428 lines
56 KiB
XML
428 lines
56 KiB
XML
<document id="F538ED1CE5504867BF5C3F837533D336" ID-CLB-Dataset="56018" ID-DOI="10.11646/zootaxa.5125.5.2" ID-GBIF-Dataset="6a677d3b-0f91-4f11-8cf2-64f4ad2a47d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6457259" ID-ZooBank="39EBA798-29B1-4450-9085-A4FB6D1DB331" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649834539484" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A., Semenchenko, Alexander A. & Palatov, Dmitry M." docDate="2022" docId="721787E39444FFF9F5C9600AFD016EF7" docLanguage="en" docName="zootaxa.5125.5.2.pdf" docOrigin="Zootaxa 5125 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diamesa kownackii Makarchenko & Semenchenko & Palatov 2022, sp. nov." docType="treatment" docUuid="4601893D-170C-427C-8F19-AEED2CA46FB2" docUuidSource="ZooBank" docVersion="7" lastPageNumber="489" masterDocId="8E2EFF9B9440FFFFF55E6618FFBE6F7D" masterDocTitle="Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of new species. I. Subgroups steinboecki and longipes" masterLastPageNumber="512" masterPageNumber="483" pageNumber="487" updateTime="1699328882303" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="57270C1DB72913A1E8321BE8206846C6">
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<mods:title id="524AFF32E31C93D21AE679F3E3D7BA26">Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of new species. I. Subgroups steinboecki and longipes</mods:title>
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<mods:name id="EA6D984BA9A83A2CF588FAFCE06FFFD4" type="personal">
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<mods:roleTerm id="F98ACA4CE27DF4C40F51B127C7187ED9">Author</mods:roleTerm>
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<mods:namePart id="4DA26B045A254BD4E661227A82973042">Makarchenko, Eugenyi A.</mods:namePart>
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<mods:affiliation id="88E9DDFA721504832410C641D20F2E5F">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia</mods:affiliation>
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<mods:namePart id="DB4478A8827A5472E53383812BC75BE4">Semenchenko, Alexander A.</mods:namePart>
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<mods:affiliation id="4220D78D1B372A1928B44C5750496741">Laboratory of Ecology and Evolutionary Biology of Aquatic Organisms, Far Eastern Federal University, Suhanova St. 8, 690950 Vladivostok, Russia</mods:affiliation>
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<mods:namePart id="E409B9FEBA75FEDF48595B5D39F85B3A">Palatov, Dmitry M.</mods:namePart>
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<mods:nameIdentifier id="7054A2448AE578819C860CABC2D0F038" type="ORCID">0000-0002-8826-9316</mods:nameIdentifier>
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<mods:affiliation id="1527FDC3DC4E0F2540D9624A9A34D2F9">A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninskij prosp. 33, 119071 Moscow, Russia triops @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8826 - 9316</mods:affiliation>
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<mods:nameIdentifier id="1836025B5C83F9627F348C2207B5BF58" type="email">triops@yandex.ru</mods:nameIdentifier>
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<mods:title id="7F09A116303B2AD751192BB5FC79FA9D">Zootaxa</mods:title>
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<mods:part id="2DEE13592205CA955A918302624C7AD4">
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<mods:date id="08A2306CD0BE6526D1547028E433D681">2022</mods:date>
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<mods:number id="9EBEE071AA368421786C611986D7EA1C">2022-04-13</mods:number>
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<mods:number id="3B4D7AB47DBCAE646B38A773C88799C3">5125</mods:number>
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<mods:number id="9F4DA582857885891C3D85070F2A1EA3">5</mods:number>
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<treatment id="721787E39444FFF9F5C9600AFD016EF7" ID-DOI="http://doi.org/10.5281/zenodo.6458131" ID-GBIF-Taxon="194591612" ID-Zenodo-Dep="6458131" LSID="urn:lsid:zoobank.org:act:4601893D-170C-427C-8F19-AEED2CA46FB2" httpUri="http://treatment.plazi.org/id/721787E39444FFF9F5C9600AFD016EF7" lastPageId="6" lastPageNumber="489" pageId="4" pageNumber="487">
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<emphasis id="C8CAEAE79444FFFBF5C9600AFD196951" bold="true" box="[151,679,1553,1580]" pageId="4" pageNumber="487">
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<taxonomicName id="3DBE4D769444FFFBF5C9600AFD836951" ID-CoL="9J2PK" authority="Makarchenko" authorityName="Makarchenko & Semenchenko & Palatov" authorityYear="2022" box="[151,573,1553,1580]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="kownackii" status="sp. nov.">
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<emphasis id="C8CAEAE79444FFFBF5C9600AFE3A6951" bold="true" box="[151,388,1553,1580]" italics="true" pageId="4" pageNumber="487">Diamesa kownackii</emphasis>
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Makarchenko
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</taxonomicName>
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,
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<taxonomicNameLabel id="D3F9579C9444FFFBF714600AFD196951" box="[586,679,1554,1580]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="FA0136F59444FFFBF5C9602EFBD6692D" blockId="4.[151,1128,1553,1653]" box="[151,1128,1590,1616]" pageId="4" pageNumber="487">
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http://zoobank.org/NomenclaturalActs/
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<subSubSection id="B2A4657E9444FFFBF5C96042FE206908" box="[151,414,1626,1653]" pageId="4" pageNumber="487" type="description">
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<paragraph id="FA0136F59444FFFBF5C96042FE206908" blockId="4.[151,1128,1553,1653]" box="[151,414,1626,1653]" pageId="4" pageNumber="487">
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(
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<figureCitation id="62852A709444FFFBF5C16042FEA76909" box="[159,281,1626,1652]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="4" pageNumber="487">Figs. 9–14</figureCitation>
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,
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<figureCitation id="62852A709444FFFBF47B6042FEFD6909" box="[293,323,1626,1652]" captionStart="FIGURES 30–36" captionStartId="13.[151,264,1503,1527]" captionTargetBox="[151,1436,758,1472]" captionTargetId="figure-286@13.[151,1436,758,1480]" captionTargetPageId="13" captionText="FIGURES 30–36. Wings of Diamesa dragani sp. nov. (30), D. kownackii sp. nov. (31), D. maisaraensis sp. nov. (32), D. marinskiyi sp. nov. (33) and D. steinboecki Goetghebuer from Swiss Alps (34), French Alps (35) and Pamir Mountains (36)." figureDoi="http://doi.org/10.5281/zenodo.6457271" httpUri="https://zenodo.org/record/6457271/files/figure.png" pageId="4" pageNumber="487">31</figureCitation>
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,
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<figureCitation id="62852A709444FFFBF4116042FE296908" box="[335,407,1626,1653]" captionStart="FIGURES 68–74" captionStartId="23.[151,264,1617,1641]" captionTargetBox="[151,1435,181,1592]" captionTargetId="figure-18@23.[151,1435,181,1593]" captionTargetPageId="23" captionText="FIGURES 68–74. Type localities of Diamesa dragani sp. nov. (68), D. kownackii sp. nov. (69–70), D. maisaraensis sp. nov. (71), D. marinskiyi sp. nov. (72), D. steinboecki Goetghebuer (73) and D. sakartvella Kownacki et Kownacka (74). 68, unnamed stream of Bolshoi On River basin, alt. 2063 m a. sea l., Sayan Mountains (photo by S.V. Dragan); 69, unnamed stream of Somnitelnaya River basin, Wrangel Island (photo by E.A. Makarchenko); 70, Somnitelnaya River, upper stream, alt. 205–210 m a. sea l., Wrangel Island (photo by O.A. Khruleva); 71, Maisara River near Maisara Pass, alt. 4168 m a. sea l., Pamir Mountains (photo by D.M. Palatov); 72, Zienzan River, Tien Shan Mountains, alt. 1923 m a. sea l., Xinjiang Uygur Autonomous Region, China (photo by D.M. Palatov); 73, Balm stream, Massif of Aiguilles–Rouges, French Alps (photo by Pierre Clévenot); 74, Skazdon River near the Tsey (Tseyskoe gorge), alt. 1943 m a. sea l., North Caucasus (photo by D.M. Palatov)." figureDoi="http://doi.org/10.5281/zenodo.6457281" httpUri="https://zenodo.org/record/6457281/files/figure.png" pageId="4" pageNumber="487">69–70</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="B2A4657E9444FFFBF5C96087FAC369CB" box="[151,1405,1694,1718]" pageId="4" pageNumber="487" type="reference_group">
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<paragraph id="FA0136F59444FFFBF5C96087FAC369CB" blockId="4.[151,1405,1694,1718]" box="[151,1405,1694,1718]" pageId="4" pageNumber="487">
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<treatmentCitationGroup id="DAAE11DB9444FFFBF5C96087FAC369CB" box="[151,1405,1694,1718]" pageId="4" pageNumber="487">
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<taxonomicName id="3DBE4D769444FFFBF5C96087FE4869CB" ID-CoL="359H8" authority="Goetghebuer" authorityName="Goetghebuer" authorityYear="1933" box="[151,502,1694,1718]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="steinboecki">
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<emphasis id="C8CAEAE79444FFFBF5C96087FED469CB" box="[151,362,1694,1718]" italics="true" pageId="4" pageNumber="487">Diamesa steinboecki</emphasis>
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Goetghebuer
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</taxonomicName>
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;
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<treatmentCitation id="7B1F10E49444FFFBF75E6086FCA469CB" author="Makarchenko, E. A. & Levanidova I. M. & Zhiltsova, L. A." box="[512,794,1694,1718]" page="6" pageId="4" pageNumber="487" year="1980">
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<bibRefCitation id="9E2F4B049444FFFBF75E6086FCA469CB" author="Makarchenko, E. A. & Levanidova I. M. & Zhiltsova, L. A." box="[512,794,1694,1718]" pageId="4" pageNumber="487" pagination="3 - 12" refId="ref19176" refString="Makarchenko, E. A., Levanidova I. M. & Zhiltsova, L. A. (1980) Preliminary data on the fauna of aquatic invertebrates of Wrangel Island. In: Freshwater fauna of the Far East. DVNC AN SSSR Press, Vladivostok, pp. 3 - 12. [in Russian]" type="book chapter" year="1980">
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||
Makarchenko
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||
<emphasis id="C8CAEAE79444FFFBF7CC6087FD0069CB" box="[658,702,1694,1718]" italics="true" pageId="4" pageNumber="487">et al</emphasis>
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, 1980: 6
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</bibRefCitation>
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</treatmentCitation>
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; 1981: 96; 1985: 71; 2006: 261, 472, 614, misidentification.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="FA0136F59444FFFBF5C960FDFDA56887" blockId="4.[151,1436,1764,2043]" pageId="4" pageNumber="487">
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<emphasis id="C8CAEAE79444FFFBF5C960FDFEFA6983" bold="true" box="[151,324,1764,1791]" pageId="4" pageNumber="487">Type material.</emphasis>
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<materialsCitation id="4AD63CA89444FFFBF41760FCFE32683B" ID-GBIF-Occurrence="3751071304" collectingDate="1978-07-22" collectorName="E. Makarchenko." country="Russia" county="Autonomous Region" elevation="208" latitude="70.99578" location="Somnitelnaya River" longLatPrecision="1" longitude="-179.53912" municipality="Wrangel Island" pageId="4" pageNumber="487" specimenCount="1" stateProvince="Chukotka" typeStatus="holotype">
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<typeStatus id="250588579444FFFBF41760FCFE0A6983" box="[329,436,1764,1790]" pageId="4" pageNumber="487" type="holotype">Holotype</typeStatus>
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, adult male,
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:
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<collectingCounty id="13604E799444FFFBF67F60FDFBB56983" box="[801,1035,1764,1791]" pageId="4" pageNumber="487">Autonomous Region</collectingCounty>
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,
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<collectingMunicipality id="1A65AC8F9444FFFBF14A60FCFB7E6983" box="[1044,1216,1764,1790]" pageId="4" pageNumber="487">Wrangel Island</collectingMunicipality>
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, unnamed stream in upper part of
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<location id="FF61602E9444FFFBF46E6110FDAF685F" LSID="urn:lsid:plazi:treatment:721787E39444FFF9F5C9600AFD016EF7:FF61602E9444FFFBF46E6110FDAF685F" box="[304,529,1800,1826]" country="Russia" county="Autonomous Region" latitude="70.99578" longLatPrecision="1" longitude="-179.53912" municipality="Wrangel Island" name="Somnitelnaya River" pageId="4" pageNumber="487" stateProvince="Chukotka">Somnitelnaya River</location>
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, altitude
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<elevation id="7193D1C69444FFFBF7266110FC13685F" box="[632,941,1800,1827]" metricMagnitude="2" metricUnit="m" metricValue="2.075" metricValueMax="2.1" metricValueMin="2.05" pageId="4" pageNumber="487" unit="m" value="207.5" valueMax="210.0" valueMin="205.0">
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above sea level
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</elevation>
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,
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<date id="8E0010359444FFFBF6E66111FBFA685E" box="[952,1092,1801,1827]" pageId="4" pageNumber="487" value="1978-07-22">
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<collectingDate id="9E44E9DD9444FFFBF6E66111FBFA685E" box="[952,1092,1801,1827]" pageId="4" pageNumber="487" value="1978-07-22">22.VII.1978</collectingDate>
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</date>
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,
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<geoCoordinate id="9F8A50329444FFFBF10E6111FB5B685E" box="[1104,1253,1800,1827]" degrees="70.995783" direction="north" orientation="latitude" pageId="4" pageNumber="487" precision="1" value="70.99578">N 70.995783</geoCoordinate>
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,
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<geoCoordinate id="9F8A50329444FFFBF1AE6111FA26685E" box="[1264,1432,1800,1827]" degrees="179.539117" direction="west" orientation="longitude" pageId="4" pageNumber="487" precision="1" value="-179.53912">W 179.539117</geoCoordinate>
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||
, leg.
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<collectorName id="574B53239444FFFBF5976135FE32683B" box="[201,396,1836,1863]" pageId="4" pageNumber="487">E. Makarchenko.</collectorName>
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</materialsCitation>
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<typeStatus id="250588579444FFFBF4CA6135FDBB683A" box="[404,517,1837,1863]" pageId="4" pageNumber="487" type="paratype">Paratypes</typeStatus>
|
||
:
|
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<specimenCount id="ECB8FD7C9444FFFBF74C6134FDD1683B" box="[530,623,1836,1863]" pageId="4" pageNumber="487" type="adult">52 adult</specimenCount>
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||
males,
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||
<materialsCitation id="4AD63CA89444FFFBF7976135FE926816" ID-GBIF-Occurrence="3751071323" collectingDate="1978-07-22" collectingDateMax="1978-08-15" collectingDateMin="1978-07-22" collectorName="E. Makarchenko." country="Russia" county="Autonomous Region" elevation="208" latitude="70.99578" location="Somnitelnaya River" longLatPrecision="1" longitude="-179.53912" municipality="Wrangel Island" pageId="4" pageNumber="487" specimenCount="51" specimenCount-larva="28" specimenCount-pupa="23" stateProvince="Chukotka" typeStatus="paratype">
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<specimenCount id="ECB8FD7C9444FFFBF7976135FC8A683A" box="[713,820,1837,1863]" pageId="4" pageNumber="487" type="pupa">23 pupae</specimenCount>
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,
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<specimenCount id="ECB8FD7C9444FFFBF61F6135FC13683B" box="[833,941,1836,1863]" pageId="4" pageNumber="487" type="larva">28 larvae</specimenCount>
|
||
, the same data as holotype, except
|
||
<date id="8E0010359444FFFBF01A6135FE926816" pageId="4" pageNumber="487" value="1978-07-22" valueMax="1978-08-15" valueMin="1978-07-22">
|
||
<collectingDate id="9E44E9DD9444FFFBF01A6135FE926816" pageId="4" pageNumber="487" value="1978-07-22" valueMax="1978-08-15" valueMin="1978-07-22">22.VII– 15.VIII.1978</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="ECB8FD7C9444FFFBF4666149FE3A6817" box="[312,388,1872,1899]" pageId="4" pageNumber="487" type="adult">9 adult</specimenCount>
|
||
males,
|
||
<materialsCitation id="4AD63CA89444FFFBF4846149FB4B6816" ID-GBIF-Occurrence="3751071305" box="[474,1269,1872,1899]" collectingDate="1979-08-07" collectingDateMax="1979-08-10" collectingDateMin="1979-08-07" collectorName="E. Makarchenko." country="Russia" county="Autonomous Region" elevation="208" latitude="70.99578" location="Somnitelnaya River" longLatPrecision="1" longitude="-179.53912" municipality="Wrangel Island" pageId="4" pageNumber="487" specimenCount="47" specimenCount-larva="33" specimenCount-pupa="14" stateProvince="Chukotka" typeStatus="paratype">
|
||
<specimenCount id="ECB8FD7C9444FFFBF4846149FDFD6816" box="[474,579,1873,1899]" pageId="4" pageNumber="487" type="pupa">14 pupae</specimenCount>
|
||
,
|
||
<specimenCount id="ECB8FD7C9444FFFBF7136149FD096817" box="[589,695,1872,1899]" pageId="4" pageNumber="487" type="larva">33 larvae</specimenCount>
|
||
, the same data as holotype, except
|
||
<date id="8E0010359444FFFBF1636149FB4B6816" box="[1085,1269,1873,1899]" pageId="4" pageNumber="487" value="1979-08-07" valueMax="1979-08-10" valueMin="1979-08-07">
|
||
<collectingDate id="9E44E9DD9444FFFBF1636149FB4B6816" box="[1085,1269,1873,1899]" pageId="4" pageNumber="487" value="1979-08-07" valueMax="1979-08-10" valueMin="1979-08-07">7–10.VIII. 1979</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4AD63CA89444FFFBF05F6148FB3568F3" ID-GBIF-Occurrence="3751071324" collectingDate="1978-08-17" collectorName="E. Makarchenko" country="Russia" county="Autonomous Region" elevation="208" latitude="70.99578" location="Khishnikov River" longLatPrecision="1" longitude="-179.53912" municipality="Wrangel Island" pageId="4" pageNumber="487" specimenCount="5" specimenCount-adult="5" stateProvince="Chukotka" typeStatus="paratype">
|
||
<specimenCount id="ECB8FD7C9444FFFBF05F6148FAF36817" box="[1281,1357,1872,1899]" pageId="4" pageNumber="487" type="adult">5 adult</specimenCount>
|
||
males, the same data as holotype, except
|
||
<location id="FF61602E9444FFFBF77E616CFD5568F3" LSID="urn:lsid:plazi:treatment:721787E39444FFF9F5C9600AFD016EF7:FF61602E9444FFFBF77E616CFD5568F3" box="[544,747,1908,1934]" name="Khishnikov River" pageId="4" pageNumber="487">Khishnikov River</location>
|
||
,
|
||
<date id="8E0010359444FFFBF7A6616DFC3368F2" box="[760,909,1909,1935]" pageId="4" pageNumber="487" value="1978-08-17">
|
||
<collectingDate id="9E44E9DD9444FFFBF7A6616DFC3368F2" box="[760,909,1909,1935]" pageId="4" pageNumber="487" value="1978-08-17">17.VIII.1978</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="574B53239444FFFBF692616DFB3568F3" box="[972,1163,1908,1935]" pageId="4" pageNumber="487">E. Makarchenko</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4AD63CA89444FFFBF1C6616DFDA56887" ID-GBIF-Occurrence="3751071301" collectingDate="1980-07-29" collectorName="E. Makarchenko." country="Russia" county="Autonomous Region" elevation="300" latitude="62.962223" location="Koryak Highlands" longLatPrecision="1" longitude="171.80583" municipality="Anadyr District" pageId="4" pageNumber="487" specimenCount="2" specimenCount-adult="2" stateProvince="Chukotka" typeStatus="paratype">
|
||
<specimenCount id="ECB8FD7C9444FFFBF1C6616DFB5968F3" box="[1176,1255,1908,1935]" pageId="4" pageNumber="487" type="adult">2 adult</specimenCount>
|
||
males,
|
||
<collectingRegion id="387AF8179444FFFBF01E616CFF0F68CF" country="Russia" name="Chukotka" pageId="4" pageNumber="487">Chukotka</collectingRegion>
|
||
<collectingCounty id="13604E799444FFFBF5E46181FE1768CF" box="[186,425,1944,1971]" pageId="4" pageNumber="487">Autonomous Region</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="1A65AC8F9444FFFBF4EB6180FDD468CF" box="[437,618,1944,1970]" pageId="4" pageNumber="487">Anadyr District</collectingMunicipality>
|
||
, northeastern spurs of the
|
||
<location id="FF61602E9444FFFBF6FE6180FBCA68CF" LSID="urn:lsid:plazi:treatment:721787E39444FFF9F5C9600AFD016EF7:FF61602E9444FFFBF6FE6180FBCA68CF" box="[928,1140,1944,1970]" country="Russia" county="Autonomous Region" latitude="62.962223" longLatPrecision="1" longitude="171.80583" municipality="Anadyr District" name="Koryak Highlands" pageId="4" pageNumber="487" stateProvince="Chukotka">Koryak Highlands</location>
|
||
, unnamed stream of
|
||
<location id="FF61602E9444FFFBF0376181FECD68AB" LSID="urn:lsid:plazi:treatment:721787E39444FFF9F5C9600AFD016EF7:FF61602E9444FFFBF0376181FECD68AB" country="Russia" county="Autonomous Region" latitude="62.962223" longLatPrecision="1" longitude="171.80583" municipality="Anadyr District" name="Gytgylveirgytgyn Lake" pageId="4" pageNumber="487" stateProvince="Chukotka">Gytgylveirgytgyn Lake</location>
|
||
basin (
|
||
<location id="FF61602E9444FFFBF49B61A4FDD168AB" LSID="urn:lsid:plazi:treatment:721787E39444FFF9F5C9600AFD016EF7:FF61602E9444FFFBF49B61A4FDD168AB" box="[453,623,1980,2006]" country="Russia" county="Autonomous Region" latitude="62.962223" longLatPrecision="1" longitude="171.80583" municipality="Anadyr District" name="Velikaya River" pageId="4" pageNumber="487" stateProvince="Chukotka">Velikaya River</location>
|
||
basin),
|
||
<date id="8E0010359444FFFBF79461A5FCEB68AA" box="[714,853,1981,2007]" pageId="4" pageNumber="487" value="1980-07-29">
|
||
<collectingDate id="9E44E9DD9444FFFBF79461A5FCEB68AA" box="[714,853,1981,2007]" pageId="4" pageNumber="487" value="1980-07-29">29.VII.1980</collectingDate>
|
||
</date>
|
||
, altitude
|
||
<emphasis id="C8CAEAE79444FFFBF6E361A5FC6968AB" box="[957,983,1981,2006]" italics="true" pageId="4" pageNumber="487">ca</emphasis>
|
||
<elevation id="7193D1C69444FFFBF68161A5FB6368AB" box="[991,1245,1980,2007]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="487" unit="m" value="300.0">
|
||
<quantity id="3D469B109444FFFBF68161A5FB9968AA" box="[991,1063,1981,2007]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="487" unit="m" value="300.0">300 m</quantity>
|
||
above sea level
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="9F8A50329444FFFBF1B761A5FAC168AB" box="[1257,1407,1980,2007]" degrees="62.962222" direction="north" orientation="latitude" pageId="4" pageNumber="487" precision="1" value="62.962223">N 62.962222</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9F8A50329444FFFBF0D561A5FEA16886" degrees="171.805833" direction="east" orientation="longitude" pageId="4" pageNumber="487" precision="1" value="171.80583">E 171.805833</geoCoordinate>
|
||
, leg.
|
||
<collectorName id="574B53239444FFFBF40461F9FDA56887" box="[346,539,2016,2043]" pageId="4" pageNumber="487">E. Makarchenko.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2A4657E9445FFFAF599668EFA8B6FA8" pageId="5" pageNumber="488" type="etymology">
|
||
<paragraph id="FA0136F59445FFFAF599668EFA8B6FA8" blockId="5.[151,1437,150,933]" pageId="5" pageNumber="488">
|
||
<emphasis id="C8CAEAE79445FFFAF599668EFE1A6FCD" bold="true" box="[199,420,150,176]" pageId="5" pageNumber="488">Derivatio nominis.</emphasis>
|
||
The species is named in honor of the Polish chironomid taxonomist and hydrobiologist Dr. Andrzej Kownacki, a leading specialist on Caucasian
|
||
<taxonomicName id="3DBE4D769445FFFAF7AD66A2FCC46FA9" authorityName="Kieffer" authorityYear="1922" box="[755,890,186,212]" class="Insecta" family="Chironomidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="subFamily" subFamily="Diamesinae">Diamesinae</taxonomicName>
|
||
, namely of
|
||
<taxonomicName id="3DBE4D769445FFFAF6A266A3FB5B6FA9" authorityName="Goetghebuer" authorityYear="1933" box="[1020,1253,186,212]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="steinboecki">
|
||
<emphasis id="C8CAEAE79445FFFAF6A266A3FB5B6FA9" box="[1020,1253,186,212]" italics="true" pageId="5" pageNumber="488">Diamesa steinboecki</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2A4657E9445FFFAF59966C6FB216830" pageId="5" pageNumber="488" type="description">
|
||
<paragraph id="FA0136F59445FFFAF59966C6FEEF6F85" blockId="5.[151,1437,150,933]" box="[199,337,222,248]" pageId="5" pageNumber="488">
|
||
<emphasis id="C8CAEAE79445FFFAF59966C6FEEF6F85" bold="true" box="[199,337,222,248]" pageId="5" pageNumber="488">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="FA0136F59445FFFAF599671AFF6E6E3C" blockId="5.[151,1437,150,933]" pageId="5" pageNumber="488">
|
||
<emphasis id="C8CAEAE79445FFFAF599671AFEF96E60" bold="true" box="[199,327,258,285]" italics="true" pageId="5" pageNumber="488">Adult male</emphasis>
|
||
(n = 10, except when otherwise stated). Total length
|
||
<quantity id="3D469B109445FFFAF6F6671AFB8A6E60" box="[936,1076,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="5.2" metricValueMin="4.2" pageId="5" pageNumber="488" unit="mm" value="4.7" valueMax="5.2" valueMin="4.2">4.2–5.2 mm</quantity>
|
||
. Total length/wing length 1.27– 1.75.
|
||
</paragraph>
|
||
<paragraph id="FA0136F59445FFFAF5996752FEA06EF4" blockId="5.[151,1437,150,933]" pageId="5" pageNumber="488">Coloration. Dark brown to black. Head brown. Thorax black. Legs brown. Abdomen dark brown to black. Wings grey.</paragraph>
|
||
<paragraph id="FA0136F59445FFFAF599678AFB0C6DF9" blockId="5.[151,1437,150,933]" pageId="5" pageNumber="488">
|
||
Head (
|
||
<figureCitation id="62852A709445FFFAF44A678AFEE46ED1" box="[276,346,402,428]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Fig. 9</figureCitation>
|
||
). Eyes hairy, reniform. Temporal setae including 0 frontals, 4–9 orbitals, 6–9 verticals and 5–6 postorbitals; frontals absent. Clypeus with 2–5 setae. Antenna with 8 flagellomeres and reduced plume of setae; number and length of these setae on 1–7 flagellomeres respectively: 2–3 (26–28 μm), 1–2 (18–32 μm), 3–5 (40–68 μm), 3 (32–44 μm); terminal flagellomere with 2 setae, 20–24 μm long in subapical and apical area. Length of 1–8 flagellomeres (μm): 105–118, 42–53, 42–46, 29–38, 29–38, 25–34, 29–38, 130–160; AR 0.38–0.50. Palpomere length (μm): 330–348, 64–84, 105–126, 78–92, 109–134. Palpomere
|
||
<quantity id="3D469B109445FFFAF6D1645EFC076D1D" box="[911,953,582,608]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="488" unit="in" value="3.0">3 in</quantity>
|
||
distal part with sensilla capitata with diameter 16–20 μm. Head width/palpal length 1.20–1.25. Antennal length/palpal length 1.12–1.40.
|
||
</paragraph>
|
||
<paragraph id="FA0136F59445FFFAF5996496FBAF6DB0" blockId="5.[151,1437,150,933]" pageId="5" pageNumber="488">Thorax. Antepronotum with 11–18 ventrolateral setae, 48–68 μm long. Dorsocentrals 7–12, 68–84 μm long; prealars 5–10, 60–76 μm long. Scutellum with 32–42 setae. AAII without setae.</paragraph>
|
||
<paragraph id="FA0136F59445FFFAF59964CEFE3D6C45" blockId="5.[151,1437,150,933]" pageId="5" pageNumber="488">
|
||
Wing (
|
||
<figureCitation id="62852A709445FFFAF44664CEFED16D8D" box="[280,367,726,752]" captionStart="FIGURES 30–36" captionStartId="13.[151,264,1503,1527]" captionTargetBox="[151,1436,758,1472]" captionTargetId="figure-286@13.[151,1436,758,1480]" captionTargetPageId="13" captionText="FIGURES 30–36. Wings of Diamesa dragani sp. nov. (30), D. kownackii sp. nov. (31), D. maisaraensis sp. nov. (32), D. marinskiyi sp. nov. (33) and D. steinboecki Goetghebuer from Swiss Alps (34), French Alps (35) and Pamir Mountains (36)." figureDoi="http://doi.org/10.5281/zenodo.6457271" httpUri="https://zenodo.org/record/6457271/files/figure.png" pageId="5" pageNumber="488">Fig. 31</figureCitation>
|
||
). Length
|
||
<quantity id="3D469B109445FFFAF4BC64CEFD356D8C" box="[482,651,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.54" metricValueMin="2.86" pageId="5" pageNumber="488" unit="mm" value="3.2" valueMax="3.54" valueMin="2.86">2.86–3.54 mm</quantity>
|
||
, width
|
||
<quantity id="3D469B109445FFFAF7BB64CEFC316D8C" box="[741,911,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.015" metricValueMax="1.1" metricValueMin="0.93" pageId="5" pageNumber="488" unit="mm" value="1.015" valueMax="1.1" valueMin="0.93">0.93–1.10 mm</quantity>
|
||
. Costal extension 90–98 µm long. Anal lobe slightly reduced, rounded. Squama with 4–16 setae, 36–44 μm long. R and R
|
||
<subScript id="663A34B09445FFFAF1536515FBAB6C66" attach="left" box="[1037,1045,781,795]" fontSize="6" pageId="5" pageNumber="488">1</subScript>
|
||
with 10–16 setae, R
|
||
<subScript id="663A34B09445FFFAF05D6514FAA36C67" attach="left" box="[1283,1309,780,794]" fontSize="6" pageId="5" pageNumber="488">4+5</subScript>
|
||
with 8–17. RM/MCu 1.33–2.27.
|
||
</paragraph>
|
||
<paragraph id="FA0136F59445FFFAF599655BFEEB6CD9" blockId="5.[151,1437,150,933]" pageId="5" pageNumber="488">
|
||
Legs. Spur of front tibia 50–59 µm long. Spurs of mid tibia 46–59 µm and 42–63 µm long. Spurs of hind tibia 88–118 µm and 55–71 µm long. Hind tibial comb with 18–25 setae. Length (μm) and proportions of leg segments are as in
|
||
<tableCitation id="B73C034E9445FFFAF5A56592FEEF6CD9" box="[251,337,906,932]" captionStart="TABLE 2" captionStartId="5.[151,239,980,1004]" captionTargetPageId="5" captionText="TABLE 2. Lengths (in µm) and proportions of leg segments of Diamesa kownackii sp. nov., male (n=12)" httpUri="http://table.plazi.org/id/AEC1667D9445FFFAF5C965CCFB436C91" pageId="5" pageNumber="488" tableUuid="AEC1667D9445FFFAF5C965CCFB436C91">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="AEC1667D9445FFFAF5C965CCFB436C91" ID-Table-UUID="AEC1667D9445FFFAF5C965CCFB436C91" box="[151,1277,980,1004]" httpUri="http://table.plazi.org/id/AEC1667D9445FFFAF5C965CCFB436C91" pageId="5" pageNumber="488" startId="5.[151,239,980,1004]" targetBox="[151,1368,1020,1405]" targetIsTable="true" targetPageId="5">
|
||
<paragraph id="FA0136F59445FFFAF5C965CCFB436C91" blockId="5.[151,1277,980,1004]" box="[151,1277,980,1004]" pageId="5" pageNumber="488">
|
||
<emphasis id="C8CAEAE79445FFFAF5C965CCFEB46C91" bold="true" box="[151,266,980,1004]" pageId="5" pageNumber="488">TABLE 2.</emphasis>
|
||
Lengths (in µm) and proportions of leg segments of
|
||
<taxonomicName id="3DBE4D769445FFFAF66165CDFBAF6C91" authority="Makarchenko & Semenchenko & Palatov, 2022" authorityName="Makarchenko & Semenchenko & Palatov" authorityYear="2022" box="[831,1041,980,1004]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="kownackii" status="sp. nov.">
|
||
<emphasis id="C8CAEAE79445FFFAF66165CDFBAF6C91" box="[831,1041,980,1004]" italics="true" pageId="5" pageNumber="488">Diamesa kownackii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C8CAEAE79445FFFAF14965CCFBD36C91" bold="true" box="[1047,1133,980,1004]" pageId="5" pageNumber="488">
|
||
<taxonomicNameLabel id="D3F9579C9445FFFAF14965CCFBD36C91" box="[1047,1133,980,1004]" pageId="5" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (n=12)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FA0136F59445FFFAF5B165E4FB426A0A" pageId="5" pageNumber="488">
|
||
<table id="88BEC45594450000F5C965E4FAE66A00" box="[151,1368,1020,1405]" colsContinueIn="9.[151,1365,1597,1981]" gridcols="9" gridrows="9" pageId="5" pageNumber="488">
|
||
<tr id="448E34B794450000F5C965E4FAE66B66" box="[151,1368,1020,1051]" gridrow="0" pageId="5" pageNumber="488" rowspan-0="1" rowspan-7="1">
|
||
<th id="075F5DCB94450000F5B165E4FEDF6B66" box="[239,353,1020,1051]" gridcol="1" gridrow="0" pageId="5" pageNumber="488">fe</th>
|
||
<th id="075F5DCB94450000F4C365E4FDB16B66" box="[413,527,1020,1051]" gridcol="2" gridrow="0" pageId="5" pageNumber="488">ti</th>
|
||
<th id="075F5DCB94450000F71465E4FD056B66" box="[586,699,1020,1051]" gridcol="3" gridrow="0" pageId="5" pageNumber="488">ta1</th>
|
||
<th id="075F5DCB94450000F7A965E4FCEE6B66" box="[759,848,1020,1051]" gridcol="4" gridrow="0" pageId="5" pageNumber="488">ta2</th>
|
||
<th id="075F5DCB94450000F6DC65E4FC436B66" box="[898,1021,1020,1051]" gridcol="5" gridrow="0" pageId="5" pageNumber="488">ta3</th>
|
||
<th id="075F5DCB94450000F10C65E4FBD56B66" box="[1106,1131,1020,1051]" gridcol="6" gridrow="0" pageId="5" pageNumber="488">ta4</th>
|
||
<th id="075F5DCB94450000F1A165E4FAE66B66" box="[1279,1368,1020,1051]" gridcol="8" gridrow="0" pageId="5" pageNumber="488">ta 5</th>
|
||
</tr>
|
||
<tr id="448E34B794450000F5C9623CFAE66B3F" box="[151,1368,1060,1090]" gridrow="1" pageId="5" pageNumber="488">
|
||
<th id="075F5DCB94450000F5C9623CFF056B3F" box="[151,187,1060,1090]" gridcol="0" gridrow="1" pageId="5" pageNumber="488">P1</th>
|
||
<td id="075F5DCB94450000F5B1623CFEDF6B3F" box="[239,353,1060,1090]" gridcol="1" gridrow="1" pageId="5" pageNumber="488">1820–2132</td>
|
||
<td id="075F5DCB94450000F4C3623CFDB16B3F" box="[413,527,1060,1090]" gridcol="2" gridrow="1" pageId="5" pageNumber="488">1664–1976</td>
|
||
<td id="075F5DCB94450000F714623CFD056B3F" box="[586,699,1060,1090]" gridcol="3" gridrow="1" pageId="5" pageNumber="488">986–1190</td>
|
||
<td id="075F5DCB94450000F7A9623CFCEE6B3F" box="[759,848,1060,1090]" gridcol="4" gridrow="1" pageId="5" pageNumber="488">374–440</td>
|
||
<td id="075F5DCB94450000F6DC623CFC436B3F" box="[898,1021,1060,1090]" gridcol="5" gridrow="1" pageId="5" pageNumber="488">221–281</td>
|
||
<td id="075F5DCB94450000F10C623CFB6F6B3F" box="[1106,1233,1060,1090]" colspan="2" colspanRight="1" gridcol="6" gridrow="1" pageId="5" pageNumber="488">111–160</td>
|
||
<td id="075F5DCB94450000F1A1623CFAE66B3F" box="[1279,1368,1060,1090]" gridcol="8" gridrow="1" pageId="5" pageNumber="488">145–200</td>
|
||
</tr>
|
||
<tr id="448E34B794450000F5C96255FAE66B17" box="[151,1368,1101,1130]" gridrow="2" pageId="5" pageNumber="488">
|
||
<th id="075F5DCB94450000F5C96255FF056B17" box="[151,187,1101,1130]" gridcol="0" gridrow="2" pageId="5" pageNumber="488">P2</th>
|
||
<td id="075F5DCB94450000F5B16255FEDF6B17" box="[239,353,1101,1130]" gridcol="1" gridrow="2" pageId="5" pageNumber="488">1612–1924</td>
|
||
<td id="075F5DCB94450000F4C36255FDB16B17" box="[413,527,1101,1130]" gridcol="2" gridrow="2" pageId="5" pageNumber="488">1248–1508</td>
|
||
<td id="075F5DCB94450000F7146255FD056B17" box="[586,699,1101,1130]" gridcol="3" gridrow="2" pageId="5" pageNumber="488">527–612</td>
|
||
<td id="075F5DCB94450000F7A96255FCEE6B17" box="[759,848,1101,1130]" gridcol="4" gridrow="2" pageId="5" pageNumber="488">204–255</td>
|
||
<td id="075F5DCB94450000F6DC6255FC436B17" box="[898,1021,1101,1130]" gridcol="5" gridrow="2" pageId="5" pageNumber="488">160–187</td>
|
||
<td id="075F5DCB94450000F10C6255FB6F6B17" box="[1106,1233,1101,1130]" colspan="2" colspanRight="1" gridcol="6" gridrow="2" pageId="5" pageNumber="488">102–128</td>
|
||
<td id="075F5DCB94450000F1A16255FAE66B17" box="[1279,1368,1101,1130]" gridcol="8" gridrow="2" pageId="5" pageNumber="488">136–160</td>
|
||
</tr>
|
||
<tr id="448E34B794450000F5C9626DFAE66BEE" box="[151,1368,1141,1171]" gridrow="3" pageId="5" pageNumber="488">
|
||
<th id="075F5DCB94450000F5C9626DFF056BEE" box="[151,187,1141,1171]" gridcol="0" gridrow="3" pageId="5" pageNumber="488">P 3</th>
|
||
<td id="075F5DCB94450000F5B1626DFEDF6BEE" box="[239,353,1141,1171]" gridcol="1" gridrow="3" pageId="5" pageNumber="488">1872–2236</td>
|
||
<td id="075F5DCB94450000F4C3626DFDB16BEE" box="[413,527,1141,1171]" gridcol="2" gridrow="3" pageId="5" pageNumber="488">1612–2028</td>
|
||
<td id="075F5DCB94450000F714626DFD056BEE" box="[586,699,1141,1171]" gridcol="3" gridrow="3" pageId="5" pageNumber="488">1020–1190</td>
|
||
<td id="075F5DCB94450000F7A9626DFCEE6BEE" box="[759,848,1141,1171]" gridcol="4" gridrow="3" pageId="5" pageNumber="488">493–544</td>
|
||
<td id="075F5DCB94450000F6DC626DFC436BEE" box="[898,1021,1141,1171]" gridcol="5" gridrow="3" pageId="5" pageNumber="488">221–323</td>
|
||
<td id="075F5DCB94450000F10C626DFB6F6BEE" box="[1106,1233,1141,1171]" colspan="2" colspanRight="1" gridcol="6" gridrow="3" pageId="5" pageNumber="488">119–160</td>
|
||
<td id="075F5DCB94450000F1A1626DFAE66BEE" box="[1279,1368,1141,1171]" gridcol="8" gridrow="3" pageId="5" pageNumber="488">153–200</td>
|
||
</tr>
|
||
<tr id="448E34B794450000F5C962A6FAE66BAB" box="[151,1368,1214,1238]" gridrow="4" pageId="5" pageNumber="488">
|
||
<th id="075F5DCB94450000F5C962A6FAE66BAB" box="[151,1368,1214,1238]" colspan="9" colspanRight="8" gridcol="0" gridrow="4" pageId="5" pageNumber="488">continued</th>
|
||
</tr>
|
||
<tr id="448E34B794450000F5C962FFFAE66B82" box="[151,1368,1255,1279]" gridrow="5" pageId="5" pageNumber="488" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
|
||
<td id="075F5DCB94450000F5B162FFFEDF6B82" box="[239,353,1255,1279]" gridcol="1" gridrow="5" pageId="5" pageNumber="488">LR</td>
|
||
<td id="075F5DCB94450000F71462FFFD056B82" box="[586,699,1255,1279]" gridcol="3" gridrow="5" pageId="5" pageNumber="488">BV</td>
|
||
<td id="075F5DCB94450000F6DC62FFFC436B82" box="[898,1021,1255,1279]" gridcol="5" gridrow="5" pageId="5" pageNumber="488">SV</td>
|
||
<td id="075F5DCB94450000F1F162FFFB6F6B82" box="[1199,1233,1255,1279]" gridcol="7" gridrow="5" pageId="5" pageNumber="488">BR</td>
|
||
</tr>
|
||
<tr id="448E34B794450000F5C96317FAE66A50" box="[151,1368,1295,1325]" gridrow="6" pageId="5" pageNumber="488" rowspan-4="1" rowspan-6="1">
|
||
<th id="075F5DCB94450000F5C96317FF056A50" box="[151,187,1295,1325]" gridcol="0" gridrow="6" pageId="5" pageNumber="488">P 1</th>
|
||
<td id="075F5DCB94450000F5B16317FDB16A50" box="[239,527,1295,1325]" colspan="2" colspanRight="1" gridcol="1" gridrow="6" pageId="5" pageNumber="488">0.58–0.63</td>
|
||
<td id="075F5DCB94450000F7146317FD056A50" box="[586,699,1295,1325]" gridcol="3" gridrow="6" pageId="5" pageNumber="488">4.70–5.62</td>
|
||
<td id="075F5DCB94450000F6DC6317FC436A50" box="[898,1021,1295,1325]" gridcol="5" gridrow="6" pageId="5" pageNumber="488">3.32–3.78</td>
|
||
<td id="075F5DCB94450000F1F16317FAE66A50" box="[1199,1368,1295,1325]" colspan="2" colspanRight="1" gridcol="7" gridrow="6" pageId="5" pageNumber="488">0.9–1.9</td>
|
||
</tr>
|
||
<tr id="448E34B794450000F5C9632FFAE66A28" box="[151,1368,1335,1365]" gridrow="7" pageId="5" pageNumber="488" rowspan-4="1" rowspan-6="1">
|
||
<th id="075F5DCB94450000F5C9632FFF056A28" box="[151,187,1335,1365]" gridcol="0" gridrow="7" pageId="5" pageNumber="488">P2</th>
|
||
<td id="075F5DCB94450000F5B1632FFDB16A28" box="[239,527,1335,1365]" colspan="2" colspanRight="1" gridcol="1" gridrow="7" pageId="5" pageNumber="488">0.37–0.42</td>
|
||
<td id="075F5DCB94450000F714632FFD056A28" box="[586,699,1335,1365]" gridcol="3" gridrow="7" pageId="5" pageNumber="488">5.28–6.02</td>
|
||
<td id="075F5DCB94450000F6DC632FFC436A28" box="[898,1021,1335,1365]" gridcol="5" gridrow="7" pageId="5" pageNumber="488">5.38–6.11</td>
|
||
<td id="075F5DCB94450000F1F1632FFAE66A28" box="[1199,1368,1335,1365]" colspan="2" colspanRight="1" gridcol="7" gridrow="7" pageId="5" pageNumber="488">0.9–1.6</td>
|
||
</tr>
|
||
<tr id="448E34B794450000F5C96347FAE66A00" box="[151,1368,1375,1405]" gridrow="8" pageId="5" pageNumber="488" rowspan-4="1" rowspan-6="1">
|
||
<th id="075F5DCB94450000F5C96347FF056A00" box="[151,187,1375,1405]" gridcol="0" gridrow="8" pageId="5" pageNumber="488">P3</th>
|
||
<td id="075F5DCB94450000F5B16347FDB16A00" box="[239,527,1375,1405]" colspan="2" colspanRight="1" gridcol="1" gridrow="8" pageId="5" pageNumber="488">0.53–0.65</td>
|
||
<td id="075F5DCB94450000F7146347FD056A00" box="[586,699,1375,1405]" gridcol="3" gridrow="8" pageId="5" pageNumber="488">4.17–4.77</td>
|
||
<td id="075F5DCB94450000F6DC6347FC436A00" box="[898,1021,1375,1405]" gridcol="5" gridrow="8" pageId="5" pageNumber="488">3.32–3.78</td>
|
||
<td id="075F5DCB94450000F1F16347FAE66A00" box="[1199,1368,1375,1405]" colspan="2" colspanRight="1" gridcol="7" gridrow="8" pageId="5" pageNumber="488">0.8–1.5</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="FA0136F59445FFFAF59963BFFCC16857" blockId="5.[151,1437,1447,2014]" pageId="5" pageNumber="488">
|
||
Hypopygium (
|
||
<figureCitation id="62852A709445FFFAF43463BFFE4A6ABC" box="[362,500,1447,1473]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Figs. 10–14</figureCitation>
|
||
). Tergite IX with 15–18 setae, 32–40 µm long and with narrow (6–8 µm), pale, and naked anal point, 76–116 µm long (
|
||
<figureCitation id="62852A709445FFFAF77163D3FD3C6A98" box="[559,642,1483,1509]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Fig. 10</figureCitation>
|
||
) which in lateral view apically slightly curved down (
|
||
<figureCitation id="62852A709445FFFAF1B263D3FAFF6A98" box="[1260,1345,1483,1509]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Fig. 14</figureCitation>
|
||
); posterior edge of tergite IX slightly concave, anal-lateral angles protrude noticeably beyond edge (
|
||
<figureCitation id="62852A709445FFFAF1E563F7FAAE6974" box="[1211,1296,1519,1545]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Fig. 10</figureCitation>
|
||
); anal tergal bands Y-type; median tergal band 76–104 µm long. Laterosternite IX with 6–10 setae, 32–48 µm long. Transverse sternopodeme (TSA) triangular, peaked (
|
||
<figureCitation id="62852A709445FFFAF735602FFD4C692C" box="[619,754,1591,1617]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Figs. 11–12</figureCitation>
|
||
) and sometimes with spire-shaped apex (
|
||
<figureCitation id="62852A709445FFFAF191602FFA9D692C" box="[1231,1315,1591,1617]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Fig. 11</figureCitation>
|
||
), 160–184 µm high, 216–268 µm wide at the base; TSA height/TSA width 0.71–0.86. Aedeagal lobe as in
|
||
<figureCitation id="62852A709445FFFAF1E56043FAFF6908" box="[1211,1345,1627,1653]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Figs. 11–12</figureCitation>
|
||
, weakly chitinized, leaf-shaped; phallapodeme sclerotized, 208–336 µm long. Gonocoxite 574–640 µm long, in middle part of dorso-internal position with small flattened transverse lobe, 112 µm long, 32–36 µm wide, directed backwards and with longitudinal lobe in middle, anterior outer margin of which is slightly rounded and not projecting (
|
||
<figureCitation id="62852A709445FFFAF06360DFFA31699C" box="[1341,1423,1735,1761]" captionStart="FIGURES 9–14" captionStartId="6.[151,264,1973,1997]" captionTargetBox="[163,1421,417,1943]" captionTargetId="figure-115@6.[160,1426,417,1950]" captionTargetPageId="6" captionText="FIGURES 9–14. Diamesa kownackii sp. nov., male. 9, head; 10, tergite IX; 11–12, transverse sternapodeme, phallapodeme and aedeagal lobe; 13, gonocoxite and gonostylus; 14, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457263" httpUri="https://zenodo.org/record/6457263/files/figure.png" pageId="5" pageNumber="488">Fig. 13</figureCitation>
|
||
). Gonostylus 246–264 µm long, curved, megaseta in form of wide terminal spine 10–12 μm long, next to it there is tooth and two setae approximately of the same long. HR 2.0–2.53.
|
||
</paragraph>
|
||
<paragraph id="FA0136F59445FFFAF599612BFB216830" blockId="5.[151,1437,1447,2014]" box="[199,1183,1842,1869]" pageId="5" pageNumber="488">
|
||
<emphasis id="C8CAEAE79445FFFAF599612BFEBA6830" bold="true" box="[199,260,1843,1869]" italics="true" pageId="5" pageNumber="488">Pupa</emphasis>
|
||
and
|
||
<emphasis id="C8CAEAE79445FFFAF464612AFECB6830" bold="true" box="[314,373,1842,1869]" italics="true" pageId="5" pageNumber="488">larva</emphasis>
|
||
were described as
|
||
<taxonomicName id="3DBE4D769445FFFAF714612CFC3D6830" authority="Goetghebuer" authorityName="Goetghebuer" authorityYear="1933" box="[586,899,1843,1869]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="steinboecki">
|
||
<emphasis id="C8CAEAE79445FFFAF714612CFD556830" box="[586,747,1843,1869]" italics="true" pageId="5" pageNumber="488">D. steinboecki</emphasis>
|
||
Goetghebuer
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="9E2F4B049445FFFAF6F2612BFB256830" author="Makarchenko, E. A. & Makarchenko, M. A." box="[940,1179,1843,1869]" pageId="5" pageNumber="488" pagination="44 - 50" refId="ref19236" refString="Makarchenko, E. A. & Makarchenko, M. A. (1981) Biomass and community structure of benthic invertebrates in the Somnitelnaya River (Wrangel Island). In: Invertebrate animals in ecosystems of salmon rivers of the Far East. DVNC AN SSSR Press, Vladivostok, pp. 44 - 50. [in Russian]" type="book chapter" year="1981">Makarchenko (1981)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2A4657E9445FFF9F599614EFE1D6FA8" lastPageId="6" lastPageNumber="489" pageId="5" pageNumber="488" type="diagnosis">
|
||
<paragraph id="FA0136F59445FFF9F599614EFE1D6FA8" blockId="5.[151,1437,1447,2014]" lastBlockId="6.[151,1436,151,394]" lastPageId="6" lastPageNumber="489" pageId="5" pageNumber="488">
|
||
<emphasis id="C8CAEAE79445FFFAF599614EFEFC680D" bold="true" box="[199,322,1878,1904]" pageId="5" pageNumber="488">Diagnosis.</emphasis>
|
||
The male of
|
||
<taxonomicName id="3DBE4D769445FFFAF48F6140FDDA680C" authority="Makarchenko & Semenchenko & Palatov, 2022" authorityName="Makarchenko & Semenchenko & Palatov" authorityYear="2022" box="[465,612,1879,1905]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="kownackii" status="sp. nov.">
|
||
<emphasis id="C8CAEAE79445FFFAF48F6140FDDA680C" box="[465,612,1879,1905]" italics="true" pageId="5" pageNumber="488">D. kownackii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C8CAEAE79445FFFAF737614FFD7F680C" bold="true" box="[617,705,1879,1905]" pageId="5" pageNumber="488">
|
||
<taxonomicNameLabel id="D3F9579C9445FFFAF737614FFD7F680C" box="[617,705,1879,1905]" pageId="5" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most closely related in the subgroup to the
|
||
<taxonomicName id="3DBE4D769445FFFAF1E96140FAE8680C" authorityName="Goetghebuer" authorityYear="1933" box="[1207,1366,1879,1905]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="steinboecki">
|
||
<emphasis id="C8CAEAE79445FFFAF1E96140FAE8680C" box="[1207,1366,1879,1905]" italics="true" pageId="5" pageNumber="488">D. steinboecki</emphasis>
|
||
</taxonomicName>
|
||
, since both have anal tergal bands Y-type, but the male of first species is larger (
|
||
<quantity id="3D469B109445FFFAF6B96163FBC868E8" box="[999,1142,1915,1942]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="5.2" metricValueMin="4.2" pageId="5" pageNumber="488" unit="mm" value="4.7" valueMax="5.2" valueMin="4.2">4.2–5.2 mm</quantity>
|
||
), it lacks the frontal setae of the head, posterior edge of tergite IX slightly concave and anal-lateral angles protrude noticeably beyond edge, median tergal band 76–104 µm long. The male of
|
||
<taxonomicName id="3DBE4D769445FFFAF78C61DCFCCD68A0" authorityName="Goetghebuer" authorityYear="1933" box="[722,883,1987,2013]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="488" phylum="Arthropoda" rank="species" species="steinboecki">
|
||
<emphasis id="C8CAEAE79445FFFAF78C61DCFCCD68A0" box="[722,883,1987,2013]" italics="true" pageId="5" pageNumber="488">D. steinboecki</emphasis>
|
||
</taxonomicName>
|
||
2.0–
|
||
<quantity id="3D469B109445FFFAF6F361DBFBBD68A0" box="[941,1027,1987,2014]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="488" unit="mm" value="3.5">3.5 mm</quantity>
|
||
long, with some frontal setae of the head, posterior edge of tergite IX almost straight and anal-lateral angles not protrude beyond edge, median tergal band 108–116 µm long.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2A4657E9446FFF9F59966C7FB166E3C" pageId="6" pageNumber="489" type="biology_ecology">
|
||
<paragraph id="FA0136F59446FFF9F59966C7FB166E3C" blockId="6.[151,1436,151,394]" pageId="6" pageNumber="489">
|
||
<emphasis id="C8CAEAE79446FFF9F59966C7FE926F84" bold="true" box="[199,300,223,249]" pageId="6" pageNumber="489">Ecology.</emphasis>
|
||
Adults, pupae and larvae were collected from stones and boulders in high arctic river, located at an altitude of
|
||
<quantity id="3D469B109446FFF9F451671BFE316E60" box="[271,399,259,285]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="489" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</quantity>
|
||
, at a flow rate of
|
||
<quantity id="3D469B109446FFF9F713671AFD016E60" box="[589,703,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="6" pageNumber="489" unit="m" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 m</quantity>
|
||
/s, with water temperatures 7.4–9.0°C. The density of larvae in the Somnitelnaya River was 8.3–16.7 ind./m
|
||
<superScript id="0DCB9BBD9446FFF9F73F673EFDD76E49" attach="left" box="[609,617,294,308]" fontSize="6" pageId="6" pageNumber="489">2</superScript>
|
||
(
|
||
<date id="8E0010359446FFF9F726673FFD566E3C" box="[632,744,295,321]" pageId="6" pageNumber="489" value="1979-07">July 1979</date>
|
||
) (
|
||
<bibRefCitation id="9E2F4B049446FFF9F7A1673FFB226E3C" author="Makarchenko, E. A. & Makarchenko, M. A." box="[767,1180,295,321]" pageId="6" pageNumber="489" pagination="44 - 50" refId="ref19236" refString="Makarchenko, E. A. & Makarchenko, M. A. (1981) Biomass and community structure of benthic invertebrates in the Somnitelnaya River (Wrangel Island). In: Invertebrate animals in ecosystems of salmon rivers of the Far East. DVNC AN SSSR Press, Vladivostok, pp. 44 - 50. [in Russian]" type="book chapter" year="1981">Makarchenko & Makarchenko 1981</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B2A4657E9446FFF9F5996752FD016EF7" pageId="6" pageNumber="489" type="distribution">
|
||
<paragraph id="FA0136F59446FFF9F5996752FD016EF7" blockId="6.[151,1436,151,394]" pageId="6" pageNumber="489">
|
||
<emphasis id="C8CAEAE79446FFF9F5996752FEDF6E19" bold="true" box="[199,353,330,356]" pageId="6" pageNumber="489">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="250588579446FFF9F7016753FD316E18" box="[607,655,331,357]" pageId="6" pageNumber="489">type</typeStatus>
|
||
locality—Somnitelnaya River basin of Wrangel Island and northeastern spurs of the Koryak Highlands (
|
||
<figureCitation id="62852A709446FFF9F7776777FD0D6EF7" box="[553,691,367,394]" captionStart="FIGURES 68–74" captionStartId="23.[151,264,1617,1641]" captionTargetBox="[151,1435,181,1592]" captionTargetId="figure-18@23.[151,1435,181,1593]" captionTargetPageId="23" captionText="FIGURES 68–74. Type localities of Diamesa dragani sp. nov. (68), D. kownackii sp. nov. (69–70), D. maisaraensis sp. nov. (71), D. marinskiyi sp. nov. (72), D. steinboecki Goetghebuer (73) and D. sakartvella Kownacki et Kownacka (74). 68, unnamed stream of Bolshoi On River basin, alt. 2063 m a. sea l., Sayan Mountains (photo by S.V. Dragan); 69, unnamed stream of Somnitelnaya River basin, Wrangel Island (photo by E.A. Makarchenko); 70, Somnitelnaya River, upper stream, alt. 205–210 m a. sea l., Wrangel Island (photo by O.A. Khruleva); 71, Maisara River near Maisara Pass, alt. 4168 m a. sea l., Pamir Mountains (photo by D.M. Palatov); 72, Zienzan River, Tien Shan Mountains, alt. 1923 m a. sea l., Xinjiang Uygur Autonomous Region, China (photo by D.M. Palatov); 73, Balm stream, Massif of Aiguilles–Rouges, French Alps (photo by Pierre Clévenot); 74, Skazdon River near the Tsey (Tseyskoe gorge), alt. 1943 m a. sea l., North Caucasus (photo by D.M. Palatov)." figureDoi="http://doi.org/10.5281/zenodo.6457281" httpUri="https://zenodo.org/record/6457281/files/figure.png" pageId="6" pageNumber="489">Figs. 69–70</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |