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<document id="444DA2C6358CA483FAE9D5E6BFBF9B8F" ID="10.11646/zootaxa.4126.2.4" ID-CLB-Dataset="38797" ID-DOI="10.11646/zootaxa.4126.2.4" ID-GBIF-Dataset="46cf6acd-87dc-4165-80fa-e2b5330cd74c" ID-ISSN="1175-5326" ID-Zenodo-Dep="258598" ID-ZooBank="54F4A01E-2BB2-4802-9B93-8593DA846008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466149131865" checkinUser="plazi" docAuthor="Li, Chun-Lin, Yang, Ping-Shih &amp; Wang, Chuan-Chan" docDate="2016" docId="720A87CC8836FFDA2EC1F918FAC1FB40" docLanguage="en" docName="zootaxa.4126.2.4.pdf" docOrigin="Zootaxa 4126 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megistophylla xitoui Li &amp; Wang, new species" docType="treatment" docVersion="9" lastPageNumber="249" masterDocId="8E33FFB48830FFD32E56FFA9FFD8FFE4" masterDocTitle="Two new species of Megistophylla Burmeister (Coleoptera: Scarabaeidae: Melolonthinae) from Taiwan, with notes on the genus" masterLastPageNumber="250" masterPageNumber="240" pageNumber="246" updateTime="1698663021023" updateUser="ExternalLinkService">
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<mods:title id="425A31FE5C77F98CA06132114E74142C">Two new species of Megistophylla Burmeister (Coleoptera: Scarabaeidae: Melolonthinae) from Taiwan, with notes on the genus</mods:title>
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<mods:namePart id="9A793951EF741252472E09266EE2EADB">Li, Chun-Lin</mods:namePart>
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<mods:namePart id="1D6A6CE66C6C0ECBBD8CC650E9A04EF5">Yang, Ping-Shih</mods:namePart>
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<mods:namePart id="97B9E48F5E7FEF0C9FD575EA755D0CAF">Wang, Chuan-Chan</mods:namePart>
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<treatment id="720A87CC8836FFDA2EC1F918FAC1FB40" ID-DOI="http://doi.org/10.5281/zenodo.6059523" ID-GBIF-Taxon="120829032" ID-Zenodo-Dep="6059523" LSID="urn:lsid:plazi:treatment:720A87CC8836FFDA2EC1F918FAC1FB40" httpUri="http://treatment.plazi.org/id/720A87CC8836FFDA2EC1F918FAC1FB40" lastPageId="9" lastPageNumber="249" pageId="6" pageNumber="246">
<subSubSection id="B2B965518836FFD52EC1F918FE49F909" pageId="6" pageNumber="246" type="nomenclature">
<paragraph id="FA1C36DA8836FFD52EC1F918FD19F928" blockId="6.[151,705,1713,1773]" box="[151,705,1713,1740]" pageId="6" pageNumber="246">
<heading id="A15481B68836FFD52EC1F918FD19F928" bold="true" box="[151,705,1713,1740]" fontSize="11" level="1" pageId="6" pageNumber="246" reason="1">
<emphasis id="C8D7EAC88836FFD52EC1F918FD19F928" bold="true" box="[151,705,1713,1740]" pageId="6" pageNumber="246">
<taxonomicName id="3DA34D598836FFD52EC1F918FDFCF928" authority="Li &amp; Wang" authorityName="Li &amp; Wang" box="[151,548,1713,1740]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="246" phylum="Arthropoda" rank="species" species="xitoui" status="sp. nov.">
<emphasis id="C8D7EAC88836FFD52EC1F918FE57F92F" bold="true" box="[151,399,1713,1739]" italics="true" pageId="6" pageNumber="246">Megistophylla xitoui</emphasis>
Li &amp; Wang
</taxonomicName>
,
<taxonomicNameLabel id="D3E457B38836FFD52C67F91BFD19F928" box="[561,705,1714,1740]" pageId="6" pageNumber="246" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FA1C36DA8836FFD52EC1F97DFE49F909" blockId="6.[151,705,1713,1773]" box="[151,401,1748,1773]" pageId="6" pageNumber="246">
<figureCitation id="62982A5F8836FFD52EC1F97DFF3DF909" box="[151,229,1748,1773]" captionStart="FIGURES 1 2" captionStartId="5.[151,264,1963,1986]" captionTargetBox="[213,1374,1039,1934]" captionTargetId="figure@5.[197,1390,1027,1942]" captionTargetPageId="5" captionText="FIGURES 1 2. Dorsal habitus of male Megistophylla. 1. Megistophylla formosana new species, holotype; 2. Megistophylla xitoui new species, holotype." httpUri="https://zenodo.org/record/258599/files/figure.png" pageId="6" pageNumber="246">Figs. 2</figureCitation>
,
<figureCitation id="62982A5F8836FFD52EA7F97DFEC5F909" box="[241,285,1748,1773]" captionStart="FIGURES 3 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 4. Megistophylla formosana new species; 5 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="6" pageNumber="246">56</figureCitation>
,
<figureCitation id="62982A5F8836FFD52F7FF97CFEE1F909" box="[297,313,1749,1773]" captionStart="FIGURES 7 8" captionStartId="6.[151,264,1585,1608]" captionTargetBox="[153,1436,1148,1564]" captionTargetId="figure@6.[151,1436,1148,1564]" captionTargetPageId="6" captionText="FIGURES 7 8. Ventral view of protarsomeres 2 3 of Megistophylla. 7. Megistophylla formosana new species; 8. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258601/files/figure.png" pageId="6" pageNumber="246">8</figureCitation>
,
<figureCitation id="62982A5F8836FFD52F13F97CFE55F909" box="[325,397,1749,1773]" captionStart="FIGURES 9 12" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[166,1427,928,1704]" captionTargetId="figure@7.[151,1436,916,1709]" captionTargetPageId="7" captionText="FIGURES 9 12. Parameres of Megistophylla. (9, 11, lateral view; 10, 12, dorsal view). 9 10. Megistophylla formosana new species; 11 12. Megistophylla xitoui new species. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/258602/files/figure.png" pageId="6" pageNumber="246">1112</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B965518836FFD42EC1F8B5FB13FCB8" lastPageId="7" lastPageNumber="247" pageId="6" pageNumber="246" type="materials_examined">
<paragraph id="FA1C36DA8836FFD42EC1F8B5FB13FCB8" blockId="6.[151,1437,1820,2025]" lastBlockId="7.[151,1437,151,861]" lastPageId="7" lastPageNumber="247" pageId="6" pageNumber="246">
<emphasis id="C8D7EAC88836FFD52EC1F8B5FE9DF8D1" bold="true" box="[151,325,1820,1845]" pageId="6" pageNumber="246">
<typeStatus id="251888788836FFD52EC1F8B5FEDCF8D1" box="[151,260,1820,1845]" pageId="6" pageNumber="246" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="62982A5F8836FFD52F03F8B5FE43F8D1" box="[341,411,1820,1845]" captionStart="FIGURES 1 2" captionStartId="5.[151,264,1963,1986]" captionTargetBox="[213,1374,1039,1934]" captionTargetId="figure@5.[197,1390,1027,1942]" captionTargetPageId="5" captionText="FIGURES 1 2. Dorsal habitus of male Megistophylla. 1. Megistophylla formosana new species, holotype; 2. Megistophylla xitoui new species, holotype." httpUri="https://zenodo.org/record/258599/files/figure.png" pageId="6" pageNumber="246">Fig. 2</figureCitation>
). Body length
<quantity id="3D5B9B3F8836FFD52C14F8B5FD72F8D0" box="[578,682,1820,1845]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="6" pageNumber="246" unit="mm" value="18.5">18.5 mm</quantity>
; width across humeri
<quantity id="3D5B9B3F8836FFD52DF4F8B4FC23F8D0" box="[930,1019,1821,1845]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="6" pageNumber="246" unit="mm" value="8.8">8.8 mm</quantity>
. Dorsal color blackish brown; venter and legs moderately shiny. Basal antennomeres reddish brown, antennal clubs blackish brown. Dorsum overall setiferous; setae tiny (high magnification for observation required); setae on disc of pronotum and elytra far smaller than diameter of puncture with exception of a row of long, sparse setae at base of head, anterior and posterior margins of pronotum, base of elytra, and scutellum, respectively; long setae brown in color with length 17 times longer than diameter of puncture; long setae varied in appearance.
<emphasis id="C8D7EAC88836FFD52DD9F879FC0CF80D" box="[911,980,2000,2025]" italics="true" pageId="6" pageNumber="246">Head:</emphasis>
Surface densely rugopunctate, puncture large. Clypeus (
<figureCitation id="62982A5F8837FFD42F02FF3EFE44FF54" box="[340,412,151,176]" captionStart="FIGURES 3 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 4. Megistophylla formosana new species; 5 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="7" pageNumber="247">Fig. 5</figureCitation>
) with apex distinctly bilobed, anterior margin moderately reflexed, narrowed at base. Clypeofrontal suture clearly defined. Frons moderately raised. Basal carina slightly crescent, strongly raised. Antenna with 10 antennomeres, antennal club with 6 antennomeres, length of club subequal to basal antennomeres 1-4 combined, first antennomere of club slightly shorter than remaining antennomeres of club; fourth basal antennomere greatly enlarged.
<emphasis id="C8D7EAC88837FFD42C58FE8EFDB0FEA4" box="[526,616,295,320]" italics="true" pageId="7" pageNumber="247">Thorax:</emphasis>
Pronotum widest at middle, anterolateral angles (
<figureCitation id="62982A5F8837FFD42A8CFE8EFAFEFEA4" box="[1242,1318,295,320]" captionStart="FIGURES 3 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 4. Megistophylla formosana new species; 5 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="7" pageNumber="247">Fig. 5</figureCitation>
) strongly protuberant with basolateral angles obtuse, basal half of lateral margin coarsely serrate; lateral declivities with weak gibbosity at middle; anterior margin completely beaded with basal margin smooth; punctures same as those on clypeus, rarely confluent; punctures unevenly distributed along midline (
<figureCitation id="62982A5F8837FFD42DB9FE3DFBECFE49" box="[1007,1076,404,429]" captionStart="FIGURES 3 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 4. Megistophylla formosana new species; 5 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="7" pageNumber="247">Fig. 6</figureCitation>
). Scutellum triangular in shape, with a puncture bearing a long seta (
<figureCitation id="62982A5F8837FFD42C7AFE1EFDA8FE34" box="[556,624,439,464]" captionStart="FIGURES 3 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 4. Megistophylla formosana new species; 5 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="7" pageNumber="247">Fig. 6</figureCitation>
). Prosternal process flat, weakly bilobed in shape.
<emphasis id="C8D7EAC88837FFD42AF1FE1EFB2EFE34" box="[1191,1270,439,464]" italics="true" pageId="7" pageNumber="247">Elytra:</emphasis>
Discal surface densely rugopunctate, densely confluent at sides of suture; sutural costa gradually broadening apically, densely punctate at margins of costa, rugopunctures strongly and transversally developed over sutural costa, costa becoming flat at apex.
<emphasis id="C8D7EAC88837FFD42FEEFD8DFDFAFDD9" box="[440,546,548,573]" italics="true" pageId="7" pageNumber="247">Abdomen</emphasis>
: Pygidium wider than long; surface with setiferous punctures in uneven distribution, tiny setae same as those of elytral disc. Sternites 12, 56 with surface densely punctate; sternites 34 with sparser setiferous punctures; setae on sternites 34 tiny, intermixed on sternites 56 with much longer setae.
<emphasis id="C8D7EAC88837FFD42EC1FD38FF14FD4C" box="[151,204,657,680]" italics="true" pageId="7" pageNumber="247">Legs</emphasis>
: Protibia tridentate. Protarsomeres 23 (
<figureCitation id="62982A5F8837FFD42CC4FD26FD0CFD4C" box="[658,724,655,680]" captionStart="FIGURES 7 8" captionStartId="6.[151,264,1585,1608]" captionTargetBox="[153,1436,1148,1564]" captionTargetId="figure@6.[151,1436,1148,1564]" captionTargetPageId="6" captionText="FIGURES 7 8. Ventral view of protarsomeres 2 3 of Megistophylla. 7. Megistophylla formosana new species; 8. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258601/files/figure.png" pageId="7" pageNumber="247">Fig. 8</figureCitation>
) with setae sparsely distributed at apices. Surface of profemora and mesofemora moderately distributed with hair-like, long setae; setae on metafemora shorter and more robust, metafemora slightly broadened at middle. Metatibia with upper apical spur reaching basal one-third of metatarsomere 2; lower apical spur slightly shorter than upper spur.
<emphasis id="C8D7EAC88837FFD42DECFD55FBFCFCF1" box="[954,1060,764,789]" italics="true" pageId="7" pageNumber="247">Genitalia</emphasis>
: Parameres in lateral view with central concave area concentrically wrinkled densely, apical upper margin smooth with tip gradually narrowed apically (
<figureCitation id="62982A5F8837FFD42EA9FCEDFE88FCB8" box="[255,336,836,861]" captionStart="FIGURES 9 12" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[166,1427,928,1704]" captionTargetId="figure@7.[151,1436,916,1709]" captionTargetPageId="7" captionText="FIGURES 9 12. Parameres of Megistophylla. (9, 11, lateral view; 10, 12, dorsal view). 9 10. Megistophylla formosana new species; 11 12. Megistophylla xitoui new species. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/258602/files/figure.png" pageId="7" pageNumber="247">Fig. 11</figureCitation>
); upper margin of parameres gradually swollen apically at one-fourth (
<figureCitation id="62982A5F8837FFD42A3CFCEDFB66FCB8" box="[1130,1214,836,861]" captionStart="FIGURES 9 12" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[166,1427,928,1704]" captionTargetId="figure@7.[151,1436,916,1709]" captionTargetPageId="7" captionText="FIGURES 9 12. Parameres of Megistophylla. (9, 11, lateral view; 10, 12, dorsal view). 9 10. Megistophylla formosana new species; 11 12. Megistophylla xitoui new species. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/258602/files/figure.png" pageId="7" pageNumber="247">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="AEDC66528837FFD42EC1F968FCBBF913" httpUri="https://zenodo.org/record/258602/files/figure.png" pageId="7" pageNumber="247" targetBox="[166,1427,928,1704]" targetPageId="7">
<paragraph id="FA1C36DA8837FFD42EC1F968FCBBF913" blockId="7.[151,1436,1729,1783]" pageId="7" pageNumber="247">
<emphasis id="C8D7EAC88837FFD42EC1F968FE90F933" bold="true" box="[151,328,1729,1752]" pageId="7" pageNumber="247">FIGURES 912.</emphasis>
Parameres of
<taxonomicName id="3DA34D598837FFD42F8AF96BFDB5F93C" box="[476,621,1730,1752]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="C8D7EAC88837FFD42F8AF96BFDB5F93C" box="[476,621,1730,1752]" italics="true" pageId="7" pageNumber="247">Megistophylla</emphasis>
</taxonomicName>
. (9, 11, lateral view; 10, 12, dorsal view). 910.
<taxonomicName id="3DA34D598837FFD42A30F96BFAB3F93C" box="[1126,1387,1730,1752]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="247" phylum="Arthropoda" rank="species" species="formosana" status="sp. nov.">
<emphasis id="C8D7EAC88837FFD42A30F96BFAB3F93C" box="[1126,1387,1730,1752]" italics="true" pageId="7" pageNumber="247">Megistophylla formosana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3E457B38837FFD42B24F96AFF3BF913" pageId="7" pageNumber="247" rank="species">new species</taxonomicNameLabel>
; 1112.
<taxonomicName id="3DA34D598837FFD42F6EF948FDD1F913" box="[312,521,1761,1783]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="247" phylum="Arthropoda" rank="species" species="xitoui" status="sp. nov.">
<emphasis id="C8D7EAC88837FFD42F6EF948FDD1F913" box="[312,521,1761,1783]" italics="true" pageId="7" pageNumber="247">Megistophylla xitoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3E457B38837FFD42C59F94BFD54F913" box="[527,652,1761,1783]" pageId="7" pageNumber="247" rank="species">new species</taxonomicNameLabel>
. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="AEDC66528838FFDB2EC1F8F3FEACF86B" httpUri="https://zenodo.org/record/258603/files/figure.png" pageId="8" pageNumber="248" targetBox="[151,1436,195,1858]" targetPageId="8">
<paragraph id="FA1C36DA8838FFDB2EC1F8F3FEACF86B" blockId="8.[151,1436,1882,1935]" pageId="8" pageNumber="248">
<emphasis id="C8D7EAC88838FFDB2EC1F8F3FEFBF88B" bold="true" box="[151,291,1882,1904]" pageId="8" pageNumber="248">FIGURE 13.</emphasis>
Distribution map of
<taxonomicName id="3DA34D598838FFDB2C51F8F3FD40F894" box="[519,664,1882,1904]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="C8D7EAC88838FFDB2C51F8F3FD40F894" box="[519,664,1882,1904]" italics="true" pageId="8" pageNumber="248">Megistophylla</emphasis>
</taxonomicName>
species in Taiwan:
<taxonomicName id="3DA34D598838FFDB2D39F8F3FBAFF894" box="[879,1143,1882,1904]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="248" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88838FFDB2D39F8F3FBAFF894" box="[879,1143,1882,1904]" italics="true" pageId="8" pageNumber="248">Megistophylla formosana</emphasis>
</taxonomicName>
(open circle),
<taxonomicName id="3DA34D598838FFDB2B5DF8F3FF09F86B" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="248" phylum="Arthropoda" rank="species" species="xitoui">
<emphasis id="C8D7EAC88838FFDB2B5DF8F3FF09F86B" italics="true" pageId="8" pageNumber="248">Megistophylla xitoui</emphasis>
</taxonomicName>
(open triangle).
</paragraph>
</caption>
<subSubSection id="B2B965518839FFDA2E91FF3EFBCBFEDB" pageId="9" pageNumber="249" type="description">
<paragraph id="FA1C36DA8839FFDA2E91FF3EFE7AFF54" blockId="9.[151,1436,151,1188]" box="[199,418,151,176]" pageId="9" pageNumber="249">
<emphasis id="C8D7EAC88839FFDA2E91FF3EFEC6FF54" bold="true" box="[199,286,151,176]" pageId="9" pageNumber="249">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="FA1C36DA8839FFDA2E91FF15FBCBFEDB" blockId="9.[151,1436,151,1188]" pageId="9" pageNumber="249">
<emphasis id="C8D7EAC88839FFDA2E91FF15FEE3FF31" bold="true" box="[199,315,188,213]" pageId="9" pageNumber="249">Variation.</emphasis>
Body length
<quantity id="3D5B9B3F8839FFDA2FB4FF15FD54FF30" box="[482,652,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" metricValueMax="2.09" metricValueMin="1.75" pageId="9" pageNumber="249" unit="mm" value="19.2" valueMax="20.9" valueMin="17.5">17.520.9 mm</quantity>
; width across humeri
<quantity id="3D5B9B3F8839FFDA2DC7FF15FBC5FF31" box="[913,1053,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.1" metricValueMin="8.4" pageId="9" pageNumber="249" unit="mm" value="8.75" valueMax="9.1" valueMin="8.4">8.49.1 mm</quantity>
.Dorsal body color mainly black, rarely dark reddish brown; pronotum with anterolateral angles moderately to strongly protuberant, punctures rarely evenly distributed; scutellum usually impunctate or with less than 10 punctures, punctures sometimes with long setae; elytral sutural costa sometimes not transversally rugopunctate over costa.
</paragraph>
</subSubSection>
<subSubSection id="B2B965518839FFDA2E91FEE5FC6CFDD8" pageId="9" pageNumber="249" type="diagnosis">
<paragraph id="FA1C36DA8839FFDA2E91FEE5FC6CFDD8" blockId="9.[151,1436,151,1188]" pageId="9" pageNumber="249">
<emphasis id="C8D7EAC88839FFDA2E91FEE5FE99FE81" bold="true" box="[199,321,332,357]" pageId="9" pageNumber="249">Diagnosis.</emphasis>
<taxonomicName id="3DA34D598839FFDA2F1EFEE5FDF6FE80" box="[328,558,332,357]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="xitoui">
<emphasis id="C8D7EAC88839FFDA2F1EFEE5FDF6FE80" box="[328,558,332,357]" italics="true" pageId="9" pageNumber="249">Megistophylla xitoui</emphasis>
</taxonomicName>
is morphologically most close to
<taxonomicName id="3DA34D598839FFDA2DFEFEE4FB9DFE80" box="[936,1093,332,356]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88839FFDA2DFEFEE4FC67FE80" box="[936,959,333,356]" italics="true" pageId="9" pageNumber="249">M</emphasis>
.
<emphasis id="C8D7EAC88839FFDA2D9BFEE5FB9DFE80" box="[973,1093,332,356]" italics="true" pageId="9" pageNumber="249">formosana</emphasis>
</taxonomicName>
and can be distinguished from the later by its distinctly black body color (mainly rufotestaceous to dark reddish brown in
<taxonomicName id="3DA34D598839FFDA2ABEFED8FA53FE6C" box="[1256,1419,368,392]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88839FFDA2ABEFED8FB27FE6C" box="[1256,1279,369,392]" italics="true" pageId="9" pageNumber="249">M</emphasis>
.
<emphasis id="C8D7EAC88839FFDA2B45FED9FA53FE6C" box="[1299,1419,368,392]" italics="true" pageId="9" pageNumber="249">formosana</emphasis>
</taxonomicName>
); punctures usually unevenly and sparsely distributed along midline of pronotum (usually evenly and densely distributed in
<taxonomicName id="3DA34D598839FFDA2F60FE10FE0DFE34" box="[310,469,440,464]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88839FFDA2F60FE10FE95FE34" box="[310,333,441,464]" italics="true" pageId="9" pageNumber="249">M</emphasis>
.
<emphasis id="C8D7EAC88839FFDA2F0BFE11FE0DFE34" box="[349,469,440,464]" italics="true" pageId="9" pageNumber="249">formosana</emphasis>
</taxonomicName>
); scutellum usually impunctate or with less than 10 punctures (usually exceeding 10 punctures in
<taxonomicName id="3DA34D598839FFDA2F60FE74FE03FE10" box="[310,475,476,500]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88839FFDA2F60FE74FE95FE10" box="[310,333,477,500]" italics="true" pageId="9" pageNumber="249">M</emphasis>
.
<emphasis id="C8D7EAC88839FFDA2F35FE75FE03FE10" box="[355,475,476,500]" italics="true" pageId="9" pageNumber="249">formosana</emphasis>
</taxonomicName>
); protarsomeres 23 with setae sparsely distributed at apices (with dense setae aggregated at apices in
<taxonomicName id="3DA34D598839FFDA2FF4FDA8FD98FDFC" box="[418,576,512,536]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88839FFDA2FF4FDA8FE61FDFC" box="[418,441,513,536]" italics="true" pageId="9" pageNumber="249">M</emphasis>
.
<emphasis id="C8D7EAC88839FFDA2F9EFDA9FD98FDFC" box="[456,576,512,536]" italics="true" pageId="9" pageNumber="249">formosana</emphasis>
</taxonomicName>
) and apical upper margin of parameres smooth with tip gradually narrowed apically (distinctly swollen with tip broadly rounded in
<taxonomicName id="3DA34D598839FFDA2D50FD8CFC7CFDD8" box="[774,932,548,572]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88839FFDA2D50FD8CFCC5FDD8" box="[774,797,549,572]" italics="true" pageId="9" pageNumber="249">M</emphasis>
.
<emphasis id="C8D7EAC88839FFDA2D7AFD8DFC7CFDD8" box="[812,932,548,572]" italics="true" pageId="9" pageNumber="249">formosana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B965518839FFDA2E91FDEEFCF4FCB8" pageId="9" pageNumber="249" type="materials_examined">
<paragraph id="FA1C36DA8839FFDA2E91FDEEFCF4FCB8" blockId="9.[151,1436,151,1188]" pageId="9" pageNumber="249">
<emphasis id="C8D7EAC88839FFDA2E91FDEEFEAFFD84" bold="true" box="[199,375,583,608]" pageId="9" pageNumber="249">
<typeStatus id="251888788839FFDA2E91FDEEFEDAFD84" box="[199,258,583,608]" pageId="9" pageNumber="249">Type</typeStatus>
material.
</emphasis>
Male
<typeStatus id="251888788839FFDA2F92FDEEFDFEFD84" box="[452,550,583,608]" pageId="9" pageNumber="249" type="holotype">holotype</typeStatus>
and
<specimenCount id="ECA5FD538839FFDA2C37FDE1FD65FD84" box="[609,701,583,608]" pageId="9" pageNumber="249" type="male">17 male</specimenCount>
<typeStatus id="251888788839FFDA2C91FDE1FCEEFD84" box="[711,822,584,608]" pageId="9" pageNumber="249" type="paratype">paratypes</typeStatus>
. The male
<typeStatus id="251888788839FFDA2DEEFDEEFBC2FD84" box="[952,1050,583,608]" pageId="9" pageNumber="249" type="holotype">holotype</typeStatus>
is pinned and with the following information on the label: “C.
<collectingCountry id="82B4764A8839FFDA2FBDFDC5FD9AFD61" box="[491,578,620,645]" name="Taiwan" pageId="9" pageNumber="249">Taiwan</collectingCountry>
: Nantou, Bird Watching Trail, Xitou,
<geoCoordinate id="9F97501D8839FFDA2DA3FDC5FB8EFD60" box="[1013,1110,620,644]" direction="east" orientation="longitude" pageId="9" pageNumber="249" precision="555" value="120.47">120.47E</geoCoordinate>
,
<geoCoordinate id="9F97501D8839FFDA2A35FDC5FB61FD60" box="[1123,1209,620,644]" direction="north" orientation="latitude" pageId="9" pageNumber="249" precision="555" value="23.4">23.40N</geoCoordinate>
, alt. ca.
<quantity id="3D5B9B3F8839FFDA2B4DFDC5FAABFD61" box="[1307,1395,620,645]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="249" unit="m" value="1250.0">1,250m</quantity>
, by FIT,
<date id="8E1D101A8839FFDA2E87FD26FE7AFD4C" box="[209,418,655,680]" pageId="9" pageNumber="249" value="2013-03-05" valueMax="2013-04-09" valueMin="2013-03-05">5/Mar-9/Apr/2013</date>
, collr. C. -L. Li”. The
<typeStatus id="251888788839FFDA2CE0FD26FCC0FD4C" box="[694,792,655,680]" pageId="9" pageNumber="249" type="holotype">holotype</typeStatus>
is deposited at the National Natural Science Museum, Taichung,
<collectingCountry id="82B4764A8839FFDA2F5BFD1DFEBBFD29" box="[269,355,692,717]" name="Taiwan" pageId="9" pageNumber="249">Taiwan</collectingCountry>
.
<typeStatus id="251888788839FFDA2F3BFD1CFE06FD29" box="[365,478,693,717]" pageId="9" pageNumber="249" type="paratype">Paratypes</typeStatus>
: Three males with the same data as
<typeStatus id="251888788839FFDA2D39FD1DFC09FD29" box="[879,977,692,717]" pageId="9" pageNumber="249" type="holotype">holotype</typeStatus>
in
<collectionCode id="9CB2AE1F8839FFDA2DA2FD1DFBEFFD28" box="[1012,1079,692,716]" pageId="9" pageNumber="249">CCLI</collectionCode>
; same locality as
<typeStatus id="251888788839FFDA2AAAFD1DFAB9FD29" box="[1276,1377,692,717]" pageId="9" pageNumber="249" type="holotype">holotype</typeStatus>
, “
<date id="8E1D101A8839FFDA2B21FD1DFE87FD14" pageId="9" pageNumber="249" value="2014-03-25" valueMax="2014-04-27" valueMin="2014-03-25">25/ Mar-27/Apr/2014</date>
” (
<specimenCount id="ECA5FD538839FFDA2FD6FD71FE08FD14" box="[384,464,727,752]" pageId="9" pageNumber="249" type="male">1 male</specimenCount>
,
<collectionCode id="9CB2AE1F8839FFDA2FB6FD71FDFDFD14" box="[480,549,728,752]" pageId="9" pageNumber="249">CCLI</collectionCode>
); same locality, “
<date id="8E1D101A8839FFDA2CB9FD7EFC08FD14" box="[751,976,727,752]" pageId="9" pageNumber="249" value="2015-05-07" valueMax="2015-06-15" valueMin="2015-05-07">7/May-15/Jun/2015</date>
” (
<specimenCount id="ECA5FD538839FFDA2DA7FD71FB99FD14" box="[1009,1089,727,752]" pageId="9" pageNumber="249" type="male">1 male</specimenCount>
,
<collectionCode id="9CB2AE1F8839FFDA2A06FD71FB4DFD14" box="[1104,1173,728,752]" pageId="9" pageNumber="249">CCLI</collectionCode>
); “C.
<collectingCountry id="82B4764A8839FFDA2A88FD7EFAEDFD14" box="[1246,1333,727,752]" name="Taiwan" pageId="9" pageNumber="249">Taiwan</collectingCountry>
: Nantou, Fong Hwan Shan, by FIT,
<geoCoordinate id="9F97501D8839FFDA2F9AFD55FDF5FCF0" box="[460,557,764,788]" direction="east" orientation="longitude" pageId="9" pageNumber="249" precision="555" value="120.48">120.48E</geoCoordinate>
,
<geoCoordinate id="9F97501D8839FFDA2C6CFD55FD48FCF0" box="[570,656,764,788]" direction="north" orientation="latitude" pageId="9" pageNumber="249" precision="555" value="23.39">23.39N</geoCoordinate>
, alt. ca.
<quantity id="3D5B9B3F8839FFDA2CA5FD55FC8BFCF0" box="[755,851,764,788]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="249" unit="m" value="1800.0">1,800 m</quantity>
,
<date id="8E1D101A8839FFDA2D37FD55FBBBFCF1" box="[865,1123,764,789]" pageId="9" pageNumber="249" value="2013-04-10" valueMax="2013-05-23" valueMin="2013-04-10">10/April-23/May/2013</date>
, collr. C.-L. Li” (
<specimenCount id="ECA5FD538839FFDA2B67FD55FAA5FCF1" box="[1329,1405,764,789]" pageId="9" pageNumber="249" type="male">1 male</specimenCount>
in
<collectionCode id="9CB2AE1F8839FFDA2EC1FC89FF33FCD3" box="[151,235,800,823]" pageId="9" pageNumber="249">ZMHB</collectionCode>
;
<specimenCount id="ECA5FD538839FFDA2EA1FC89FE96FCDC" box="[247,334,799,824]" pageId="9" pageNumber="249" type="male">2 males</specimenCount>
in
<collectionCode id="9CB2AE1F8839FFDA2F23FC89FE15FCD3" box="[373,461,800,823]" pageId="9" pageNumber="249">BMNH</collectionCode>
;
<specimenCount id="ECA5FD538839FFDA2F8EFC89FDFCFCDC" box="[472,548,799,824]" pageId="9" pageNumber="249" type="male">1 male</specimenCount>
in
<collectionCode id="9CB2AE1F8839FFDA2C1CFC89FD54FCD3" box="[586,652,800,823]" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="9" pageNumber="249">MTD</collectionCode>
;
<specimenCount id="ECA5FD538839FFDA2CCFFC89FD3DFCDC" box="[665,741,799,824]" pageId="9" pageNumber="249" type="male">1 male</specimenCount>
in
<collectionCode id="9CB2AE1F8839FFDA2D5DFC89FC93FCD3" box="[779,843,800,823]" httpUri="http://grbio.org/cool/fitg-90ys" name="Taiwan Fisheries Research Institute" pageId="9" pageNumber="249">TFRI</collectionCode>
;
<specimenCount id="ECA5FD538839FFDA2D01FC89FC7BFCDC" box="[855,931,799,824]" pageId="9" pageNumber="249" type="male">1 male</specimenCount>
in
<collectionCode id="9CB2AE1F8839FFDA2D9FFC89FBC7FCDC" box="[969,1055,800,824]" pageId="9" pageNumber="249">CMNC</collectionCode>
;
<specimenCount id="ECA5FD538839FFDA2A7AFC89FB5BFCDC" box="[1068,1155,799,824]" pageId="9" pageNumber="249" type="male">2 males</specimenCount>
in
<collectionCode id="9CB2AE1F8839FFDA2AFFFC89FB36FCDC" box="[1193,1262,800,824]" pageId="9" pageNumber="249">CCLI</collectionCode>
); same locality, “
<date id="8E1D101A8839FFDA2EF7FCEDFE4DFCB9" box="[161,405,836,861]" pageId="9" pageNumber="249" value="2014-04-27" valueMax="2014-05-26" valueMin="2014-04-27">27/Apr-26/May/2014</date>
” (
<specimenCount id="ECA5FD538839FFDA2FE7FCEDFE22FCB9" box="[433,506,836,861]" pageId="9" pageNumber="249" type="male">1 male</specimenCount>
in ZNS,
<specimenCount id="ECA5FD538839FFDA2C37FCEDFD6FFCB9" box="[609,695,836,861]" pageId="9" pageNumber="249" type="male">3 males</specimenCount>
in
<collectionCode id="9CB2AE1F8839FFDA2C8DFCEDFCF8FCB8" box="[731,800,836,860]" pageId="9" pageNumber="249">CCLI</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B965518839FFDA2E91FCCEFD5AFC64" box="[199,642,871,896]" pageId="9" pageNumber="249" type="distribution">
<paragraph id="FA1C36DA8839FFDA2E91FCCEFD5AFC64" blockId="9.[151,1436,151,1188]" box="[199,642,871,896]" pageId="9" pageNumber="249">
<emphasis id="C8D7EAC88839FFDA2E91FCCEFEB8FC64" bold="true" box="[199,352,871,896]" pageId="9" pageNumber="249">Distribution.</emphasis>
Central
<collectingCountry id="82B4764A8839FFDA2F96FCCEFDCAFC64" box="[448,530,871,896]" name="Taiwan" pageId="9" pageNumber="249">Taiwan</collectingCountry>
(
<figureCitation id="62982A5F8839FFDA2C74FCCEFDADFC64" box="[546,629,871,896]" captionStart="FIGURE 13" captionStartId="8.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1858]" captionTargetId="figure@8.[151,1436,195,1858]" captionTargetPageId="8" captionText="FIGURE 13. Distribution map of Megistophylla species in Taiwan: Megistophylla formosana (open circle), Megistophylla xitoui (open triangle)." httpUri="https://zenodo.org/record/258603/files/figure.png" pageId="9" pageNumber="249">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B965518839FFDA2E91FC25FD86FC14" pageId="9" pageNumber="249" type="etymology">
<paragraph id="FA1C36DA8839FFDA2E91FC25FD05FC2C" blockId="9.[151,1436,151,1188]" pageId="9" pageNumber="249">
<emphasis id="C8D7EAC88839FFDA2E91FC25FE96FC41" bold="true" box="[199,334,908,933]" pageId="9" pageNumber="249">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="251888788839FFDA2CE3FC24FD3DFC41" box="[693,741,909,933]" pageId="9" pageNumber="249">type</typeStatus>
locality, Xitou, a mid-elevated (approximately
<quantity id="3D5B9B3F8839FFDA2AAEFC25FA4CFC40" box="[1272,1428,908,933]" metricMagnitude="3" metricUnit="m" metricValue="1.525" metricValueMax="1.8" metricValueMin="1.25" pageId="9" pageNumber="249" unit="m" value="1525.0" valueMax="1800.0" valueMin="1250.0">12501800 m</quantity>
) humid throughout the year valley in central
<collectingCountry id="82B4764A8839FFDA2CD5FC06FD01FC2C" box="[643,729,943,968]" name="Taiwan" pageId="9" pageNumber="249">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="FA1C36DA8839FFDA2E91FC7EFD86FC14" blockId="9.[151,1436,151,1188]" box="[199,606,983,1008]" pageId="9" pageNumber="249">
<emphasis id="C8D7EAC88839FFDA2E91FC7EFEAFFC14" bold="true" box="[199,375,983,1008]" pageId="9" pageNumber="249">Chinese name:</emphasis>
溪頭脊頭多鰓金龜
</paragraph>
</subSubSection>
<subSubSection id="B2B965518839FFDA2E91FC52FAC1FB40" pageId="9" pageNumber="249" type="discussion">
<paragraph id="FA1C36DA8839FFDA2E91FC52FAC1FB40" blockId="9.[151,1436,151,1188]" pageId="9" pageNumber="249">
<emphasis id="C8D7EAC88839FFDA2E91FC52FEE3FBF0" bold="true" box="[199,315,1019,1044]" pageId="9" pageNumber="249">Remarks.</emphasis>
This new species is only known from the
<typeStatus id="251888788839FFDA2D70FC55FC8EFBF0" box="[806,854,1020,1044]" pageId="9" pageNumber="249">type</typeStatus>
locality. The
<typeStatus id="251888788839FFDA2DA1FC55FBFFFBF0" box="[1015,1063,1020,1044]" pageId="9" pageNumber="249">type</typeStatus>
locality is approximately
<quantity id="3D5B9B3F8839FFDA2B05FC52FA44FBF0" box="[1363,1436,1019,1044]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="249" unit="km" value="15.0">15 km</quantity>
south of a
<taxonomicName id="3DA34D598839FFDA2F44FB88FE6AFBDC" box="[274,434,1056,1080]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88839FFDA2F44FB88FEF1FBDC" box="[274,297,1057,1080]" italics="true" pageId="9" pageNumber="249">M</emphasis>
.
<emphasis id="C8D7EAC88839FFDA2F6CFB89FE6AFBDC" box="[314,434,1056,1080]" italics="true" pageId="9" pageNumber="249">formosana</emphasis>
</taxonomicName>
locality (approximately
<quantity id="3D5B9B3F8839FFDA2C9AFBB6FCCDFBD3" box="[716,789,1055,1080]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="249" unit="m" value="600.0">600 m</quantity>
in elevation), without any significant geographic barrier between these localities. Interestingly, elevation is probably not an important factor separating these species because
<taxonomicName id="3DA34D598839FFDA2EAEFBC0FE4EFB64" box="[248,406,1128,1152]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88839FFDA2EAEFBC0FED7FB64" box="[248,271,1129,1152]" italics="true" pageId="9" pageNumber="249">M</emphasis>
.
<emphasis id="C8D7EAC88839FFDA2F48FBC1FE4EFB64" box="[286,406,1128,1152]" italics="true" pageId="9" pageNumber="249">formosana</emphasis>
</taxonomicName>
is found from
<quantity id="3D5B9B3F8839FFDA2C14FBCEFD0AFB64" box="[578,722,1127,1152]" metricMagnitude="3" metricUnit="m" metricValue="1.02" metricValueMax="1.44" metricValueMin="0.6" pageId="9" pageNumber="249" unit="m" value="1020.0" valueMax="1440.0" valueMin="600.0">6001440 m</quantity>
, which partly overlaps the known elevation of
<taxonomicName id="3DA34D598839FFDA2ABCFBC0FA88FB64" box="[1258,1360,1129,1152]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="species" species="xitoui">
<emphasis id="C8D7EAC88839FFDA2ABCFBC0FA88FB64" box="[1258,1360,1129,1152]" italics="true" pageId="9" pageNumber="249">M. xitoui</emphasis>
</taxonomicName>
. More field investigations are needed to see if these species of
<taxonomicName id="3DA34D598839FFDA2D5FFB22FC71FB40" box="[777,937,1163,1188]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="C8D7EAC88839FFDA2D5FFB22FC71FB40" box="[777,937,1163,1188]" italics="true" pageId="9" pageNumber="249">Megistophylla</emphasis>
</taxonomicName>
are sympatric in central
<collectingCountry id="82B4764A8839FFDA2AE9FB22FACDFB40" box="[1215,1301,1163,1188]" name="Taiwan" pageId="9" pageNumber="249">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>