177 lines
15 KiB
XML
177 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.791.28137" ID-GBIF-Dataset="e674f085-58c8-4315-8aad-09c132d5f7d0" ID-PMC="PMC6205991" ID-Pensoft-Pub="1313-2970-791-107" ID-PubMed="30386156" ID-ZBK="95940307D4494EEEA21E3A4D8256FBEF" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-791-107" ModsDocOrigin="ZooKeys 791" ModsDocTitle="Sixteen new species of the genus Pseudopoda Jäger, 2000 from China, Myanmar, and Thailand (Sparassidae, Heteropodinae)" checkinTime="1540260181599" checkinUser="pensoft" docAuthor="Jiang, Tongyao, Zhao, Qingyuan & Li, Shuqiang" docDate="2018" docId="7219089FF1EA2C8821CDECE941A017B8" docLanguage="en" docName="ZooKeys 791: 107-161" docOrigin="ZooKeys 791" docSource="http://dx.doi.org/10.3897/zookeys.791.28137" docTitle="Pseudopoda colubrina Zhao & Li, sp. n." docType="treatment" docUuid="8EDAFE92-8991-4BD0-A68F-C3895F419AB5" docUuidSource="ZooBank" docVersion="5" lastPageNumber="113" masterDocId="FFE8FF8EFF828C0EFFDCFFA76D287419" masterDocTitle="Sixteen new species of the genus Pseudopoda Jaeger, 2000 from China, Myanmar, and Thailand (Sparassidae, Heteropodinae)" masterLastPageNumber="161" masterPageNumber="107" pageNumber="110" updateTime="1668166307399" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Sixteen new species of the genus Pseudopoda Jaeger, 2000 from China, Myanmar, and Thailand (Sparassidae, Heteropodinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Jiang, Tongyao</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Zhao, Qingyuan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Li, Shuqiang</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2018</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>791</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>107</mods:start>
|
||
<mods:end>161</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.791.28137</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.791.28137</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-791-107</mods:identifier>
|
||
<mods:identifier type="ZBK">95940307D4494EEEA21E3A4D8256FBEF</mods:identifier>
|
||
<mods:identifier type="ZooBank">95940307D4494EEEA21E3A4D8256FBEF</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="148892017" LSID="urn:lsid:zoobank.org:act:8EDAFE92-8991-4BD0-A68F-C3895F419AB5" httpUri="http://treatment.plazi.org/id/7219089FF1EA2C8821CDECE941A017B8" lastPageId="6" lastPageNumber="113" pageId="3" pageNumber="110">
|
||
<subSubSection pageId="3" pageNumber="110" type="nomenclature">
|
||
<paragraph pageId="3" pageNumber="110">
|
||
<taxonomicName LSID="http://zoobank.org/8EDAFE92-8991-4BD0-A68F-C3895F419AB5" authority="Zhao & Li" class="Arachnida" family="Sparassidae" genus="Pseudopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopoda colubrina" order="Araneae" pageId="3" pageNumber="110" phylum="Arthropoda" rank="species" species="colubrina">Pseudopoda colubrina Zhao & Li</taxonomicName>
|
||
<taxonomicNameLabel pageId="3" pageNumber="110">sp. n.</taxonomicNameLabel>
|
||
Figs 3, 4, 5, 37
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="111" type="type material">
|
||
<paragraph pageId="4" pageNumber="111">
|
||
<pageBreakToken pageId="4" pageNumber="111" start="start">Type</pageBreakToken>
|
||
material.
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="111">
|
||
Holotype ♂: Myanmar, Kachin State, Putao, road to Ziradum Village,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.560284">27°33.617'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="1" value="97.10945">97°06.567'E</geoCoordinate>
|
||
, 1003 m, 8 V 2017, J. Wu & Z. Chen. Paratype: 1 ♀, same locality as holotype, 13 XII 2016, J. Wu.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="111" type="etymology">
|
||
<paragraph pageId="4" pageNumber="111">Etymology.</paragraph>
|
||
<paragraph pageId="4" pageNumber="111">
|
||
The specific name is derived from the Latin word colubrinus, -a, -um, meaning 'serpentine,
|
||
<normalizedToken originalValue="winding’">winding'</normalizedToken>
|
||
, and referring to the shape of embolus in this species, which coils at the basal part and erects distally and looks like an alarmed snake; adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="5" lastPageNumber="112" pageId="4" pageNumber="111" type="diagnosis">
|
||
<paragraph pageId="4" pageNumber="111">Diagnosis.</paragraph>
|
||
<paragraph pageId="4" pageNumber="111">
|
||
Small to median-sized
|
||
<taxonomicName class="Arachnida" family="Sparassidae" genus="Pseudopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopoda" order="Araneae" pageId="4" pageNumber="111" phylum="Arthropoda" rank="genus">Pseudopoda</taxonomicName>
|
||
species. Male resembles
|
||
<taxonomicName lsidName="P. wu" pageId="4" pageNumber="111" rank="species" species="wu">P. wu</taxonomicName>
|
||
<normalizedToken originalValue="Jäger">Jaeger</normalizedToken>
|
||
, Li & Krehenwinkel, 2015 (see
|
||
<bibRefCitation author="Jaeger, P" journalOrPublisher="Zootaxa" pageId="33" pageNumber="140" pagination="363 - 392" title="Morphological and molecular taxonomic analysis of Pseudopoda Jaeger, 2000 (Araneae: Sparassidae: Heteropodinae) in Sichuan Province, China." url="https://doi.org/10.11646/zootaxa.3999.3.3" volume="3999" year="2015">
|
||
<normalizedToken originalValue="Jäger">Jaeger</normalizedToken>
|
||
et al. 2015
|
||
</bibRefCitation>
|
||
: 384, figs 115-129) and
|
||
<taxonomicName lsidName="P. tji" pageId="4" pageNumber="111" rank="species" species="tji">P. tji</taxonomicName>
|
||
<normalizedToken originalValue="Jäger">Jaeger</normalizedToken>
|
||
, 2015 (see
|
||
<bibRefCitation author="Jaeger, P" journalOrPublisher="Arachnology" pageId="33" pageNumber="140" pagination="333 - 350" title="Conductor-less and vertically niched: new species of the genus Pseudopoda (Araneae: Sparassidae: Heteropodinae) from Myanmar." url="https://doi.org/10.13156/arac.2015.16.9.333" volume="16" year="2015">
|
||
<normalizedToken originalValue="Jäger">Jaeger</normalizedToken>
|
||
2015
|
||
</bibRefCitation>
|
||
: 333, figs 1-15, 91) by: 1. embolus robust but twisted, forming loops (Figure 4A, B; rarely seem in other
|
||
<taxonomicName class="Arachnida" family="Sparassidae" genus="Pseudopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopoda" order="Araneae" pageId="4" pageNumber="111" phylum="Arthropoda" rank="genus">Pseudopoda</taxonomicName>
|
||
spp.); 2. conductor absent (Figure 4A, B). It can be easily distinguished from the two congeners by the following combination of characters: 1. only basal part of embolus twisted, distal part elongated and mildly bent (Figure 4A, B; distal part coiled in
|
||
<taxonomicName lsidName="P. tji" pageId="4" pageNumber="111" rank="species" species="tji">P. tji</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="P. wu" pageId="4" pageNumber="111" rank="species" species="wu">P. wu</taxonomicName>
|
||
); 2. tegulum occupying two third of alveolus (Figure 3B; covering whole or most of alveolus in
|
||
<taxonomicName lsidName="P. tji" pageId="4" pageNumber="111" rank="species" species="tji">P. tji</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="P. wu" pageId="4" pageNumber="111" rank="species" species="wu">P. wu</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption pageId="4" pageNumber="111">
|
||
<paragraph pageId="4" pageNumber="111">
|
||
Figure 3.
|
||
<taxonomicName class="Arachnida" family="Sparassidae" genus="Pseudopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopoda colubrina" order="Araneae" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="colubrina">Pseudopoda colubrina</taxonomicName>
|
||
Zhao & Li, sp. n., left palp of male holotype. A Prolateral view B Ventral view C Retrolateral view. Scale bar equal for A, B, C.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="4" pageNumber="111">
|
||
<paragraph pageId="4" pageNumber="111">
|
||
Figure 4.
|
||
<taxonomicName class="Arachnida" family="Sparassidae" genus="Pseudopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopoda colubrina" order="Araneae" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="colubrina">Pseudopoda colubrina</taxonomicName>
|
||
Zhao & Li, sp. n., male holotype. A Bulb, ventral view B Bulb, dorsal view C Habitus, dorsal view D Habitus, ventral view. Scale bar equal for A, B.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph lastPageId="5" lastPageNumber="112" pageId="4" pageNumber="111">
|
||
Female resembles
|
||
<taxonomicName lsidName="P. hyatti" pageId="4" pageNumber="111" rank="species" species="hyatti">P. hyatti</taxonomicName>
|
||
<normalizedToken originalValue="Jäger">Jaeger</normalizedToken>
|
||
, 2001 (see
|
||
<bibRefCitation author="Jaeger, P" journalOrPublisher="Courier Forschungsinstitut Senckenberg" pageId="33" pageNumber="140" pagination="1 - 136" title="Diversitaet der Riesenkrabbenspinnen im Himalaya - die Radiation zweier Gattungen in den Schneetropen (Araneae, Sparassidae, Heteropodinae)." volume="232" year="2001">
|
||
<normalizedToken originalValue="Jäger">Jaeger</normalizedToken>
|
||
2001
|
||
</bibRefCitation>
|
||
: 72, figs 41
|
||
<normalizedToken originalValue="j–m">j-m</normalizedToken>
|
||
, 84) by: 1. posterior part of lateral lobes surpassing the epigastric furrow; 2. loops of internal duct system mainly winding near the central axis, running transversally (Figure 5A, B, E).
|
||
<pageBreakToken pageId="5" pageNumber="112" start="start">It</pageBreakToken>
|
||
can be distinguished from the latter by the following combination of characters: 1. copulatory opening located at the middle to posterior part of epigyne (Figure 5A; located near the anterior margin of lateral lobe in
|
||
<taxonomicName lsidName="P. hyatti" pageId="5" pageNumber="112" rank="species" species="hyatti">P. hyatti</taxonomicName>
|
||
); 2. anterior margin of epigynal field truncated, anterior bands absent (Figure 5A; anterior margin of epigynal field trilobate with short anterior bands in
|
||
<taxonomicName lsidName="P. hyatti" pageId="5" pageNumber="112" rank="species" species="hyatti">P. hyatti</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption pageId="5" pageNumber="112">
|
||
<paragraph pageId="5" pageNumber="112">
|
||
Figure 5.
|
||
<taxonomicName class="Arachnida" family="Sparassidae" genus="Pseudopoda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudopoda colubrina" order="Araneae" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="colubrina">Pseudopoda colubrina</taxonomicName>
|
||
Zhao & Li, sp. n., paratype female. A Epigyne, ventral view B Vulva, dorsal view C Habitus, dorsal view D Habitus, ventral view E Schematic course of internal duct system, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112" type="description">
|
||
<paragraph pageId="5" pageNumber="112">Description.</paragraph>
|
||
<paragraph lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112">
|
||
Male (holotype). Body length 8.8, DS length 4.3, DS width 4.1, OS length 4.5, OS width 4.0. Eyes: AME 0.17, ALE 0.34, PME 0.29, PLE 0.28, AME-AME 0.23, AME-ALE 0.09, PME-PME 0.16, PME-PLE 0.33, AME-PME 0.37, ALE-PLE 0.32, CH AME 0.60, CH ALE 0.39. Leg formula: II-I-IV-III. Spination: palp 131, 101, 2101; legs: femur I-II 323, III 322, IV 321; patella I-IV 101; tibia I-II 2026, III 2126, IV 2026; metatarsus I-II 1014, III 2025, IV 3036. Measurements of palp and legs: palp 6.3 (2.0, 0.9, 1.1, -, 2.3), leg I 20.4 (5.8, 2.0, 5.9, 4.9, 1.8), leg II 22.1 (6.1, 2.3, 6.4, 5.4, 1.9), leg III 16.7 (5.0, 1.8, 4.5, 4.0, 1.4), leg IV 19.6 (5.6, 1.7,
|
||
<pageBreakToken pageId="6" pageNumber="113" start="start">5.0</pageBreakToken>
|
||
, 5.5, 1.8). Promargin of chelicerae with three teeth, retromargin with four teeth. Cheliceral furrow with ca. 21 denticles.
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">
|
||
Palp as in diagnosis. Cymbium slender, tip slightly bent prolaterally, with a distinct retrobasal bulge. RTA arising basally from tibia, simple but elongated, blunt in retrolateral view (Figure 3
|
||
<normalizedToken originalValue="A–C">A-C</normalizedToken>
|
||
). Sperm duct bending near the top of tegulum, then running submarginally retrolaterally in tegulum. Basal part of embolus with distinct double rims. Tip of embolus pointing distally prolaterally. Conductor completely absent, like a few other species (e.g.
|
||
<taxonomicName lsidName="P. ashcharya" pageId="6" pageNumber="113" rank="species" species="ashcharya">P. ashcharya</taxonomicName>
|
||
<normalizedToken originalValue="Jäger">Jaeger</normalizedToken>
|
||
& Kulkarni, 2016) (Figure 4A, B).
|
||
</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">Coloration in ethanol: carapace yellowish. Radial furrows and fovea darker brown. Dorsal opisthosoma reddish brown. Legs yellowish, with randomly distributed brown dots (Figure 4C, D).</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">Female (paratype). Body length 10.0, DS length 4.9, DS width 4.3, OS length 5.1, OS width 3.2. Eyes: AME 0.22, ALE 0.33, PME 0.25, PLE 0.31, AME-AME 0.20, AME-ALE 0.04, PME-PME 0.20, PME-PLE 0.41, AME-PME 0.40, ALE-PLE 0.37, CH AME 0.51, CH ALE 0.41. Leg formula: II-IV-I-III. Spination: palp 131, 101, 2121, 1014; legs: femur I-II 323, III 322, IV 321; patella I-IV 101; tibia I-IV 2026; metatarsus I 1014, II-III 2024, IV 3036. Measurements of palp and legs: palp 5.3 (1.6, 0.7, 1.0, -, 2.0), leg I 17.6 (4.9, 2.0, 5.0, 4.2, 1.5), leg II 19.2 (5.5, 2.2, 5.5, 4.4, 1.6), leg III 14.9 (4.4, 1.8, 3.9, 3.4, 1.4), leg IV 18.1 (5.5, 1.8, 4.5, 4.6, 1.7). Promargin of chelicerae with three teeth, retromargin with four teeth. Cheliceral furrow with ca. 20 denticles.</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">Epigyne as in diagnosis. Epigynal field with nearly equal length in transverse and longitudinal axis. Lateral lobes longer in longitudinal axis. Median margin of lateral lobes touching each other medially. Internal duct system with loops looming through the lateral lobes in ventral view (Figure 5A). A pair of small appendages present (Figure 5E).</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">Coloration in ethanol: As in male, but generally darker with more dots and patches (Figure 5C, D).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="113" type="distribution">
|
||
<paragraph pageId="6" pageNumber="113">Distribution.</paragraph>
|
||
<paragraph pageId="6" pageNumber="113">Known only from the type locality.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |