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<document id="09E2F6B8D940D301073A2BCF6B292BE9" ID-DOI="10.5281/zenodo.5374196" ID-ISSN="1638-9395" ID-Zenodo-Dep="5374196" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630276223888" checkinUser="marcus" docAuthor="Baciu, Dorin-Sorin, Bannikov, Alexandre F. &amp; Santini, Francesco" docDate="2005" docId="720BF774FFD66B1EFF38F9CD8E6ED5D7" docLanguage="en" docName="Geodiversitas.2005.27.3.381-390.pdf" docOrigin="Geodiversitas 27 (3)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Capros arambourgi Baciu &amp; Bannikov &amp; Santini 2005, n. sp." docType="treatment" docVersion="3" lastPageNumber="388" masterDocId="8E328F0CFFD46B19FFC9FFC18A28D712" masterDocTitle="A new species of Caproidae (Acanthomorpha, Teleostei) from the Messinian (upper Miocene) of Oran (Algeria)" masterLastPageNumber="390" masterPageNumber="381" pageNumber="383" updateTime="1699209536617" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="8B13E21F6E0BAB4FED3B287653921CEF">A new species of Caproidae (Acanthomorpha, Teleostei) from the Messinian (upper Miocene) of Oran (Algeria)</mods:title>
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<mods:namePart id="99CE0DC94A19687DBAC3EC7EBA225E32">Baciu, Dorin-Sorin</mods:namePart>
<mods:affiliation id="539948EB34CF136F0667AB13180216C8">Muzeul de Știint, e ale Naturii, Str. Petru Rareș, 26, 5600 Piatra Neamt, (Romania) dsbaciu @ ambra. ro</mods:affiliation>
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<mods:namePart id="28C8A315BFAFCB044F386FFD39372E28">Bannikov, Alexandre F.</mods:namePart>
<mods:affiliation id="C927CB4CA7BADF9DC312A5A808BBE9D5">Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya 123, Moscow 117997 (Russia) aban @ paleo. ru</mods:affiliation>
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<mods:namePart id="DB6093679A23FE58CE436639C9E36F99">Santini, Francesco</mods:namePart>
<mods:affiliation id="7DA49D7A94861244346FC57040A45820">Département Systématique et Évolution, UMR 7138 CNRS, Muséum national dHistoire naturelle, case postale 26, 43 rue Cuvier, F- 75231 Paris cedex 05 (France) fsantini @ mnhn. fr</mods:affiliation>
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<mods:title id="4A1AFE5FF8F8283975D7A43CE79A58A5">Geodiversitas</mods:title>
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<mods:date id="5E4D6E68EF3F5445C7C59F003B4B6007">2005</mods:date>
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<paragraph id="FA1D4662FFD66B1BFF38F9CD8B98D157" blockId="2.[241,518,1548,1605]" pageId="2" pageNumber="383">
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<taxonomicName id="3DA23DE1FFD66B1BFF38F9CD8BECD137" authority="Baciu &amp; Bannikov &amp; Santini, 2005" authorityName="Baciu &amp; Bannikov &amp; Santini" authorityYear="2005" box="[241,452,1548,1573]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="383" phylum="Chordata" rank="species" species="arambourgi" status="sp. nov.">
<emphasis id="C8D69A70FFD66B1BFF38F9CD8BECD137" bold="true" box="[241,452,1548,1573]" italics="true" pageId="2" pageNumber="383">Capros arambourgi</emphasis>
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<taxonomicNameLabel id="D3E5270BFFD66B1BFE02F9CD882ED137" box="[459,518,1548,1573]" pageId="2" pageNumber="383" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="62995AE7FFD66B1BFE87F9ED8B8ED157" box="[334,422,1580,1605]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="3.[183,194,910,927]" captionStartId-1="4.[121,132,910,927]" captionStartId-2="5.[120,131,947,964]" captionTargetBox-0="[132,1190,204,876]" captionTargetBox-1="[132,1190,218,877]" captionTargetBox-2="[132,1190,198,911]" captionTargetId-0="figure-848@3.[132,1190,204,880]" captionTargetId-1="figure-535@4.[132,1190,218,877]" captionTargetId-2="figure-742@5.[132,1190,198,911]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIG. 1. — Capros arambourgi n. sp., holotype (MNHN ORA 85 D), Messinian of Raz-el-Aïn, Oran. Scale bar: 10 mm." captionText-1="FIG. 2. — Capros arambourgi n. sp., counterpart of holotype (MNHN ORA 85 G), Messinian of Raz-el-Aïn, Oran. Scale bar: 10 mm." captionText-2="FIG. 3. — Capros arambourgi n. sp., reconstruction of the skeleton based on the holotype and referred specimens, scales omitted." figureDoi-0="http://doi.org/10.5281/zenodo.5374198" figureDoi-1="http://doi.org/10.5281/zenodo.5374200" figureDoi-2="http://doi.org/10.5281/zenodo.5374202" httpUri-0="https://zenodo.org/record/5374198/files/figure.png" httpUri-1="https://zenodo.org/record/5374200/files/figure.png" httpUri-2="https://zenodo.org/record/5374202/files/figure.png" pageId="2" pageNumber="383">Figs 1-3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
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<paragraph id="FA1D4662FFD66B1BFFB8F9B488C6D607" blockId="2.[113,646,1653,1701]" lastBlockId="2.[676,1210,204,1360]" pageId="2" pageNumber="383">
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<taxonomicName id="3DA23DE1FFD66B1BFFB8F9B48AC1D19E" ID-CoL="QSYF" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[113,233,1653,1676]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="383" phylum="Chordata" rank="species" species="aper">
<emphasis id="C8D69A70FFD66B1BFFB8F9B48AC1D19E" box="[113,233,1653,1676]" italics="true" pageId="2" pageNumber="383">Capros aper</emphasis>
</taxonomicName>
<treatmentCitation id="7B036073FFD66B1BFEC1F9B4882AD19E" author="ARAMBOURG C." box="[264,514,1653,1676]" page="1243" pageId="2" pageNumber="383" year="1921">
<bibRefCitation id="9E333B93FFD66B1BFEC1F9B4882AD19E" author="ARAMBOURG C." box="[264,514,1653,1676]" pageId="2" pageNumber="383" pagination="1243 - 1245" refId="ref5175" refString="ARAMBOURG C. 1921. - Sur la faune ichthyologique du Sahelien de la region d'Oran. Comptes Rendus hebdomadaires des Seances de l'Academie des Sciences 172 (20): 1243 - 1245." type="journal article" year="1921">Arambourg 1921: 1243</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="7B036073FFD66B1BFDDBF9B48857D19E" author="ARAMBOURG C." box="[530,639,1653,1676]" page="148" pageId="2" pageNumber="383" year="1927">
<bibRefCitation id="9E333B93FFD66B1BFDDBF9B48857D19E" author="ARAMBOURG C." box="[530,639,1653,1676]" pageId="2" pageNumber="383" pagination="1 - 298" refId="ref5210" refString="ARAMBOURG C. 1927. - Les poissons fossiles d'Oran. Materiaux pour la Carte geologique de l'Algerie, 1 re ser., Paleontologie 6: 1 - 298." type="journal article" year="1927">1927: 148</bibRefCitation>
</treatmentCitation>
, pl. XXVI, figs 2-4, pl. XXVII, fig. 1, text-figs 33, 34. —
<bibRefCitation id="9E333B93FFD66B1BFD6DFF0D896ED7F1" author="D'ERASMO G." box="[676,838,204,227]" pageId="2" pageNumber="383" pagination="1 - 118" refId="ref5562" refString="D'ERASMO G. 1930. - Studi sui pesci neogenici d'Italia. Parte III. L'ittiofauna fossile del Gabbro. Atti della R. Accademia delle Scienze Fisiche e Matematiche di Napoli, serie 2, 18 (6): 1 - 118." type="journal article" year="1930">DErasmo 1930</bibRefCitation>
: III, figs 1, 2. —
<treatmentCitation id="7B036073FFD66B1BFC35FF0D8929D7EE" author="BRADLEY F. &amp; LANDINI W." page="30" pageId="2" pageNumber="383" year="1984">
<bibRefCitation id="9E333B93FFD66B1BFC35FF0D8929D7EE" author="BRADLEY F. &amp; LANDINI W." pageId="2" pageNumber="383" pagination="5 - 33" refId="ref5479" refString="BRADLEY F. &amp; LANDINI W. 1984. - I fossili del &quot; Tripoli &quot; messiniano di Gabbro (Livorno). Palaeontographia Italica 73 (n. s. vol. 43): 5 - 33." type="book chapter" year="1984">Bradley &amp; Landini 1984: 30</bibRefCitation>
</treatmentCitation>
, pl. III, figs 4-6 (nec
<taxonomicName id="3DA23DE1FFD66B1BFC13FF2488C2D607" ID-CoL="QSYF" authority="(Linnaeus, 1758))" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="383" phylum="Chordata" rank="species" species="aper">
<emphasis id="C8D69A70FFD66B1BFC13FF248E62D7EE" box="[986,1098,229,252]" italics="true" pageId="2" pageNumber="383">Capros aper</emphasis>
(Linnaeus, 1758))
</taxonomicName>
.
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</paragraph>
<paragraph id="FA1D4662FFD66B1BFD6DFEE889C5D661" blockId="2.[676,1210,204,1360]" pageId="2" pageNumber="383">
<treatmentCitationGroup id="DAB2614CFFD66B1BFD6DFEE889C5D661" pageId="2" pageNumber="383">
<taxonomicName id="3DA23DE1FFD66B1BFD6DFEE88955D652" baseAuthorityName="Kramberger" baseAuthorityYear="1882" box="[676,893,297,320]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="383" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="C8D69A70FFD66B1BFD6DFEE88955D652" box="[676,893,297,320]" italics="true" pageId="2" pageNumber="383">Capros steindachneri</emphasis>
</taxonomicName>
<treatmentCitation id="7B036073FFD66B1BFC55FEE88EA5D652" author="ARAMBOURG C." box="[924,1165,297,320]" page="152" pageId="2" pageNumber="383" year="1927">
<bibRefCitation id="9E333B93FFD66B1BFC55FEE88EA5D652" author="ARAMBOURG C." box="[924,1165,297,320]" pageId="2" pageNumber="383" pagination="1 - 298" refId="ref5210" refString="ARAMBOURG C. 1927. - Les poissons fossiles d'Oran. Materiaux pour la Carte geologique de l'Algerie, 1 re ser., Paleontologie 6: 1 - 298." type="journal article" year="1927">Arambourg 1927: 152</bibRefCitation>
</treatmentCitation>
, pl. XXVII, figs 2, 3, text-fig. 35 (nec
<taxonomicName id="3DA23DE1FFD66B1BFC3FFE828E91D648" baseAuthorityName="Kramberger" baseAuthorityYear="1882" box="[1014,1209,323,346]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="383" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="C8D69A70FFD66B1BFC3FFE828E91D648" box="[1014,1209,323,346]" italics="true" pageId="2" pageNumber="383">Capros steindachneri</emphasis>
</taxonomicName>
(Gorjanović- Kramberger, 1882)).
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</paragraph>
</subSubSection>
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<paragraph id="FA1D4662FFD66B1BFD6DFE4688D8D6F8" blockId="2.[676,1210,204,1360]" pageId="2" pageNumber="383">
<materialsCitation id="4ACA4C3FFFD66B1BFD6DFE4688D8D6F8" collectionCode="MNHN, SL" country="Oran, Algeria" location="upper Miocene" municipality="Messinian of Raz-el-Ain" pageId="2" pageNumber="383" specimenCount="1" typeStatus="holotype">
<typeStatus id="2519F8C0FFD66B1BFD6DFE468927D68E" box="[676,783,391,414]" pageId="2" pageNumber="383" type="holotype">HOLOTYPE</typeStatus>
. —
<collectionCode id="9CB3DEA7FFD66B1BFCF3FE4689A5D68C" box="[826,909,391,414]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="383">MNHN</collectionCode>
ORA 85 D-G, part and counterpart, complete skeleton,
<quantity id="3D5AEB87FFD66B1BFC23FE618E14D6A5" box="[1002,1084,416,439]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="383" unit="mm" value="77.0">77 mm</quantity>
<collectionCode id="9CB3DEA7FFD66B1BFB81FE618E4CD6A5" box="[1096,1124,416,439]" pageId="2" pageNumber="383">SL</collectionCode>
;
<location id="FF7D10B9FFD66B1BFBB1FE6188D4D6C2" LSID="urn:lsid:plazi:treatment:720BF774FFD66B1EFF38F9CD8E6ED5D7:FF7D10B9FFD66B1BFBB1FE6188D4D6C2" country="Oran, Algeria" municipality="Messinian of Raz-el-Ain" name="upper Miocene" pageId="2" pageNumber="383">upper Miocene</location>
,
<collectingMunicipality id="1A79DC18FFD66B1BFCC0FE7889D1D6C2" box="[777,1017,441,464]" pageId="2" pageNumber="383">Messinian of Raz-el-Aïn</collectingMunicipality>
(5.6-5 Ma),
<collectingCountry id="82B506F2FFD66B1BFBB6FE7888C4D6F8" isEnumeration="true" pageId="2" pageNumber="383">Oran, Algeria</collectingCountry>
.
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<paragraph id="FA1D4662FFD66B1BFD6DFE3C8E2FD572" blockId="2.[676,1210,204,1360]" pageId="2" pageNumber="383">
ETYMOLOGY. — The species is named in honour of Dr. Camille Arambourg, pre-eminent French paleontologist, in recognition of his contribution to the study of the Messinian fish fauna of
<collectingCountry id="82B506F2FFD66B1BFC04FD888E29D572" box="[973,1025,585,608]" name="Algeria" pageId="2" pageNumber="383">Oran</collectingCountry>
.
</paragraph>
<paragraph id="FA1D4662FFD66B1BFD6DFDB58956D4FC" blockId="2.[676,1210,204,1360]" pageId="2" pageNumber="383">
<materialsCitation id="4ACA4C3FFFD66B1BFD6DFDB58914D5E2" collectionCode="MNHN" collectorName="In" country="Algeria" location="ORA" pageId="2" pageNumber="383" specimenCount="1">
REFERRED SPECIMENS. —
<collectorName id="575723B4FFD66B1BFC69FDB5899ED599" box="[928,950,628,651]" pageId="2" pageNumber="383">In</collectorName>
the
<collectionCode id="9CB3DEA7FFD66B1BFC28FDB58E1CD599" box="[993,1076,628,651]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="383">MNHN</collectionCode>
collection: in part and counterpart:
<collectingCountry id="82B506F2FFD66B1BFC4FFD4C8993D5B6" box="[902,955,653,676]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
17 D-G,
<collectingCountry id="82B506F2FFD66B1BFBEDFD4C8E71D5B6" box="[1060,1113,653,676]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
98 D-G,
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733 D-G,
<collectingCountry id="82B506F2FFD66B1BFC90FD6689B8D5AC" box="[857,912,679,702]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
741 D-G,
<collectingCountry id="82B506F2FFD66B1BFBC7FD668E6DD5AC" box="[1038,1093,679,702]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
743 D-G,
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747 D-G,
<collectingCountry id="82B506F2FFD66B1BFC81FD018954D5C5" box="[840,892,704,727]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
749 D-G,
<collectingCountry id="82B506F2FFD66B1BFC22FD018E37D5C5" box="[1003,1055,704,727]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
1227 D-G and
<collectingCountry id="82B506F2FFD66B1BFD6DFD1888F2D5E2" box="[676,730,729,752]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
729/742
</materialsCitation>
; in single plate:
<collectingCountry id="82B506F2FFD66B1BFC39FD188E0ED5E2" box="[1008,1062,729,752]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
16,
<collectingCountry id="82B506F2FFD66B1BFB90FD188EA7D5E2" box="[1113,1167,729,752]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
86,
<collectingCountry id="82B506F2FFD66B1BFD6DFD3288F0D418" box="[676,728,755,778]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
90,
<collectingCountry id="82B506F2FFD66B1BFCCEFD328913D418" box="[775,827,755,778]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
103,
<collectingCountry id="82B506F2FFD66B1BFCBEFD328983D418" box="[887,939,755,778]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
395,
<collectingCountry id="82B506F2FFD66B1BFC2EFD328E33D418" box="[999,1051,755,778]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
563-566,
<collectingCountry id="82B506F2FFD66B1BFB4CFD328E91D418" box="[1157,1209,755,778]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
730,
<collectingCountry id="82B506F2FFD66B1BFD14FCCD893CD431" box="[733,788,780,803]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
731,
<collectingCountry id="82B506F2FFD66B1BFC9EFCCD89A6D431" box="[855,910,780,803]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
736,
<collectingCountry id="82B506F2FFD66B1BFC19FCCD8E2FD431" box="[976,1031,780,803]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
739,
<collectingCountry id="82B506F2FFD66B1BFB80FCCD8EA8D431" box="[1097,1152,780,803]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
740,
<collectingCountry id="82B506F2FFD66B1BFD6DFCE488F0D42E" box="[676,728,805,828]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
744,
<collectingCountry id="82B506F2FFD66B1BFCDCFCE48961D42E" box="[789,841,805,828]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
746,
<collectingCountry id="82B506F2FFD66B1BFC4EFCE48993D42E" box="[903,955,805,828]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
747 D,
<collectingCountry id="82B506F2FFD66B1BFBDDFCE48E60D42E" box="[1044,1096,805,828]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
748,
<collectingCountry id="82B506F2FFD66B1BFB4CFCE48E91D42E" box="[1157,1209,805,828]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
752,
<collectingCountry id="82B506F2FFD66B1BFD1EFCFE8923D444" box="[727,779,831,854]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
1351;
<materialsCitation id="4ACA4C3FFFD66B1BFC9BFCFE89BAD49A" collectionCode="SL" country="Algeria" location="ORA" pageId="2" pageNumber="383" specimenCount="1">
<collectingCountry id="82B506F2FFD66B1BFC9BFCFE89AED444" box="[850,902,831,854]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
751 (
<specimenCount id="ECA48DEBFFD66B1BFC09FCFE8E78D444" box="[960,1104,831,854]" pageId="2" pageNumber="383" type="generic">two specimens</specimenCount>
in a single slab), skeletons and fragments, 46 to
<emphasis id="C8D69A70FFD66B1BFBDCFC998E36D47D" box="[1045,1054,856,879]" italics="true" pageId="2" pageNumber="383">c</emphasis>
.
<quantity id="3D5AEB87FFD66B1BFBE2FC998E5BD47D" box="[1067,1139,856,879]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="383" unit="mm" value="90.0">90 mm</quantity>
<collectionCode id="9CB3DEA7FFD66B1BFBB3FC998EBFD47D" box="[1146,1175,856,879]" pageId="2" pageNumber="383">SL</collectionCode>
, all from the
<typeStatus id="2519F8C0FFD66B1BFCC5FCB08912D49A" box="[780,826,881,904]" pageId="2" pageNumber="383">type</typeStatus>
locality
</materialsCitation>
;
<materialsCitation id="4ACA4C3FFFD66B1BFC6DFCB08952D4FC" country="Algeria" location="upper Miocene" municipality="Messinian" pageId="2" pageNumber="383" specimenCount="2">
<specimenCount id="ECA48DEBFFD66B1BFC6DFCB08E0FD49A" box="[932,1063,881,904]" pageId="2" pageNumber="383" type="generic">2 specimens</specimenCount>
described by
<bibRefCitation id="9E333B93FFD66B1BFD6DFC4A894CD4B0" author="ARAMBOURG C." box="[676,868,907,930]" pageId="2" pageNumber="383" pagination="1 - 298" refId="ref5210" refString="ARAMBOURG C. 1927. - Les poissons fossiles d'Oran. Materiaux pour la Carte geologique de l'Algerie, 1 re ser., Paleontologie 6: 1 - 298." type="journal article" year="1927">Arambourg (1927)</bibRefCitation>
as
<taxonomicName id="3DA23DE1FFD66B1BFC41FC4A8E7BD4B0" baseAuthorityName="Kramberger" baseAuthorityYear="1882" box="[904,1107,907,930]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="383" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="C8D69A70FFD66B1BFC41FC4A8E7BD4B0" box="[904,1107,907,930]" italics="true" pageId="2" pageNumber="383">Capros steindachneri</emphasis>
</taxonomicName>
, ORA 15 (single plate) and
<collectingCountry id="82B506F2FFD66B1BFC93FC6589A6D4A9" box="[858,910,932,955]" name="Algeria" pageId="2" pageNumber="383">ORA</collectingCountry>
396/1857 (part and counterpart), from the
<location id="FF7D10B9FFD66B1BFCF3FC7C89F8D4C6" LSID="urn:lsid:plazi:treatment:720BF774FFD66B1EFF38F9CD8E6ED5D7:FF7D10B9FFD66B1BFCF3FC7C89F8D4C6" box="[826,976,957,980]" country="Algeria" municipality="Messinian" name="upper Miocene" pageId="2" pageNumber="383">upper Miocene</location>
,
<collectingMunicipality id="1A79DC18FFD66B1BFC13FC7C8E14D4C6" box="[986,1084,957,980]" pageId="2" pageNumber="383">Messinian</collectingMunicipality>
of St-Denisdu-Sig, Oran,
<collectingCountry id="82B506F2FFD66B1BFCFBFC168952D4FC" box="[818,890,983,1006]" name="Algeria" pageId="2" pageNumber="383">Algeria</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="FA1D4662FFD66B1BFD6DFC3188DBD216" blockId="2.[676,1210,204,1360]" pageId="2" pageNumber="383">
In the MSNTUP collection:
<specimenCount id="ECA48DEBFFD66B1BFC06FC318E70D315" box="[975,1112,1008,1031]" pageId="2" pageNumber="383" type="generic">16 specimens</specimenCount>
from the upper Miocene, Messinian of Gabbro,
<collectingRegion id="38668880FFD66B1BFBEEFBC88E54D332" box="[1063,1148,1033,1056]" country="Italy" name="Toscana" pageId="2" pageNumber="383">Tuscany</collectingRegion>
,
<collectingCountry id="82B506F2FFD66B1BFB4EFBC88E9ED332" box="[1159,1206,1033,1056]" name="Italy" pageId="2" pageNumber="383">Italy</collectingCountry>
: MSNTUP
<accessionNumber id="E5F1DB81FFD66B1BFCD2FBE28958D328" box="[795,880,1059,1082]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/I10964" pageId="2" pageNumber="383" type="EnaNcbi">I 10964</accessionNumber>
(3 skeletons on a single slab),
<accessionNumber id="E5F1DB81FFD66B1BFD6DFBFD88D2D341" box="[676,762,1084,1107]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/I10965" pageId="2" pageNumber="383" type="EnaNcbi">I 10965</accessionNumber>
-
<accessionNumber id="E5F1DB81FFD66B1BFCCDFBFD8972D341" box="[772,858,1084,1107]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/I10976" pageId="2" pageNumber="383" type="EnaNcbi">I 10976</accessionNumber>
, I 11564, I 11588 and I 13994 (
<accessionNumber id="E5F1DB81FFD66B1BFD65FB9488D2D37E" box="[684,762,1109,1132]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/I10967" pageId="2" pageNumber="383" type="EnaNcbi">I 10967</accessionNumber>
/
<accessionNumber id="E5F1DB81FFD66B1BFCCAFB94897AD37E" box="[771,850,1109,1132]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/I10973" pageId="2" pageNumber="383" type="EnaNcbi">I 10973</accessionNumber>
and
<accessionNumber id="E5F1DB81FFD66B1BFC4DFB9489FCD37E" box="[900,980,1109,1132]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/I10971" pageId="2" pageNumber="383" type="EnaNcbi">I 10971</accessionNumber>
/I 11564 represent part and counterpart), their size ranges from
<quantity id="3D5AEB87FFD66B1BFBFDFBAE8E91D394" box="[1076,1209,1135,1158]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="5.5" metricValueMin="1.0" pageId="2" pageNumber="383" unit="mm" value="32.5" valueMax="55.0" valueMin="10.0">10 to 55 mm</quantity>
SL (most of the skeletons are more or less incomplete). According to
<bibRefCitation id="9E333B93FFD66B1BFCE0FB6089F9D3AA" author="D'ERASMO G." box="[809,977,1185,1208]" pageId="2" pageNumber="383" pagination="1 - 118" refId="ref5562" refString="D'ERASMO G. 1930. - Studi sui pesci neogenici d'Italia. Parte III. L'ittiofauna fossile del Gabbro. Atti della R. Accademia delle Scienze Fisiche e Matematiche di Napoli, serie 2, 18 (6): 1 - 118." type="journal article" year="1930">dErasmo (1930)</bibRefCitation>
,
<specimenCount id="ECA48DEBFFD66B1BFC12FB608E76D3AA" box="[987,1118,1185,1208]" pageId="2" pageNumber="383" type="generic">10 specimens</specimenCount>
from the same locality of Gabbro are in the Museo Geologico di Bologna, and
<specimenCount id="ECA48DEBFFD66B1BFCE6FB15898ED3F9" box="[815,934,1236,1259]" pageId="2" pageNumber="383" type="generic">2 specimens</specimenCount>
in the Museo Geologico di Firenze.
</paragraph>
<paragraph id="FA1D4662FFD66B1BFD6DFAC68EAAD242" blockId="2.[676,1210,204,1360]" pageId="2" pageNumber="383">
In the MPM collection: 1 uncatalogued specimen
<quantity id="3D5AEB87FFD66B1BFD6DFAE188D8D225" box="[676,752,1312,1335]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="383" unit="mm" value="46.0">46 mm</quantity>
SL, single plate, from the upper Miocene, Messinian of Mondaino,
<collectingRegion id="38668880FFD66B1BFC55FAF88E6DD242" box="[924,1093,1337,1360]" country="Italy" name="Emilia-Romagna" pageId="2" pageNumber="383">Emilia-Romagna</collectingRegion>
,
<collectingCountry id="82B506F2FFD66B1BFB86FAF88E56D242" box="[1103,1150,1337,1360]" name="Italy" pageId="2" pageNumber="383">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B815E9FFD66B1BFD6DFAA48E38D106" pageId="2" pageNumber="383" type="materials_examined">
<paragraph id="FA1D4662FFD66B1BFD6DFAA48E38D106" blockId="2.[676,1210,1381,1701]" pageId="2" pageNumber="383">
TYPE LOCALITY AND HORIZON. — Most specimens examined originate from the
<typeStatus id="2519F8C0FFD66B1BFC05FABF89DED287" box="[972,1014,1406,1429]" pageId="2" pageNumber="383">type</typeStatus>
locality Raz-el-Aïn near
<collectingCountry id="82B506F2FFD66B1BFD1FFA598925D2BD" box="[726,781,1432,1455]" name="Algeria" pageId="2" pageNumber="383">Oran</collectingCountry>
, NW
<collectingCountry id="82B506F2FFD66B1BFC85FA5989BCD2BD" box="[844,916,1432,1455]" name="Algeria" pageId="2" pageNumber="383">Algeria</collectingCountry>
. This locality is placed in the Djebel Murdjadjo carbonate platform in the Chelif Basin. The fish-bearing layers consist of light marls intercalated basally with limestone and are of Messinian (uppermost Miocene) age.
</paragraph>
</subSubSection>
<subSubSection id="B2B815E9FFD66B1AFD6DF9E98872D3A5" lastPageId="3" lastPageNumber="384" pageId="2" pageNumber="383" type="diagnosis">
<paragraph id="FA1D4662FFD66B1AFD6DF9E98872D3A5" blockId="2.[676,1210,1381,1701]" lastBlockId="3.[113,646,1007,1207]" lastPageId="3" lastPageNumber="384" pageId="2" pageNumber="383">
DIAGNOSIS. — Head length
<quantity id="3D5AEB87FFD66B1BFC76F9E98E0AD12D" box="[959,1058,1576,1599]" metricMagnitude="-2" metricUnit="m" metricValue="6.096" metricValueMax="6.604" metricValueMin="5.588" pageId="2" pageNumber="383" unit="in" value="2.4" valueMax="2.6" valueMin="2.2">2.2-2.6 in</quantity>
SL. Maximum body depth 1.8-2.0 in SL. The anterior angle of the supraoccipital crest extends to the vertical line through the posterior edge of the orbit or slightly anterior to it. There are 22 (10 + 12) vertebrae. The dorsal fin has nine spines and 20-21 soft rays; the third spine is the longest, 28-35% SL and 0.5-0.6 of maximum body depth. The sixth interneural space is vacant. The anal fin has three spines (the second spine being the longest) and 19- 20 soft rays. The soft dorsal- and anal-fin rays have spinules. The length of pelvic-fin spine is slightly greater than the distance between the posterior end of pelvis and the anterior edge of the first anal-fin pterygiophore.
</paragraph>
</subSubSection>
<caption id="AEDD16EAFFD76B1AFF7EFC4F8E5DD48D" ID-DOI="http://doi.org/10.5281/zenodo.5374198" ID-Zenodo-Dep="5374198" box="[183,1141,910,927]" httpUri="https://zenodo.org/record/5374198/files/figure.png" pageId="3" pageNumber="384" startId="3.[183,194,910,927]" targetBox="[132,1190,204,876]" targetPageId="3">
<paragraph id="FA1D4662FFD76B1AFF7EFC4F8E5DD48D" blockId="3.[183,1141,910,927]" box="[183,1141,910,927]" pageId="3" pageNumber="384">
FIG. 1. —
<taxonomicName id="3DA23DE1FFD76B1AFECEFC4F8B8FD48D" authority="Baciu &amp; Bannikov &amp; Santini, 2005" authorityName="Baciu &amp; Bannikov &amp; Santini" authorityYear="2005" box="[263,423,910,927]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="384" phylum="Chordata" rank="species" species="arambourgi" status="sp. nov.">
<emphasis id="C8D69A70FFD76B1AFECEFC4F8B8FD48D" box="[263,423,910,927]" italics="true" pageId="3" pageNumber="384">Capros arambourgi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3E5270BFFD76B1AFE65FC4F8BF6D48D" box="[428,478,910,927]" pageId="3" pageNumber="384" rank="species">n. sp.</taxonomicNameLabel>
, holotype (MNHN ORA 85 D), Messinian of Raz-el-Aïn, Oran. Scale bar: 10 mm.
</paragraph>
</caption>
<subSubSection id="B2B815E9FFD76B1EFFB8FB2D8856D3F0" lastPageId="7" lastPageNumber="388" pageId="3" pageNumber="384" type="description">
<paragraph id="FA1D4662FFD76B1AFFB8FB2D8B2AD216" blockId="3.[113,646,1260,1701]" box="[113,258,1260,1285]" pageId="3" pageNumber="384">DESCRIPTION</paragraph>
<paragraph id="FA1D4662FFD76B1AFFB8FACD8840D117" blockId="3.[113,646,1260,1701]" pageId="3" pageNumber="384">The body is deep, and has a short caudal peduncle. The caudal peduncle depth is about 0.10- 0.20 of the body depth. The head is relatively large, its length is about 0.6-0.8 of the body depth. The head length is contained 2.2- 2.6 times in SL. The maximum body depth is contained 1.8-2.0 times in SL. The dorsal and ventral profiles of the body are equally convex.</paragraph>
<paragraph id="FA1D4662FFD76B1AFFB8F9CD89D8D3F7" blockId="3.[113,646,1260,1701]" lastBlockId="3.[676,1209,1004,1253]" pageId="3" pageNumber="384">The head is deep, but its depth is not greater than its length. The eye is of moderate size, the horizontal diameter of the orbit being longer than the postorbital distance. The mouth is terminal, and presumably highly protractile. The lower jaw articulation is situated on the level vertical to the anterior edge of the orbit or slightly anterior to it. The infraorbital bones usually are not recognizable with confidence, but the lacrymal is often distinguishable. The lacrymal is thick, subrhomboid, and bears multiple spines and radiating ridges on its lateral surface, and there are strong serrations on the lower border.</paragraph>
<paragraph id="FA1D4662FFD76B1AFD6DFACD8915D237" blockId="3.[676,1209,1292,1701]" box="[676,829,1292,1317]" pageId="3" pageNumber="384">
<emphasis id="C8D69A70FFD76B1AFD6DFACD8915D237" box="[676,829,1292,1317]" italics="true" pageId="3" pageNumber="384">Neurocranium</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD76B1AFD6DFAED8E68D1B7" blockId="3.[676,1209,1292,1701]" pageId="3" pageNumber="384">The neurocranium is deep, and the ethmoid region is moderately long. The supraoccipital crest is high, triangular, thorn-like, and pointed; its posterior edge is slightly concave, and the leading edge is thickened and lacks prominent ornamentation. The anterior angle of the supraoccipital crest extends to the level of the posterior edge of the orbit or slightly anterior to it. The frontal is highly ornamented over the orbit with radiating spinous crests. The parasphenoid is exposed in the lowermost part of the orbit; it is slender and almost straight, without a ventral flange.</paragraph>
<paragraph id="FA1D4662FFD06B1DFCDFFF8E8E82D772" blockId="4.[790,1194,79,96]" box="[790,1194,79,96]" pageId="4" pageNumber="385">
New
<taxonomicName id="3DA23DE1FFD06B1DFCF4FF8E89F1D772" authority="(Teleostei)" authorityName="Lowe" authorityYear="1843" baseAuthorityName="Teleostei" box="[829,985,79,96]" class="Actinopterygii" family="Caproidae" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="family">Caproidae (Teleostei)</taxonomicName>
from the Miocene of
<collectingCountry id="82B506F2FFD06B1DFBB1FF8E8E82D772" box="[1144,1194,79,96]" name="Algeria" pageId="4" pageNumber="385">Algeria</collectingCountry>
</paragraph>
<caption id="AEDD16EAFFD06B1DFFB0FC4F8E99D48D" ID-DOI="http://doi.org/10.5281/zenodo.5374200" ID-Zenodo-Dep="5374200" box="[121,1201,910,927]" httpUri="https://zenodo.org/record/5374200/files/figure.png" pageId="4" pageNumber="385" startId="4.[121,132,910,927]" targetBox="[132,1190,218,877]" targetPageId="4">
<paragraph id="FA1D4662FFD06B1DFFB0FC4F8E99D48D" blockId="4.[121,1201,910,927]" box="[121,1201,910,927]" pageId="4" pageNumber="385">
FIG. 2. —
<taxonomicName id="3DA23DE1FFD06B1DFF00FC4F8B41D48D" authority="Baciu &amp; Bannikov &amp; Santini, 2005" authorityName="Baciu &amp; Bannikov &amp; Santini" authorityYear="2005" box="[201,361,910,927]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="385" phylum="Chordata" rank="species" species="arambourgi" status="sp. nov.">
<emphasis id="C8D69A70FFD06B1DFF00FC4F8B41D48D" box="[201,361,910,927]" italics="true" pageId="4" pageNumber="385">Capros arambourgi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3E5270BFFD06B1DFEA7FC4F8B88D48D" box="[366,416,910,927]" pageId="4" pageNumber="385" rank="species">n. sp.</taxonomicNameLabel>
, counterpart of holotype (MNHN ORA 85 G), Messinian of Raz-el-Aïn, Oran. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="FA1D4662FFD06B1DFFB8FC2D8AFED317" blockId="4.[113,647,1004,1541]" box="[113,214,1004,1029]" pageId="4" pageNumber="385">
<emphasis id="C8D69A70FFD06B1DFFB8FC2D8AFED317" box="[113,214,1004,1029]" italics="true" pageId="4" pageNumber="385">Oral jaws</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD06B1DFFB8FBCD8AC9D3F7" blockId="4.[113,647,1004,1541]" pageId="4" pageNumber="385">The premaxilla has a long ascending process; the ascending process is evidently longer than the alveolar process. The alveolar process bears a well developed postmaxillary process. Small premaxillary teeth are distinguishable in some specimens. The maxilla is tapered and not expanded at its distal end.</paragraph>
<paragraph id="FA1D4662FFD06B1DFFB8FB2D8BABD117" blockId="4.[113,647,1004,1541]" pageId="4" pageNumber="385">The mandible is moderately long and low at the symphysis; its length being greater than its depth by about 2.0 times. The lower mandibular margin is slightly concave. There are small spines on the lower border of the dentary posterior to the symphysis. The retroarticular process of the angulo-articular has a posterior process directed along the lower border of the mandible. The lower jaw teeth are small.</paragraph>
<paragraph id="FA1D4662FFD06B1DFFB8F9ED8AD6D157" blockId="4.[113,646,1580,1701]" box="[113,254,1580,1605]" pageId="4" pageNumber="385">
<emphasis id="C8D69A70FFD06B1DFFB8F9ED8AD6D157" box="[113,254,1580,1605]" italics="true" pageId="4" pageNumber="385">Suspensorium</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD06B1DFFB8F98D88AED1B7" blockId="4.[113,646,1580,1701]" pageId="4" pageNumber="385">The hyomandibular shaft is inclined anteroventrally. The head of the hyomandibula is double in lateral view. The quadrate is triangular in shape,</paragraph>
<paragraph id="FA1D4662FFD06B1DFD6DFC2D8E3AD337" blockId="4.[676,1209,1004,1061]" pageId="4" pageNumber="385">with a small articular condyle. The ectopterygoid is curved, incompletely preserved.</paragraph>
<paragraph id="FA1D4662FFD06B1DFD6DFB8D897BD377" blockId="4.[676,1210,1100,1573]" box="[676,851,1100,1125]" pageId="4" pageNumber="385">
<emphasis id="C8D69A70FFD06B1DFD6DFB8D897BD377" box="[676,851,1100,1125]" italics="true" pageId="4" pageNumber="385">Opercular region</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD06B1DFD6DFBAD8E08D137" blockId="4.[676,1210,1100,1573]" pageId="4" pageNumber="385">The opercular region is narrow. The limits of the subopercle and interopercle are unclear. The opercle is vertically elongate; its lateral surface has a ridge which runs parallel to the anterior border of the bone. The opercle is indented posterodorsally, and it is ornamented laterally with radiating rows of spines. The preopercle is only slightly curved along its anterior edge, but the ventral margin of the dorsal limb (which is longer, albeit slightly) meets the posterior margin of the ventral limb almost at a right angle. The lateral surface of the preopercle is covered with spines. The peripheral border of the preopercle bears serrations and lacks a spine at the angle.</paragraph>
<paragraph id="FA1D4662FFD06B1DFD6DF98D89EAD177" blockId="4.[676,1209,1612,1701]" box="[676,962,1612,1637]" pageId="4" pageNumber="385">
<emphasis id="C8D69A70FFD06B1DFD6DF98D89EAD177" box="[676,962,1612,1637]" italics="true" pageId="4" pageNumber="385">Hyoid and branchial arches</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD06B1DFD6DF9AD8E91D1B7" blockId="4.[676,1209,1612,1701]" pageId="4" pageNumber="385">The epihyal is obscured in most specimens. The ceratohyal is partly recognizable in some specimens,</paragraph>
<paragraph id="FA1D4662FFD16B1CFF49FF8E8AD3D772" blockId="5.[128,251,79,96]" box="[128,251,79,96]" pageId="5" pageNumber="386">
Baciu D.-S.
<emphasis id="C8D69A70FFD16B1CFF10FF8E8ADED772" box="[217,246,79,96]" italics="true" pageId="5" pageNumber="386">et al</emphasis>
.
</paragraph>
<caption id="AEDD16EAFFD16B1CFFB1FC728E9CD4D6" ID-DOI="http://doi.org/10.5281/zenodo.5374202" ID-Zenodo-Dep="5374202" box="[120,1204,947,964]" httpUri="https://zenodo.org/record/5374202/files/figure.png" pageId="5" pageNumber="386" startId="5.[120,131,947,964]" targetBox="[132,1190,198,911]" targetPageId="5">
<paragraph id="FA1D4662FFD16B1CFFB1FC728E9CD4D6" blockId="5.[120,1204,947,964]" box="[120,1204,947,964]" pageId="5" pageNumber="386">
FIG. 3. —
<taxonomicName id="3DA23DE1FFD16B1CFF01FC728B40D4D6" authority="Baciu &amp; Bannikov &amp; Santini, 2005" authorityName="Baciu &amp; Bannikov &amp; Santini" authorityYear="2005" box="[200,360,947,964]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="arambourgi" status="sp. nov.">
<emphasis id="C8D69A70FFD16B1CFF01FC728B40D4D6" box="[200,360,947,964]" italics="true" pageId="5" pageNumber="386">Capros arambourgi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3E5270BFFD16B1CFEA4FC728BB7D4D6" box="[365,415,947,964]" pageId="5" pageNumber="386" rank="species">n. sp.</taxonomicNameLabel>
, reconstruction of the skeleton based on the holotype and referred specimens, scales omitted.
</paragraph>
</caption>
<paragraph id="FA1D4662FFD16B1CFFB8FBCD8AC9D3B7" blockId="5.[113,646,1036,1189]" pageId="5" pageNumber="386">and it has a distinctly concave ventral border. There are about six sabre-like branchiostegal rays. The pharyngeal teeth and branchial arch bones are not distinguishable in the material examined.</paragraph>
<paragraph id="FA1D4662FFD16B1CFFB8FB0D8AD6D3F7" blockId="5.[113,647,1228,1701]" box="[113,254,1228,1253]" pageId="5" pageNumber="386">
<emphasis id="C8D69A70FFD16B1CFFB8FB0D8AD6D3F7" box="[113,254,1228,1253]" italics="true" pageId="5" pageNumber="386">Axial skeleton</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD16B1CFFB8FB2D8E03D297" blockId="5.[113,647,1228,1701]" lastBlockId="5.[676,1209,1036,1413]" pageId="5" pageNumber="386">There are 10 abdominal and 12 caudal vertebrae. The first vertebra articulates closely with the skull (being “plastered” or tightly adherent to it). The vertebral column is almost straight, being slightly elevated anteriorly. The vertebral centra are almost square, being as long as high, and that of the first vertebra is fore-shortened. The neural spines of the abdominal vertebrae are fairly short, pointed, and either straight or slightly curved. The neural spines of the caudal vertebrae are slightly longer than those of the abdominal vertebrae, slender, and slightly curved; the spines become shorter posteriorly in the series. The haemal spine of the first caudal vertebra is somewhat stout, whereas the spines more posteriorly are slender. The neural and haemal spines of PU3 are slightly expanded distally. All the neural and haemal spines are inclined posteriorly, but those near the abdominal/caudal transition are close to vertical. Parapophyses are present on four of the posteriormost abdominal vertebrae. Pleural ribs are associated with the third to 10th abdominal vertebrae; they are very short and slender, increasing in length from the first to fifth; the penultimate rib is somewhat stouter than the preceding ones and the ultimate rib.</paragraph>
<paragraph id="FA1D4662FFD16B1CFD6DFA6D89A3D2D7" blockId="5.[676,1209,1452,1701]" box="[676,907,1452,1477]" pageId="5" pageNumber="386">
<emphasis id="C8D69A70FFD16B1CFD6DFA6D89A3D2D7" box="[676,907,1452,1477]" italics="true" pageId="5" pageNumber="386">Pectoral fin and girdle</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD16B1FFD6DFA0D8B0DD5D7" blockId="5.[676,1209,1452,1701]" lastBlockId="6.[113,646,204,709]" lastPageId="6" lastPageNumber="387" pageId="5" pageNumber="386">The posttemporal is not preserved. The supracleithrum is elongate and slender. The cleithrum is an elongate, robust and very slightly sigmoid bone situated just below the first vertebra. The ventral limb of the cleithrum is broad and spatulate; the dorsal limb tapers to a blunt point distally and it is expanded posteriorly as a broad plate ornamented laterally with fine serrate crests. The single postcleithrum is long and rod-like ventrally; the ventral end of the postcleithrum is close to the ventral profile of the body, and posterior to the pelvic bone. The coracoid is subtriangular and projects anteroventrally as a deep process which reaches the ventral limb of the cleithrum; the postcoracoid process seems short. There are four pectoral radials; the radials are slightly constricted in the middle and increase in length from above to below. The pectoral fin is inserted low on the flank, near the midpoint between the vertebral column and the ventral profile of the body. The pectoral fin probably is rounded distally; 15 pectoral-fin rays are recognizable in some specimens.</paragraph>
<paragraph id="FA1D4662FFD26B1FFFB8FD2D8B6AD417" blockId="6.[113,647,748,1125]" box="[113,322,748,773]" pageId="6" pageNumber="387">
<emphasis id="C8D69A70FFD26B1FFFB8FD2D8B6AD417" box="[113,322,748,773]" italics="true" pageId="6" pageNumber="387">Pelvic fin and girdle</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD26B1FFFB8FCCD8B1DD377" blockId="6.[113,647,748,1125]" pageId="6" pageNumber="387">The pelvic bone is relatively long and wedge-like; it is oriented posteroventrally from its attachment to the cleithra. The posterior process of the pelvis is short. The pelvic fin is moderately long, with a strong spine and about five branched rays. The pelvic-fin spine and soft rays bear small spinules. The pelvic-fin origin is situated behind the pectoral-fin base. The pelvic-fin spine length slightly exceeds the distance between the posterior end of the pelvis and the anterior edge of the first analfin pterygiophore.</paragraph>
<paragraph id="FA1D4662FFD26B1FFFB8FB4D8AF1D3B7" blockId="6.[113,646,1164,1701]" box="[113,217,1164,1189]" pageId="6" pageNumber="387">
<emphasis id="C8D69A70FFD26B1FFFB8FB4D8AF1D3B7" box="[113,217,1164,1189]" italics="true" pageId="6" pageNumber="387">Dorsal fin</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD26B1FFFB8FB6D88C5D657" blockId="6.[113,646,1164,1701]" lastBlockId="6.[676,1210,204,869]" pageId="6" pageNumber="387">The dorsal fin is continuous, with a slight indentation between the spiny and soft portions. There are nine dorsal-fin spines; the first spine is the shortest. The first two spines are supernumerary on the first dorsal-fin pterygiophore. The third dorsal-fin spine is the strongest and longest; the succeeding spines gradually decrease in length posteriorly in the series. The third spine is 1.2 times longer than the fourth spine, and the fourth spine is somewhat longer than the pelvicfin spine. The second spine is less than one-half the length of the fourth spine. The six anteriormost dorsal-fin spines bear longitudinal striations. There are 20 or 21 soft segmented rays in the dorsal fin. The length of the base of the soft portion of the dorsal fin is slightly longer than the length of the base of the spiny portion. There are spinules on the soft rays. The soft dorsal-fin rays are more closely spaced than the dorsal-fin spines.</paragraph>
<paragraph id="FA1D4662FFD26B1FFD6DFE8D8E05D477" blockId="6.[676,1210,204,869]" pageId="6" pageNumber="387">The pterygiophores of the spiny dorsal fin are large and sturdy, expanded anteroposteriorly, and each bears a longitudinal ridge. The first pterygiophore is the longest; its ventral shaft is situated in front of the first interneural space, probably being inserted between the halves of the bifid neural spine of the first vertebra. There is a point- ed procumbent spine directed anteroventrally from the upper part of the first dorsal-fin pterygiophore. The sixth interneural space is vacant. The ventral shafts of the dorsal-fin pterygiophores are inclined posteriorly. The pterygiophores of the soft dorsal fin are slender and decrease in length posteriorly in the series. Each interneural space between the 11th to 18th vertebrae contains two or three ventral shafts of the pterygiophores of the soft dorsal fin.</paragraph>
<paragraph id="FA1D4662FFD26B1FFD6DFC4D88D2D4B7" blockId="6.[676,1210,908,1573]" box="[676,762,908,933]" pageId="6" pageNumber="387">
<emphasis id="C8D69A70FFD26B1FFD6DFC4D88D2D4B7" box="[676,762,908,933]" italics="true" pageId="6" pageNumber="387">Anal fin</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD26B1FFD6DFC6D89BED397" blockId="6.[676,1210,908,1573]" pageId="6" pageNumber="387">There are three spines and 19 or 20 soft segment- ed rays in the anal fin. The first two anal-fin spines are supernumerary on the first pterygiophore. The second spine is the longest, and the third spine is the shortest. The middle anal-fin soft rays appear to be the longest. The anal-fin soft rays bear spinules.</paragraph>
<paragraph id="FA1D4662FFD26B1FFD6DFB4D8961D137" blockId="6.[676,1210,908,1573]" pageId="6" pageNumber="387">The first anal-fin pterygiophore is long and stout; its dorsal shaft is slightly inclined, being situated along the anterior edge of the haemal spine of the first caudal vertebra. The ventral end of this pterygiophore is slightly expanded anteriorly. All the other anal-fin pterygiophores are slender, most of them are almost as equally long as opposite soft dorsal-fin pterygiophores. The anal-fin pterygiophores are inclined posteriorly; their lengths gradually decrease posteriorly in the series. The interhaemal spaces above the anal fin have the dorsal shafts of two to four pterygiophores present.</paragraph>
<paragraph id="FA1D4662FFD26B1FFD6DF98D89B2D177" blockId="6.[676,1209,1612,1701]" box="[676,922,1612,1637]" pageId="6" pageNumber="387">
<emphasis id="C8D69A70FFD26B1FFD6DF98D89B2D177" box="[676,922,1612,1637]" italics="true" pageId="6" pageNumber="387">Caudal fin and skeleton</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD26B1EFD6DF9AD881ED657" blockId="6.[676,1209,1612,1701]" lastBlockId="7.[113,647,204,421]" lastPageId="7" lastPageNumber="388" pageId="6" pageNumber="387">The hypurals and parhypural are autogenous. The short neural spine of PU2 forms a low triangular crest. There are three epurals; uroneurals are not visible in the material examined. The centrum of PU2 is only slightly foreshortened; its haemal spine is autogenous.</paragraph>
<paragraph id="FA1D4662FFD36B1EFFB8FE8D8BB0D6B7" blockId="7.[113,647,204,421]" pageId="7" pageNumber="388">The caudal fin is long and rounded; it has 14 principal rays (I,6 + 6,I), and two procurrent rays both above and below.</paragraph>
<paragraph id="FA1D4662FFD36B1EFFB8FE0D8AC7D6F7" blockId="7.[113,646,460,709]" box="[113,239,460,485]" pageId="7" pageNumber="388">
<emphasis id="C8D69A70FFD36B1EFFB8FE0D8AC7D6F7" box="[113,239,460,485]" italics="true" pageId="7" pageNumber="388">Squamation</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD36B1EFFB8FE2D8824D5D7" blockId="7.[113,646,460,709]" pageId="7" pageNumber="388">The body is completely covered with scales which bear numerous (up to 20) needle-like laterally projecting spines on their surfaces. The scales are relatively small and either round or oval, with even margins. The lateral line is not evident in most of the specimens; a few specimens show the arched anterior part of the lateral line.</paragraph>
<paragraph id="FA1D4662FFD36B1EFFB8FD2D8BA7D417" blockId="7.[113,646,748,824]" box="[113,399,748,773]" pageId="7" pageNumber="388">
<emphasis id="C8D69A70FFD36B1EFFB8FD2D8BA7D417" box="[113,399,748,773]" italics="true" pageId="7" pageNumber="388">Measurements (see Table 1)</emphasis>
</paragraph>
<paragraph id="FA1D4662FFD36B1EFFB8FCD3887FD42A" blockId="7.[113,646,748,824]" pageId="7" pageNumber="388">
TABLE 1. — Measurements taken from
<specimenCount id="ECA48DEBFFD36B1EFE7FFCD38805D431" box="[438,557,786,803]" pageId="7" pageNumber="388" type="generic">six specimens</specimenCount>
of
<taxonomicName id="3DA23DE1FFD36B1EFD83FCD38AF8D42A" authority="Baciu &amp; Bannikov &amp; Santini, 2005" authorityName="Baciu &amp; Bannikov &amp; Santini" authorityYear="2005" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="arambourgi" status="sp. nov.">
<emphasis id="C8D69A70FFD36B1EFD83FCD38AF8D42A" italics="true" pageId="7" pageNumber="388">Capros arambourgi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3E5270BFFD36B1EFF1CFCE68B2DD42A" box="[213,261,807,824]" pageId="7" pageNumber="388" rank="species">n. sp.</taxonomicNameLabel>
of
<quantity id="3D5AEB87FFD36B1EFED7FCE68B5CD42A" box="[286,372,807,824]" metricMagnitude="-2" metricUnit="m" metricValue="6.55" metricValueMax="8.5" metricValueMin="4.6" pageId="7" pageNumber="388" unit="mm" value="65.5" valueMax="85.0" valueMin="46.0">46-85 mm</quantity>
standard body length (SL).
</paragraph>
<paragraph id="FA1D4662FFD36B1EFDFDFCA68856D3F0" blockId="7.[113,644,871,1250]" pageId="7" pageNumber="388">
<emphasis id="C8D69A70FFD36B1EFDFDFCA688ACD469" bold="true" box="[564,644,871,891]" pageId="7" pageNumber="388">% ofSL</emphasis>
Head length from tip of snout to posterior border of opercle 38-45 Maximum body depth 50-58 Depth of caudal peduncle 9-12 Distance between origins of pelvic and anal fins 17-22 Length of first spine of dorsal fin 2.5-3.5 Length of longest spine (third) of dorsal fin 27-38 Length of base of spiny dorsal fin 22-26 Length of base of soft dorsal fin 23-27 Length of longest spine (second) of anal fin 7-14 Length of base of anal fin 30-33 Length of spine of pelvic fin 22-28 Length of longest ray of caudal fin 28-30 Length of lower jaw 15-19
</paragraph>
</subSubSection>
<subSubSection id="B2B815E9FFD36B1EFFB8FAED8E6ED5D7" pageId="7" pageNumber="388" type="discussion">
<paragraph id="FA1D4662FFD36B1EFFB8FAED8AFDD256" blockId="7.[113,646,1324,1701]" box="[113,213,1324,1349]" pageId="7" pageNumber="388">REMARKS</paragraph>
<paragraph id="FA1D4662FFD36B1EFFB8FA8D8AF8D1B7" blockId="7.[113,646,1324,1701]" pageId="7" pageNumber="388">
Our investigation of the MNHN Messinian
<taxonomicName id="3DA23DE1FFD36B1EFFB8FAAD8A94D297" authorityName="Lacepede" authorityYear="1802" box="[113,188,1388,1413]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="388" phylum="Chordata" rank="genus">
<emphasis id="C8D69A70FFD36B1EFFB8FAAD8A94D297" box="[113,188,1388,1413]" italics="true" pageId="7" pageNumber="388">Capros</emphasis>
</taxonomicName>
materials from
<collectingCountry id="82B506F2FFD36B1EFEBEFAAD8B9BD297" box="[375,435,1388,1413]" name="Algeria" pageId="7" pageNumber="388">Oran</collectingCountry>
(
<collectingCountry id="82B506F2FFD36B1EFE0FFAAD883ED297" box="[454,534,1388,1413]" name="Algeria" pageId="7" pageNumber="388">Algeria</collectingCountry>
) revealed several incorrect interpretations of characters made by
<bibRefCitation id="9E333B93FFD36B1EFF1AFA6D8BB7D2D7" author="ARAMBOURG C." box="[211,415,1452,1477]" pageId="7" pageNumber="388" pagination="1 - 298" refId="ref5210" refString="ARAMBOURG C. 1927. - Les poissons fossiles d'Oran. Materiaux pour la Carte geologique de l'Algerie, 1 re ser., Paleontologie 6: 1 - 298." type="journal article" year="1927">Arambourg (1927)</bibRefCitation>
in his descriptions of
<taxonomicName id="3DA23DE1FFD36B1EFFB8FA0D8A95D2F7" authorityName="" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[113,189,1484,1509]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="aper">
<emphasis id="C8D69A70FFD36B1EFFB8FA0D8AABD2F7" box="[113,131,1484,1509]" italics="true" pageId="7" pageNumber="388">C</emphasis>
.
<emphasis id="C8D69A70FFD36B1EFF58FA0D8A95D2F7" box="[145,189,1484,1509]" italics="true" pageId="7" pageNumber="388">aper</emphasis>
</taxonomicName>
. For example, the vertebral formula of the Messinian boarfish is 10 + 12 rather than 9(10) + 13 as indicated by
<bibRefCitation id="9E333B93FFD36B1EFEF5F9CD886DD137" author="ARAMBOURG C." box="[316,581,1548,1573]" pageId="7" pageNumber="388" pagination="1 - 298" refId="ref5210" refString="ARAMBOURG C. 1927. - Les poissons fossiles d'Oran. Materiaux pour la Carte geologique de l'Algerie, 1 re ser., Paleontologie 6: 1 - 298." type="journal article" year="1927">Arambourg (1927: 151)</bibRefCitation>
. Also, none of the specimens available have more than 21 soft rays in both the dorsal and anal fins (Arambourg indicated 23 soft dorsal- and analfin rays).
</paragraph>
<paragraph id="FA1D4662FFD36B1EFD6DFF0D8E6ED5D7" blockId="7.[676,1210,204,709]" pageId="7" pageNumber="388">
I n a d d i t i o n t o
<emphasis id="C8D69A70FFD36B1EFCA8FF0D895BD7F7" box="[865,883,204,229]" italics="true" pageId="7" pageNumber="388">C</emphasis>
.
<emphasis id="C8D69A70FFD36B1EFC4EFF0D8992D7F7" box="[903,954,204,229]" italics="true" pageId="7" pageNumber="388">a p e r</emphasis>
, A r a m b o u r g (1 9 2 7) described from the Messinian of
<collectingCountry id="82B506F2FFD36B1EFBCEFF2D8E69D617" box="[1031,1089,236,261]" name="Algeria" pageId="7" pageNumber="388">Oran</collectingCountry>
the species
<taxonomicName id="3DA23DE1FFD36B1EFD6DFECD8E62D637" authority="(Kramberger, 1882)" baseAuthorityName="Kramberger" baseAuthorityYear="1882" box="[676,1098,268,293]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="C8D69A70FFD36B1EFD6DFECD889ED637" box="[676,694,268,293]" italics="true" pageId="7" pageNumber="388">C</emphasis>
.
<emphasis id="C8D69A70FFD36B1EFD01FECD8971D637" box="[712,857,268,293]" italics="true" pageId="7" pageNumber="388">steindachneri</emphasis>
(Kramberger, 1882)
</taxonomicName>
based on
<specimenCount id="ECA48DEBFFD36B1EFD6DFEED896CD657" box="[676,836,300,325]" pageId="7" pageNumber="388" type="generic">two specimens</specimenCount>
from the site Saint-Denis-du-Sig. We refer both specimens to
<taxonomicName id="3DA23DE1FFD36B1EFC13FE8D8E5DD677" authority="Baciu &amp; Bannikov &amp; Santini, 2005" authorityName="Baciu &amp; Bannikov &amp; Santini" authorityYear="2005" box="[986,1141,332,357]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="arambourgi" status="sp. nov.">
<emphasis id="C8D69A70FFD36B1EFC13FE8D89C4D677" box="[986,1004,332,357]" italics="true" pageId="7" pageNumber="388">C</emphasis>
.
<emphasis id="C8D69A70FFD36B1EFC35FE8D8E5DD677" box="[1020,1141,332,357]" italics="true" pageId="7" pageNumber="388">arambourgi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3E5270BFFD36B1EFBB4FE8D8E91D677" box="[1149,1209,332,357]" pageId="7" pageNumber="388" rank="species">n. sp.</taxonomicNameLabel>
because the supposedly distinctive characters of “
<taxonomicName id="3DA23DE1FFD36B1EFD78FE4D8943D6B7" baseAuthorityName="Kramberger" baseAuthorityYear="1882" box="[689,875,396,421]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="C8D69A70FFD36B1EFD78FE4D88EBD6B7" box="[689,707,396,421]" italics="true" pageId="7" pageNumber="388">C</emphasis>
.
<emphasis id="C8D69A70FFD36B1EFD1CFE4D8943D6B7" box="[725,875,396,421]" italics="true" pageId="7" pageNumber="388">steindachneri</emphasis>
</taxonomicName>
” either lie within the range recorded in the new species (body depth, relative lengths of the third dorsal-fin spine and pelvic- and anal-fin spines) or can be explained by a slight distortion of the neurocranium (the “profile fronto-facial” relatively more abruptly oblique). The species name
<taxonomicName id="3DA23DE1FFD36B1EFC14FD8D8E4FD577" baseAuthorityName="Kramberger" baseAuthorityYear="1882" box="[989,1127,588,613]" class="Actinopterygii" family="Caproidae" genus="Capros" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="steindachneri">
<emphasis id="C8D69A70FFD36B1EFC14FD8D8E4FD577" box="[989,1127,588,613]" italics="true" pageId="7" pageNumber="388">steindachneri</emphasis>
</taxonomicName>
should be regarded as a synonym of the Sarmatian species
<taxonomicName id="3DA23DE1FFD36B1EFD37FD4D88F4D5D7" authority="Kramberger, 1882" authorityName="Kramberger" authorityYear="1882" genus="Proantigonia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="radobojana">
<emphasis id="C8D69A70FFD36B1EFD37FD4D8E34D5B7" box="[766,1052,652,677]" italics="true" pageId="7" pageNumber="388">Proantigonia radobojana</emphasis>
Kramberger, 1882
</taxonomicName>
from
<collectingCountry id="82B506F2FFD36B1EFCD6FD6D8958D5D7" box="[799,880,684,709]" name="Croatia" pageId="7" pageNumber="388">Croatia</collectingCountry>
(
<bibRefCitation id="9E333B93FFD36B1EFCB7FD6D8E12D5D7" author="BACIU D. - S. &amp; BANNIKOV A. F. &amp; TYLER J. C." box="[894,1082,684,709]" pageId="7" pageNumber="388" pagination="7 - 74" refId="ref5241" refString="BACIU D. - S., BANNIKOV A. F. &amp; TYLER J. C. 2005. - Revision of the fossil fishes of the family Caproidae (Acanthomorpha). Miscellanea Paleontologica, Studi e Ricerche sui Giacimenti Terziari di Bolca 11: 7 - 74." type="journal article" year="2005">
Baciu
<emphasis id="C8D69A70FFD36B1EFC0DFD6D89C7D5D7" box="[964,1007,684,709]" italics="true" pageId="7" pageNumber="388">et al</emphasis>
. 2005
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</treatment>
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