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<document ID-CLB-Dataset="6744" ID-DOI="http://dx.doi.org/10.3897/dez.68.66200" ID-GBIF-Dataset="22670907-a451-4089-a3b0-e16edba09a28" ID-Pensoft-Pub="1860-1324-1-189" ID-Pensoft-UUID="84A6EF91377850DF803DE5E2A4062204" ID-ZooBank="F88F5F2BEC5A408E98190188E87DB024" ModsDocID="1860-1324-1-189" checkinTime="1621365470254" checkinUser="pensoft" docAuthor="Yang, Zhen-ming, Jia, Fenglong, Jiang, Lu &amp; Guo, Qiang" docDate="2021" docId="7207B547081750D6A276E84D147BA280" docLanguage="en" docName="DeutEntomolZeit 68(1): 189-205" docOrigin="Deutsche Entomologische Zeitschrift 68 (1)" docPubDate="2021-05-18" docSource="http://dx.doi.org/10.3897/dez.68.66200" docTitle="Agraphydrus komareki Yang &amp; Jia 2021, sp. nov." docType="treatment" docUuid="C40E8204-E492-4147-B202-CF76A8871E2C" docUuidSource="ZooBank" docVersion="5" id="84A6EF91377850DF803DE5E2A4062204" lastPageNumber="189" masterDocId="84A6EF91377850DF803DE5E2A4062204" masterDocTitle="Four new species of Agraphydrus Regimbart, 1903 with additional faunastic record from China (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="205" masterPageNumber="189" pageNumber="189" updateTime="1732802310598" updateUser="ExternalLinkService">
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<mods:title id="7A7F41596BA023C88FAC52B680993ADD">Four new species of Agraphydrus Regimbart, 1903 with additional faunastic record from China (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
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<mods:namePart id="DA212603730B5E9E87A00E4909B6FCDD">Yang, Zhen-ming</mods:namePart>
<mods:nameIdentifier id="2D5A9014CAEF8D010634F434DCDBF7AB" type="ORCID">https://orcid.org/0000-0001-7838-8605</mods:nameIdentifier>
<mods:affiliation id="ED56357906D2B97401370D8AE4CB8CE2">Institute of Entomology, Life Sciences School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China</mods:affiliation>
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<mods:namePart id="5A9AAFD6C15F70CDA182F2ECF9CD4901">Jia, Fenglong</mods:namePart>
<mods:affiliation id="A01E8CE292E5F92C0E76286D85727310">Institute of Entomology, Life Sciences School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China</mods:affiliation>
<mods:nameIdentifier id="F5A579F86B92EA2768678A290472BE2E" type="email">lssjfl@mail.sysu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="16B1F7D1FA75F81B3A7DE0147C966AED">Jiang, Lu</mods:namePart>
<mods:affiliation id="75D3817798E086BBFC62F4E3D5D4105F">Shenzhen Wilde life Conservation Division, Shenzhen, China</mods:affiliation>
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<mods:namePart id="FB15233DD08C85F97284B95799CF3238">Guo, Qiang</mods:namePart>
<mods:affiliation id="CA73FBF916CBF944CB37007097247426">Shenzhen Wilde life Conservation Division, Shenzhen, China</mods:affiliation>
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<mods:title id="E3F26E751160A4ED1AF06C8FDA40FDC6">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="7E69AE6C77C4AB259D5957458D7FD0B9">2021</mods:date>
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<mods:number id="7F7A3C60188D229DC9FD515F071CDD0F">2021-05-18</mods:number>
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<mods:number id="C39C316CC87FD54E72E081EF20AB6FB3">68</mods:number>
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<mods:number id="C94C3871A0C6C147DC60DACE4262F731">1</mods:number>
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<mods:url id="895765166950CC0623C9AA48A5B64F4B">http://dx.doi.org/10.3897/dez.68.66200</mods:url>
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<mods:classification id="470FD122BE5906294E1F4774C1412F7B">journal article</mods:classification>
<mods:identifier id="484A3E806874EC8CB3C2F1DBF9F1DC8B" type="DOI">http://dx.doi.org/10.3897/dez.68.66200</mods:identifier>
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<treatment id="7207B547081750D6A276E84D147BA280" ID-GBIF-Taxon="182223088" LSID="urn:lsid:zoobank.org:act:C40E8204-E492-4147-B202-CF76A8871E2C" httpUri="http://treatment.plazi.org/id/7207B547081750D6A276E84D147BA280" lastPageNumber="189" pageId="0" pageNumber="189" scope_family="Hydrophilidae" scope_order="Coleoptera" scope_subFamily="Acidocerinae">
<subSubSection id="EFBCECD85A02E612ED53E8D91C3EF0D7" pageId="0" pageNumber="189" type="nomenclature">
<paragraph id="A263F3DFB347E4767F8B660AD5F27C5E" pageId="0" pageNumber="189">
<taxonomicName id="BF08F9E431F3A9A7C2B755362A6129EC" ID-CoL="8XT8R" LSID="http://zoobank.org/C40E8204-E492-4147-B202-CF76A8871E2C" authority="Yang &amp; Jia" authorityName="Yang &amp; Jia" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus komareki" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="komareki" status="sp. nov.">Agraphydrus komareki Yang &amp; Jia</taxonomicName>
<taxonomicNameLabel id="64DF993051BD673271450643E2B6FBB5" pageId="0" pageNumber="189">sp. nov.</taxonomicNameLabel>
<figureCitation id="2A59C6C129B725C84225F44880724098" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus: A, B. Agraphydrus dapengensis sp. nov. C, D. Agraphydrus komareki sp. nov." figureDoi="10.3897/dez.68.66200.figure1" httpUri="https://binary.pensoft.net/fig/544289" pageId="0" pageNumber="189">Figs 1C, D</figureCitation>
<figureCitation id="7A788EB0E122AB09524C854484B1ED59" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agraphydrus komareki sp. nov.: A. Clypeus; B. Antennae; C. Maxillary palpi; D. Mentum; E. Mesoventrite; F. Ventrite 5; G. Profemora; H. Mesofemora; I. Metafemora." figureDoi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189">; 4</figureCitation>
<figureCitation id="C5F32D4F02E772E7A618600E58B7365D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Aedeagi: A. Agraphydrus dapengensis sp. nov.; B. Agraphydrus komareki sp. nov.; C. Agraphydrus pseudoniger sp. nov.; D. Agraphydrus sabulosus sp. nov." figureDoi="10.3897/dez.68.66200.figure7" httpUri="https://binary.pensoft.net/fig/544295" pageId="0" pageNumber="189">; 7B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8FA4C39E7216A66041DD33CD12A7A724" pageId="0" pageNumber="189" type="type locality">
<paragraph id="781D5B569CF13F6455D2CBFE89319E92" pageId="0" pageNumber="189">Type locality.</paragraph>
<paragraph id="2E70EEF0C54B7B7B137D1E2485469FE4" pageId="0" pageNumber="189">China, Guangdong Province, Taishan County, Shangchuan Island.</paragraph>
</subSubSection>
<subSubSection id="31BFEECFB135F36EFF3AF2E270E3B848" pageId="0" pageNumber="189" type="materials_examined">
<paragraph id="6F64F7EFD1117B6774E87B8F00219A13" pageId="0" pageNumber="189">Material examined.</paragraph>
<paragraph id="B1C5D3CFAA6A244042DA7C92F825580C" pageId="0" pageNumber="189">
<materialsCitation id="2BCF29B423F8B3F163D31F00BA96726A" collectingDate="2019-01-19" collectorName="Fenglong Jia, Zu-long Liang" county="Rhesus" elevation="10" latitude="21.77" location="Guangdong Province" longLatPrecision="21" longitude="112.86083" municipality="Taishan" specimenCount="1" typeStatus="Holotype">
<emphasis id="8424647D21425402B28CB401659DDC4C" bold="true" pageId="0" pageNumber="189">
<emphasis id="A803D2E40B9FF0CC12C2CB614ACF4FA2" italics="true" pageId="0" pageNumber="189">
<typeStatus id="65650AD467E17BE2C73CE6467007E99D">Holotype</typeStatus>
</emphasis>
:
</emphasis>
male (SYSU),
<location id="21B405A01B1CA2DBBADB62DBA68C7854" LSID="urn:lsid:plazi:treatment:7207B547081750D6A276E84D147BA280:21B405A01B1CA2DBBADB62DBA68C7854" county="Rhesus" latitude="21.77" longLatPrecision="21" longitude="112.86083" municipality="Taishan" name="Guangdong Province">Guangdong Province</location>
,
<collectingMunicipality id="0B414B5E92D968683D2EB502267416CF">Taishan</collectingMunicipality>
, shangchuan island,
<collectingCounty id="3D29502580983A06CAA188F5978B9343">Rhesus</collectingCounty>
monkey provincial national nature reserve, ca
<geoCoordinate id="375E4E51895C90AD5D7E6B10827ACB75" degrees="21" direction="north" minutes="46" orientation="latitude" precision="15" seconds="12" value="21.77">21°46'12&quot;N</geoCoordinate>
,
<geoCoordinate id="4E72750749243E7730D8970109BDA4CF" degrees="112" direction="east" minutes="51" orientation="longitude" precision="15" seconds="39" value="112.86083">112°51'39&quot;E</geoCoordinate>
,
<elevation id="8228F505101BB7E93EAB98DD87BF1523" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">
<quantity id="EC5B6F56CB80EDF135A174467739AB9A" metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="ED54E4D735FA3151A582CD7C9CD7364F" value="2019-01-19">19.I.2019</collectingDate>
,
<collectorName id="A3575CF149BB7B169BFDBC7531D83194">Fenglong Jia</collectorName>
&amp;
<collectorName id="D07ADE9CE43893C85B5053F548975C23">Zu-long Liang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="403BBE988074026557A565B311D3E213" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Fenglong Jia &amp; Zu-long Liang" county="Rhesus" elevation="10" latitude="21.77" location="Guangdong Province" longLatPrecision="21" longitude="112.86083" municipality="Taishan" specimenCount="13" typeStatus="Paratypes">
<emphasis id="A369C376DE232586CCED38BCA3777A13" bold="true" italics="true" pageId="0" pageNumber="189">
<typeStatus id="DFB4DCF92695144C6C8FE7773FF17068">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="D189487E4D980EEAB47E3459603084B7" type="generic">13 exs.</specimenCount>
, IZCAS, SYSU): same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="246C470EA750142EFA2EF5FBB8E72910" pageId="0" pageNumber="189" type="diagnosis">
<paragraph id="878682655FB68B5EE89E2C6899245F6B" pageId="0" pageNumber="189">Diagnosis.</paragraph>
<paragraph id="6CBE329AA73E4CC3DA9706659EA7F0A9" pageId="0" pageNumber="189">
This species is distinguished from the other species of
<taxonomicName id="E5480AA4614EA47C2871076F29B40E5A" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="genus">
<emphasis id="FDF1CDEC46CC689E9BE9889D23C73FFC" italics="true" pageId="0" pageNumber="189">Agraphydrus</emphasis>
</taxonomicName>
by the following combination of characters: Labrum, clypeus and frons black; preocular patches present; clypeus entirely microsculptured; maxillary palpi with apical palpomere about 1.5-1.6
<normalizedToken id="D22CA0F33317310AD74C8659C9DFB1F6" originalValue="×">x</normalizedToken>
as long as penultimate, palpomere 4 without apical infuscation; antennae with eight antennomeres; elytra with four systematic punctures; meso- and metafemora pubescent in about basal 3/4; aedeagus with median lobe wider than parameres, corona large, located at basal third; parameres with indistinct subapical constriction. Differs from
<taxonomicName id="A52E88876522AE534B0889F9ED5C90C5" lsidName="A. gracilipalpis" pageId="0" pageNumber="189" rank="species" species="gracilipalpis">
<emphasis id="33643778AFA837DBBAADBC282A57E866" italics="true" pageId="0" pageNumber="189">A. gracilipalpis</emphasis>
</taxonomicName>
by eight antennomeres and features of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="663D4C2AACB87A060ABE8BE5AD06050A" pageId="0" pageNumber="189" type="description">
<paragraph id="7A965B31118F76D1E607041E6339A920" pageId="0" pageNumber="189">Description.</paragraph>
<paragraph id="CCC2003EDD3BD807E92360C17CB03303" pageId="0" pageNumber="189">
<emphasis id="6B8EF907416CCDAC8F62372671545F72" bold="true" pageId="0" pageNumber="189">
<emphasis id="11BDE18A28CE01C7BE8C9A4534D6ED1F" italics="true" pageId="0" pageNumber="189">Form and color</emphasis>
.
</emphasis>
Total length: 1.6-2.0mm; elytral width: 0.8-1.0mm; E.I.:1.1-1.3, P.I.:2.1-2.4, elytra 2.6-3.0
<normalizedToken id="6B892C6574B6A97B5506B84B3320BCC1" originalValue="×">x</normalizedToken>
as long as pronotum. Body oval, convex dorsally. Labrum, clypeus and frons dark reddish-brown, clypeus with reddish-brown preocular patches; maxillary palpi unicolored yellow; pronotum dark reddish-brown mesally with light red brown margin; elytral dark reddish-brown, lighter posteriorly; ventrites reddish-brown or brown; legs light brown or light reddish-brown.
</paragraph>
<paragraph id="31DD7B8148989464E6E93B8AA6EE7D7B" pageId="0" pageNumber="189">
<emphasis id="939C9DF38296B90ABB2C1114BBD3DD97" bold="true" pageId="0" pageNumber="189">
<emphasis id="F45FEF387AC73721FD28204E10385613" italics="true" pageId="0" pageNumber="189">Head</emphasis>
.
</emphasis>
Labrum entirely microsculptured, with few punctures at the anterior margin. Clypeus (Fig.
<figureCitation id="C53CAF22C5FA03EBC8752346024FAA8B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agraphydrus komareki sp. nov.: A. Clypeus; B. Antennae; C. Maxillary palpi; D. Mentum; E. Mesoventrite; F. Ventrite 5; G. Profemora; H. Mesofemora; I. Metafemora." figureDoi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189">4A</figureCitation>
) with angularly excised anterior margin, entirely microsculptured, ground punctures sparse, interspaces 3-4
<normalizedToken id="F6BB654D8274B0EC867FA30541A1DABB" originalValue="×">x</normalizedToken>
as large as punctures, systematic punctures distinct. Frons with dense ground punctures, interspaces 2-4
<normalizedToken id="7C3EDC6D30E5CB29773ECCE1F4AB79F2" originalValue="×">x</normalizedToken>
as large as punctures, systematic punctures distinct. Eyes large, not protruding. Antennae (Fig.
<figureCitation id="9096223FA0FD8A25F8947E3E57866215" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agraphydrus komareki sp. nov.: A. Clypeus; B. Antennae; C. Maxillary palpi; D. Mentum; E. Mesoventrite; F. Ventrite 5; G. Profemora; H. Mesofemora; I. Metafemora." figureDoi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189">4B</figureCitation>
) with eight antennomeres. Maxillary palpi (Fig.
<figureCitation id="CA05A24EB257F87CC4276241ED18CBAA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agraphydrus komareki sp. nov.: A. Clypeus; B. Antennae; C. Maxillary palpi; D. Mentum; E. Mesoventrite; F. Ventrite 5; G. Profemora; H. Mesofemora; I. Metafemora." figureDoi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189">4C</figureCitation>
) 1.2-1.3
<normalizedToken id="950739984265BDED3813C9FC10CA6BA2" originalValue="×">x</normalizedToken>
as long as pronotum in midline, about as long as maximum width of clypeus; length ratio palpomere 4:3 =1.5-1.6, palpomere 4 asymmetrical. Mentum (Fig.
<figureCitation id="CB43F357F2941D4E6B3E08C9EF9AF292" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agraphydrus komareki sp. nov.: A. Clypeus; B. Antennae; C. Maxillary palpi; D. Mentum; E. Mesoventrite; F. Ventrite 5; G. Profemora; H. Mesofemora; I. Metafemora." figureDoi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189">4D</figureCitation>
) with sparse coarse punctures, microsculpture existing on lateral portions.
</paragraph>
<caption id="1CC70B8D99A2C136BC883C829B8A19D6" doi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189" start="Figure 4" startId="F4">
<paragraph id="89464013799AEB7CE2EBABC69C1A6032" pageId="0" pageNumber="189">
<emphasis id="91D5792362B4EF162368E3379F78C244" bold="true" pageId="0" pageNumber="189">Figure 4.</emphasis>
<taxonomicName id="66F23D0040FFCEC0FAE5C00BEBCDD48A" authorityName="Yang &amp; Jia" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus komareki" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="species" species="komareki">
<emphasis id="9241875463C9CADFAFE200610553284B" italics="true" pageId="0" pageNumber="189">Agraphydrus komareki</emphasis>
</taxonomicName>
sp. nov.:
<emphasis id="E58B47156D326DBBD2523AB7F29E92EB" bold="true" pageId="0" pageNumber="189">A.</emphasis>
Clypeus;
<emphasis id="0030F2F8CF0DE935BB4EBF4EE3A45137" bold="true" pageId="0" pageNumber="189">B.</emphasis>
Antennae;
<emphasis id="0650331C721CD84D1B60E806F88243CB" bold="true" pageId="0" pageNumber="189">C.</emphasis>
Maxillary palpi;
<emphasis id="BAADB9A88E574F190A9BC4158CF29B13" bold="true" pageId="0" pageNumber="189">D.</emphasis>
Mentum;
<emphasis id="349681264DCB58C523004890E49AD9C9" bold="true" pageId="0" pageNumber="189">E.</emphasis>
Mesoventrite;
<emphasis id="B91A975EB2DD9A83B97C34452D4A57A5" bold="true" pageId="0" pageNumber="189">F.</emphasis>
Ventrite 5;
<emphasis id="031311DEF349EDC34B74B6629C1350C2" bold="true" pageId="0" pageNumber="189">G.</emphasis>
Profemora;
<emphasis id="8082A45B2DBBDED63E9268CAC750C887" bold="true" pageId="0" pageNumber="189">H.</emphasis>
Mesofemora;
<emphasis id="8C246C607A2DAA8F54C4BE1CDDCAC83D" bold="true" pageId="0" pageNumber="189">I.</emphasis>
Metafemora.
</paragraph>
</caption>
<paragraph id="F24BC8D877626A7FC29E3AAE6195AAA4" pageId="0" pageNumber="189">
<emphasis id="20477F84A2269CDEAA96AF169B4F3718" bold="true" pageId="0" pageNumber="189">
<emphasis id="35F90563F5341A55DBBF31FB10D34834" italics="true" pageId="0" pageNumber="189">Thorax</emphasis>
.
</emphasis>
Pronotum ca. 1.1-1.3
<normalizedToken id="7ADEC2A39D4979D442D4BD540CD9AB20" originalValue="×">x</normalizedToken>
as wide as long, pronotal ground punctures as on frons, surface between ground punctures with shallow microsculpture; systematic punctures indistinct, only several coarse punctures existing on the middle of the lateral part. Elytral ground punctation slightly larger than that on head and pronotum, interspaces 1.4-1.6
<normalizedToken id="D5D47702B8AE60A1812942E99613E4C1" originalValue="×">x</normalizedToken>
as large as punctures, surface between ground punctures as on the pronotum; 4 rows of systematic punctures distinct, with strongly reduced number of punctures, mesal row not reaching anterior margin, intervals without coarse punctures. Prosternum slightly convex, without carina medially. Mesoventrite (Fig.
<figureCitation id="D15F7A908AE6D91A170CC641E8999B00" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agraphydrus komareki sp. nov.: A. Clypeus; B. Antennae; C. Maxillary palpi; D. Mentum; E. Mesoventrite; F. Ventrite 5; G. Profemora; H. Mesofemora; I. Metafemora." figureDoi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189">4E</figureCitation>
) with distinct, flexuose and narrow mesal carina. Metaventrite distinctly convex mesally, with a small glabrous area posteromedially.
</paragraph>
<paragraph id="D42DCD323C05921DCA2F71D6494D2CFD" pageId="0" pageNumber="189">
<emphasis id="93BC826881747A4A23A208B1735540C5" bold="true" pageId="0" pageNumber="189">
<emphasis id="DBF2D7683078C70C9F7F16C38853B9EF" italics="true" pageId="0" pageNumber="189">Legs</emphasis>
.
</emphasis>
Pubescence present on about 2/3 of femora (Fig.
<figureCitation id="EDE4482903E0A28BB11222E7B2DD027E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agraphydrus komareki sp. nov.: A. Clypeus; B. Antennae; C. Maxillary palpi; D. Mentum; E. Mesoventrite; F. Ventrite 5; G. Profemora; H. Mesofemora; I. Metafemora." figureDoi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189">4G, H, I</figureCitation>
), hairline oblique on pro- and mesofemora, slightly oblique on metafemora.
</paragraph>
<paragraph id="2B4A0C4E464E8971F967FC27A6EC1512" pageId="0" pageNumber="189">
<emphasis id="7022C63C5C5FC9178F3BE1C641D09884" bold="true" pageId="0" pageNumber="189">
<emphasis id="33FB737E5F88FA6839061DFA669C32CC" italics="true" pageId="0" pageNumber="189">Abdomen</emphasis>
.
</emphasis>
Ventrite 5 (Fig.
<figureCitation id="6CD8A8E474C4E2E91E0C1A4012545A3A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agraphydrus komareki sp. nov.: A. Clypeus; B. Antennae; C. Maxillary palpi; D. Mentum; E. Mesoventrite; F. Ventrite 5; G. Profemora; H. Mesofemora; I. Metafemora." figureDoi="10.3897/dez.68.66200.figure4" httpUri="https://binary.pensoft.net/fig/544292" pageId="0" pageNumber="189">4F</figureCitation>
) densely pubescent, apical emargination semicircular, with a row of strong setae.
</paragraph>
<paragraph id="D5A4480A0C801AA45FB20BAE095DF573" pageId="0" pageNumber="189">
<emphasis id="AA972731C36332F0407614B86DF50BB2" bold="true" italics="true" pageId="0" pageNumber="189">Aedeagus</emphasis>
(Fig.
<figureCitation id="08F951747258CED52A5CD113F22C951F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Aedeagi: A. Agraphydrus dapengensis sp. nov.; B. Agraphydrus komareki sp. nov.; C. Agraphydrus pseudoniger sp. nov.; D. Agraphydrus sabulosus sp. nov." figureDoi="10.3897/dez.68.66200.figure7" httpUri="https://binary.pensoft.net/fig/544295" pageId="0" pageNumber="189">7B</figureCitation>
). Length: 0.33mm. Phallobase about 2/3
<normalizedToken id="D1D822AEFE54048DE0A06644ADB09A2F" originalValue="×">x</normalizedToken>
as long as the parameres, manubrium blunt basally. Apex of parameres truncate, almost flat, weakly inclining toward midline; lateral margin straight, with indistinct subapical constriction; inner margin slightly concave subapically; basal portion extending into about half of phallobase. Median lobe wider than paramere, apical third gradually narrowing apicad, apex bluntly rounded; corona large, locating at the basal third; basal apophyses moderately short, bending laterad; reaching third of the phallobase.
</paragraph>
</subSubSection>
<subSubSection id="40FF112F394079F1C41B7BA927C51A7B" pageId="0" pageNumber="189" type="etymology">
<paragraph id="513E1A22222C2559D28EB883FA8AC121" pageId="0" pageNumber="189">Etymology.</paragraph>
<paragraph id="77A7E9B2A25BC08BEA11DD1FB0FCC02D" pageId="0" pageNumber="189">
This species is named after Dr. Albrecht Komarek, a specialist in
<taxonomicName id="ABB29E29E8E4134311C4920329835EDE" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Hydrophilidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="189" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
, who works at Naturhistorisches Museum, Wien, Austria.
</paragraph>
</subSubSection>
<subSubSection id="72CE71BE6E7553E7B79ACE8700E464F7" pageId="0" pageNumber="189" type="distribution">
<paragraph id="531B78BA08D1D85BB6B4CDDABEB92E17" pageId="0" pageNumber="189">Distribution.</paragraph>
<paragraph id="5D2AF69B47935A381E9981DEBAE5F249" pageId="0" pageNumber="189">Only known type locality.</paragraph>
</subSubSection>
<subSubSection id="4D9B01CC5BA238F2094E2C13239D897B" pageId="0" pageNumber="189" type="remarks">
<paragraph id="372400B2EFF1DF7B08399D50D4E94987" pageId="0" pageNumber="189">Remark.</paragraph>
<paragraph id="20E2B97E13BF64D80A3B59B6720F051F" pageId="0" pageNumber="189">
This species belongs to the group of species entirely chagrinate clypeus and unicolorous maxillary palpomere 4, together with
<taxonomicName id="F827C7936B1D3139395E8CC1F033FD84" lsidName="A. igneus" pageId="0" pageNumber="189" rank="species" species="igneus">
<emphasis id="A5584E5704D6B47FC9F340E4FCD524AA" italics="true" pageId="0" pageNumber="189">A. igneus</emphasis>
</taxonomicName>
Komarek &amp; Hebauer,
<taxonomicName id="E9F1EC32EB197BB48FFA1C74CD8D188D" lsidName="A. umbrosus" pageId="0" pageNumber="189" rank="species" species="umbrosus">
<emphasis id="DE154288FBC36867DCDFF9112B4A261E" italics="true" pageId="0" pageNumber="189">A. umbrosus</emphasis>
</taxonomicName>
Komarek &amp; Hebauer,
<taxonomicName id="19693BACCD9675973309CA69573BE2E8" lsidName="A. gracilipalpis" pageId="0" pageNumber="189" rank="species" species="gracilipalpis">
<emphasis id="F62D549D227CA57812945967E23176F3" italics="true" pageId="0" pageNumber="189">A. gracilipalpis</emphasis>
</taxonomicName>
Komarek &amp; Hebauer and
<taxonomicName id="67B30F67BBC1DF049DBDD94AB48D494F" lsidName="A. schoenmanni" pageId="0" pageNumber="189" rank="species" species="schoenmanni">
<emphasis id="2572C1F9198B9D35BBA0FE3900F291FC" italics="true" pageId="0" pageNumber="189">A. schoenmanni</emphasis>
</taxonomicName>
Komarek &amp; Hebauer. It differs from
<taxonomicName id="95F86E1AE8921602590BD3B3B357CD7C" lsidName="A. igneus" pageId="0" pageNumber="189" rank="species" species="igneus">
<emphasis id="11CD789A144A80CFE21FC67476F1CBED" italics="true" pageId="0" pageNumber="189">A. igneus</emphasis>
</taxonomicName>
by eight antennomeres (nine antennomeres in
<taxonomicName id="4BB468B0DC74567CD5198DC16BB38BC1" lsidName="A. igneus" pageId="0" pageNumber="189" rank="species" species="igneus">
<emphasis id="E550A8C29A40E3E06EEA95EC790EE7B7" italics="true" pageId="0" pageNumber="189">A. igneus</emphasis>
</taxonomicName>
), pronotum without anterolateral chagrination, lateral margin of parameres straight and parameres slightly constricted subapically (lateral margin of parameres strongly sigmoid and with distinct subapical constriction in
<taxonomicName id="AE0F3437B9DF7F34E3B0B39AECC315A6" lsidName="A. igneus" pageId="0" pageNumber="189" rank="species" species="igneus">
<emphasis id="D54A73A2A1870429CFC9049FA028DFDF" italics="true" pageId="0" pageNumber="189">A. igneus</emphasis>
</taxonomicName>
), from
<taxonomicName id="082880C97B03DBC453B364F67273A182" lsidName="A. umbrosus" pageId="0" pageNumber="189" rank="species" species="umbrosus">
<emphasis id="9A563706822B76489EA77B0770A164DD" italics="true" pageId="0" pageNumber="189">A. umbrosus</emphasis>
</taxonomicName>
by eight antennomeres (nine antennomeres in
<taxonomicName id="046B8D75A86473932D8A5DABA43B0F11" lsidName="A. umbrosus" pageId="0" pageNumber="189" rank="species" species="umbrosus">
<emphasis id="5971F5FB428DB772C9C23B5ECADE906E" italics="true" pageId="0" pageNumber="189">A. umbrosus</emphasis>
</taxonomicName>
) and parameres moderately narrow, lateral margin straight with indistinct subapical constriction (Parameres moderately wide; lateral margins very slightly curved with distinct subapical constriction in
<taxonomicName id="F6E2792C087D5DD210972B355CE14114" lsidName="A. umbrosus" pageId="0" pageNumber="189" rank="species" species="umbrosus">
<emphasis id="7E055E7EBCC52431BCDED7EEC4C84DF6" italics="true" pageId="0" pageNumber="189">A. umbrosus</emphasis>
</taxonomicName>
), from
<taxonomicName id="B88650A5C8639A7331ABEF427B394BE6" lsidName="A. gracilipalpis" pageId="0" pageNumber="189" rank="species" species="gracilipalpis">
<emphasis id="0504EC201D05D41A0B7696536C51F7D6" italics="true" pageId="0" pageNumber="189">A. gracilipalpis</emphasis>
</taxonomicName>
by eight antennomeres and lateral margin of parameres with indistinct subapical constriction and median lobe with apical third gradually narrowing apicad (lateral margin of parameres with distinct supapical constriction and median lobe with margins of ventral face converging to narrow, parallel-sided apical half in
<taxonomicName id="9395594D9D67F0277A732F393BECF919" lsidName="A. gracilipalpis" pageId="0" pageNumber="189" rank="species" species="gracilipalpis">
<emphasis id="B8F4A7B638FBDA891BF03FCCF2F4A9C5" italics="true" pageId="0" pageNumber="189">A. gracilipalpis</emphasis>
</taxonomicName>
), from
<taxonomicName id="C0DD192F5FB94F5449EE30C89C352010" lsidName="A. schoenmanni" pageId="0" pageNumber="189" rank="species" species="schoenmanni">
<emphasis id="DF0203FD402F6C5853CEDE83C236C82C" italics="true" pageId="0" pageNumber="189">A. schoenmanni</emphasis>
</taxonomicName>
by eight antennomeres (nine antennomeres in
<taxonomicName id="F6159BDC9E043A1255DB636B45A55396" lsidName="A. schoenmanni" pageId="0" pageNumber="189" rank="species" species="schoenmanni">
<emphasis id="D514574C456B6A9E44DEE7D5F92C3C82" italics="true" pageId="0" pageNumber="189">A. schoenmanni</emphasis>
</taxonomicName>
), lateral margin of parameres straight with indistinct subapical constriction and median lobe with corona locating at the basal third (lateral margins of aedeagus distinctly sigmoid with strong subapical constriction and median lobe with corona located at apical third in
<taxonomicName id="852A7459AA1BB1252D2F56F782E5647C" lsidName="A. schoenmanni" pageId="0" pageNumber="189" rank="species" species="schoenmanni">
<emphasis id="20DD72342119D2655B6F539ED2F9E6A0" italics="true" pageId="0" pageNumber="189">A. schoenmanni</emphasis>
</taxonomicName>
). Very similar to
<taxonomicName id="8C96FC059E8F25F2DB85BF24BD783930" lsidName="A. reticuliceps" pageId="0" pageNumber="189" rank="species" species="reticuliceps">
<emphasis id="6FEDD3322CA691668D8220827650C321" italics="true" pageId="0" pageNumber="189">A. reticuliceps</emphasis>
</taxonomicName>
, differs in its small size, entire chagrinate clypeus, eight antennomeres (nine antennomeres in
<taxonomicName id="B8AB4E84DD5BF1802D8F0418047D7A58" lsidName="A. reticuliceps" pageId="0" pageNumber="189" rank="species" species="reticuliceps">
<emphasis id="08C3F75756E53FC28AFBC538ADFA46B9" italics="true" pageId="0" pageNumber="189">A. reticuliceps</emphasis>
</taxonomicName>
), median lobe with apical third gradually narrowing apicad and bluntly rounded apically (median lobe apex globular, with subapical constriction in
<taxonomicName id="15EC82AC877ED06F168CDFEE1C71946D" lsidName="A. reticuliceps" pageId="0" pageNumber="189" rank="species" species="reticuliceps">
<emphasis id="2CE047EF11703EBDEED40ED30A536E8F" italics="true" pageId="0" pageNumber="189">A. reticuliceps</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>