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<document id="94E4C29E3F016645D94D99387ED8DBEF" ID="10.11646/zootaxa.4126.2.4" ID-CLB-Dataset="38797" ID-DOI="10.11646/zootaxa.4126.2.4" ID-GBIF-Dataset="46cf6acd-87dc-4165-80fa-e2b5330cd74c" ID-ISSN="1175-5326" ID-Zenodo-Dep="258598" ID-ZooBank="54F4A01E-2BB2-4802-9B93-8593DA846008" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466149131865" checkinUser="plazi" docAuthor="Li, Chun-Lin, Yang, Ping-Shih &amp; Wang, Chuan-Chan" docDate="2016" docId="720A87CC8834FFD62EC1FE72FBA0FC28" docLanguage="en" docName="zootaxa.4126.2.4.pdf" docOrigin="Zootaxa 4126 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megistophylla formosana Wang &amp; Li, new species" docType="treatment" docVersion="9" lastPageNumber="245" masterDocId="8E33FFB48830FFD32E56FFA9FFD8FFE4" masterDocTitle="Two new species of Megistophylla Burmeister (Coleoptera: Scarabaeidae: Melolonthinae) from Taiwan, with notes on the genus" masterLastPageNumber="250" masterPageNumber="240" pageNumber="244" updateTime="1698663021023" updateUser="ExternalLinkService">
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<mods:title id="E03E1E4D8185BD1491575C4C10C5C3EB">Two new species of Megistophylla Burmeister (Coleoptera: Scarabaeidae: Melolonthinae) from Taiwan, with notes on the genus</mods:title>
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<mods:namePart id="2E51E579B90EE5E6DDD5BE6A2F009C1B">Li, Chun-Lin</mods:namePart>
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<mods:namePart id="B94448603FC1EA85310EB1A453C15848">Yang, Ping-Shih</mods:namePart>
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<mods:namePart id="4FBD36DDB0BDF05EE0D344A63FFE1AA2">Wang, Chuan-Chan</mods:namePart>
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<mods:date id="FAA781019E9370937ED4F8D0522C91D9">2016</mods:date>
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<mods:identifier id="93D5A92E9CA7325F35FE1911D6098FE5" type="DOI">10.11646/zootaxa.4126.2.4</mods:identifier>
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<treatment id="720A87CC8834FFD62EC1FE72FBA0FC28" ID-DOI="http://doi.org/10.5281/zenodo.6059521" ID-GBIF-Taxon="120829033" ID-Zenodo-Dep="6059521" LSID="urn:lsid:plazi:treatment:720A87CC8834FFD62EC1FE72FBA0FC28" httpUri="http://treatment.plazi.org/id/720A87CC8834FFD62EC1FE72FBA0FC28" lastPageId="5" lastPageNumber="245" pageId="4" pageNumber="244">
<subSubSection id="B2B965518834FFD72EC1FE72FE5DFDF3" pageId="4" pageNumber="244" type="nomenclature">
<paragraph id="FA1C36DA8834FFD72EC1FE72FD25FE12" blockId="4.[151,765,475,535]" box="[151,765,475,502]" pageId="4" pageNumber="244">
<heading id="A15481B68834FFD72EC1FE72FD25FE12" bold="true" box="[151,765,475,502]" fontSize="11" level="1" pageId="4" pageNumber="244" reason="1">
<emphasis id="C8D7EAC88834FFD72EC1FE72FD25FE12" bold="true" box="[151,765,475,502]" pageId="4" pageNumber="244">
<taxonomicName id="3DA34D598834FFD72EC1FE72FDB8FE12" authority="Wang &amp; Li" authorityName="Wang &amp; Li" box="[151,608,475,502]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="244" phylum="Arthropoda" rank="species" species="formosana" status="sp. nov.">
<emphasis id="C8D7EAC88834FFD72EC1FE72FE13FE11" bold="true" box="[151,459,475,501]" italics="true" pageId="4" pageNumber="244">Megistophylla formosana</emphasis>
Wang &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="D3E457B38834FFD72C3BFE75FD25FE12" box="[621,765,476,502]" pageId="4" pageNumber="244" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FA1C36DA8834FFD72EC1FE57FE5DFDF3" blockId="4.[151,765,475,535]" box="[151,389,510,535]" pageId="4" pageNumber="244">
<figureCitation id="62982A5F8834FFD72EC1FE57FF3BFDF3" box="[151,227,510,535]" captionStart="FIGURES 1 – 2" captionStartId="5.[151,264,1963,1986]" captionTargetBox="[213,1374,1039,1934]" captionTargetId="figure@5.[197,1390,1027,1942]" captionTargetPageId="5" captionText="FIGURES 1 – 2. Dorsal habitus of male Megistophylla. 1. Megistophylla formosana new species, holotype; 2. Megistophylla xitoui new species, holotype." httpUri="https://zenodo.org/record/258599/files/figure.png" pageId="4" pageNumber="244">Figs. 1</figureCitation>
,
<figureCitation id="62982A5F8834FFD72EA7FE56FEC5FDF3" box="[241,285,511,535]" captionStart="FIGURES 3 – 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 – 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 – 4. Megistophylla formosana new species; 5 – 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="4" pageNumber="244">3–4</figureCitation>
,
<figureCitation id="62982A5F8834FFD72F7FFE56FEE1FDF2" box="[297,313,511,534]" captionStart="FIGURES 7 – 8" captionStartId="6.[151,264,1585,1608]" captionTargetBox="[153,1436,1148,1564]" captionTargetId="figure@6.[151,1436,1148,1564]" captionTargetPageId="6" captionText="FIGURES 7 – 8. Ventral view of protarsomeres 2 – 3 of Megistophylla. 7. Megistophylla formosana new species; 8. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258601/files/figure.png" pageId="4" pageNumber="244">7</figureCitation>
,
<figureCitation id="62982A5F8834FFD72F13FE56FE59FDF3" box="[325,385,511,535]" captionStart="FIGURES 9 – 12" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[166,1427,928,1704]" captionTargetId="figure@7.[151,1436,916,1709]" captionTargetPageId="7" captionText="FIGURES 9 – 12. Parameres of Megistophylla. (9, 11, lateral view; 10, 12, dorsal view). 9 – 10. Megistophylla formosana new species; 11 – 12. Megistophylla xitoui new species. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/258602/files/figure.png" pageId="4" pageNumber="244">9–10</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B965518834FFD72EC1FDEFFD00FA07" pageId="4" pageNumber="244" type="materials_examined">
<paragraph id="FA1C36DA8834FFD72EC1FDEFFD00FA07" blockId="4.[151,1437,582,2011]" pageId="4" pageNumber="244">
<emphasis id="C8D7EAC88834FFD72EC1FDEFFE9EFDBB" bold="true" box="[151,326,582,607]" pageId="4" pageNumber="244">
<typeStatus id="251888788834FFD72EC1FDEFFEDCFDBB" box="[151,260,582,607]" pageId="4" pageNumber="244" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<figureCitation id="62982A5F8834FFD72F03FDEFFE4FFDBB" box="[341,407,582,607]" captionStart="FIGURES 1 – 2" captionStartId="5.[151,264,1963,1986]" captionTargetBox="[213,1374,1039,1934]" captionTargetId="figure@5.[197,1390,1027,1942]" captionTargetPageId="5" captionText="FIGURES 1 – 2. Dorsal habitus of male Megistophylla. 1. Megistophylla formosana new species, holotype; 2. Megistophylla xitoui new species, holotype." httpUri="https://zenodo.org/record/258599/files/figure.png" pageId="4" pageNumber="244">Fig. 1</figureCitation>
). Body length
<quantity id="3D5B9B3F8834FFD72C15FDEFFD73FDBA" box="[579,683,582,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="4" pageNumber="244" unit="mm" value="19.5">19.5 mm</quantity>
; width across humeri
<quantity id="3D5B9B3F8834FFD72DF2FDEEFC26FDBA" box="[932,1022,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="4" pageNumber="244" unit="mm" value="8.7">8.7 mm</quantity>
. Color dark reddish brown on dorsal surface, venter and legs moderately shiny. Antennomeres reddish brown. Dorsum overall setiferous, setae tiny (high magnification for observation required); setae on disc of pronotum and elytra far smaller than diameter of puncture with exception of a row of long; sparse setae at base of head, anterior, and posterior margins of pronotum and base of elytra, respectively; long setae brown in color, 1–7 times longer than diameter of puncture.
<emphasis id="C8D7EAC88834FFD72B00FD7FFA44FD0B" box="[1366,1436,726,751]" italics="true" pageId="4" pageNumber="244">Head:</emphasis>
Surface densely rugopunctate, punctures large. Clypeus (
<figureCitation id="62982A5F8834FFD72D65FD53FCA3FCF7" box="[819,891,762,787]" captionStart="FIGURES 3 – 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 – 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 – 4. Megistophylla formosana new species; 5 – 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="4" pageNumber="244">Fig. 3</figureCitation>
) with apex distinctly bilobed, anterior margin moderately reflexed, narrowed at base. Clypeofrontal suture clearly defined. Frons moderately raised. Basal carina slightly crescent, strongly raised. Antenna with 10 antennomeres, antennal club with 6 antennomeres, length of club subequal to antennomeres 1–4 combined, first antennomere of club slightly shorter than remaining club antennomeres; fourth basal antennomere greatly enlarged.
<emphasis id="C8D7EAC88834FFD72D73FC23FCA7FC47" box="[805,895,906,931]" italics="true" pageId="4" pageNumber="244">Thorax:</emphasis>
Pronotum widest at middle, anterolateral angles (
<figureCitation id="62982A5F8834FFD72EC9FC07FF3AFC23" box="[159,226,942,967]" captionStart="FIGURES 3 – 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 – 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 – 4. Megistophylla formosana new species; 5 – 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="4" pageNumber="244">Fig. 3</figureCitation>
) moderately to strongly protuberant with basolateral angles obtuse, basal half of lateral margin moderately serrate; lateral declivities with weak gibbosity at middle; anterior margin completely beaded with basal margin smooth; punctures same as those on clypeus, rarely confluent, punctures evenly distributed along midline (
<figureCitation id="62982A5F8834FFD72B1FFC5FFA57FBEB" box="[1353,1423,1014,1039]" captionStart="FIGURES 3 – 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 – 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 – 4. Megistophylla formosana new species; 5 – 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="4" pageNumber="244">Fig. 4</figureCitation>
). Scutellum (
<figureCitation id="62982A5F8834FFD72F4FFBB3FE86FBD7" box="[281,350,1050,1075]" captionStart="FIGURES 3 – 6" captionStartId="6.[151,264,1045,1068]" captionTargetBox="[151,1436,193,1023]" captionTargetId="figure@6.[151,1436,193,1024]" captionTargetPageId="6" captionText="FIGURES 3 – 6. Close view of dorsum of male Megistophylla (3, 5 head and anterior half of pronotum; 4, 6 basal half of pronotum, scutellum. and elytra). 3 – 4. Megistophylla formosana new species; 5 – 6. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258600/files/figure.png" pageId="4" pageNumber="244">Fig. 4</figureCitation>
) triangular in shape with 10 punctures in uneven distribution, punctures rarely with long setae. Prosternal process flat, weakly bilobed in shape.
<emphasis id="C8D7EAC88834FFD72CE8FB97FCD5FBB3" box="[702,781,1086,1111]" italics="true" pageId="4" pageNumber="244">Elytra:</emphasis>
Discal surface densely rugopunctate, densely confluent at sides of suture; sutural costa gradually broadening apically, densely punctate at margins or transversally rugopunctate over costa; costa becoming flat at apex.
<emphasis id="C8D7EAC88834FFD72CA5FB2FFC85FB7B" box="[755,861,1158,1183]" italics="true" pageId="4" pageNumber="244">Abdomen</emphasis>
: Pygidium wider than long; surface with setiferous punctures in uneven distribution, tiny setae same as those of elytral disc. Sternites 1–2, 5–6 with surface densely punctate; punctures in sternites 3–4 sparsely distributed, punctures setiferous; setae on sternites 3–4 tiny, intermixed on sternites 5–6 with much longer setae.
<emphasis id="C8D7EAC88834FFD72D57FB5DFCEEFAEF" box="[769,822,1268,1291]" italics="true" pageId="4" pageNumber="244">Legs</emphasis>
: Protibia tridentate. Protarsomeres 2–3 (
<figureCitation id="62982A5F8834FFD72B46FB5BFA8FFAEF" box="[1296,1367,1266,1291]" captionStart="FIGURES 7 – 8" captionStartId="6.[151,264,1585,1608]" captionTargetBox="[153,1436,1148,1564]" captionTargetId="figure@6.[151,1436,1148,1564]" captionTargetPageId="6" captionText="FIGURES 7 – 8. Ventral view of protarsomeres 2 – 3 of Megistophylla. 7. Megistophylla formosana new species; 8. Megistophylla xitoui new species." httpUri="https://zenodo.org/record/258601/files/figure.png" pageId="4" pageNumber="244">Fig. 7</figureCitation>
) with dense setae aggregated at apices. Surface of profemora and mesofemora moderately distributed with hair-like, long setae; setae on metafemora shorter and more robust, metafemora slightly broadened at middle. Metatibia with upper apical spur reaching basal one-fourth of metatarsomere 2; lower apical spur slightly shorter than upper one.
<emphasis id="C8D7EAC88834FFD72EC1FA2BFED9FA7F" box="[151,257,1410,1435]" italics="true" pageId="4" pageNumber="244">Genitalia</emphasis>
: Parameres in lateral view with central concave area extended and concentrically wrinkled, apical upper margin distinctly swollen with tip broadly rounded (
<figureCitation id="62982A5F8834FFD72D52FA0FFC96FA5B" box="[772,846,1446,1471]" captionStart="FIGURES 9 – 12" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[166,1427,928,1704]" captionTargetId="figure@7.[151,1436,916,1709]" captionTargetPageId="7" captionText="FIGURES 9 – 12. Parameres of Megistophylla. (9, 11, lateral view; 10, 12, dorsal view). 9 – 10. Megistophylla formosana new species; 11 – 12. Megistophylla xitoui new species. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/258602/files/figure.png" pageId="4" pageNumber="244">Fig. 9</figureCitation>
); upper margin of parameres gradually narrowing apically at apical one-third in dorsal view (
<figureCitation id="62982A5F8834FFD72C21FA63FD13FA07" box="[631,715,1482,1507]" captionStart="FIGURES 9 – 12" captionStartId="7.[151,264,1729,1752]" captionTargetBox="[166,1427,928,1704]" captionTargetId="figure@7.[151,1436,916,1709]" captionTargetPageId="7" captionText="FIGURES 9 – 12. Parameres of Megistophylla. (9, 11, lateral view; 10, 12, dorsal view). 9 – 10. Megistophylla formosana new species; 11 – 12. Megistophylla xitoui new species. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/258602/files/figure.png" pageId="4" pageNumber="244">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B965518834FFD72E91FA47FE72F95E" pageId="4" pageNumber="244" type="description">
<paragraph id="FA1C36DA8834FFD72E91FA47FE7AF9E3" blockId="4.[151,1437,582,2011]" box="[199,418,1518,1543]" pageId="4" pageNumber="244">
<emphasis id="C8D7EAC88834FFD72E91FA47FEC6F9E3" bold="true" box="[199,286,1518,1543]" pageId="4" pageNumber="244">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="FA1C36DA8834FFD72E91F9BBFE72F95E" blockId="4.[151,1437,582,2011]" pageId="4" pageNumber="244">
<emphasis id="C8D7EAC88834FFD72E91F9BBFEEDF9CF" bold="true" box="[199,309,1554,1579]" pageId="4" pageNumber="244">Variation</emphasis>
. Body length
<quantity id="3D5B9B3F8834FFD72FB4F9BBFD55F9CE" box="[482,653,1554,1579]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" metricValueMax="2.15" metricValueMin="1.5699999999999998" pageId="4" pageNumber="244" unit="mm" value="18.6" valueMax="21.5" valueMin="15.7">15.7–21.5 mm</quantity>
; width across humeri
<quantity id="3D5B9B3F8834FFD72DC4F9BBFBF8F9CE" box="[914,1056,1554,1579]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="9.5" metricValueMin="7.8" pageId="4" pageNumber="244" unit="mm" value="8.65" valueMax="9.5" valueMin="7.8">7.8–9.5 mm</quantity>
. Dorsal body color dark reddish brown to rufotestaceous; antennae reddish brown to blackish brown; anterior margin of clypeus moderately to weakly reflexed; pronotal punctures usually evenly distributed along midline; scutellum usually with 10 or more punctures in uneven distribution, punctures rarely with long setae; elytral surface sometimes transversally rugopunctate over costa.
</paragraph>
</subSubSection>
<subSubSection id="B2B965518834FFD72E91F96FFD7BF88B" pageId="4" pageNumber="244" type="diagnosis">
<paragraph id="FA1C36DA8834FFD72E91F96FFD7BF88B" blockId="4.[151,1437,582,2011]" pageId="4" pageNumber="244">
<emphasis id="C8D7EAC88834FFD72E91F96FFE99F93B" bold="true" box="[199,321,1734,1759]" pageId="4" pageNumber="244">Diagnosis.</emphasis>
<taxonomicName id="3DA34D598834FFD72F1AF96FFDB7F93B" box="[332,623,1734,1759]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="244" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88834FFD72F1AF96FFDB7F93B" box="[332,623,1734,1759]" italics="true" pageId="4" pageNumber="244">Megistophylla formosana</emphasis>
</taxonomicName>
is distinguished from the other
<taxonomicName id="3DA34D598834FFD72DBAF96FFB54F93B" box="[1004,1164,1734,1759]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="C8D7EAC88834FFD72DBAF96FFB54F93B" box="[1004,1164,1734,1759]" italics="true" pageId="4" pageNumber="244">Megistophylla</emphasis>
</taxonomicName>
species (except for
<taxonomicName id="3DA34D598834FFD72B2BF96EFF0FF8E7" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="244" phylum="Arthropoda" rank="species" species="xitoui">
<emphasis id="C8D7EAC88834FFD72B2BF96EFA4CF93A" box="[1405,1428,1735,1758]" italics="true" pageId="4" pageNumber="244">M</emphasis>
.
<emphasis id="C8D7EAC88834FFD72EC1F945FF0FF8E7" box="[151,215,1772,1795]" italics="true" pageId="4" pageNumber="244">xitoui</emphasis>
</taxonomicName>
) by its distinctly straight antennal club with length subequal to basal antennomeres (antennal club slightly to strongly curved and the length of club longer than basal antennomeres in other species); basal half of the pronotal lateral margins serrate (smooth in other species); and central concave area of the parameres wrinkled concentrically (non-concentrically wrinkled in other species).
</paragraph>
</subSubSection>
<subSubSection id="B2B965518834FFD62E91F8D3FC8DFD14" lastPageId="5" lastPageNumber="245" pageId="4" pageNumber="244" type="materials_examined">
<paragraph id="FA1C36DA8834FFD62E91F8D3FC8DFD14" blockId="4.[151,1437,582,2011]" lastBlockId="5.[151,1437,151,972]" lastPageId="5" lastPageNumber="245" pageId="4" pageNumber="244">
<emphasis id="C8D7EAC88834FFD72E91F8D3FEAFF877" bold="true" box="[199,375,1914,1939]" pageId="4" pageNumber="244">
<typeStatus id="251888788834FFD72E91F8D3FEDAF877" box="[199,258,1914,1939]" pageId="4" pageNumber="244">Type</typeStatus>
material.
</emphasis>
Male
<typeStatus id="251888788834FFD72F92F8D3FDFEF877" box="[452,550,1914,1939]" pageId="4" pageNumber="244" type="holotype">holotype</typeStatus>
and
<specimenCount id="ECA5FD538834FFD72C37F8D3FD65F877" box="[609,701,1914,1939]" pageId="4" pageNumber="244" type="male">61 male</specimenCount>
<typeStatus id="251888788834FFD72C91F8D2FCEEF877" box="[711,822,1915,1939]" pageId="4" pageNumber="244" type="paratype">paratypes</typeStatus>
. The male
<typeStatus id="251888788834FFD72DEEF8D3FBC2F877" box="[952,1050,1914,1939]" pageId="4" pageNumber="244" type="holotype">holotype</typeStatus>
is pinned and with the following information on the label: “N.
<collectingCountry id="82B4764A8834FFD72FB4F837FDE0F853" box="[482,568,1950,1975]" name="Taiwan" pageId="4" pageNumber="244">Taiwan</collectingCountry>
: Taipei County (presently New Taipei City), Wulai, Nei Dong Logging Road, alt. ca.
<quantity id="3D5B9B3F8834FFD72EB9F86BFEE4F83E" box="[239,316,1986,2011]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="244" unit="m" value="850.0">850 m</quantity>
,
<date id="8E1D101A8834FFD72F1AF86BFE14F83F" box="[332,460,1986,2011]" pageId="4" pageNumber="244" value="2003-04-16">16/IV/2003</date>
by FIT, collr. C.-L. Li”. The
<typeStatus id="251888788834FFD72D79F86BFC49F83F" box="[815,913,1986,2011]" pageId="4" pageNumber="244" type="holotype">holotype</typeStatus>
is deposited at the National Natural Science Museum, Taichung,
<collectingCountry id="82B4764A8835FFD62F28FF3EFE0CFF54" box="[382,468,151,176]" name="Taiwan" pageId="5" pageNumber="245">Taiwan</collectingCountry>
.
<typeStatus id="251888788835FFD62F88FF31FD88FF54" box="[478,592,152,176]" pageId="5" pageNumber="245" type="paratype">Paratypes</typeStatus>
: Locality same as
<typeStatus id="251888788835FFD62D76FF3EFC5DFF54" box="[800,901,151,176]" pageId="5" pageNumber="245" type="holotype">holotype</typeStatus>
, “
<date id="8E1D101A8835FFD62DCDFF31FBC8FF54" box="[923,1040,152,176]" pageId="5" pageNumber="245" value="2003-04-07">7/IV/2003</date>
by FIT, collr. C.-L. Li” (
<specimenCount id="ECA5FD538835FFD62B7BFF31FAA2FF54" box="[1325,1402,151,176]" pageId="5" pageNumber="245" type="male">1 male</specimenCount>
, in
<collectionCode id="9CB2AE1F8835FFD62EC1FF15FF04FF30" box="[151,220,188,212]" pageId="5" pageNumber="245">CCLI</collectionCode>
); “
<date id="8E1D101A8835FFD62EAAFF15FE68FF31" box="[252,432,188,213]" pageId="5" pageNumber="245" value="2005-04-05" valueMax="2005-05-10" valueMin="2005-04-05">5/IV-10/V/2005</date>
by FIT, collr. C.-L. Li” (
<specimenCount id="ECA5FD538835FFD62C8CFF15FCF1FF31" box="[730,809,188,213]" pageId="5" pageNumber="245" type="male">1 male</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62D61FF15FCA4FF30" box="[823,892,188,212]" pageId="5" pageNumber="245">CCLI</collectionCode>
); “
<date id="8E1D101A8835FFD62DCBFF15FBC7FF30" box="[925,1055,188,212]" pageId="5" pageNumber="245" value="2003-03-17">17/III/2003</date>
by FIT, collr. C.-L. Li” (
<specimenCount id="ECA5FD538835FFD62B1FFF15FA40FF31" box="[1353,1432,188,213]" pageId="5" pageNumber="245" type="male">1 male</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62EC1FF49FF04FF1C" box="[151,220,224,248]" pageId="5" pageNumber="245">CCLI</collectionCode>
); “
<date id="8E1D101A8835FFD62EAFFF49FEA4FF1C" box="[249,380,224,248]" pageId="5" pageNumber="245" value="2003-04-11">11/IV/2003</date>
by FIT, collr. C.-L. Li” (
<specimenCount id="ECA5FD538835FFD62CCEFF49FD3DFF1C" box="[664,741,223,248]" pageId="5" pageNumber="245" type="male">1 male</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62CA6FF49FCEDFF1C" box="[752,821,224,248]" pageId="5" pageNumber="245">CCLI</collectionCode>
); “
<date id="8E1D101A8835FFD62D05FF49FC0FFF1C" box="[851,983,224,248]" pageId="5" pageNumber="245" value="2003-04-18">18/IV/2003</date>
by FIT, collr. C.-L. Li” (
<specimenCount id="ECA5FD538835FFD62AA5FF49FAE7FF1C" box="[1267,1343,223,248]" pageId="5" pageNumber="245" type="male">1 male</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62B1DFF49FA48FF1C" box="[1355,1424,224,248]" pageId="5" pageNumber="245">CCLI</collectionCode>
); “
<date id="8E1D101A8835FFD62EF7FEADFEB6FEF9" box="[161,366,260,285]" pageId="5" pageNumber="245" value="2004-03-31" valueMax="2004-04-26" valueMin="2004-03-31">31/III-26/IV/2004</date>
by FIT, collr. C.-L. Li” (
<specimenCount id="ECA5FD538835FFD62CC1FEADFD2CFEF9" box="[663,756,260,285]" pageId="5" pageNumber="245" type="male">2 males</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62D55FEADFC90FEF8" box="[771,840,260,284]" pageId="5" pageNumber="245">CCLI</collectionCode>
); “
<date id="8E1D101A8835FFD62D3EFEADFBC4FEF9" box="[872,1052,260,285]" pageId="5" pageNumber="245" value="2005-04-05" valueMax="2005-05-10" valueMin="2005-04-05">5/IV-10/V/2005</date>
by FIT, collr. C.-L. Li” (
<specimenCount id="ECA5FD538835FFD62B1EFEADFA40FEF9" box="[1352,1432,260,285]" pageId="5" pageNumber="245" type="male">1 male</specimenCount>
, deposited at the Natural History Museum, London,
<collectingCountry id="82B4764A8835FFD62CA6FE8EFC68FEA4" box="[752,944,295,320]" name="United Kingdom" pageId="5" pageNumber="245">United Kingdom</collectingCountry>
(
<collectionCode id="9CB2AE1F8835FFD62D94FE81FBC4FEA4" box="[962,1052,296,320]" pageId="5" pageNumber="245">BMNH</collectionCode>
); “Taoyuan County: Fu-Fu Shan,
<geoCoordinate id="9F97501D8835FFD62EC1FEE5FEF6FE80" box="[151,302,332,356]" direction="east" orientation="longitude" pageId="5" pageNumber="245" precision="15" value="121.37417">121°22’27’’E</geoCoordinate>
/
<geoCoordinate id="9F97501D8835FFD62F6EFEE5FE1EFE80" box="[312,454,332,356]" direction="north" orientation="latitude" pageId="5" pageNumber="245" precision="15" value="24.726112">24°43’34’’N</geoCoordinate>
, alt. ca.
<quantity id="3D5B9B3F8835FFD62C7FFEE5FD52FE80" box="[553,650,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="5" pageNumber="245" unit="m" value="1440.0">1,440 m</quantity>
, by FIT,
<date id="8E1D101A8835FFD62CA3FEE5FC47FE81" box="[757,927,332,357]" pageId="5" pageNumber="245" value="2004-04-15" valueMax="2004-04-28" valueMin="2004-04-15">15-28/IV/2004</date>
, collr. C-L. Li” (
<specimenCount id="ECA5FD538835FFD62A37FEE5FB66FE81" box="[1121,1214,332,357]" pageId="5" pageNumber="245" type="male">4 males</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62A9DFEE5FAC8FE80" box="[1227,1296,332,356]" pageId="5" pageNumber="245">CCLI</collectionCode>
); “Shih Lin,
<geoCoordinate id="9F97501D8835FFD62EC1FEC6FEF6FE6C" box="[151,302,367,392]" direction="east" orientation="longitude" pageId="5" pageNumber="245" precision="15" value="121.42222">121°25’20’’E</geoCoordinate>
/
<geoCoordinate id="9F97501D8835FFD62F6EFEC6FE1EFE6C" box="[312,454,367,392]" direction="north" orientation="latitude" pageId="5" pageNumber="245" precision="15" value="24.649445">24°38’58’’N</geoCoordinate>
, alt. ca.
<quantity id="3D5B9B3F8835FFD62C75FED9FDA2FE6C" box="[547,634,368,392]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="5" pageNumber="245" unit="m" value="1180.0">1,180m</quantity>
, by FIT,
<date id="8E1D101A8835FFD62C89FEC6FC4BFE6C" box="[735,915,367,392]" pageId="5" pageNumber="245" value="2005-04-09" valueMax="2005-05-17" valueMin="2005-04-09">9/IV-17/V/2005</date>
, collr. C-L. Li” (
<specimenCount id="ECA5FD538835FFD62A19FEC6FB7DFE6C" box="[1103,1189,367,392]" pageId="5" pageNumber="245" type="male">5 males</specimenCount>
in Canadian Museum of Nature, Ottawa,
<collectingCountry id="82B4764A8835FFD62F38FE3DFE1AFE49" box="[366,450,404,429]" name="Canada" pageId="5" pageNumber="245">Canada</collectingCountry>
(
<collectionCode id="9CB2AE1F8835FFD62F87FE3DFDF1FE48" box="[465,553,404,428]" pageId="5" pageNumber="245">CMNC</collectionCode>
),
<specimenCount id="ECA5FD538835FFD62C6BFE3DFD4EFE49" box="[573,662,404,429]" pageId="5" pageNumber="245" type="male">6 males</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62CF7FE3DFD3DFE48" box="[673,741,404,428]" pageId="5" pageNumber="245">CCLI</collectionCode>
); same locality, “
<date id="8E1D101A8835FFD62DF3FE3DFBEFFE48" box="[933,1079,404,428]" pageId="5" pageNumber="245" value="2004-05-01" valueMax="2004-05-17" valueMin="2004-05-01">1-17/V/2004</date>
” (
<specimenCount id="ECA5FD538835FFD62A07FE3DFB6EFE49" box="[1105,1206,404,429]" pageId="5" pageNumber="245" type="male">20 males</specimenCount>
in
<collectionCode id="9CB2AE1F8835FFD62A8CFE3DFAC5FE48" box="[1242,1309,404,428]" pageId="5" pageNumber="245">CCLI</collectionCode>
;
<specimenCount id="ECA5FD538835FFD62B7FFE3DFAA7FE49" box="[1321,1407,404,429]" pageId="5" pageNumber="245" type="male">4 males</specimenCount>
in Museum für Naturkunde der Humboldt Universitat, Berlin,
<collectingCountry id="82B4764A8835FFD62D60FE11FC46FE34" box="[822,926,440,464]" name="Germany" pageId="5" pageNumber="245">Germany</collectingCountry>
(
<collectionCode id="9CB2AE1F8835FFD62DFBFE11FBDCFE34" box="[941,1028,440,464]" pageId="5" pageNumber="245">ZMHB</collectionCode>
);
<specimenCount id="ECA5FD538835FFD62A4EFE11FBB6FE34" box="[1048,1134,439,464]" pageId="5" pageNumber="245" type="male">2 males</specimenCount>
in Museum für Tierkunde, Senckenberg Naturhistorische Sammlungen Dresden,
<collectingCountry id="82B4764A8835FFD62D14FE75FC72FE10" box="[834,938,476,500]" name="Germany" pageId="5" pageNumber="245">Germany</collectingCountry>
(
<collectionCode id="9CB2AE1F8835FFD62D9BFE75FBC9FE10" box="[973,1041,476,500]" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="5" pageNumber="245">MTD</collectionCode>
);
<specimenCount id="ECA5FD538835FFD62A6EFE75FB79FE11" box="[1080,1185,476,501]" pageId="5" pageNumber="245" type="male">2 males</specimenCount>
in Zentralmagazin Naturwissenschaftlicher Sammlungen der Martin-Luther-Universität Zoologische Sammlung, Halle (Saale),
<collectingCountry id="82B4764A8835FFD62EC1FD8DFF27FDD8" box="[151,255,548,572]" name="Germany" pageId="5" pageNumber="245">Germany</collectingCountry>
(ZNS)); same locality, “
<date id="8E1D101A8835FFD62C4FFD8DFD05FDD8" box="[537,733,548,572]" pageId="5" pageNumber="245" value="2004-05-20" valueMax="2004-06-14" valueMin="2004-05-20">20/V-14/VI/2004</date>
” (
<specimenCount id="ECA5FD538835FFD62CAFFD8DFC8DFDD9" box="[761,853,548,573]" pageId="5" pageNumber="245" type="male">2 males</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62D34FD8DFC63FDD8" box="[866,955,548,572]" pageId="5" pageNumber="245">BMNH</collectionCode>
); “Ilan County: Fushan Botanical Garden,
<date id="8E1D101A8835FFD62EC1FDE1FEC4FD84" box="[151,284,584,608]" pageId="5" pageNumber="245" value="1991-03-13">III-13-1991</date>
, Y. B. Fan” (
<specimenCount id="ECA5FD538835FFD62F94FDE1FDCCFD84" box="[450,532,583,608]" pageId="5" pageNumber="245" type="male">1 male</specimenCount>
,
<collectingCountry id="82B4764A8835FFD62C72FDEEFDAEFD84" box="[548,630,583,608]" name="Taiwan" pageId="5" pageNumber="245">Taiwan</collectingCountry>
Forest Research Institute, Taipei,
<collectingCountry id="82B4764A8835FFD62A5BFDEEFB87FD84" box="[1037,1119,583,608]" name="Taiwan" pageId="5" pageNumber="245">Taiwan</collectingCountry>
(
<collectionCode id="9CB2AE1F8835FFD62A25FDE1FB6DFD84" box="[1139,1205,584,608]" httpUri="http://grbio.org/cool/fitg-90ys" name="Taiwan Fisheries Research Institute" pageId="5" pageNumber="245">TFRI</collectionCode>
)); “Nantou County: Lienhuachih,
<date id="8E1D101A8835FFD62F64FDC5FE31FD61" box="[306,489,620,645]" pageId="5" pageNumber="245" value="2002-05-06" valueMax="2002-06-10" valueMin="2002-05-06">V/6-VI/10/2002</date>
, C.S. Lin &amp; W.T. Yang, Malaise trap (KCN)” (
<specimenCount id="ECA5FD538835FFD62A56FDC5FB83FD61" box="[1024,1115,620,645]" pageId="5" pageNumber="245" type="male">2 males</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62A3EFDC5FB66FD60" box="[1128,1214,620,644]" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="5" pageNumber="245">NMNS</collectionCode>
); same locality, “
<date id="8E1D101A8835FFD62BD6FDC5FF34FD4C" bridgedPair="/" pageId="5" pageNumber="245">II/ 14-III/7</date>
/2 0 0 5, C.S. Lin &amp; W.T. Yang, Malaise trap (KCN)” (
<specimenCount id="ECA5FD538835FFD62D05FD39FC77FD4C" box="[851,943,655,680]" pageId="5" pageNumber="245" type="male">2 males</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62DEBFD39FBCBFD4C" box="[957,1043,656,680]" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="5" pageNumber="245">NMNS</collectionCode>
); same locality, “
<date id="8E1D101A8835FFD62A8EFD26FA40FD4C" box="[1240,1432,655,680]" pageId="5" pageNumber="245" value="2005-03-07" valueMax="2005-04-11" valueMin="2005-03-07">III/7-IV/11/2005</date>
, C.S. Lin &amp; W.T. Yang, Malaise trap (KCN)” (
<specimenCount id="ECA5FD538835FFD62CF3FD1DFCD8FD29" box="[677,768,692,717]" pageId="5" pageNumber="245" type="male">2 males</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62D5BFD1DFCBBFD28" box="[781,867,692,716]" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="5" pageNumber="245">NMNS</collectionCode>
); same locality, “Watersher No. 3,
<date id="8E1D101A8835FFD62AA6FD1DFA40FD29" box="[1264,1432,692,717]" pageId="5" pageNumber="245" value="2003-03-04" valueMax="2003-05-06" valueMin="2003-03-04">III/4-V/6/2003</date>
, C.S. Lin &amp; W.T. Yang, Malaise trap (KCN)” (
<specimenCount id="ECA5FD538835FFD62CCDFD71FD3FFD14" box="[667,743,727,752]" pageId="5" pageNumber="245" type="male">1 male</specimenCount>
,
<collectionCode id="9CB2AE1F8835FFD62CA5FD71FC91FD14" box="[755,841,728,752]" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="5" pageNumber="245">NMNS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B965518835FFD62E91FD55FCC0FCF0" box="[199,792,764,789]" pageId="5" pageNumber="245" type="distribution">
<paragraph id="FA1C36DA8835FFD62E91FD55FCC0FCF0" blockId="5.[151,1437,151,972]" box="[199,792,764,789]" pageId="5" pageNumber="245">
<emphasis id="C8D7EAC88835FFD62E91FD55FEB8FCF1" bold="true" box="[199,352,764,789]" pageId="5" pageNumber="245">Distribution.</emphasis>
Northern and central
<collectingCountry id="82B4764A8835FFD62C00FD55FD70FCF1" box="[598,680,764,789]" name="Taiwan" pageId="5" pageNumber="245">Taiwan</collectingCountry>
(
<figureCitation id="62982A5F8835FFD62CE1FD55FCD2FCF0" box="[695,778,764,789]" captionStart="FIGURE 13" captionStartId="8.[151,250,1882,1904]" captionTargetBox="[151,1436,195,1858]" captionTargetId="figure@8.[151,1436,195,1858]" captionTargetPageId="8" captionText="FIGURE 13. Distribution map of Megistophylla species in Taiwan: Megistophylla formosana (open circle), Megistophylla xitoui (open triangle)." httpUri="https://zenodo.org/record/258603/files/figure.png" pageId="5" pageNumber="245">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B965518835FFD62E91FCB6FD86FC60" pageId="5" pageNumber="245" type="etymology">
<paragraph id="FA1C36DA8835FFD62E91FCB6FC5BFCB9" blockId="5.[151,1437,151,972]" pageId="5" pageNumber="245">
<emphasis id="C8D7EAC88835FFD62E91FCB6FE96FCDC" bold="true" box="[199,334,799,824]" pageId="5" pageNumber="245">Etymology.</emphasis>
The specific name refers to the historic name of
<collectingCountry id="82B4764A8835FFD62D25FCB6FC1DFCDC" box="[883,965,799,824]" name="Taiwan" pageId="5" pageNumber="245">Taiwan</collectingCountry>
by the Spanish sailors in the 16th century indicating the first
<taxonomicName id="3DA34D598835FFD62F3DFCEDFDD3FCB9" box="[363,523,836,861]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="C8D7EAC88835FFD62F3DFCEDFDD3FCB9" box="[363,523,836,861]" italics="true" pageId="5" pageNumber="245">Megistophylla</emphasis>
</taxonomicName>
species recognized from
<collectingCountry id="82B4764A8835FFD62D7FFCEDFCA7FCB9" box="[809,895,836,861]" name="Taiwan" pageId="5" pageNumber="245">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="FA1C36DA8835FFD62E91FCC2FD86FC60" blockId="5.[151,1437,151,972]" box="[199,606,875,900]" pageId="5" pageNumber="245">
<emphasis id="C8D7EAC88835FFD62E91FCC2FEAFFC60" bold="true" box="[199,375,875,900]" pageId="5" pageNumber="245">Chinese name:</emphasis>
蓬萊脊頭多鰓金龜
</paragraph>
</subSubSection>
<subSubSection id="B2B965518835FFD62E91FC26FBA0FC28" pageId="5" pageNumber="245" type="discussion">
<paragraph id="FA1C36DA8835FFD62E91FC26FBA0FC28" blockId="5.[151,1437,151,972]" pageId="5" pageNumber="245">
<emphasis id="C8D7EAC88835FFD62E91FC26FEE3FC4C" bold="true" box="[199,315,911,936]" pageId="5" pageNumber="245">Remarks.</emphasis>
<taxonomicName id="3DA34D598835FFD62F10FC26FDB2FC4C" box="[326,618,911,936]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="245" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="C8D7EAC88835FFD62F10FC26FDB2FC4C" box="[326,618,911,936]" italics="true" pageId="5" pageNumber="245">Megistophylla formosana</emphasis>
</taxonomicName>
has wider range in distribution, both in elevation (
<quantity id="3D5B9B3F8835FFD62A9DFC26FAB8FC4C" box="[1227,1376,911,936]" metricMagnitude="3" metricUnit="m" metricValue="1.02" metricValueMax="1.44" metricValueMin="0.6" pageId="5" pageNumber="245" unit="m" value="1020.0" valueMax="1440.0" valueMin="600.0">600–1440 m</quantity>
) and latitude (north throughout central
<collectingCountry id="82B4764A8835FFD62C44FC1AFDBEFC28" box="[530,614,947,972]" name="Taiwan" pageId="5" pageNumber="245">Taiwan</collectingCountry>
) than
<taxonomicName id="3DA34D598835FFD62CFAFC1CFCC9FC28" box="[684,785,948,972]" class="Insecta" family="Melolonthidae" genus="Megistophylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="245" phylum="Arthropoda" rank="species" species="xitoui">
<emphasis id="C8D7EAC88835FFD62CFAFC1CFD1BFC28" box="[684,707,949,972]" italics="true" pageId="5" pageNumber="245">M</emphasis>
.
<emphasis id="C8D7EAC88835FFD62C87FC1CFCC9FC28" box="[721,785,949,972]" italics="true" pageId="5" pageNumber="245">xitoui</emphasis>
</taxonomicName>
, the other species from
<collectingCountry id="82B4764A8835FFD62A48FC1AFBACFC28" box="[1054,1140,947,972]" name="Taiwan" pageId="5" pageNumber="245">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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