,840,619,645]" italics="true" pageId="8" pageNumber="198">Agostenus</emphasis>
</taxonomicName>
other than the widespread
<emphasis id="C8E9F26A9E7BFFFBFB7CFD84FEAACAD4" italics="true" pageId="8" pageNumber="198">
<taxonomicName id="3D9D55FB9E7BFFFBFB7CFD84FAC6CAF8" baseAuthorityName="Paykull" baseAuthorityYear="1790" box="[1162,1381,619,645]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="quadrisulcatus">Ch. quadrisulcatus</taxonomicName>
,
<taxonomicName id="3D9D55FB9E7BFFFBFA83FD84FEAACAD4" baseAuthorityName="Paykull" baseAuthorityYear="1798" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="sulcicollis">Ch. sulcicollis</taxonomicName>
</emphasis>
and
<taxonomicName id="3D9D55FB9E7BFFFBFEC9FD60FE7ACAD4" authorityName="Gibler" authorityYear="1830" box="[319,473,655,681]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="alutaceus">
<emphasis id="C8E9F26A9E7BFFFBFEC9FD60FE7ACAD4" box="[319,473,655,681]" italics="true" pageId="8" pageNumber="198">Ch. alutaceus</emphasis>
</taxonomicName>
live in the Far East.
</paragraph>
<paragraph id="FA222E789E7BFFFBFF31FD5CFBCCCBFC" blockId="8.[151,1437,151,969]" pageId="8" pageNumber="198">
We can assume that the cause of the confusion seems to have been a typographical error in the second volume of the Faune de
<collectingCountry id="828A6EE89E7BFFFBFEBBFD38FE3ACA8C" box="[333,409,727,753]" name="France" pageId="8" pageNumber="198">France</collectingCountry>
(
<bibRefCitation id="9E0C53899E7BFFFBFE5EFD38FDEBCA8C" author="Jeannel, R." box="[424,584,727,753]" pageId="8" pageNumber="198" refId="ref7846" refString="Jeannel, R. (1942) Coleopteres carabiques. Deuxieme partie. Faune de France. Vol. 40. Librairie de la Faculte des Sciences, Paris, pp. 572 - 1173." type="book" year="1942">Jeannel, 1942</bibRefCitation>
), where actually it was the aedeagus of
<taxonomicName id="3D9D55FB9E7BFFFBFBFEFD38FB0CCA8C" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[1032,1199,727,753]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="sulcicollus">
<emphasis id="C8E9F26A9E7BFFFBFBFEFD38FB0CCA8C" box="[1032,1199,727,753]" italics="true" pageId="8" pageNumber="198">Ch. sulcicollus</emphasis>
</taxonomicName>
that was depicted on page
<quantity id="3D65839D9E7BFFFBFF22FD14FEB8CB68" box="[212,283,763,789]" metricMagnitude="1" metricUnit="m" metricValue="2.48158" pageId="8" pageNumber="198" unit="in" value="977.0">977 in</quantity>
figures 323a and 323b., not
<taxonomicName id="3D9D55FB9E7BFFFBFDA9FD14FC96CB68" baseAuthorityName="Paykull" baseAuthorityYear="1790" box="[607,821,763,789]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="quadrisulcatus">
<emphasis id="C8E9F26A9E7BFFFBFDA9FD14FC96CB68" box="[607,821,763,789]" italics="true" pageId="8" pageNumber="198">Ch. quadrisulcatus</emphasis>
</taxonomicName>
. This has been confirmed by comparing with museum specimens from Europe.
<bibRefCitation id="9E0C53899E7BFFFBFE45FCF0FD58CB44" author="Vertyankin, A. V. &amp; Lafer, G. Sh." box="[435,763,799,825]" pageId="8" pageNumber="198" pagination="433 - 436" refId="ref9508" refString="Vertyankin, A. V. &amp; Lafer, G. Sh. (2012) New records of ground beetles (Coleoptera, Carabidae) from Sakhalin and Moneron Islands of the Russian Far East. Euroasian Entomological Journal, 11 (5), 433 - 436. [in Russian]" type="journal article" year="2012">Vertyankin and Lafer (2012)</bibRefCitation>
, comparing the aedeagus of the
<taxonomicName id="3D9D55FB9E7BFFFBFB9BFCF0FB43CB44" authorityName="Fischer von Waldheim" authorityYear="1829" box="[1133,1248,799,825]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="subGenus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E7BFFFBFB9BFCF0FB43CB44" box="[1133,1248,799,825]" italics="true" pageId="8" pageNumber="198">Agostenus</emphasis>
</taxonomicName>
specimens from Sakhalin with the illustrations in the monograph by R.
<bibRefCitation id="9E0C53899E7BFFFBFD03FCACFC3FCB20" author="Jeannel, R." box="[757,924,835,861]" pageId="8" pageNumber="198" refId="ref7846" refString="Jeannel, R. (1942) Coleopteres carabiques. Deuxieme partie. Faune de France. Vol. 40. Librairie de la Faculte des Sciences, Paris, pp. 572 - 1173." type="book" year="1942">Jeannel (1942)</bibRefCitation>
, made an incorrect conclusion of its similarity to
<emphasis id="C8E9F26A9E7BFFFBFF42FC88FEFFCBFC" box="[180,348,871,897]" italics="true" pageId="8" pageNumber="198">
<taxonomicName id="3D9D55FB9E7BFFFBFF42FC88FEFBCBFC" baseAuthorityName="Paykull" baseAuthorityYear="1798" box="[180,344,871,897]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="sulcicollis">Ch. sulcicollis</taxonomicName>
.
</emphasis>
Based on this, they resurrected
<taxonomicName id="3D9D55FB9E7BFFFBFD34FC88FC9CCBFC" authorityName="Ganglbauer" authorityYear="1891" box="[706,831,871,897]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="gebleri">
<emphasis id="C8E9F26A9E7BFFFBFD34FC88FC9CCBFC" box="[706,831,871,897]" italics="true" pageId="8" pageNumber="198">Ch. gebleri</emphasis>
</taxonomicName>
as an independent species.
</paragraph>
<paragraph id="FA222E789E7BFFFBFF31FC64FCA7CBB4" blockId="8.[151,1437,151,969]" pageId="8" pageNumber="198">
Thus, we believe that the Far Eastern islands (
<collectingRegion id="3859E09A9E7BFFFBFD2CFC64FCE3CBD8" box="[730,832,907,933]" country="Russia" name="Sakhalin" pageId="8" pageNumber="198">Sakhalin</collectingRegion>
, Kunashir,
<collectingRegion id="3859E09A9E7BFFFBFC35FC64FB97CBD8" box="[963,1076,907,933]" country="Japan" name="Hokkaido" pageId="8" pageNumber="198">Hokkaido</collectingRegion>
, and Honshu) support only one species of the subgenus
<taxonomicName id="3D9D55FB9E7BFFFBFE50FC40FDBACBB4" authorityName="Fischer von Waldheim" authorityYear="1829" box="[422,537,943,969]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="subGenus" subGenus="Agostenus">
<emphasis id="C8E9F26A9E7BFFFBFE50FC40FDBACBB4" box="[422,537,943,969]" italics="true" pageId="8" pageNumber="198">Agostenus</emphasis>
</taxonomicName>
:
<taxonomicName id="3D9D55FB9E7BFFFBFDDEFC40FD5FCBB4" baseAuthorityName="Paykull" baseAuthorityYear="1790" box="[552,764,943,969]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="quadrisulcatus">
<emphasis id="C8E9F26A9E7BFFFBFDDEFC40FD5FCBB4" box="[552,764,943,969]" italics="true" pageId="8" pageNumber="198">Ch. quadrisulcatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>