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<mods:title id="A0AE488DD396B6E74DAA7CDD063874CF">Revision of the shell-carrying crab genus Conchoecetes Stimpson, 1858 (Crustacea: Brachyura: Dromiidae)</mods:title>
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<bibRefCitation id="9E3E7B893B12C534315FFF11E104FF54" author="Stimpson, W." box="[325,516,225,251]" pageId="2" pageNumber="3" pagination="225 - 252" refId="ref25872" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. W. Stimpson Pars VII: Crustacea Anomura. Proceedings of the Academy of Natural Science, 10 (4), 225 - 252. https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
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<treatmentCitation id="7B0E20693B12C5343212FED6E172FE91" author="Stimpson, W." box="[520,626,294,318]" page="180" pageId="2" pageNumber="3" year="1907">
<bibRefCitation id="9E3E7B893B12C5343212FED6E172FE91" author="Stimpson, W." box="[520,626,294,318]" pageId="2" pageNumber="3" pagination="1 - 240" refId="ref25948" refString="Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) - collected by the North Pacific Exploring Expedition 1853 - 1856. Smithsonian Miscellaneous Collections, 49 (1717), 1 - 240. https: // doi. org / 10.5962 / bhl. title. 51448" type="journal article" year="1907">1907: 180</bibRefCitation>
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; Haswell 1882: 141;
<treatmentCitation id="7B0E20693B12C5343354FED5E72DFE91" author="Henderson, J. R." box="[846,1069,293,318]" page="407" pageId="2" pageNumber="3" year="1893">
<bibRefCitation id="9E3E7B893B12C5343354FED5E72DFE91" author="Henderson, J. R." box="[846,1069,293,318]" pageId="2" pageNumber="3" pagination="325 - 458" refId="ref23837" refString="Henderson, J. R. (1893) A contribution to Indian Carcinology. Transactions of the Linnaean Society London, Zoology, 5 (10), 325 - 458, https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal article" year="1893">Henderson 1893: 407</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C534342DFED5E7E5FE91" author="Alcock, A." box="[1079,1253,293,318]" page="41" pageId="2" pageNumber="3" year="1899">
<bibRefCitation id="9E3E7B893B12C534342DFED5E7E5FE91" author="Alcock, A." box="[1079,1253,293,318]" pageId="2" pageNumber="3" refId="ref22859" refString="Alcock, A. (1899) An account of the deep-sea Brachyura collected by the marine survey ship &quot; Investigator &quot;. Trustees of the Indian Museum, Calcutta, 85 pp. https: // doi. org / 10.5962 / bhl. title. 10330" type="book" year="1899">Alcock 1899: 41</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C53434EBFED6E64EFE91" author="Alcock, A." box="[1265,1358,294,318]" page="40" pageId="2" pageNumber="3" year="1901">
<bibRefCitation id="9E3E7B893B12C53434EBFED6E64EFE91" author="Alcock, A." box="[1265,1358,294,318]" pageId="2" pageNumber="3" refId="ref22954" refString="Alcock, A. (1901) Catalogue of the Indian Decapod Crustacea in the collection of the Indian Museum. Part I. Brachyura. Fasc. I. Introduction and Dromides or Dromiacea (Brachyura Primigenia). Trustees of the Indian Museum, Calcutta, 80 pp." type="book" year="1901">1901: 40</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C5343540FED6E266FEF3" author="Borradaile, L. B." page="301" pageId="2" pageNumber="3" year="1903">
<bibRefCitation id="9E3E7B893B12C5343540FED6E266FEF3" author="Borradaile, L. B." pageId="2" pageNumber="3" pagination="297 - 303" refId="ref23088" refString="Borradaile, L. B. (1903) On the genera of the Dromiidae. Annals and Magazine of Natural History, Series 7, 11 (63), 297 - 303. https: // doi. org / 10.1080 / 00222930308678771" type="journal article" year="1903">Borradaile 1903: 301</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C5343175FEB4E137FEF2" author="Stebbing, T. R. R." box="[367,567,324,349]" page="346" pageId="2" pageNumber="3" year="1910">
<bibRefCitation id="9E3E7B893B12C5343175FEB4E137FEF2" author="Stebbing, T. R. R." box="[367,567,324,349]" pageId="2" pageNumber="3" pagination="281 - 593" refId="ref25751" refString="Stebbing, T. R. R. (1910) General catalogue of South African Crustacea (Part V of S. A. Crustacea for the Marine Investigations in South Africa). Annals of the South African Museum, 6, 281 - 593. https: // doi. org / 10.5962 / bhl. part. 15558" type="journal article" year="1910">Stebbing 1910: 346</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C534325AFEB4E1A9FEF2" author="Stebbing, T. R. R." box="[576,681,324,349]" page="253" pageId="2" pageNumber="3" year="1920">
<bibRefCitation id="9E3E7B893B12C534325AFEB4E1A9FEF2" author="Stebbing, T. R. R." box="[576,681,324,349]" pageId="2" pageNumber="3" pagination="23" refId="ref25813" refString="Stebbing, T. R. R. (1920) South African Crustacea (Part X of S. A. Crustacea for the Marine Investigations in South Africa). Annals of the South African Museum, 17, 23 I- 272. https: // doi. org / 10.5962 / bhl. part. 22316" type="journal article" year="1920">1920: 253</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C53432A8FEB4E03AFEF2" author="Ihle, J. E. W." box="[690,826,324,349]" page="59" pageId="2" pageNumber="3" year="1913">
<bibRefCitation id="9E3E7B893B12C53432A8FEB4E03AFEF2" author="Ihle, J. E. W." box="[690,826,324,349]" pageId="2" pageNumber="3" pagination="1 - 96" refId="ref24208" refString="Ihle, J. E. W. (1913) Die Decapoda Brachyura der Siboga-Expedition I. Dromiacea. Siboga Expeditie, Monograph, 39 (b), Livr., 71, 1 - 96. https: // doi. org / 10.5962 / bhl. title. 13009" type="journal article" year="1913">Ihle 1913: 59</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C534335EFEB4E703FEF3" author="Barnard, K. H." box="[836,1027,324,349]" page="308" pageId="2" pageNumber="3" year="1950">
<bibRefCitation id="9E3E7B893B12C534335EFEB4E703FEF3" author="Barnard, K. H." box="[836,1027,324,349]" pageId="2" pageNumber="3" pagination="1 - 837" refId="ref23053" refString="Barnard, K. H. (1950) Descriptive catalogue of South African decapod Crustacea (Crabs and Shrimps). Annals of the South African Museum, 38, 1 - 837." type="journal article" year="1950">Barnard 1950: 308</bibRefCitation>
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; Serène &amp; Lohavanijaya 1974: 12;
<treatmentCitation id="7B0E20693B12C534357EFEB4E223FED4" author="Sakai, T." page="26" pageId="2" pageNumber="3" year="1976">
<bibRefCitation id="9E3E7B893B12C534357EFEB4E223FED4" author="Sakai, T." pageId="2" pageNumber="3" refId="ref25569" refString="Sakai, T. (1976) Crabs of Japan and Adjacent Seas. Kodansha Ltd, Tokyo, 3 vols, 773 pp." type="book" year="1976">Sakai 1976: 26</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="7B0E20693B12C5343134FE92E108FED4" author="Dai, A. &amp; Yang, S. - L." box="[302,520,354,379]" page="30" pageId="2" pageNumber="3" year="1991">
<bibRefCitation id="9E3E7B893B12C5343134FE92E108FED4" author="Dai, A. &amp; Yang, S. - L." box="[302,520,354,379]" pageId="2" pageNumber="3" refId="ref23292" refString="Dai, A. &amp; Yang, S. - L. (1991) Crabs of the China Seas. Springer-Verlag, Berlin, 608 pp." type="book" year="1991">Dai &amp; Yang 1991: 30</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C5343209FE92E1F1FED4" author="Lewinsohn, C." box="[531,753,354,379]" page="119" pageId="2" pageNumber="3" year="1984">
<bibRefCitation id="9E3E7B893B12C5343209FE92E1F1FED4" author="Lewinsohn, C." box="[531,753,354,379]" pageId="2" pageNumber="3" pagination="89 - 129" refId="ref24377" refString="Lewinsohn, C. (1984) Dromiidae from Madagascar and the Seychelles (Crustacea Decapoda Brachyura). Bulletin Museum nationale Histoire naturelle, Paris, Series 4, 6 (Section A), 1, 89 - 129." type="journal article" year="1984">Lewinsohn 1984: 119</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C53432E6FE93E0B4FED4" author="McLay, C. L." box="[764,948,355,379]" page="174" pageId="2" pageNumber="3" year="1993">
<bibRefCitation id="9E3E7B893B12C53432E6FE93E0B4FED4" author="McLay, C. L." box="[764,948,355,379]" pageId="2" pageNumber="3" pagination="111 - 251" refId="ref24509" refString="McLay, C. L. (1993) Crustacea Decapoda: the sponge crabs (Dromiidae) of New Caledonia and the Philippines with a review of the genera. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM. Vol. 10. Memoires du Museum national d'Histoire naturelle, Paris, 156, pp. 111 - 251." type="journal article" year="1993">McLay 1993: 174</bibRefCitation>
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;
<treatmentCitation id="7B0E20693B12C53433A4FE92E7E2FED4" author="Guinot, D. &amp; Tavares, M. S." box="[958,1250,354,379]" page="301" pageId="2" pageNumber="3" year="2000">
<bibRefCitation id="9E3E7B893B12C53433A4FE92E7E2FED4" author="Guinot, D. &amp; Tavares, M. S." box="[958,1250,354,379]" pageId="2" pageNumber="3" pagination="301 - 309" refId="ref23564" refString="Guinot, D. &amp; Tavares, M. S. (2000) Conchoeodromia alcocki Chopra, 1934: megalopa of Conchoecetes artificiosus (Fabricius, 1798) (Decapoda, Brachyura, Dromiidae). Journal of Crustacean Biology, 20 (Special Number 2), 301 - 309. https: // doi. org / 10.1163 / 1937240 X- 90000032" type="book chapter" year="2000">Guinot &amp; Tavares 2000: 301</bibRefCitation>
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; Davie 2002: 159;
<treatmentCitation id="7B0E20693B12C53430DDFE71E2AEFE35" author="Ahyong, S. T. &amp; Naruse, T. &amp; Tan, S. H. &amp; Ng, P. K. L." box="[199,430,385,410]" page="30" pageId="2" pageNumber="3" year="2009">
<bibRefCitation id="9E3E7B893B12C53430DDFE71E2AEFE35" author="Ahyong, S. T. &amp; Naruse, T. &amp; Tan, S. H. &amp; Ng, P. K. L." box="[199,430,385,410]" pageId="2" pageNumber="3" pagination="27 - 198" refId="ref22745" refString="Ahyong, S. T., Naruse, T., Tan, S. H. &amp; Ng, P. K. L. (2009) Part II. Infraorder Brachyura: Sections Dromiacea, Raninoida, Cyclodorippoida. In: Chan, T. - Y., Ng, P. K. L., Ahyong, S. T. &amp; Tan, S. H. (Eds.), Crustacean Fauna of Taiwan: Brachyuran Crabs, Volume 1. Carcinology in Taiwan and Dromiacea, Raninoida, Cyclodorippoida. National Taiwan Ocean University, Keelung, pp. 27 - 198." type="book chapter" year="2009">
Ahyong
<emphasis id="C8DBDA6A3B12C5343105FE72E24CFE35" box="[287,332,386,410]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
2009: 30
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;
<treatmentCitation id="7B0E20693B12C53431A3FE71E1EDFE35" author="Guinot, D. &amp; Wicksten, M. K." box="[441,749,385,410]" page="591" pageId="2" pageNumber="3" year="2015">
<bibRefCitation id="9E3E7B893B12C53431A3FE71E1EDFE35" author="Guinot, D. &amp; Wicksten, M. K." box="[441,749,385,410]" pageId="2" pageNumber="3" pagination="583 - 638" refId="ref23629" refString="Guinot, D. &amp; Wicksten, M. K. (2015) Chapter 71 - 11. Camouflage: carrying behaviour and other modalities of concealment in Brachyura. In: Castro, P., Davie, P., Guinot, D., Schram, F. R. &amp; von Vaupel Klein, J. C. (Eds.), Decapoda: Brachyura, Treatise on Zoology-Anatomy, Taxonomy, Biology. Vol. 9 C-I. Brill, Leiden and Boston, pp. 583 - 638. https: // doi. org / 10.1163 / 9789004190832 _ 013" type="book chapter" year="2015">Guinot &amp; Wicksten 2015: 591</bibRefCitation>
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.
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<treatmentCitation id="7B0E20693B12C534317AFE54E131FE11" author="Chopra, B." box="[352,561,420,446]" page="477" pageId="2" pageNumber="3" year="1934">
<bibRefCitation id="9E3E7B893B12C534317AFE54E131FE11" author="Chopra, B." box="[352,561,420,446]" pageId="2" pageNumber="3" pagination="477 - 481" refId="ref23241" refString="Chopra, B. (1934) Further notes on Crustacea Decapoda in the Indian Museum. VI. On a New dromiid and a rare oxystomous crab from the Sandheads off the mouth of the Hooghly River. Records of the Indian Museum, 36, 477 - 481." type="journal article" year="1934">Chopra, 1934: 477</bibRefCitation>
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<taxonomicName id="3DAF7DFB3B12C53432CAFE54E78DFE11" authority="Chopra, 1934" authorityName="Chopra" authorityYear="1934" box="[720,1165,420,446]" genus="Conchoeodromia" kingdom="Animalia" pageId="2" pageNumber="3" rank="species" species="alcocki">
<emphasis id="C8DBDA6A3B12C53432CAFE54E0EAFE11" box="[720,1002,420,446]" italics="true" pageId="2" pageNumber="3">Conchoeodromia alcocki</emphasis>
<bibRefCitation id="9E3E7B893B12C53433EAFE54E78DFE11" author="Chopra, B." box="[1008,1165,420,446]" pageId="2" pageNumber="3" pagination="477 - 481" refId="ref23241" refString="Chopra, B. (1934) Further notes on Crustacea Decapoda in the Indian Museum. VI. On a New dromiid and a rare oxystomous crab from the Sandheads off the mouth of the Hooghly River. Records of the Indian Museum, 36, 477 - 481." type="journal article" year="1934">Chopra, 1934</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B12C534308DFE18E050FDAE" box="[151,848,488,513]" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="FA1006783B12C534308DFE18E050FDAE" blockId="2.[151,848,488,513]" box="[151,848,488,513]" pageId="2" pageNumber="3">
<emphasis id="C8DBDA6A3B12C534308DFE18E223FDAE" bold="true" box="[151,291,488,513]" pageId="2" pageNumber="3">
<typeStatus id="2514B8DA3B12C534308DFE18E3CCFDAF" box="[151,204,488,512]" pageId="2" pageNumber="3">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="3DAF7DFB3B12C5343130FE19E18CFDAF" ID-CoL="37PML" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" box="[298,652,488,513]" class="Malacostraca" family="Dromiidae" genus="Dromia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="artificiosa">
<emphasis id="C8DBDA6A3B12C5343130FE19E2E6FDAF" box="[298,486,488,512]" italics="true" pageId="2" pageNumber="3">Dromia artificiosa</emphasis>
<bibRefCitation id="9E3E7B893B12C53431F6FE18E18CFDAF" author="Fabricius, J. C." box="[492,652,488,513]" pageId="2" pageNumber="3" refId="ref23486" refString="Fabricius, J. C. (1798) Supplementum Entomologiae Systematicae. Proft et Storch, Hafniae, 572 pp." type="book" year="1798">Fabricius, 1798</bibRefCitation>
</taxonomicName>
, gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B12C534308DFDDEE18FFC56" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="FA1006783B12C534308DFDDEE18FFC56" blockId="2.[151,1437,558,1413]" pageId="2" pageNumber="3">
<emphasis id="C8DBDA6A3B12C534308DFDDEE212FDE6" bold="true" box="[151,274,558,585]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Carapace as wide as or slightly wider than long; mostly flattened, subpentagonal, sometimes posterior cardiac and branchial areas poorly calcified; cervical and branchial grooves usually distinct; surface granular to almost smooth, with fine tomentum; front narrow, tridentate, lateral teeth well developed, median rostral tooth lower than lateral teeth; anterolateral margin variable, may be unarmed not marked by a distinct edge or with an edge that may bear small teeth and interrupted by one or more notches; cervical and branchial grooves usually marked by distinct teeth on margin. Chelipeds granular; with epipod. P2 and P3 dactyli fringed with setae and many minute adpressed spines on ventral margin; propodi without distal spine. P4 larger than P5; dactylus enlarged, talon-like, occluding with stout proximal propodal extension; used for carrying bivalve shell. P5 small; segments flattened, dactylus small simple, upturned; propodus without distal spine occluding with dactylus. Pleon of 6 free somites and telson. Uropod plates well-developed, visible externally; forming pleon holding mechanism by locking against strong prominence on P2 coxae. Male gonopods consisting of three articles: G1 stout, semi-rolled, setose tube; G2 as long as G1 needle-like. Female 7/8 sternal sutures normally end apart on erect tube-like structures between or near chelipeds (modified after
<bibRefCitation id="9E3E7B893B12C53431F5FC2FE183FC56" author="McLay, C. L." box="[495,643,991,1017]" pageId="2" pageNumber="3" pagination="111 - 251" refId="ref24509" refString="McLay, C. L. (1993) Crustacea Decapoda: the sponge crabs (Dromiidae) of New Caledonia and the Philippines with a review of the genera. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM. Vol. 10. Memoires du Museum national d'Histoire naturelle, Paris, 156, pp. 111 - 251." type="journal article" year="1993">McLay 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B555F33B12C53430DDFBF3E3F0FA2A" pageId="2" pageNumber="3" type="discussion">
<paragraph id="FA1006783B12C53430DDFBF3E3F0FA2A" blockId="2.[151,1437,558,1413]" pageId="2" pageNumber="3">
<emphasis id="C8DBDA6A3B12C53430DDFBF3E23BFBB2" bold="true" box="[199,315,1027,1053]" pageId="2" pageNumber="3">Remarks.</emphasis>
The species of
<taxonomicName id="3DAF7DFB3B12C5343219FBF3E19FFBB2" authorityName="Stimpson" authorityYear="1858" box="[515,671,1027,1053]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="C8DBDA6A3B12C5343219FBF3E19FFBB2" box="[515,671,1027,1053]" italics="true" pageId="2" pageNumber="3">Conchoecetes</emphasis>
</taxonomicName>
are most easily distinguished by carapace shape and characters associated with variation in the anterolateral margin. The degree of curvature of the carapace surface has an effect on the definition of the cervical and branchial grooves and shape of the anterolateral margin. We can contrast
<taxonomicName id="3DAF7DFB3B12C5343599FBBBE21BFB26" authorityName="Lewinsohn" authorityYear="1984" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="C8DBDA6A3B12C5343599FBBBE21BFB26" italics="true" pageId="2" pageNumber="3">C. intermedius</emphasis>
</taxonomicName>
which has a gently curved carapace surface, on which the cervical and branchial grooves are scarcely marked, which merges gradually into the hepatic area without a distinct margin or teeth, with
<taxonomicName id="3DAF7DFB3B12C53434A8FB63E651FB02" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1202,1361,1171,1197]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="artificiosus">
<emphasis id="C8DBDA6A3B12C53434A8FB63E651FB02" box="[1202,1361,1171,1197]" italics="true" pageId="2" pageNumber="3">C. artificiosus</emphasis>
</taxonomicName>
which has a flat carapace surface deeply marked by the grooves and pronounced structured anterolateral margin with teeth and notches. Here we find the
<typeStatus id="2514B8DA3B12C534322DFB2BE167FB5A" box="[567,615,1243,1269]" pageId="2" pageNumber="3">type</typeStatus>
specimen invaluable as a reference point with which all the former “
<taxonomicName id="3DAF7DFB3B12C5343599FB2BE215FAB6" baseAuthorityName="Fabricius" baseAuthorityYear="1798" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="artificiosus">
<emphasis id="C8DBDA6A3B12C5343599FB2BE215FAB6" italics="true" pageId="2" pageNumber="3">C. artificiosus</emphasis>
</taxonomicName>
” material can be compared to demonstrate differences which warrant recognition of several new species. The result is that
<typeStatus id="2514B8DA3B12C5343178FAD3E292FA92" box="[354,402,1315,1341]" pageId="2" pageNumber="3">type</typeStatus>
species
<taxonomicName id="3DAF7DFB3B12C53431ECFAD3E018FA92" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[502,792,1315,1341]" class="Malacostraca" family="Dromiidae" genus="Conchoecetes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="artificiosus">
<emphasis id="C8DBDA6A3B12C53431ECFAD3E018FA92" box="[502,792,1315,1341]" italics="true" pageId="2" pageNumber="3">Conchoecetes artificiosus</emphasis>
</taxonomicName>
appears to be restricted to the Western Indian Ocean in the vicinity of the Indian sub-continent. We re-describe and provide photographs of the
<typeStatus id="2514B8DA3B12C5343493FAB7E7B9FACE" box="[1161,1209,1351,1377]" pageId="2" pageNumber="3">type</typeStatus>
specimens for each species.
</paragraph>
</subSubSection>
</treatment>
</document>