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<document id="1CAB6EC9C8977C6C0ACD3423D5698F84" ID-CLB-Dataset="23869" ID-DOI="10.11646/zootaxa.4742.3.1" ID-GBIF-Dataset="84ef451f-d1a1-48f6-871e-f688204fd959" ID-ISSN="1175-5326" ID-Zenodo-Dep="3677869" ID-ZooBank="44ECBE3C-DEA4-4A6B-87D8-D7065591577C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582272866112" checkinUser="plazi" docAuthor="Nunes, Benedito Mendes, Campos, Lourival Dias, Mendonca, Maria Thayane Da Silva, Cunha, Eduardo Victor De Paiva &amp; Fernandes, Jose Antonio Marin" docDate="2020" docId="720587AC3166D90344B26DF4FCAE1999" docLanguage="en" docName="zootaxa.4742.3.1.pdf" docOrigin="Zootaxa 4742 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hypoxys brachyacanthus Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes 2020, comb. nov." docType="treatment" docVersion="12" lastPageNumber="429" masterDocId="8E3CFFD4317DD91F44256D6EFF9C1A3E" masterDocTitle="Revision of Hypoxys Amyot &amp; Serville, 1843 stat. rest. (Heteroptera Pentatomidae)" masterLastPageNumber="441" masterPageNumber="401" pageNumber="428" updateTime="1732689660721" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="45D54BDCEEC4643EFBB044029CD44B81" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4FF92CB146DE9BB0F7B6C879ECDE1F43">Revision of Hypoxys Amyot &amp; Serville, 1843 stat. rest. (Heteroptera Pentatomidae)</mods:title>
</mods:titleInfo>
<mods:name id="0B537BE75B70FD379EE6CD65800C9F38" type="personal">
<mods:role id="EAAD78FE783D9768B05AE54AD68FF5D2">
<mods:roleTerm id="A16713843DF1B080BD555D7E438F137E">Author</mods:roleTerm>
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<mods:namePart id="8A4EBA81FBE48352125C9668588F5C66">Nunes, Benedito Mendes</mods:namePart>
</mods:name>
<mods:name id="E05B49DF990861BB9357BED54B2AA007" type="personal">
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<mods:namePart id="5C7E6630041F4E435DD9EFF9546A1FDD">Campos, Lourival Dias</mods:namePart>
</mods:name>
<mods:name id="6C4A0FFC08E9F6FC7EC474C8A5A8C624" type="personal">
<mods:role id="468E7DA7FBF19CE40B23901A3CD005C4">
<mods:roleTerm id="C277C3857497DE376B3730706F50D2FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="51771D4723C8912DD9303F1681439022">Mendonca, Maria Thayane Da Silva</mods:namePart>
</mods:name>
<mods:name id="1A27567D2959587307A59FA23FF93BA5" type="personal">
<mods:role id="E11523BA05A353972853CC3C8FA643F2">
<mods:roleTerm id="9479B6CF45E0E81B3DB54E582947C84D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3500F4CF9117B6389B46F39E813347B7">Cunha, Eduardo Victor De Paiva</mods:namePart>
</mods:name>
<mods:name id="233718CAEAA56A90599F0726B20B012E" type="personal">
<mods:role id="8E443A78D6A5445B4CAE074F52E19309">
<mods:roleTerm id="DDEF243F8BC1D5B9BBBA48C052C9B935">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="772F204A8546BA4DB3F8B7734F48648A">Fernandes, Jose Antonio Marin</mods:namePart>
</mods:name>
<mods:typeOfResource id="27B4E8A5976ABE3D4FE06ECA5E2FC78C">text</mods:typeOfResource>
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<mods:title id="D4E3499F1AA2B358A2361393B428F056">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AC8B925AA9DFDD5ED4770B091104425A">
<mods:date id="73EA2B4A64569FDC0E31E13AC721DE31">2020</mods:date>
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<mods:number id="C53FCA0EF7909F8ADF1D062057C02204">2020-02-21</mods:number>
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<mods:detail id="FE0FC1F50E200057F4AF07B3DE5F73A3" type="volume">
<mods:number id="ED484C5C81D3CB0DEB9A8C2109AC6CAF">4742</mods:number>
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<mods:classification id="D1AE1EA97B79AE0CE1DA0C540C21ABE9">journal article</mods:classification>
<mods:identifier id="30E8A7E73EA7BBD75A6F431D54961ECE" type="CLB-Dataset">23869</mods:identifier>
<mods:identifier id="D7C74E71C9104385B9FDEDAB69ADD23B" type="DOI">10.11646/zootaxa.4742.3.1</mods:identifier>
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<treatment id="720587AC3166D90344B26DF4FCAE1999" ID-DOI="http://doi.org/10.5281/zenodo.3684957" ID-GBIF-Taxon="162244901" ID-Zenodo-Dep="3684957" LSID="urn:lsid:plazi:treatment:720587AC3166D90344B26DF4FCAE1999" httpUri="http://treatment.plazi.org/id/720587AC3166D90344B26DF4FCAE1999" lastPageId="28" lastPageNumber="429" pageId="27" pageNumber="428" scope_family="Pentatomidae">
<subSubSection id="B2B665313166D90444B26DF4FCBD1A8A" box="[151,801,153,180]" pageId="27" pageNumber="428" type="nomenclature">
<paragraph id="FA1336BA3166D90444B26DF4FCBD1A8A" blockId="27.[151,801,153,216]" box="[151,801,153,180]" pageId="27" pageNumber="428">
<heading id="A15B81D63166D90444B26DF4FCBD1A8A" bold="true" box="[151,801,153,180]" fontSize="11" level="1" pageId="27" pageNumber="428" reason="1">
<emphasis id="C8D8EAA83166D90444B26DF4FCBD1A8A" bold="true" box="[151,801,153,180]" pageId="27" pageNumber="428">
<taxonomicName id="3DAC4D393166D90444B26DF4FD0F1A8D" authority="(Breddin, 1904)" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[151,659,153,180]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="brachyacanthus" status="comb. nov.">
<emphasis id="C8D8EAA83166D90444B26DF4FE551A8A" bold="true" box="[151,457,153,180]" italics="true" pageId="27" pageNumber="428">Hypoxys brachyacanthus</emphasis>
(Breddin, 1904)
</taxonomicName>
<taxonomicNameLabel id="D3EB57D33166D90446BF6DF4FCBD1A8A" box="[666,801,154,180]" pageId="27" pageNumber="428" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B2B665313166D90444B26DD3FE551AE9" box="[151,457,189,216]" pageId="27" pageNumber="428" type="description">
<paragraph id="FA1336BA3166D90444B26DD3FE551AE9" blockId="27.[151,801,153,216]" box="[151,457,189,216]" pageId="27" pageNumber="428">
(
<figureCitation id="62972A3F3166D90444BA6DD3FF681AE9" box="[159,244,189,215]" captionStart="FIGURE 13" captionStartId="27.[151,250,1908,1933]" captionTargetBox="[184,1412,342,1856]" captionTargetId="figure@27.[151,1436,328,1883]" captionTargetPageId="27" captionText="FIGURE 13. H. brachyacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677895" httpUri="https://zenodo.org/record/3677895/files/figure.png" pageId="27" pageNumber="428">Figs 13</figureCitation>
A–D; 20 A–B; 23)
</paragraph>
</subSubSection>
<subSubSection id="B2B665313166D90444B26D95FC231B0C" pageId="27" pageNumber="428" type="reference_group">
<paragraph id="FA1336BA3166D90444B26D95FBFA1B2D" blockId="27.[151,1126,250,306]" box="[151,1126,250,275]" pageId="27" pageNumber="428">
<treatmentCitationGroup id="DABC11943166D90444B26D95FBFA1B2D" box="[151,1126,250,275]" pageId="27" pageNumber="428">
<treatmentCitation id="7B0D10AB3166D90444B26D95FDCA1B2D" author="Breddin, G." box="[151,598,250,275]" page="140" pageId="27" pageNumber="428" year="1904">
<taxonomicName id="3DAC4D393166D90444B26D95FDCA1B2D" authority="Breddin, 1904 a: 140" authorityName="Breddin" authorityPageNumber="140" authorityYear="1904" box="[151,598,250,275]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="brachyacantha">
<emphasis id="C8D8EAA83166D90444B26D95FEE11B2D" box="[151,381,251,275]" italics="true" pageId="27" pageNumber="428">Edessa brachyacantha</emphasis>
<bibRefCitation id="9E3D4B4B3166D90445A16D94FDCA1B2D" author="Breddin, G." box="[388,598,250,275]" pageId="27" pageNumber="428" pagination="135 - 153" refId="ref24439" refString="Breddin, G. (1904 a) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1904, 135 - 153." type="journal article" year="1904">Breddin, 1904a: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; Bergroth, 1908: 178;
<treatmentCitation id="7B0D10AB3166D904471F6D94FB991B2D" author="Kirkaldy, G. W." box="[826,1029,250,275]" page="155" pageId="27" pageNumber="428" year="1909">
<bibRefCitation id="9E3D4B4B3166D904471F6D94FB991B2D" author="Kirkaldy, G. W." box="[826,1029,250,275]" pageId="27" pageNumber="428" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 155</bibRefCitation>
</treatmentCitation>
(catalog)
</treatmentCitationGroup>
</paragraph>
<paragraph id="FA1336BA3166D90444B26C74FC231B0C" blockId="27.[151,1126,250,306]" box="[151,959,281,306]" pageId="27" pageNumber="428">
<materialsCitation id="4AC43CE73166D90444B26C74FC231B0C" ID-GBIF-Occurrence="2570056031" box="[151,959,281,306]" collectionCode="SDEI" country="Bolivia" location="Bolivia" pageId="27" pageNumber="428" specimenCount="1" typeStatus="lectotype">
<typeStatus id="251788183166D90444B26C74FF601B0C" box="[151,252,282,306]" pageId="27" pageNumber="428" type="lectotype">Lectotype</typeStatus>
male.
<collectingCountry id="82BB762A3166D90445646C77FE101B0C" box="[321,396,281,306]" name="Bolivia" pageId="27" pageNumber="428">Bolivia</collectingCountry>
(
<collectionCode id="9CBDAE7F3166D90445BC6C74FE4B1B0C" box="[409,471,282,306]" pageId="27" pageNumber="428">SDEI</collectionCode>
). Designated by
<bibRefCitation id="9E3D4B4B3166D904465A6C77FCD41B0C" author="Gaedike, H." box="[639,840,281,306]" pageId="27" pageNumber="428" pagination="79 - 159" refId="ref24984" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen- V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike (1971: 81)</bibRefCitation>
. Examined.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="AED366323166D90444B26A1AFE311D8F" ID-DOI="http://doi.org/10.5281/zenodo.3677895" ID-Zenodo-Dep="3677895" httpUri="https://zenodo.org/record/3677895/files/figure.png" pageId="27" pageNumber="428" startId="27.[151,250,1908,1933]" targetBox="[184,1412,342,1856]" targetPageId="27">
<paragraph id="FA1336BA3166D90444B26A1AFE311D8F" blockId="27.[151,1436,1908,1969]" pageId="27" pageNumber="428">
<emphasis id="C8D8EAA83166D90444B26A1AFEBC1DB3" bold="true" box="[151,288,1908,1933]" pageId="27" pageNumber="428">FIGURE 13.</emphasis>
<taxonomicName id="3DAC4D393166D90445026A18FE751DB3" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[295,489,1909,1933]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="brachyacanthus">
<emphasis id="C8D8EAA83166D90445026A18FE751DB3" box="[295,489,1909,1933]" italics="true" pageId="27" pageNumber="428">H. brachyacanthus</emphasis>
</taxonomicName>
. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—metaster-nal process (Scale = 1mm).
</paragraph>
</caption>
<subSubSection id="B2B665313161D90344B26DF6FEB61B21" pageId="28" pageNumber="429" type="materials_examined">
<paragraph id="FA1336BA3161D90344B26DF6FEB61B21" blockId="28.[151,1437,152,935]" pageId="28" pageNumber="429">
<emphasis id="C8D8EAA83161D90344B26DF6FE1C1A8D" bold="true" box="[151,384,152,179]" pageId="28" pageNumber="429">Material examined.</emphasis>
<materialsCitation id="4AC43CE73161D90345A36DF6FBBD1AE9" ID-GBIF-Occurrence="2570056075" collectionCode="USMN" collectorName="From Rolle." country="Bolivia" elevation="1000" location="La Paz" pageId="28" pageNumber="429" specimenCount="1" specimenCount-male="1" stateProvince="La Paz">
<emphasis id="C8D8EAA83161D90345A36DF6FDF51A8D" bold="true" box="[390,617,152,179]" pageId="28" pageNumber="429">
<collectingCountry id="82BB762A3161D90345A36DF6FD9D1A8D" box="[390,513,152,179]" name="Bolivia" pageId="28" pageNumber="429">BOLIVIA</collectingCountry>
.
<collectingRegion id="3868F8583161D90346286DF7FDF81A8D" box="[525,612,153,179]" country="Bolivia" name="La Paz" pageId="28" pageNumber="429">La Paz</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD333161D90346556DF7FD191A8D" box="[624,645,153,179]" pageId="28" pageNumber="429" type="male">♂</specimenCount>
Yungas,
<quantity id="3D549B5F3161D90346CF6DF7FCDE1A8D" box="[746,834,153,179]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="429" unit="m" value="1000.0">
<elevation id="7181D1893161D90346CF6DF7FCDE1A8D" box="[746,834,153,179]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="429" unit="m" value="1000.0">1000 m</elevation>
</quantity>
.
<collectorName id="5759536C3161D90347696DF7FC4F1A8C" box="[844,979,152,179]" pageId="28" pageNumber="429">From Rolle.</collectorName>
EPVanDuzee collection,
<taxonomicName id="3DAC4D393161D90340CA6DF6FC8C1AE9" authority="Breddin, 1904 Fernandes, J. A. M. 1999 Comp." class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="brachyacantha">
<emphasis id="C8D8EAA83161D90340CA6DF6FF6D1AE8" italics="true" pageId="28" pageNumber="429">Edessa brachyacantha</emphasis>
Breddin, 1904 Fernandes, J. A. M. 1999 Comp.
</taxonomicName>
HOMOTYPE (
<collectionCode id="9CBDAE7F3161D90347E76DD3FB851AE9" box="[962,1049,189,215]" pageId="28" pageNumber="429">USMN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4AC43CE73161D90340086DD2FEBA1B21" ID-GBIF-Occurrence="2570056098" collectingDate="1950-09-17" collectionCode="ZSM" collectorName="W. Forster." country="Bolivia" determinerName="P. H. v. Doesburg Jr." elevation="1000" location="San Carlos" municipality="Sarampiuni" pageId="28" pageNumber="429" specimenCount="1" specimenCount-male="1" stateProvince="Santa Cruz">
<emphasis id="C8D8EAA83161D90340086DD2FB221AE9" bold="true" box="[1069,1214,188,215]" pageId="28" pageNumber="429">
<collectingRegion id="3868F8583161D90340086DD2FB261AE9" box="[1069,1210,188,215]" country="Bolivia" name="Santa Cruz" pageId="28" pageNumber="429">Santa Cruz</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD333161D90340E36DD3FB471AE9" box="[1222,1243,189,215]" pageId="28" pageNumber="429" type="male">♂</specimenCount>
<collectingMunicipality id="1A77ACC03161D90340C66DD2FAFB1AE8" box="[1251,1383,188,214]" pageId="28" pageNumber="429">Sarampiuni</collectingMunicipality>
,
<location id="FF7360613161D90341576DD3FF7F1AC4" LSID="urn:lsid:plazi:treatment:720587AC3166D90344B26DF4FCAE1999:FF7360613161D90341576DD3FF7F1AC4" country="Bolivia" municipality="Sarampiuni" name="San Carlos" pageId="28" pageNumber="429" stateProvince="Santa Cruz">San Carlos</location>
,
<quantity id="3D549B5F3161D90344CA6D8FFED61AC5" box="[239,330,225,251]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="429" unit="m" value="1000.0">
<elevation id="7181D1893161D90344CA6D8FFED61AC5" box="[239,330,225,251]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="429" unit="m" value="1000.0">1000 m</elevation>
</quantity>
.,
<date id="8E12107A3161D903457F6D8EFE471AC5" box="[346,475,224,251]" pageId="28" pageNumber="429" value="1950-09-17">
<collectingDate id="9E56E9923161D903457F6D8EFE471AC5" box="[346,475,224,251]" pageId="28" pageNumber="429" value="1950-09-17">17–9–1950</collectingDate>
</date>
, leg.
<collectorName id="5759536C3161D90346326D8FFD0F1AC5" box="[535,659,225,251]" pageId="28" pageNumber="429">W. Forster.</collectorName>
<taxonomicName id="3DAC4D393161D90346BF6D8EFB8A1AC5" authority="Br. Comp." authorityName="Br. Comp." box="[666,1046,224,251]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="brachyacantha">
<emphasis id="C8D8EAA83161D90346BF6D8EFC051AC4" box="[666,921,224,250]" italics="true" pageId="28" pageNumber="429">Edessa brachyacantha</emphasis>
Br. Comp.
</taxonomicName>
w.
<typeStatus id="251788183161D903401B6D8FFB181AC5" box="[1086,1156,225,251]" pageId="28" pageNumber="429">TYPE</typeStatus>
<determinerName id="0FCFDA7E3161D90340A96D8FFAEF1AC5" box="[1164,1395,224,251]" pageId="28" pageNumber="429">P. H. v. Doesburg Jr.</determinerName>
det 1978 (
<collectionCode id="9CBDAE7F3161D90344FB6C6BFE811B21" box="[222,285,261,287]" country="Germany" name="Bavarian State Collection of Zoology" pageId="28" pageNumber="429" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313161D90344E26C47FAD61BB5" pageId="28" pageNumber="429" type="description">
<paragraph id="FA1336BA3161D90344E26C47FAD61BB5" blockId="28.[151,1437,152,935]" pageId="28" pageNumber="429">
<emphasis id="C8D8EAA83161D90344E26C47FEE21B7D" bold="true" box="[199,382,297,323]" pageId="28" pageNumber="429">Measurements.</emphasis>
total length: 15.3–18.1; head length: 1.9–2.2; head width: 3.0–3.6; pronotal width: 9.7–12.3; abdominal width: 8.0–10.3; length of antennal segments. I: 0.7–1.0; II: 1.6–2.0; III: 1.1–1.6; IV: 3.3–4.8; V: 4.3–5.0; interocular distance: 1.4–1.7; scutellum length: 7.1–8.6; scutellum width: 5.2–6.7; pronotal length: 3.3–4.0.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313161D90344E26CFAFDDA18F0" pageId="28" pageNumber="429" type="diagnosis">
<paragraph id="FA1336BA3161D90344E26CFAFDDA18F0" blockId="28.[151,1437,152,935]" pageId="28" pageNumber="429">
<emphasis id="C8D8EAA83161D90344E26CFAFEDE1B91" bold="true" box="[199,322,404,431]" pageId="28" pageNumber="429">Diagnosis.</emphasis>
Anterolateral margin of pronotum without solid black stripe. Pronotum with small, elongated, lateral black spots including posterolateral angles. Connexivum green without spots (
<figureCitation id="62972A3F3161D90340366CD6FBE21BED" box="[1043,1150,440,467]" captionStart="FIGURE 20" captionStartId="36.[151,250,1749,1774]" captionTargetBox="[181,1415,196,1699]" captionTargetId="figure@36.[151,1436,181,1723]" captionTargetPageId="36" captionText="FIGURE 20. Dorsal and ventral view. H. brachyacanthus A–B; H. infulatus C–D; H. leptacanthus E–F; H. tragelaphus G–H (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677909" httpUri="https://zenodo.org/record/3677909/files/figure.png" pageId="28" pageNumber="429">Fig. 20A</figureCitation>
). Anterolateral margin of pronotum not punctured. Anterior margin of scutellum concolorous (
<figureCitation id="62972A3F3161D90347AB6CB2FC6A1BC9" box="[910,1014,476,503]" captionStart="FIGURE 20" captionStartId="36.[151,250,1749,1774]" captionTargetBox="[181,1415,196,1699]" captionTargetId="figure@36.[151,1436,181,1723]" captionTargetPageId="36" captionText="FIGURE 20. Dorsal and ventral view. H. brachyacanthus A–B; H. infulatus C–D; H. leptacanthus E–F; H. tragelaphus G–H (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677909" httpUri="https://zenodo.org/record/3677909/files/figure.png" pageId="28" pageNumber="429">Fig. 20A</figureCitation>
). Black spot of costal margin reaching approximately 1/3 of corial extension. Dorsal surface of abdomen yellow to olive green with brown longitudinal stripe. Ventrally end of pseudosutures with rounded dark spots (
<figureCitation id="62972A3F3161D903475F6F4AFC7F1801" box="[890,995,548,575]" captionStart="FIGURE 20" captionStartId="36.[151,250,1749,1774]" captionTargetBox="[181,1415,196,1699]" captionTargetId="figure@36.[151,1436,181,1723]" captionTargetPageId="36" captionText="FIGURE 20. Dorsal and ventral view. H. brachyacanthus A–B; H. infulatus C–D; H. leptacanthus E–F; H. tragelaphus G–H (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677909" httpUri="https://zenodo.org/record/3677909/files/figure.png" pageId="28" pageNumber="429">Fig. 20B</figureCitation>
). Male (
<figureCitation id="62972A3F3161D90340606F4AFB0A1801" box="[1093,1174,548,575]" captionStart="FIGURE 13" captionStartId="27.[151,250,1908,1933]" captionTargetBox="[184,1412,342,1856]" captionTargetId="figure@27.[151,1436,328,1883]" captionTargetPageId="27" captionText="FIGURE 13. H. brachyacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677895" httpUri="https://zenodo.org/record/3677895/files/figure.png" pageId="28" pageNumber="429">Fig. 13</figureCitation>
A–C). Pygophore with dorsal rim sinuous; median part almost straight (
<figureCitation id="62972A3F3161D90346E06F26FCB3185D" box="[709,815,584,611]" captionStart="FIGURE 13" captionStartId="27.[151,250,1908,1933]" captionTargetBox="[184,1412,342,1856]" captionTargetId="figure@27.[151,1436,328,1883]" captionTargetPageId="27" captionText="FIGURE 13. H. brachyacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677895" httpUri="https://zenodo.org/record/3677895/files/figure.png" pageId="28" pageNumber="429">Fig. 13A</figureCitation>
). Paramere with anterior lobe truncate, reaching superior process of genital cup; posterior lobe triangular and laterally bent (
<figureCitation id="62972A3F3161D903479E6F02FBBE18B9" box="[955,1058,620,647]" captionStart="FIGURE 13" captionStartId="27.[151,250,1908,1933]" captionTargetBox="[184,1412,342,1856]" captionTargetId="figure@27.[151,1436,328,1883]" captionTargetPageId="27" captionText="FIGURE 13. H. brachyacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677895" httpUri="https://zenodo.org/record/3677895/files/figure.png" pageId="28" pageNumber="429">Fig. 13B</figureCitation>
). Proctiger with widely separated tufts of short setae; posterior face ogival and flat (
<figureCitation id="62972A3F3161D90346E26FFEFCB21895" box="[711,814,656,683]" captionStart="FIGURE 13" captionStartId="27.[151,250,1908,1933]" captionTargetBox="[184,1412,342,1856]" captionTargetId="figure@27.[151,1436,328,1883]" captionTargetPageId="27" captionText="FIGURE 13. H. brachyacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677895" httpUri="https://zenodo.org/record/3677895/files/figure.png" pageId="28" pageNumber="429">Fig. 13B</figureCitation>
). Ventral rim with expansions little developed, flat, and rounded (
<figureCitation id="62972A3F3161D90345276FDAFEF418F1" box="[258,360,692,719]" captionStart="FIGURE 13" captionStartId="27.[151,250,1908,1933]" captionTargetBox="[184,1412,342,1856]" captionTargetId="figure@27.[151,1436,328,1883]" captionTargetPageId="27" captionText="FIGURE 13. H. brachyacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677895" httpUri="https://zenodo.org/record/3677895/files/figure.png" pageId="28" pageNumber="429">Fig. 13C</figureCitation>
). Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313161D90344E26FB6FD5319BD" pageId="28" pageNumber="429" type="discussion">
<paragraph id="FA1336BA3161D90344E26FB6FD5319BD" blockId="28.[151,1437,152,935]" pageId="28" pageNumber="429">
<emphasis id="C8D8EAA83161D90344E26FB6FED318CD" bold="true" box="[199,335,728,755]" pageId="28" pageNumber="429">Comments.</emphasis>
<bibRefCitation id="9E3D4B4B3161D90345726FB6FD8C18CD" author="Breddin, G." box="[343,528,728,755]" pageId="28" pageNumber="429" pagination="135 - 153" refId="ref24439" refString="Breddin, G. (1904 a) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1904, 135 - 153." type="journal article" year="1904">Breddin (1904a)</bibRefCitation>
related
<taxonomicName id="3DAC4D393161D903464E6FB6FCF618CC" box="[619,874,728,754]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="brachyacantha">
<emphasis id="C8D8EAA83161D903464E6FB6FCF618CC" box="[619,874,728,754]" italics="true" pageId="28" pageNumber="429">Edessa brachyacantha</emphasis>
</taxonomicName>
to
<taxonomicName id="3DAC4D393161D90347AA6FB7FBD018CC" authorityName="Breddin" authorityYear="1904" box="[911,1100,728,754]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="pachyacantha">
<emphasis id="C8D8EAA83161D90347AA6FB7FBD018CC" box="[911,1100,728,754]" italics="true" pageId="28" pageNumber="429">E. pachyacantha</emphasis>
</taxonomicName>
. Here, the species
<taxonomicName id="3DAC4D393161D90341076FB7FF7F1928" authorityName="Breddin" authorityYear="1904" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="pachyacantha">
<emphasis id="C8D8EAA83161D90341076FB7FAAF18CC" box="[1314,1331,729,754]" italics="true" pageId="28" pageNumber="429">E</emphasis>
.
<emphasis id="C8D8EAA83161D90341676FB6FF7F1928" italics="true" pageId="28" pageNumber="429">pachyacantha</emphasis>
</taxonomicName>
is considered a junior synonym of
<taxonomicName id="3DAC4D393161D90346506F93FC821928" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1903" box="[629,798,764,790]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="tragelaphus">
<emphasis id="C8D8EAA83161D90346506F93FC821928" box="[629,798,764,790]" italics="true" pageId="28" pageNumber="429">H. tragelaphus</emphasis>
</taxonomicName>
. Both species are very different;
<taxonomicName id="3DAC4D393161D90340BD6F93FADE1928" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1903" box="[1176,1346,764,790]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="tragelaphus">
<emphasis id="C8D8EAA83161D90340BD6F93FADE1928" box="[1176,1346,764,790]" italics="true" pageId="28" pageNumber="429">H. tragelaphus</emphasis>
</taxonomicName>
has humeral angles developed and globose (
<figureCitation id="62972A3F3161D903461A6E4EFD341905" box="[575,680,800,827]" captionStart="FIGURE 20" captionStartId="36.[151,250,1749,1774]" captionTargetBox="[181,1415,196,1699]" captionTargetId="figure@36.[151,1436,181,1723]" captionTargetPageId="36" captionText="FIGURE 20. Dorsal and ventral view. H. brachyacanthus A–B; H. infulatus C–D; H. leptacanthus E–F; H. tragelaphus G–H (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677909" httpUri="https://zenodo.org/record/3677909/files/figure.png" pageId="28" pageNumber="429">Fig. 20G</figureCitation>
).
<taxonomicName id="3DAC4D393161D90346996E4FFC4A1904" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[700,982,800,826]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="brachyacanthus">
<emphasis id="C8D8EAA83161D90346996E4FFC4A1904" box="[700,982,800,826]" italics="true" pageId="28" pageNumber="429">Hypoxys brachyacanthus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="3DAC4D393161D903404F6E4FFA801904" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[1130,1308,800,826]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="leptacanthus">
<emphasis id="C8D8EAA83161D903404F6E4FFA801904" box="[1130,1308,800,826]" italics="true" pageId="28" pageNumber="429">H. leptacanthus</emphasis>
</taxonomicName>
in size and spots on pronotum, but
<taxonomicName id="3DAC4D393161D90345B86E2BFDD21960" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[413,590,836,862]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="leptacanthus">
<emphasis id="C8D8EAA83161D90345B86E2BFDD21960" box="[413,590,836,862]" italics="true" pageId="28" pageNumber="429">H. leptacanthus</emphasis>
</taxonomicName>
has dorsal rim of the pygophore projected beyond posterolateral angles (not projected in
<taxonomicName id="3DAC4D393161D90345066E07FE6419BC" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[291,504,872,898]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="brachyacanthus">
<emphasis id="C8D8EAA83161D90345066E07FE6419BC" box="[291,504,872,898]" italics="true" pageId="28" pageNumber="429">H. brachyacanthus</emphasis>
</taxonomicName>
) (
<figureCitation id="62972A3F3161D90346346E06FD1F19BD" box="[529,643,872,899]" captionStart="FIGURE 15" captionStartId="30.[151,250,1667,1692]" captionTargetBox="[171,1415,201,1606]" captionTargetId="figure@30.[151,1436,181,1641]" captionTargetPageId="30" captionText="FIGURE 15. H. leptacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677899" httpUri="https://zenodo.org/record/3677899/files/figure.png" pageId="28" pageNumber="429">Figs. 15A</figureCitation>
,
<figureCitation id="62972A3F3161D90346AB6E07FD5F19BD" box="[654,707,873,899]" captionStart="FIGURE 13" captionStartId="27.[151,250,1908,1933]" captionTargetBox="[184,1412,342,1856]" captionTargetId="figure@27.[151,1436,328,1883]" captionTargetPageId="27" captionText="FIGURE 13. H. brachyacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677895" httpUri="https://zenodo.org/record/3677895/files/figure.png" pageId="28" pageNumber="429">13A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2B665313161D90344E26EE2FCAE1999" box="[199,818,908,935]" pageId="28" pageNumber="429" type="distribution">
<paragraph id="FA1336BA3161D90344E26EE2FCAE1999" blockId="28.[151,1437,152,935]" box="[199,818,908,935]" pageId="28" pageNumber="429">
<emphasis id="C8D8EAA83161D90344E26EE2FE571999" bold="true" box="[199,459,908,935]" pageId="28" pageNumber="429">
Distribution (
<figureCitation id="62972A3F3161D903454D6EE2FE221999" box="[360,446,908,935]" captionStart="FIGURE 23" captionStartId="39.[151,250,1206,1231]" captionTargetBox="[155,1431,192,1174]" captionTargetId="figure@39.[151,1436,181,1181]" captionTargetPageId="39" captionText="FIGURE 23. Distributional map of the species of H. oxyacanthus group." figureDoi="http://doi.org/10.5281/zenodo.3677915" httpUri="https://zenodo.org/record/3677915/files/figure.png" pageId="28" pageNumber="429">Fig. 23</figureCitation>
).
</emphasis>
<collectingCountry id="82BB762A3161D90345F76EE3FDD61999" box="[466,586,909,935]" name="Bolivia" pageId="28" pageNumber="429">BOLIVIA</collectingCountry>
:
<collectingRegion id="3868F8583161D90346716EE3FD3F1999" box="[596,675,909,935]" country="Bolivia" name="La Paz" pageId="28" pageNumber="429">La Paz</collectingRegion>
,
<collectingRegion id="3868F8583161D903468A6EE3FCB21999" box="[687,814,909,935]" country="Bolivia" name="Santa Cruz" pageId="28" pageNumber="429">Santa Cruz</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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