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<document id="92558C59275BA54F799075202193933F" ID="10.11646/zootaxa.4121.5.5" ID-CLB-Dataset="38880" ID-DOI="10.11646/zootaxa.4121.5.5" ID-GBIF-Dataset="116fb0e8-794d-402f-a4fe-e1558ba2da12" ID-ISSN="1175-5326" ID-Zenodo-Dep="258604" ID-ZooBank="5DE272A5-EDFF-468E-82F5-DC5D92880011" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465800124550" checkinUser="plazi" docAuthor="Podenas, Sigitas" docDate="2016" docId="7201A67AFFBAFFBB31A497B65D59FC64" docLanguage="en" docName="zootaxa.4121.5.5.pdf" docOrigin="Zootaxa 4121 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geranomyia piagola Podenas, 2016, n. sp." docType="treatment" docVersion="9" lastPageNumber="561" masterDocId="8E38DE02FFBEFFBD313397215F6CFF89" masterDocTitle="New Geranomyia crane flies (Diptera: Limoniidae) from Korea and Kunashir Island" masterLastPageNumber="565" masterPageNumber="555" pageNumber="559" updateTime="1698662824346" updateUser="ExternalLinkService">
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<mods:title id="771CA389FD9F7B5A849887A8DF9034E7">New Geranomyia crane flies (Diptera: Limoniidae) from Korea and Kunashir Island</mods:title>
</mods:titleInfo>
<mods:name id="5B5B679CB6880D39CAF6732FD5B22495" type="personal">
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<mods:namePart id="C06F1C891920A0B12A4D2E6008099B58">Podenas, Sigitas</mods:namePart>
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<mods:date id="E07B15DD08DA4BB60BDA9FFA7CB4A92B">2016</mods:date>
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<mods:number id="6742C87BB3CB3EC27DB802538BC45AD6">4121</mods:number>
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<mods:identifier id="213323A3E14A33F16AE3B4212172B36C" type="DOI">10.11646/zootaxa.4121.5.5</mods:identifier>
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<treatment id="7201A67AFFBAFFBB31A497B65D59FC64" ID-DOI="http://doi.org/10.5281/zenodo.6068945" ID-GBIF-Taxon="120827919" ID-Zenodo-Dep="6068945" LSID="urn:lsid:plazi:treatment:7201A67AFFBAFFBB31A497B65D59FC64" httpUri="http://treatment.plazi.org/id/7201A67AFFBAFFBB31A497B65D59FC64" lastPageId="6" lastPageNumber="561" pageId="4" pageNumber="559">
<subSubSection id="B2B244E7FFBAFFB931A497B65E45FF5A" pageId="4" pageNumber="559" type="nomenclature">
<paragraph id="FA17176CFFBAFFB931A497B65EBBFF3B" blockId="4.[151,471,151,211]" box="[151,471,151,178]" pageId="4" pageNumber="559">
<heading id="A15FA000FFBAFFB931A497B65EBBFF3B" bold="true" box="[151,471,151,178]" fontSize="11" level="1" pageId="4" pageNumber="559" reason="1">
<emphasis id="C8DCCB7EFFBAFFB931A497B65EBBFF3B" bold="true" box="[151,471,151,178]" pageId="4" pageNumber="559">
<taxonomicName id="3DA86CEFFFBAFFB931A497B65EE2FF38" ID-CoL="8H2GP" box="[151,398,151,177]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="559" phylum="Arthropoda" rank="species" species="piagola" status="sp. nov.">
<emphasis id="C8DCCB7EFFBAFFB931A497B65EE2FF38" bold="true" box="[151,398,151,177]" italics="true" pageId="4" pageNumber="559">Geranomyia piagola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3EF7605FFBAFFB930A697B95EBBFF3B" box="[405,471,152,178]" pageId="4" pageNumber="559" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FA17176CFFBAFFB931A4979B5E45FF5A" blockId="4.[151,471,151,211]" box="[151,297,186,211]" pageId="4" pageNumber="559">
<figureCitation id="62930BE9FFBAFFB931A4979B5E45FF5A" box="[151,297,186,211]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[188,1348,213,1877]" captionTargetId="figure@5.[173,1390,193,1898]" captionTargetPageId="5" captionText="FIGURES 7 11. Geranomyia piagola n. sp. 7: Wing; 8: Male genitalia, dorsal view; 9: Male genitalia, ventral view; 10: Ovipositor, lateral view; 11: Paramere of male genitalia. Abbreviations: cerc, cercus; hyp vlv, hypogynial valve; st 8, eight sternite; tg 9, ninth tergite; tg 10, tenth tergite." httpUri="https://zenodo.org/record/258606/files/figure.png" pageId="4" pageNumber="559">Figures 711</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B2B244E7FFBAFFB931A496235A41FE92" box="[151,1325,258,283]" pageId="4" pageNumber="559" type="etymology">
<paragraph id="FA17176CFFBAFFB931A496235A41FE92" blockId="4.[151,1437,258,2013]" box="[151,1325,258,283]" pageId="4" pageNumber="559">
<emphasis id="C8DCCB7EFFBAFFB931A496235E72FE92" bold="true" box="[151,286,258,283]" pageId="4" pageNumber="559">Etymology.</emphasis>
The species is named after the place name of Piagol valley (
<figureCitation id="62930BE9FFBAFFB932F796235B7AFE92" box="[964,1046,258,283]" captionStart="FIGURES 16 17" captionStartId="8.[151,264,1939,1962]" captionTargetBox="[279,1311,193,1906]" captionTargetId="figure@8.[279,1311,193,1932]" captionTargetPageId="8" captionText="FIGURES 16 17. Collecting localities of Geranomyia in South Korea. 16: Seomjin-gang (river) margin at Hadong, where first specimen of G. semjina n. sp. was collected in 2012; 17: Type locality of G. piagola n. sp. and G. semjina n. sp., Piagol valley, Jirisan National Park." httpUri="https://zenodo.org/record/258608/files/figure.png" pageId="4" pageNumber="559">Fig. 17</figureCitation>
), where it was collected.
</paragraph>
</subSubSection>
<subSubSection id="B2B244E7FFBAFFB931F496075BD4FD9F" pageId="4" pageNumber="559" type="materials_examined">
<paragraph id="FA17176CFFBAFFB931F496075BE4FEB6" blockId="4.[151,1437,258,2013]" box="[199,1160,294,319]" pageId="4" pageNumber="559">
<emphasis id="C8DCCB7EFFBAFFB931F496075EC5FEB6" bold="true" box="[199,425,294,319]" pageId="4" pageNumber="559">Material examined</emphasis>
.
<typeStatus id="2513A9CEFFBAFFB9308496065E83FEB6" box="[439,495,295,319]" pageId="4" pageNumber="559">Type</typeStatus>
specimens.
<typeStatus id="2513A9CEFFBAFFB9334B96075D8CFEB6" box="[632,736,294,319]" pageId="4" pageNumber="559" type="holotype">Holotype</typeStatus>
male,
<typeStatus id="2513A9CEFFBAFFB9321F96065CFBFEB6" box="[812,919,295,319]" pageId="4" pageNumber="559" type="paratype">paratypes</typeStatus>
<specimenCount id="ECAEDCE5FFBAFFB932AD96065C85FEB6" box="[926,1001,294,319]" pageId="4" pageNumber="559" type="male">1 male</specimenCount>
and
<specimenCount id="ECAEDCE5FFBAFFB9351396065BE8FEB6" box="[1056,1156,294,319]" pageId="4" pageNumber="559" type="female">1 female</specimenCount>
.
</paragraph>
<paragraph id="FA17176CFFBAFFB931F4966B5C12FE22" blockId="4.[151,1437,258,2013]" pageId="4" pageNumber="559">
<emphasis id="C8DCCB7EFFBAFFB931F4966B5E56FEEA" bold="true" box="[199,314,330,355]" pageId="4" pageNumber="559">
<typeStatus id="2513A9CEFFBAFFB931F4966B5E5AFEEA" box="[199,310,330,355]" pageId="4" pageNumber="559" type="holotype">Holotype</typeStatus>
,
</emphasis>
male (pinned, wing slide-mounted, genitalia in microvial filled with glycerol and pinned together).
<collectingCountry id="82BF57FCFFBAFFB931A4964E5E05FE0E" box="[151,361,367,391]" name="South Korea" pageId="4" pageNumber="559">SOUTH KOREA</collectingCountry>
, Jeollanam-do, Gurye-gun, Toji-myeon, Naeseo-ri, Jirisan National Park, Piagol valley,
<geoCoordinate id="9F9C71ABFFBAFFB931A496B35E7BFE22" box="[151,279,402,427]" direction="north" orientation="latitude" pageId="4" pageNumber="559" precision="1" value="35.2659">N35.26590</geoCoordinate>
,
<geoCoordinate id="9F9C71ABFFBAFFB9301196B35EC1FE22" box="[290,429,402,427]" direction="east" orientation="longitude" pageId="4" pageNumber="559" precision="1" value="127.58096">E127.58096</geoCoordinate>
, alt.
<quantity id="3D50BA89FFBAFFB930D196B35D40FE23" box="[482,556,402,427]" metricMagnitude="2" metricUnit="m" metricValue="4.46" pageId="4" pageNumber="559" unit="m" value="446.0">446 m</quantity>
, 2014.08.24, coll. S. Podenas.
</paragraph>
<paragraph id="FA17176CFFBAFFB931F496975BD4FD9F" blockId="4.[151,1437,258,2013]" pageId="4" pageNumber="559">
<emphasis id="C8DCCB7EFFBAFFB931F496975E24FE46" bold="true" box="[199,328,438,463]" pageId="4" pageNumber="559">
<typeStatus id="2513A9CEFFBAFFB931F496975E2FFE46" box="[199,323,438,463]" pageId="4" pageNumber="559" type="paratype">Paratypes</typeStatus>
:
</emphasis>
female (in ethanol), S.
<collectingCountry id="82BF57FCFFBAFFB9335696965DC0FE47" box="[613,684,439,462]" name="South Korea" pageId="4" pageNumber="559">Korea</collectingCountry>
, Jeollanam-do, Gurye-gun, Toji-myeon, Naedong-ri,
<geoCoordinate id="9F9C71ABFFBAFFB9342B96975AF4FE46" box="[1304,1432,438,463]" direction="north" orientation="latitude" pageId="4" pageNumber="559" precision="1" value="35.2658">N35.26580</geoCoordinate>
,
<geoCoordinate id="9F9C71ABFFBAFFB931A496FB5E4EFE7A" box="[151,290,474,499]" direction="east" orientation="longitude" pageId="4" pageNumber="559" precision="1" value="127.58128">E127.58128</geoCoordinate>
, alt.
<quantity id="3D50BA89FFBAFFB9306896FA5EC9FE7B" box="[347,421,475,499]" metricMagnitude="2" metricUnit="m" metricValue="3.78" pageId="4" pageNumber="559" unit="m" value="378.0">378 m</quantity>
, 2013.05.12, S. Podenas, net (
<collectionCode id="9CB98FA9FFBAFFB9323296FA5C26FE7A" box="[769,842,475,499]" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="4" pageNumber="559">NIBR</collectionCode>
); male (in ethanol), S.
<collectingCountry id="82BF57FCFFBAFFB9356196FA5BF5FE7B" box="[1106,1177,475,498]" name="South Korea" pageId="4" pageNumber="559">Korea</collectingCountry>
, Jeollanam-do, Guryegun, Toji-myeon, Naeseo-ri,
<geoCoordinate id="9F9C71ABFFBAFFB930E996DF5D36FD9E" box="[474,602,510,535]" direction="north" orientation="latitude" pageId="4" pageNumber="559" precision="1" value="35.27177">N35.27177</geoCoordinate>
,
<geoCoordinate id="9F9C71ABFFBAFFB9335696DF5D9DFD9E" box="[613,753,510,535]" direction="east" orientation="longitude" pageId="4" pageNumber="559" precision="1" value="127.57146">E127.57146</geoCoordinate>
, alt.
<quantity id="3D50BA89FFBAFFB9321696DE5C02FD9F" box="[805,878,511,535]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="559" unit="m" value="490.0">490 m</quantity>
, 2015.06.29, S. Podenas, net.
</paragraph>
</subSubSection>
<subSubSection id="B2B244E7FFBAFFB931F495035D9CFD0A" pageId="4" pageNumber="559" type="diagnosis">
<paragraph id="FA17176CFFBAFFB931F495035D9CFD0A" blockId="4.[151,1437,258,2013]" pageId="4" pageNumber="559">
<emphasis id="C8DCCB7EFFBAFFB931F495035E2DFDB2" bold="true" box="[199,321,546,571]" pageId="4" pageNumber="559">Diagnosis.</emphasis>
Distinguished from other species in the genus by distinct dark longitudinal stripe on pleura and details of male genitalia, especially strongly extended inner gonostylus, bearing small rostral prolongation armed with two-three short spines situated on same tubercle.
</paragraph>
</subSubSection>
<subSubSection id="B2B244E7FFBAFFBB31F495AF5BC9FDE9" lastPageId="6" lastPageNumber="561" pageId="4" pageNumber="559" type="description">
<paragraph id="FA17176CFFBAFFB931F495AF5A60F854" blockId="4.[151,1437,258,2013]" pageId="4" pageNumber="559">
<emphasis id="C8DCCB7EFFBAFFB931F495AF5E3EFD2E" bold="true" box="[199,338,654,679]" pageId="4" pageNumber="559">Description</emphasis>
(
<figureCitation id="62930BE9FFBAFFB9305295AF5EB7FD2E" box="[353,475,654,679]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[188,1348,213,1877]" captionTargetId="figure@5.[173,1390,193,1898]" captionTargetPageId="5" captionText="FIGURES 7 11. Geranomyia piagola n. sp. 7: Wing; 8: Male genitalia, dorsal view; 9: Male genitalia, ventral view; 10: Ovipositor, lateral view; 11: Paramere of male genitalia. Abbreviations: cerc, cercus; hyp vlv, hypogynial valve; st 8, eight sternite; tg 9, ninth tergite; tg 10, tenth tergite." httpUri="https://zenodo.org/record/258606/files/figure.png" pageId="4" pageNumber="559">Figs. 711</figureCitation>
). Adult, male (
<emphasis id="C8DCCB7EFFBAFFB933B795AE5DFBFD2F" box="[644,663,655,678]" italics="true" pageId="4" pageNumber="559">N</emphasis>
=2). General body coloration brown. Body length
<quantity id="3D50BA89FFBAFFB935F995AF5A38FD2F" box="[1226,1364,654,679]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.2" metricValueMin="5.9" pageId="4" pageNumber="559" unit="mm" value="6.05" valueMax="6.2" valueMin="5.9">5.96.2 mm</quantity>
. Wing length
<quantity id="3D50BA89FFBAFFB931D795935E03FD43" box="[228,367,690,715]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.2" metricValueMin="6.5" pageId="4" pageNumber="559" unit="mm" value="6.85" valueMax="7.2" valueMin="6.5">6.57.2 mm</quantity>
.
<emphasis id="C8DCCB7EFFBAFFB9304995935EDAFD42" box="[378,438,690,715]" italics="true" pageId="4" pageNumber="559">Head</emphasis>
: black, sparsely dusted with gray pruinosity, covered with scarce semi-erect black setae. Vertex very narrow. Eyes large, dorsally separated by very narrow vertex, ventrally reaching each other. Length of antenna 1.0
<quantity id="3D50BA89FFBAFFB9301B95DA5EEEFC9B" box="[296,386,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="559" unit="mm" value="1.2">1.2 mm</quantity>
. Scape cylindrical, black, dorsally bearing few short erect dark setae. Pedicel rounded, same color as scape, dorsally bearing few short erect dark setae. Both basal segments sparsely dusted with gray. Flagellum 12-segmented. Three basal flagellomeres black, remaining segments brown, turning lighter towards apex. Basal flagellomere approximately as long as pedicel, distinctly narrower; remaining flagellomeres oval, slightly decreasing in length, apical flagellomere approximately 1.5 times as long as preceding segment. Verticils short, approximately as long as respective segments. Short whitish pubescence denser on ventral side of segments. Rostrum dark brown dorsally, lighter ventrally, reaching to second flagellar segment. Mouth parts same color as rostrum, covered with short adherent brownish setae. Rostrum together with mouth parts
<quantity id="3D50BA89FFBAFFB9359A94D65A6FFB87" box="[1193,1283,1015,1039]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="559" unit="mm" value="2.3">2.3 mm</quantity>
long. Palpus dark brown, three-segmented. Basal segment very short, second segment oval, wider, longer than first segment, apical segment very small, rounded with few distinct erect setae.
<emphasis id="C8DCCB7EFFBAFFB932A5931F5C8BFBDE" box="[918,999,1086,1111]" italics="true" pageId="4" pageNumber="559">Thorax</emphasis>
: generally yellowish brown. Cervical sclerite blackened. Pronotum frontally dark brown, posteriorly brownish yellow. Ground color of mesonotal prescutum light brown. Median prescutal stripe brown, distinct, lateral stripes indistinct, slightly darker than surrounding area. Frontal margin of prescutum yellowish. Pseudosutural fovea yellowish. Scutal lobes brown frontally, dark brown posteriorly. Scutellum dark brown frontally, posteriorly and laterally brown. Mediotergite uniformly dark brown. Pleuron yellowish brown dorsally, pale yellowish ventrally with distinct wide dark brown longitudinal stripe. Wing (
<figureCitation id="62930BE9FFBAFFB930F592375D67FAA6" box="[454,523,1302,1327]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[188,1348,213,1877]" captionTargetId="figure@5.[173,1390,193,1898]" captionTargetPageId="5" captionText="FIGURES 7 11. Geranomyia piagola n. sp. 7: Wing; 8: Male genitalia, dorsal view; 9: Male genitalia, ventral view; 10: Ovipositor, lateral view; 11: Paramere of male genitalia. Abbreviations: cerc, cercus; hyp vlv, hypogynial valve; st 8, eight sternite; tg 9, ninth tergite; tg 10, tenth tergite." httpUri="https://zenodo.org/record/258606/files/figure.png" pageId="4" pageNumber="559">Fig. 7</figureCitation>
) brownish, subcostal cell yellowish. Stigma distinct, brown, egg-shaped. Small brownish spots surrounding tip of
<emphasis id="C8DCCB7EFFBAFFB93327921D5D58FADA" box="[532,564,1339,1363]" italics="true" pageId="4" pageNumber="559">R3</emphasis>
, both branches of
<emphasis id="C8DCCB7EFFBAFFB93232921A5C77FADB" box="[769,795,1339,1362]" italics="true" pageId="4" pageNumber="559">Sc</emphasis>
, base of
<emphasis id="C8DCCB7EFFBAFFB9324D921D5CF6FADA" box="[894,922,1340,1363]" italics="true" pageId="4" pageNumber="559">Rs</emphasis>
and supernumerary cross-vein in cell
<emphasis id="C8DCCB7EFFBAFFB93471921D5A35FADA" box="[1346,1369,1340,1363]" italics="true" pageId="4" pageNumber="559">sc</emphasis>
. Very weak darkenings surround cord and distal end of discal cell. Most veins brown, veins at wing base pale. Venation:
<emphasis id="C8DCCB7EFFBAFFB931A492A55FD3FA15" box="[151,191,1412,1436]" italics="true" pageId="4" pageNumber="559">Sc1</emphasis>
long, ending slightly before end of
<emphasis id="C8DCCB7EFFBAFFB9336392A55D00FA12" box="[592,620,1412,1435]" italics="true" pageId="4" pageNumber="559">Rs</emphasis>
,
<emphasis id="C8DCCB7EFFBAFFB9334992A55DCFFA15" box="[634,675,1412,1436]" italics="true" pageId="4" pageNumber="559">Sc2</emphasis>
close to
<emphasis id="C8DCCB7EFFBAFFB9323492A55C43FA15" box="[775,815,1412,1436]" italics="true" pageId="4" pageNumber="559">Sc1</emphasis>
tip. Supernumerary cross-vein in cell
<emphasis id="C8DCCB7EFFBAFFB935EF92A55B9FFA12" box="[1244,1267,1412,1435]" italics="true" pageId="4" pageNumber="559">sc</emphasis>
approximately at two thirds distance from humeral vein to base of
<emphasis id="C8DCCB7EFFBAFFB933D092895D93FA36" box="[739,767,1448,1471]" italics="true" pageId="4" pageNumber="559">Rs</emphasis>
. Radial sector slightly arched at base. Both branches of
<emphasis id="C8DCCB7EFFBAFFB934B392895AF0FA36" box="[1408,1436,1448,1471]" italics="true" pageId="4" pageNumber="559">Rs</emphasis>
slightly diverging towards wing margin. Vein
<emphasis id="C8DCCB7EFFBAFFB9339592EC5DA2FA6D" box="[678,718,1485,1508]" italics="true" pageId="4" pageNumber="559">r-m</emphasis>
well developed. Discal cell comparatively wide, slightly more than twice as long as wide. Free end of
<emphasis id="C8DCCB7EFFBAFFB9336E92D05DC8F981" box="[605,676,1520,1544]" italics="true" pageId="4" pageNumber="559">M1+2</emphasis>
slightly shorter than its basal part forming frontal margin of discal cell. Basal deflection of
<emphasis id="C8DCCB7EFFBAFFB9308491345E9BF9A4" box="[439,503,1557,1581]" italics="true" pageId="4" pageNumber="559">CuA1</emphasis>
exactly at branching point of
<emphasis id="C8DCCB7EFFBAFFB9326D91345C19F9A5" box="[862,885,1557,1580]" italics="true" pageId="4" pageNumber="559">M</emphasis>
. Both anal veins long,
<emphasis id="C8DCCB7EFFBAFFB935BF91345BC7F9A4" box="[1164,1195,1557,1581]" italics="true" pageId="4" pageNumber="559">A1</emphasis>
slightly sinuous,
<emphasis id="C8DCCB7EFFBAFFB9344E91345AF0F9A4" box="[1405,1436,1557,1581]" italics="true" pageId="4" pageNumber="559">A2</emphasis>
slightly arched at distal end. Anal angle medium-sized, posterior margin widely rounded. Halter 0.81.0 mm long, grayish, base of stem pale. Coxae yellow, frontal face of fore coxae slightly infuscated. Frontal trochanter yellow with slightly darkened distal part, second and third pair grayish. Distal margin of all trochanters narrowly blackened. Femur obscure yellow, tibiae light brown, tarsus light brown at base turning dark brown towards distal end. Femur covered with short blackish adjacent setae, similar setae on tibiae and tarsus brownish. Femur I:
<quantity id="3D50BA89FFBAFFB9345891E85F9CF88D" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.9" metricValueMin="4.4" pageId="4" pageNumber="559" unit="mm" value="4.65" valueMax="4.9" valueMin="4.4">4.4 4.9 mm</quantity>
, II: 5.0
<quantity id="3D50BA89FFBAFFB9306291CD5EC5F88D" box="[337,425,1772,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="559" unit="mm" value="5.3">5.3 mm</quantity>
, III: 5.0
<quantity id="3D50BA89FFBAFFB9332191CD5D00F88D" box="[530,620,1772,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="559" unit="mm" value="5.6">5.6 mm</quantity>
; tibiae I:
<quantity id="3D50BA89FFBAFFB933E591CD5C0CF88D" box="[726,864,1772,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.699999999999999" metricValueMax="5.8" metricValueMin="5.6" pageId="4" pageNumber="559" unit="mm" value="5.699999999999999" valueMax="5.8" valueMin="5.6">5.65.8 mm</quantity>
, II:
<quantity id="3D50BA89FFBAFFB932BE91CD5B7BF88D" box="[909,1047,1772,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="5.8" metricValueMin="5.7" pageId="4" pageNumber="559" unit="mm" value="5.75" valueMax="5.8" valueMin="5.7">5.75.8 mm</quantity>
, III:
<quantity id="3D50BA89FFBAFFB9357C91CD5BB7F88D" box="[1103,1243,1772,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="5.9" metricValueMin="5.8" pageId="4" pageNumber="559" unit="mm" value="5.85" valueMax="5.9" valueMin="5.8">5.85.9 mm</quantity>
; tarsus I:
<quantity id="3D50BA89FFBAFFB9347B91CD5FA9F8A1" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="4" pageNumber="559" unit="mm" value="4.8" valueMax="5.1" valueMin="4.5">4.55.1 mm</quantity>
, II:
<quantity id="3D50BA89FFBAFFB931C790305E22F8A1" box="[244,334,1809,1833]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="559" unit="mm" value="3.9">3.9 mm</quantity>
, III:
<quantity id="3D50BA89FFBAFFB930BB90305D79F8A1" box="[392,533,1809,1833]" metricMagnitude="-3" metricUnit="m" metricValue="3.9499999999999997" metricValueMax="4.1" metricValueMin="3.8" pageId="4" pageNumber="559" unit="mm" value="3.9499999999999997" valueMax="4.1" valueMin="3.8">3.84.1 mm</quantity>
. Claw blackish with small subbasal spine.
<emphasis id="C8DCCB7EFFBAFFB9353290315B07F8A0" box="[1025,1131,1808,1833]" italics="true" pageId="4" pageNumber="559">Abdomen</emphasis>
: tergites blackish, covered with erect golden setae. Basal sternite pale, remaining sternites brownish. Male terminalia (
<figureCitation id="62930BE9FFBAFFB9358390155A72F8C4" box="[1200,1310,1844,1869]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[188,1348,213,1877]" captionTargetId="figure@5.[173,1390,193,1898]" captionTargetPageId="5" captionText="FIGURES 7 11. Geranomyia piagola n. sp. 7: Wing; 8: Male genitalia, dorsal view; 9: Male genitalia, ventral view; 10: Ovipositor, lateral view; 11: Paramere of male genitalia. Abbreviations: cerc, cercus; hyp vlv, hypogynial valve; st 8, eight sternite; tg 9, ninth tergite; tg 10, tenth tergite." httpUri="https://zenodo.org/record/258606/files/figure.png" pageId="4" pageNumber="559">Figs. 8, 9</figureCitation>
) with dark brown ninth tergite, gonocoxite and inner gonostylus light brown. Posterior margin of ninth tergite very shallowly emarginate, lateral parts slightly raised and covered with dense short brown setae. Gonocoxite elongate, bearing large ventro-median lobe (
<figureCitation id="62930BE9FFBAFFB930FB90815D7CF830" box="[456,528,1952,1977]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[188,1348,213,1877]" captionTargetId="figure@5.[173,1390,193,1898]" captionTargetPageId="5" captionText="FIGURES 7 11. Geranomyia piagola n. sp. 7: Wing; 8: Male genitalia, dorsal view; 9: Male genitalia, ventral view; 10: Ovipositor, lateral view; 11: Paramere of male genitalia. Abbreviations: cerc, cercus; hyp vlv, hypogynial valve; st 8, eight sternite; tg 9, ninth tergite; tg 10, tenth tergite." httpUri="https://zenodo.org/record/258606/files/figure.png" pageId="4" pageNumber="559">Fig. 9</figureCitation>
). Distal part of lobe rounded, covered with long erect setae. Outer gonostylus strongly sclerotized, darkened, hook-shaped with sharply pointed distal end. Inner gonostylus very
</paragraph>
<caption id="AED747E4FFBBFFB831A490B35D05F86F" httpUri="https://zenodo.org/record/258606/files/figure.png" pageId="5" pageNumber="560" targetBox="[188,1348,213,1877]" targetPageId="5">
<paragraph id="FA17176CFFBBFFB831A490B35D05F86F" blockId="5.[151,1436,1938,2022]" pageId="5" pageNumber="560">
<emphasis id="C8DCCB7EFFBBFFB831A490B35E25F821" bold="true" box="[151,329,1938,1961]" pageId="5" pageNumber="560">FIGURES 711.</emphasis>
<taxonomicName id="3DA86CEFFFBBFFB8306190B55D4BF821" box="[338,551,1939,1961]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="560" phylum="Arthropoda" rank="species" species="piagola" status="sp. nov.">
<emphasis id="C8DCCB7EFFBBFFB8306190B55D4BF821" box="[338,551,1939,1961]" italics="true" pageId="5" pageNumber="560">Geranomyia piagola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3EF7605FFBBFFB8330290B25D02F821" box="[561,622,1939,1960]" pageId="5" pageNumber="560" rank="species">
<emphasis id="C8DCCB7EFFBBFFB8330290B25D02F821" bold="true" box="[561,622,1939,1960]" pageId="5" pageNumber="560">n. sp.</emphasis>
</taxonomicNameLabel>
7: Wing; 8: Male genitalia, dorsal view; 9: Male genitalia, ventral view; 10: Ovipositor, lateral view; 11: Paramere of male genitalia. Abbreviations: cerc, cercus; hyp vlv, hypogynial valve; st 8, eight sternite; tg 9, ninth tergite; tg 10, tenth tergite.
</paragraph>
</caption>
<paragraph id="FA17176CFFB8FFBB31A497B65E4CFEEC" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="561">
elongate, about three times as long as gonocoxite, oval, with fronto-median angle extended into small rostral prolongation. Rostral prolongation directed towards frontal end with small subapical tubercle covered with small brown setae. Tubercle armed with two-three short straight brown spines, situated closely together along entire length. Right gonostylus of
<typeStatus id="2513A9CEFFB8FFBB30E696255D5BFE94" box="[469,567,260,285]" pageId="6" pageNumber="561" type="holotype">holotype</typeStatus>
with two, left with three spines. Paramere (
<figureCitation id="62930BE9FFB8FFBB351F96255B11FE95" box="[1068,1149,260,285]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[188,1348,213,1877]" captionTargetId="figure@5.[173,1390,193,1898]" captionTargetPageId="5" captionText="FIGURES 7 11. Geranomyia piagola n. sp. 7: Wing; 8: Male genitalia, dorsal view; 9: Male genitalia, ventral view; 10: Ovipositor, lateral view; 11: Paramere of male genitalia. Abbreviations: cerc, cercus; hyp vlv, hypogynial valve; st 8, eight sternite; tg 9, ninth tergite; tg 10, tenth tergite." httpUri="https://zenodo.org/record/258606/files/figure.png" pageId="6" pageNumber="561">Fig. 11</figureCitation>
) wide at base, distal part extended into narrow lobe. Penis long, narrow in dorsal view, “S“-shaped in lateral view, with tip pointed downwards.
</paragraph>
<paragraph id="FA17176CFFB8FFBB31F4964E5D31FE7C" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="561">
Female (N=1). Generally similar to male. Body length
<quantity id="3D50BA89FFB8FFBB321E964E5CEAFE0E" box="[813,902,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="6" pageNumber="561" unit="mm" value="5.8">5.8 mm</quantity>
. Wing length
<quantity id="3D50BA89FFB8FFBB351296515B15FE0E" box="[1057,1145,368,392]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="6" pageNumber="561" unit="mm" value="7.9">7.9 mm</quantity>
. Ovipositor (
<figureCitation id="62930BE9FFB8FFBB3439964E5A37FE01" box="[1290,1371,367,392]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1938,1961]" captionTargetBox="[188,1348,213,1877]" captionTargetId="figure@5.[173,1390,193,1898]" captionTargetPageId="5" captionText="FIGURES 7 11. Geranomyia piagola n. sp. 7: Wing; 8: Male genitalia, dorsal view; 9: Male genitalia, ventral view; 10: Ovipositor, lateral view; 11: Paramere of male genitalia. Abbreviations: cerc, cercus; hyp vlv, hypogynial valve; st 8, eight sternite; tg 9, ninth tergite; tg 10, tenth tergite." httpUri="https://zenodo.org/record/258606/files/figure.png" pageId="6" pageNumber="561">Fig. 10</figureCitation>
) with cercus and hypovalva brown. Cercus nearly parallel-sided, straight, tip rounded. Hypovalva wide, comparatively short and blunt-apexed, reaching to about middle of cercus. Vaginal apodeme very extended, distal part very narrow, apex nearly reaches tip of valve.
</paragraph>
<paragraph id="FA17176CFFB8FFBB31F496DE5A0CFD91" blockId="6.[151,1437,151,1005]" box="[199,1376,511,536]" pageId="6" pageNumber="561">
<emphasis id="C8DCCB7EFFB8FFBB31F496DE5E46FD91" bold="true" box="[199,298,511,536]" pageId="6" pageNumber="561">Habitat.</emphasis>
Margins of medium-sized mountainous stream with springs on slopes, covered by mixed forest.
</paragraph>
<paragraph id="FA17176CFFB8FFBB31F495055CE4FDB5" blockId="6.[151,1437,151,1005]" box="[199,904,548,573]" pageId="6" pageNumber="561">
<emphasis id="C8DCCB7EFFB8FFBB31F495055E2CFDB4" bold="true" box="[199,320,548,573]" pageId="6" pageNumber="561">Elevation.</emphasis>
Species was found at altitudes of about
<quantity id="3D50BA89FFB8FFBB323195055CE8FDB5" box="[770,900,548,573]" metricMagnitude="2" metricUnit="m" metricValue="4.4" metricValueMax="5.0" metricValueMin="3.8" pageId="6" pageNumber="561" unit="m" value="440.0" valueMax="500.0" valueMin="380.0">
380
<specimenCount id="ECAEDCE5FFB8FFBB320995055CE8FDB5" box="[826,900,548,573]" pageId="6" pageNumber="561" type="generic">500 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="FA17176CFFB8FFBB31F495665BC9FDE9" blockId="6.[151,1437,151,1005]" box="[199,1189,583,608]" pageId="6" pageNumber="561">
<emphasis id="C8DCCB7EFFB8FFBB31F495665EF6FDE9" bold="true" box="[199,410,583,608]" pageId="6" pageNumber="561">Period of activity.</emphasis>
Adults are flying from the middle of May through the end of August.
</paragraph>
</subSubSection>
<subSubSection id="B2B244E7FFB8FFBB31F4954D5D9FFD0D" box="[199,755,620,645]" pageId="6" pageNumber="561" type="distribution">
<paragraph id="FA17176CFFB8FFBB31F4954D5D9FFD0D" blockId="6.[151,1437,151,1005]" box="[199,755,620,645]" pageId="6" pageNumber="561">
<emphasis id="C8DCCB7EFFB8FFBB31F4954D5E35FD0C" bold="true" box="[199,345,620,645]" pageId="6" pageNumber="561">Distribution</emphasis>
. Jirisan National Park,
<collectingCountry id="82BF57FCFFB8FFBB3353954D5D83FD0D" box="[608,751,620,645]" name="South Korea" pageId="6" pageNumber="561">South Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B244E7FFB8FFBB31F495AE5D59FC64" pageId="6" pageNumber="561" type="discussion">
<paragraph id="FA17176CFFB8FFBB31F495AE5D59FC64" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="561">
<emphasis id="C8DCCB7EFFB8FFBB31F495AE5E20FD21" bold="true" box="[199,332,655,680]" pageId="6" pageNumber="561">Discussion.</emphasis>
<taxonomicName id="3DA86CEFFFB8FFBB306195B15D56FD21" box="[338,570,655,680]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="piagola" status="sp. nov.">
<emphasis id="C8DCCB7EFFB8FFBB306195B15D56FD21" box="[338,570,655,680]" italics="true" pageId="6" pageNumber="561">Geranomyia piagola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3EF7605FFB8FFBB337295AE5D13FD21" box="[577,639,655,680]" pageId="6" pageNumber="561" rank="species">
<emphasis id="C8DCCB7EFFB8FFBB337295AE5D13FD21" bold="true" box="[577,639,655,680]" pageId="6" pageNumber="561">n. sp.</emphasis>
</taxonomicNameLabel>
has very distinct male genitalia, especially an extremely elongate inner gonostylus. In general, the male genitalia of
<taxonomicName id="3DA86CEFFFB8FFBB33A195945C6BFD44" box="[658,775,692,717]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="piagola" status="sp. nov.">
<emphasis id="C8DCCB7EFFB8FFBB33A195945C6BFD44" box="[658,775,692,717]" italics="true" pageId="6" pageNumber="561">G. piagola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3EF7605FFB8FFBB322395955C3DFD44" box="[784,849,692,717]" pageId="6" pageNumber="561" rank="species">
<emphasis id="C8DCCB7EFFB8FFBB322395955C3DFD44" bold="true" box="[784,849,692,717]" pageId="6" pageNumber="561">n. sp.</emphasis>
</taxonomicNameLabel>
resembles that of the Chinese species
<taxonomicName id="3DA86CEFFFB8FFBB343F95945E05FD79" authority="Alexander, 1940" authorityName="Alexander" authorityYear="1940" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="obesistyla">
<emphasis id="C8DCCB7EFFB8FFBB343F95945AF0FD44" box="[1292,1436,692,717]" italics="true" pageId="6" pageNumber="561">G. obesistyla</emphasis>
(
<bibRefCitation id="9E396A9DFFB8FFBB31AD95F65E0DFD79" author="Alexander" box="[158,353,727,752]" pageId="6" pageNumber="561" refString="Alexander, C. P. (1940) New or little-known Tipulidae from eastern Asia (Diptera). XLII. Philippine Journal of Science, 71, 169 - 204." type="journal article" year="1940">Alexander, 1940</bibRefCitation>
)
</taxonomicName>
. Genitalia of both species differ only in the small details of the rostral prolongation of inner gonostylus, the shape of ventro-median lobe of gonocoxite, the shape of penis and the shape of the paramere. Both species could be easily separated on wing venation, wing pattern and body coloration.
<taxonomicName id="3DA86CEFFFB8FFBB35A394015A64FCB1" box="[1168,1288,799,824]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="piagola" status="sp. nov.">
<emphasis id="C8DCCB7EFFB8FFBB35A394015A64FCB1" box="[1168,1288,799,824]" italics="true" pageId="6" pageNumber="561">G. piagola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3EF7605FFB8FFBB3421943E5A38FCB1" box="[1298,1364,799,824]" pageId="6" pageNumber="561" rank="species">
<emphasis id="C8DCCB7EFFB8FFBB3421943E5A38FCB1" bold="true" box="[1298,1364,799,824]" pageId="6" pageNumber="561">n. sp.</emphasis>
</taxonomicNameLabel>
has a distinct longitudinal dark stripe along the pleura, the pleura of
<taxonomicName id="3DA86CEFFFB8FFBB32AA94645B42FCD4" box="[921,1070,836,861]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="obesistyla">
<emphasis id="C8DCCB7EFFB8FFBB32AA94645CC6FCD5" box="[921,938,837,860]" italics="true" pageId="6" pageNumber="561">G</emphasis>
.
<emphasis id="C8DCCB7EFFB8FFBB328D94655B42FCD4" box="[958,1070,836,861]" italics="true" pageId="6" pageNumber="561">obesistyla</emphasis>
</taxonomicName>
is almost uniformly brownish testaceous. The wing of
<taxonomicName id="3DA86CEFFFB8FFBB309594485D58FC08" box="[422,564,872,897]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="obesistyla">
<emphasis id="C8DCCB7EFFB8FFBB309594485D58FC08" box="[422,564,872,897]" italics="true" pageId="6" pageNumber="561">G. obesistyla</emphasis>
</taxonomicName>
has distinct brown spots on the frontal margin, the basal deflection of
<emphasis id="C8DCCB7EFFB8FFBB347094495AEFFC09" box="[1347,1411,872,896]" italics="true" pageId="6" pageNumber="561">CuA1</emphasis>
is situated well before the branching point of
<emphasis id="C8DCCB7EFFB8FFBB334894AC5DFEFC2D" box="[635,658,909,932]" italics="true" pageId="6" pageNumber="561">M</emphasis>
. The wing venation and pattern of
<taxonomicName id="3DA86CEFFFB8FFBB352F94AC5BFDFC2C" box="[1052,1169,908,933]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="piagola" status="sp. nov.">
<emphasis id="C8DCCB7EFFB8FFBB352F94AC5BFDFC2C" box="[1052,1169,908,933]" italics="true" pageId="6" pageNumber="561">G. piagola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D3EF7605FFB8FFBB35AB94AD5BBBFC2C" box="[1176,1239,908,933]" pageId="6" pageNumber="561" rank="species">
<emphasis id="C8DCCB7EFFB8FFBB35AB94AD5BBBFC2C" bold="true" box="[1176,1239,908,933]" pageId="6" pageNumber="561">n. sp.</emphasis>
</taxonomicNameLabel>
is very similar to that of other Korean species
<taxonomicName id="3DA86CEFFFB8FFBB30C794905D10FC40" box="[500,636,945,969]" class="Insecta" family="Limoniidae" genus="Geranomyia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="561" phylum="Arthropoda" rank="species" species="gifuensis">
<emphasis id="C8DCCB7EFFB8FFBB30C794905D69FC41" box="[500,517,945,968]" italics="true" pageId="6" pageNumber="561">G</emphasis>
.
<emphasis id="C8DCCB7EFFB8FFBB332B94905D10FC40" box="[536,636,945,969]" italics="true" pageId="6" pageNumber="561">gifuensis</emphasis>
</taxonomicName>
, but both species are easily separated based on the male genitalia, especially shape of inner gonostylus.
</paragraph>
</subSubSection>
</treatment>
</document>