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<mods:title id="E618BCA8EFB9FA215ACBEB0D9E5F1433">Revision of Hypoxys Amyot &amp; Serville, 1843 stat. rest. (Heteroptera Pentatomidae)</mods:title>
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<mods:namePart id="42528C47EE6A8361A44A9EE1D93946C7">Nunes, Benedito Mendes</mods:namePart>
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<mods:namePart id="780121E88195FC6D050DD2ABCCF3E854">Mendonca, Maria Thayane Da Silva</mods:namePart>
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<mods:namePart id="013B0B7CB112E7313C6A98105C7545F2">Cunha, Eduardo Victor De Paiva</mods:namePart>
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<heading id="A15B81D63172D91044B269C3FD401EF9" bold="true" box="[151,732,1196,1223]" fontSize="11" level="1" pageId="15" pageNumber="416" reason="1">
<emphasis id="C8D8EAA83172D91044B269C3FD401EF9" bold="true" box="[151,732,1196,1223]" pageId="15" pageNumber="416">
<taxonomicName id="3DAC4D393172D91044B269C3FDD21EF8" authority="(Breddin, 1907)" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[151,590,1196,1223]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="offuscatus" status="comb. nov.">
<emphasis id="C8D8EAA83172D91044B269C3FE181EF8" bold="true" box="[151,388,1196,1223]" italics="true" pageId="15" pageNumber="416">Hypoxys offuscatus</emphasis>
(
<bibRefCitation id="9E3D4B4B3172D91045B669C2FDDA1EF8" author="Breddin, G." box="[403,582,1196,1222]" pageId="15" pageNumber="416" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="D3EB57D33172D910467069C3FD401EF9" box="[597,732,1197,1223]" pageId="15" pageNumber="416" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="B2B665313172D91044B269BEFE231ED5" box="[151,447,1232,1259]" pageId="15" pageNumber="416" type="description">
<paragraph id="FA1336BA3172D91044B269BEFE231ED5" blockId="15.[151,732,1196,1259]" box="[151,447,1232,1259]" pageId="15" pageNumber="416">
(
<figureCitation id="62972A3F3172D91044BA69BEFF711ED5" box="[159,237,1232,1259]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="15" pageNumber="416">Figs. 7</figureCitation>
AE; 19 AB; 21)
</paragraph>
</subSubSection>
<subSubSection id="B2B665313172D91044B2687BFB361F54" pageId="15" pageNumber="416" type="reference_group">
<paragraph id="FA1336BA3172D91044B2687BFCDA1F13" blockId="15.[151,1426,1300,1417]" box="[151,838,1300,1325]" pageId="15" pageNumber="416">
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<treatmentCitation id="7B0D10AB3172D91044B2687BFD8C1F13" author="Breddin, G." box="[151,528,1300,1325]" page="332" pageId="15" pageNumber="416" year="1907">
<taxonomicName id="3DAC4D393172D91044B2687BFD8C1F13" authority="Breddin, 1907: 332" authorityName="Breddin" authorityPageNumber="332" authorityYear="1907" box="[151,528,1300,1325]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="offuscata">
<emphasis id="C8D8EAA83172D91044B2687BFEDE1F13" box="[151,322,1301,1325]" italics="true" pageId="15" pageNumber="416">Edessa offuscata</emphasis>
<bibRefCitation id="9E3D4B4B3172D910456C687AFD8C1F13" author="Breddin, G." box="[329,528,1300,1325]" pageId="15" pageNumber="416" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907: 332</bibRefCitation>
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;
<treatmentCitation id="7B0D10AB3172D910463F687AFD791F13" author="Kirkaldy, G. W." box="[538,741,1300,1325]" page="368" pageId="15" pageNumber="416" year="1909">
<bibRefCitation id="9E3D4B4B3172D910463F687AFD791F13" author="Kirkaldy, G. W." box="[538,741,1300,1325]" pageId="15" pageNumber="416" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 368</bibRefCitation>
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(catalog)
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</paragraph>
<paragraph id="FA1336BA3172D91044B2685DFC0C1F72" blockId="15.[151,1426,1300,1417]" box="[151,912,1331,1356]" pageId="15" pageNumber="416">
<treatmentCitationGroup id="DABC11943172D91044B2685DFC0C1F72" box="[151,912,1331,1356]" pageId="15" pageNumber="416">
<treatmentCitation id="7B0D10AB3172D91044B2685DFE6B1F75" author="Breddin, G." box="[151,503,1331,1356]" page="338" pageId="15" pageNumber="416" year="1907">
<taxonomicName id="3DAC4D393172D91044B2685DFE6B1F75" authority="Breddin, 1907: 338" authorityName="Breddin" authorityPageNumber="338" authorityYear="1907" box="[151,503,1331,1356]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="rimata">
<emphasis id="C8D8EAA83172D91044B2685DFEB51F75" box="[151,297,1331,1355]" italics="true" pageId="15" pageNumber="416">Edessa rimata</emphasis>
<bibRefCitation id="9E3D4B4B3172D910450A685DFE6B1F75" author="Breddin, G." box="[303,503,1331,1356]" pageId="15" pageNumber="416" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907: 338</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="7B0D10AB3172D9104624685DFD501F72" author="Kirkaldy, G. W." box="[513,716,1331,1356]" page="368" pageId="15" pageNumber="416" year="1909">
<bibRefCitation id="9E3D4B4B3172D9104624685DFD501F72" author="Kirkaldy, G. W." box="[513,716,1331,1356]" pageId="15" pageNumber="416" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 368</bibRefCitation>
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(catalog)
<emphasis id="C8D8EAA83172D9104716685AFC0C1F72" bold="true" box="[819,912,1332,1356]" pageId="15" pageNumber="416">syn. nov.</emphasis>
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<typeStatus id="251788183172D91044B2683DFF601F55" box="[151,252,1363,1387]" pageId="15" pageNumber="416" type="lectotype">Lectotype</typeStatus>
male.
<emphasis id="C8D8EAA83172D9104564683CFE6F1F54" box="[321,499,1362,1386]" italics="true" pageId="15" pageNumber="416">
<taxonomicName id="3DAC4D393172D9104564683CFE731F54" authorityName="Breddin" authorityYear="1907" box="[321,495,1362,1386]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="offuscata">Edessa offuscata</taxonomicName>
.
</emphasis>
<collectingCountry id="82BB762A3172D91045DD683FFDAB1F54" box="[504,567,1361,1386]" name="Brazil" pageId="15" pageNumber="416">Brasil</collectingCountry>
,
<location id="FF7360613172D9104667683FFDE11F54" LSID="urn:lsid:plazi:treatment:720587AC3172D90E44B269C3FDE71B21:FF7360613172D9104667683FFDE11F54" box="[578,637,1361,1386]" country="Brazil" name="Pebas" pageId="15" pageNumber="416">Pebas</location>
(
<collectionCode id="9CBDAE7F3172D91046AF683CFD541F54" box="[650,712,1362,1386]" pageId="15" pageNumber="416">SDEI</collectionCode>
). Designated by
<bibRefCitation id="9E3D4B4B3172D9104754683FFBAE1F54" author="Gaedike, H." box="[881,1074,1361,1386]" pageId="15" pageNumber="416" pagination="79 - 159" refId="ref24984" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen- V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike (1971:94)</bibRefCitation>
. Examined.
</materialsCitation>
</paragraph>
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<subSubSection id="B2B665313172D91044B2681FFA0E1FB7" box="[151,1426,1392,1417]" pageId="15" pageNumber="416" type="description">
<paragraph id="FA1336BA3172D91044B2681FFA0E1FB7" blockId="15.[151,1426,1300,1417]" box="[151,1426,1392,1417]" pageId="15" pageNumber="416">
<materialsCitation id="4AC43CE73172D91044B2681FFA0E1FB7" ID-GBIF-Occurrence="2570056012" box="[151,1426,1392,1417]" collectionCode="SDEI" country="Peru" location="Iguapo" pageId="15" pageNumber="416" specimenCount="2" typeStatus="lectotype">
<typeStatus id="251788183172D91044B2681FFF601FB7" box="[151,252,1393,1417]" pageId="15" pageNumber="416" type="lectotype">Lectotype</typeStatus>
female.
<emphasis id="C8D8EAA83172D9104571681FFE711FB6" box="[340,493,1393,1417]" italics="true" pageId="15" pageNumber="416">
<taxonomicName id="3DAC4D393172D9104571681FFE751FB6" authorityName="Breddin" authorityYear="1907" box="[340,489,1393,1417]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="rimata">Edessa rimata</taxonomicName>
.
</emphasis>
<collectingCountry id="82BB762A3172D91045D6681FFDB81FB6" box="[499,548,1393,1416]" name="Peru" pageId="15" pageNumber="416">Peru</collectingCountry>
,
<location id="FF7360613172D910460B681FFDE81FB7" LSID="urn:lsid:plazi:treatment:720587AC3172D90E44B269C3FDE71B21:FF7360613172D910460B681FFDE81FB7" box="[558,628,1393,1417]" country="Peru" name="Iguapo" pageId="15" pageNumber="416">Iguapo</location>
(
<collectionCode id="9CBDAE7F3172D91046A7681FFD5C1FB7" box="[642,704,1393,1417]" pageId="15" pageNumber="416">SDEI</collectionCode>
). Designated by
<bibRefCitation id="9E3D4B4B3172D910474D681EFBBD1FB7" author="Gaedike, H." box="[872,1057,1392,1417]" pageId="15" pageNumber="416" pagination="79 - 159" refId="ref24984" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen- V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike (1971:97</bibRefCitation>
, by use of “
<typeStatus id="251788183172D91040BD681EFA951FB7" box="[1176,1289,1392,1417]" pageId="15" pageNumber="416" type="holotype">Holotypus</typeStatus>
”). Examined.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B2B665313172D91044B268D8FBE01C27" pageId="15" pageNumber="416" type="materials_examined">
<paragraph id="FA1336BA3172D91044B268D8FBE01C27" blockId="15.[151,1437,1462,2030]" pageId="15" pageNumber="416">
<emphasis id="C8D8EAA83172D91044B268D8FDA21FEF" bold="true" box="[151,574,1462,1489]" pageId="15" pageNumber="416">
Material examined.
<collectingCountry id="82BB762A3172D91045AC68D9FE4B1FEF" box="[393,471,1463,1489]" name="Peru" pageId="15" pageNumber="416">PERU</collectingCountry>
.
<collectingRegion id="3868F8583172D91045C168D9FDA51FEF" box="[484,569,1463,1489]" country="Peru" name="Loreto" pageId="15" pageNumber="416">Loreto</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD333172D910466368D6FDC71FEC" box="[582,603,1464,1490]" pageId="15" pageNumber="416" type="female"></specimenCount>
Iquitos, Amazon River,
<date id="8E12107A3172D910475468D9FC6A1FEF" box="[881,1014,1463,1489]" pageId="15" pageNumber="416" value="1927-02-21">II211927</date>
, H. Bassler (
<collectionCode id="9CBDAE7F3172D91040AD68D9FB431FEF" box="[1160,1247,1463,1489]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="416" type="Museum">USNM</collectionCode>
);
<specimenCount id="ECAAFD333172D91040D668D6FA941FEC" box="[1267,1288,1464,1490]" pageId="15" pageNumber="416" type="male"></specimenCount>
<specimenCount id="ECAAFD333172D910413568D6FAB91FEC" box="[1296,1317,1464,1490]" pageId="15" pageNumber="416" type="female"></specimenCount>
Confl. De los Rios Zamur e Yahuasyacu, II80, J. Becker leg.
<taxonomicName id="3DAC4D393172D91046FD68B5FBED1FCB" authority="Breddin, 1907 Comp." authorityName="Breddin" authorityYear="1907" box="[728,1137,1499,1525]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="rimata">
<emphasis id="C8D8EAA83172D91046FD68B5FCE51FCB" box="[728,889,1499,1525]" italics="true" pageId="15" pageNumber="416">Edessa rimata</emphasis>
<bibRefCitation id="9E3D4B4B3172D910475A68B5FB831FCB" author="Breddin, G." box="[895,1055,1499,1525]" pageId="15" pageNumber="416" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907</bibRefCitation>
Comp.
</taxonomicName>
w.
<typeStatus id="251788183172D91040B268B2FB411FC8" box="[1175,1245,1500,1526]" pageId="15" pageNumber="416">TYPE</typeStatus>
Fernandes, JAM 1999,
<taxonomicName id="3DAC4D393172D91044F86891FD081C27" authority="Breddin, 1907 Comp." authorityName="Breddin" authorityYear="1907" box="[221,660,1535,1561]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="offuscata">
<emphasis id="C8D8EAA83172D91044F86891FE071C27" box="[221,411,1535,1561]" italics="true" pageId="15" pageNumber="416">Edessa offuscata</emphasis>
<bibRefCitation id="9E3D4B4B3172D91045846891FDDD1C27" author="Breddin, G." box="[417,577,1535,1561]" pageId="15" pageNumber="416" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin, 1907</bibRefCitation>
Comp.
</taxonomicName>
w.
<typeStatus id="251788183172D910469E6B6EFC9D1C24" box="[699,769,1536,1562]" pageId="15" pageNumber="416">TYPE</typeStatus>
Fernandes, JAM 1999. (
<collectionCode id="9CBDAE7F3172D91040336891FBED1C27" box="[1046,1137,1535,1561]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="15" pageNumber="416" type="Museum">RMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B665313172D91044E26B4DFB561CBB" pageId="15" pageNumber="416" type="description">
<paragraph id="FA1336BA3172D91044E26B4DFB561CBB" blockId="15.[151,1437,1462,2030]" pageId="15" pageNumber="416">
<emphasis id="C8D8EAA83172D91044E26B4DFEE21C03" bold="true" box="[199,382,1571,1597]" pageId="15" pageNumber="416">Measurements.</emphasis>
total length: 17.219.0; head length: 2.3; head width: 3.43.6; pronotal width: 11.011.3; abdominal width: 9.610.3; length of antennal segments. I: 0.91.0; II: 2.4; III: 1.31.4; IV: 5.7; V: 6.3; interocular distance: 1.61.7; scutellum length: 8.49.0; scutellum width: 6.26.6; pronotal length: 3.63.9.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313172D91044E26BE0FC9F1DD3" pageId="15" pageNumber="416" type="diagnosis">
<paragraph id="FA1336BA3172D91044E26BE0FC9F1DD3" blockId="15.[151,1437,1462,2030]" pageId="15" pageNumber="416">
<emphasis id="C8D8EAA83172D91044E26BE0FEDE1C97" bold="true" box="[199,322,1678,1705]" pageId="15" pageNumber="416">Diagnosis.</emphasis>
Anterolateral margin of pronotum with solid black stripe reaching all its extension. Cicatrices with dark brown punctures delimiting anterior sulci (
<figureCitation id="62972A3F3172D910468D6BDDFC931CF3" box="[680,783,1715,1741]" captionStart="FIGURE 19" captionStartId="35.[151,250,1983,2008]" captionTargetBox="[250,1354,187,1936]" captionTargetId="figure@35.[190,1398,181,1957]" captionTargetPageId="35" captionText="FIGURE 19. Dorsal and ventral view. H. offuscatus AB; H. trabeculus CD; H. triangularis EF; H. capito GH; H. subrastratus IJ; H. oxyacanthus KL (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677907" httpUri="https://zenodo.org/record/3677907/files/figure.png" pageId="15" pageNumber="416">Fig. 19A</figureCitation>
). Anterolateral margin with few dark punctures. Male (
<figureCitation id="62972A3F3172D910414A6BDDFF391CCF" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="15" pageNumber="416">Fig. 7</figureCitation>
AC). Pygophore with dorsal rim widely excavated (
<figureCitation id="62972A3F3172D91046D26BB9FCCD1CCF" box="[759,849,1751,1777]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="15" pageNumber="416">Fig. 7A</figureCitation>
). Superior process of genital cup small, arrow-headshaped, adjacent to dorsal rim (
<figureCitation id="62972A3F3172D91045D06B95FDD21D28" box="[501,590,1787,1814]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="15" pageNumber="416">Fig. 7B</figureCitation>
). Paramere elongated, shank triangular (cross section) due to a conspicuous carina on inner side ending before enlarged part; head of the paramere spatulate (
<figureCitation id="62972A3F3172D91040386A71FBE91D04" box="[1053,1141,1823,1850]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="15" pageNumber="416">Fig. 7B</figureCitation>
). Proctiger with laterodorsal tufts of setae widely separated by medial ridge; posterior face pentagonal, flat (
<figureCitation id="62972A3F3172D910400C6A2DFB1D1D60" box="[1065,1153,1859,1886]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="15" pageNumber="416">Fig. 7B</figureCitation>
). Ventral rim with expansions rounded, wide, tumid, well-projected, clearly surpassing level of posterolateral angle in ventral view (
<figureCitation id="62972A3F3172D910414A6A09FF271D9B" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="15" pageNumber="416">Fig. 7C</figureCitation>
); base of each expansion with a conspicuous transverse carina. Female (
<figureCitation id="62972A3F3172D91047C26AE5FBDE1D98" box="[999,1090,1931,1958]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="15" pageNumber="416">Fig. 7D</figureCitation>
). Gonocoxites 8 with posterior margin sinuous, inner angle rounded projected posteriorly reaching gonocoxites 9; sutural border with elliptic, short excavation on posterior half. Gonapophyses 8 exposed.
</paragraph>
</subSubSection>
<caption id="AED36632316DD90F44B26BBCFDEF1D31" ID-DOI="http://doi.org/10.5281/zenodo.3677883" ID-Zenodo-Dep="3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="16" pageNumber="417" startId="16.[151,250,1746,1771]" targetBox="[169,1410,207,1693]" targetPageId="16">
<paragraph id="FA1336BA316DD90F44B26BBCFDEF1D31" blockId="16.[151,1436,1746,1807]" pageId="16" pageNumber="417">
<emphasis id="C8D8EAA8316DD90F44B26BBCFE881CD5" bold="true" box="[151,276,1746,1771]" pageId="16" pageNumber="417">FIGURE 7.</emphasis>
<taxonomicName id="3DAC4D39316DD90F45396BBAFE3F1CD5" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[284,419,1747,1771]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="offuscatus">
<emphasis id="C8D8EAA8316DD90F45396BBAFE3F1CD5" box="[284,419,1747,1771]" italics="true" pageId="16" pageNumber="417">H. offuscatus</emphasis>
</taxonomicName>
. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm).
</paragraph>
</caption>
<subSubSection id="B2B66531316DD90E44E26A52FAD11AC5" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417" type="discussion">
<paragraph id="FA1336BA316DD90E44E26A52FAD11AC5" blockId="16.[151,1437,1852,2024]" lastBlockId="17.[151,1436,152,287]" lastPageId="17" lastPageNumber="418" pageId="16" pageNumber="417">
<emphasis id="C8D8EAA8316DD90F44E26A52FED31D69" bold="true" box="[199,335,1852,1879]" pageId="16" pageNumber="417">Comments.</emphasis>
<taxonomicName id="3DAC4D39316DD90F45736A53FD8E1D69" authorityName="Breddin" authorityYear="1907" box="[342,530,1853,1879]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="offuscata">
<emphasis id="C8D8EAA8316DD90F45736A53FD8E1D69" box="[342,530,1853,1879]" italics="true" pageId="16" pageNumber="417">Edessa offuscata</emphasis>
</taxonomicName>
and
<taxonomicName id="3DAC4D39316DD90F46626A50FD2D1D69" authorityName="Breddin" authorityYear="1907" box="[583,689,1854,1880]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="rimata">
<emphasis id="C8D8EAA8316DD90F46626A50FDC41D69" box="[583,600,1854,1879]" italics="true" pageId="16" pageNumber="417">E</emphasis>
.
<emphasis id="C8D8EAA8316DD90F46436A50FD2D1D69" box="[614,689,1854,1879]" italics="true" pageId="16" pageNumber="417">rimata</emphasis>
</taxonomicName>
were described by
<bibRefCitation id="9E3D4B4B316DD90F47AD6A53FBAA1D69" author="Breddin, G." box="[904,1078,1853,1879]" pageId="16" pageNumber="417" pagination="327 - 348" refId="ref24495" refString="Breddin, G. (1907) Beitrage zur Systematik der Pentatomiden Sudamerikas. Sitzungs-Berichte der Gesellschaft naturforschender freunde zu Berlin, 10, 327 - 348." type="journal article" year="1907">Breddin (1907)</bibRefCitation>
. The author considered
<taxonomicName id="3DAC4D39316DD90F411A6A50FF311D45" authorityName="Breddin" authorityYear="1907" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="rimata">
<emphasis id="C8D8EAA8316DD90F411A6A50FACC1D69" box="[1343,1360,1854,1879]" italics="true" pageId="16" pageNumber="417">E</emphasis>
.
<emphasis id="C8D8EAA8316DD90F417B6A50FF311D45" italics="true" pageId="16" pageNumber="417">rimata</emphasis>
</taxonomicName>
related to
<taxonomicName id="3DAC4D39316DD90F45026A0CFE2E1D45" authorityName="Breddin" authorityYear="1907" box="[295,434,1889,1916]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="trabecula">
<emphasis id="C8D8EAA8316DD90F45026A0CFEA41D45" box="[295,312,1890,1915]" italics="true" pageId="16" pageNumber="417">E</emphasis>
.
<emphasis id="C8D8EAA8316DD90F45626A0FFE2E1D45" box="[327,434,1889,1915]" italics="true" pageId="16" pageNumber="417">trabecula</emphasis>
</taxonomicName>
and
<taxonomicName id="3DAC4D39316DD90F45CE6A0CFDEE1D45" authorityName="Breddin" authorityYear="1907" box="[491,626,1889,1916]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="offuscata">
<emphasis id="C8D8EAA8316DD90F45CE6A0CFE601D45" box="[491,508,1890,1915]" italics="true" pageId="16" pageNumber="417">E</emphasis>
.
<emphasis id="C8D8EAA8316DD90F462E6A0FFDEE1D45" box="[523,626,1889,1915]" italics="true" pageId="16" pageNumber="417">offuscata</emphasis>
</taxonomicName>
close to
<taxonomicName id="3DAC4D39316DD90F46FC6A0CFCEB1D45" authorityName=", Amyot &amp; Serville" authorityYear="1843" box="[729,887,1889,1916]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="C8D8EAA8316DD90F46FC6A0CFD761D45" box="[729,746,1890,1915]" italics="true" pageId="16" pageNumber="417">E</emphasis>
.
<emphasis id="C8D8EAA8316DD90F46DC6A0FFCEB1D45" box="[761,887,1889,1915]" italics="true" pageId="16" pageNumber="417">quadridens</emphasis>
</taxonomicName>
and
<taxonomicName id="3DAC4D39316DD90F47956A0CFBAC1D45" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[944,1072,1889,1916]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="boerneri">
<emphasis id="C8D8EAA8316DD90F47956A0CFC5D1D45" box="[944,961,1890,1915]" italics="true" pageId="16" pageNumber="417">E</emphasis>
.
<emphasis id="C8D8EAA8316DD90F47F56A0FFBAC1D45" box="[976,1072,1889,1915]" italics="true" pageId="16" pageNumber="417">boerneri</emphasis>
</taxonomicName>
. However, these similarities are based only on the external morphology of the species. Females of
<taxonomicName id="3DAC4D39316DD90F47596AE8FB8C1DA1" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[892,1040,1925,1951]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="offuscatus">
<emphasis id="C8D8EAA8316DD90F47596AE8FB8C1DA1" box="[892,1040,1925,1951]" italics="true" pageId="16" pageNumber="417">H. offuscatus</emphasis>
</taxonomicName>
have inner angles of gonocoxites 8 more protruding and sutural border less excavated (
<figureCitation id="62972A3F316DD90F46F16AC7FCB71DFA" box="[724,811,1961,1988]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="16" pageNumber="417">Fig. 7D</figureCitation>
) than
<taxonomicName id="3DAC4D39316DD90F47556AC4FC6D1DFD" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[880,1009,1961,1987]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="boerneri">
<emphasis id="C8D8EAA8316DD90F47556AC4FC6D1DFD" box="[880,1009,1961,1987]" italics="true" pageId="16" pageNumber="417">H. boerneri</emphasis>
</taxonomicName>
(
<figureCitation id="62972A3F316DD90F40246AC7FBC61DFD" box="[1025,1114,1961,1987]" captionStart="FIGURE 2" captionStartId="8.[151,250,1805,1830]" captionTargetBox="[171,1421,437,1746]" captionTargetId="figure@8.[151,1436,417,1780]" captionTargetPageId="8" captionText="FIGURE 2. H. boerneri. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677873" httpUri="https://zenodo.org/record/3677873/files/figure.png" pageId="16" pageNumber="417">Fig. 2D</figureCitation>
); both species have posterior margin of gonocoxites 8 sinuous while
<emphasis id="C8D8EAA8316DD90F466F6AA0FD721DD9" box="[586,750,1996,2024]" italics="true" pageId="16" pageNumber="417">
<taxonomicName id="3DAC4D39316DD90F466F6AA0FD7F1DD9" box="[586,739,1996,2024]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA8316DD90F466F6AA0FD7F1DD9" bold="true" box="[586,739,1996,2024]" italics="true" pageId="16" pageNumber="417">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
has posterior margin arched (
<figureCitation id="62972A3F316DD90F40136AA3FB121DD9" box="[1078,1166,1997,2023]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. AC Male, pygophore; A—dorsal view; Bposterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="16" pageNumber="417">Fig. 1D</figureCitation>
). Males of
<taxonomicName id="3DAC4D39316DD90F412F6AA0FA011DD9" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[1290,1437,1997,2023]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="417" phylum="Arthropoda" rank="species" species="offuscatus">
<emphasis id="C8D8EAA8316DD90F412F6AA0FA011DD9" box="[1290,1437,1997,2023]" italics="true" pageId="16" pageNumber="417">H. offuscatus</emphasis>
</taxonomicName>
and
<emphasis id="C8D8EAA8316CD90E44ED6DF7FEF31A8D" box="[200,367,152,179]" italics="true" pageId="17" pageNumber="418">
<taxonomicName id="3DAC4D39316CD90E44ED6DF7FEFF1A8D" box="[200,355,152,179]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA8316CD90E44ED6DF7FEFF1A8D" bold="true" box="[200,355,152,179]" italics="true" pageId="17" pageNumber="418">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
are very similar and can be separated by subtle characteristics like paramere more robust and dorsal rim more excavated in
<taxonomicName id="3DAC4D39316CD90E45C26DD0FDE01AE9" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[487,636,189,215]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="offuscatus">
<emphasis id="C8D8EAA8316CD90E45C26DD0FDE01AE9" box="[487,636,189,215]" italics="true" pageId="17" pageNumber="418">H. offuscatus</emphasis>
</taxonomicName>
(
<figureCitation id="62972A3F316CD90E46A96DD3FD7D1AE9" box="[652,737,189,215]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="17" pageNumber="418">Fig. 7B</figureCitation>
and
<figureCitation id="62972A3F316CD90E473C6DD3FCA71AE9" box="[793,827,189,215]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. AC Male, pygophore; A—dorsal view; Bposterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="17" pageNumber="418">1B</figureCitation>
); more obvious distinctive characteristic is the expansion of ventral rim more developed and rounded in
<taxonomicName id="3DAC4D39316CD90E46F26D8CFCF01AC5" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1907" box="[727,876,225,251]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="offuscatus">
<emphasis id="C8D8EAA8316CD90E46F26D8CFCF01AC5" box="[727,876,225,251]" italics="true" pageId="17" pageNumber="418">H. offuscatus</emphasis>
</taxonomicName>
(
<figureCitation id="62972A3F316CD90E475F6D8FFC4D1AC5" box="[890,977,225,251]" captionStart="FIGURE 7" captionStartId="16.[151,250,1746,1771]" captionTargetBox="[169,1410,207,1693]" captionTargetId="figure@16.[151,1436,181,1721]" captionTargetPageId="16" captionText="FIGURE 7. H. offuscatus. AC Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677883" httpUri="https://zenodo.org/record/3677883/files/figure.png" pageId="17" pageNumber="418">Fig. 7C</figureCitation>
) than in
<emphasis id="C8D8EAA8316CD90E40116D8FFB451AC5" box="[1076,1241,224,251]" italics="true" pageId="17" pageNumber="418">
<taxonomicName id="3DAC4D39316CD90E40116D8FFB521AC5" box="[1076,1230,224,251]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="418" phylum="Arthropoda" rank="species" species="quadriden">
<emphasis id="C8D8EAA8316CD90E40116D8FFB521AC5" bold="true" box="[1076,1230,224,251]" italics="true" pageId="17" pageNumber="418">H. quadriden</emphasis>
</taxonomicName>
s
</emphasis>
(
<figureCitation id="62972A3F316CD90E40CD6D8FFAA31AC5" box="[1256,1343,225,251]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1937]" captionTargetBox="[225,1364,199,1866]" captionTargetId="figure@5.[214,1374,179,1884]" captionTargetPageId="5" captionText="FIGURE 1. H. quadridens. AC Male, pygophore; A—dorsal view; Bposterior view; C—ventral view. D—Female, genital plates. E—metasternal process; F—Habitus, Lectotype male; G—specimen data. (pyg—pygophore; dr—dorsal rim; vr—ventral rim; proc—proctiger; gp—genital superior process; pa—parameres; gc8—gonocoxite 8; gc9—gonocoxites 9; la8—laterotergite 8; la9—laterotergite 9; x—abdominal segment X. Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677871" httpUri="https://zenodo.org/record/3677871/files/figure.png" pageId="17" pageNumber="418">Fig. 1C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B66531316CD90E44E26C6AFDE71B21" box="[199,635,260,287]" pageId="17" pageNumber="418" type="distribution">
<paragraph id="FA1336BA316CD90E44E26C6AFDE71B21" blockId="17.[151,1436,152,287]" box="[199,635,260,287]" pageId="17" pageNumber="418">
<emphasis id="C8D8EAA8316CD90E44E26C6AFE571B21" bold="true" box="[199,459,260,287]" pageId="17" pageNumber="418">
Distribution (
<figureCitation id="62972A3F316CD90E454D6C6AFE211B21" box="[360,445,260,287]" captionStart="FIGURE 21" captionStartId="37.[151,250,1173,1198]" captionTargetBox="[159,1429,193,1137]" captionTargetId="figure@37.[151,1436,181,1148]" captionTargetPageId="37" captionText="FIGURE 21. Distributional map of the species of H. quadridens group." figureDoi="http://doi.org/10.5281/zenodo.3677911" httpUri="https://zenodo.org/record/3677911/files/figure.png" pageId="17" pageNumber="418">Fig. 21</figureCitation>
).
</emphasis>
<collectingCountry id="82BB762A316CD90E45F76C6BFD821B21" box="[466,542,261,287]" name="Peru" pageId="17" pageNumber="418">PERU</collectingCountry>
:
<collectingRegion id="3868F858316CD90E460D6C6BFDEA1B21" box="[552,630,261,287]" country="Peru" name="Loreto" pageId="17" pageNumber="418">Loreto</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>