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<document id="CDCF38174BC3AE96E5E8D72CEA0F7F5F" ID-DOI="10.3897/zookeys.1202.121633" ID-ZooBank="F2DE6CAC-842E-4275-B3C1-C84D7CEDF5B7" ID-publisher-id="121633" URI-arpha="CB9CB628-9D1E-53A4-84B5-3B9C013566B4" URI-zoobank="http://zoobank.org/F2DE6CAC-842E-4275-B3C1-C84D7CEDF5B7" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1716826741008" checkinUser="pensoft" docAuthor="Zhang, Ludan, Wu, Zheng, Li, Shuqiang &amp; Yao, Zhiyuan" docDate="2024" docId="715AB4B4D68E591BBC30B7284CB2823C" docLanguage="en" docName="ZooKeys 1202: 255-286" docOrigin="ZooKeys 1202" docSource="https://zookeys.pensoft.net/article/121633/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Belisana tengchong Zhang, Li &amp; Yao 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="286" masterDocId="CB9CB6289D1E53A484B53B9C013566B4" masterDocTitle="Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China" masterLastPageNumber="286" masterPageNumber="255" pageNumber="255" updateTime="1716918634710" updateUser="admin">
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<mods:title id="8C3514CDBA69821A815F29A40393330B">Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China</mods:title>
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<mods:name id="A7CF138BADE169CE67FFA27E1C0DA3C3" type="personal">
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<mods:namePart id="B157BAE84843D0E8A454FB310AC64445">Zhang, Ludan</mods:namePart>
<mods:nameIdentifier id="A6DD43EEB700704E672AD46060E96887" type="ORCID">0000-0001-5657-751X</mods:nameIdentifier>
<mods:affiliation id="7948CD40FAFCFB7579F1C81E9D3BC31E">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="B86F3ECEF70622FB7919050578A22ED2">Wu, Zheng</mods:namePart>
<mods:affiliation id="7CEB4735F735EB47BF2572835F9CDDA4">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
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<mods:namePart id="44944713BDA8F3A34581E873A5EA3931">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="7FE881F4DEAD72BEAE57F64A86F0D559" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="9AAAF6373B7EB6C859D18B1C113C7070">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
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<mods:namePart id="A513835B1C7B07BA8C2330D43CEA05AC">Yao, Zhiyuan</mods:namePart>
<mods:nameIdentifier id="11D464564382CA33E9D0D7B5CDC9E3DA" type="ORCID">0000-0002-1631-0949</mods:nameIdentifier>
<mods:affiliation id="A9485C6FD4E865FFA4A977C815560484">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:typeOfResource id="2AD95E51C9073B3D07EA951D733EC9D0">text</mods:typeOfResource>
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<mods:title id="0E538BBEC47CCE9629389F8ED6AFD0F3">ZooKeys</mods:title>
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<mods:part id="30CBED6A35648A69BE3E05820A7D9BD6">
<mods:date id="EE98526798D370C1016DE6BF52262D3F">2024</mods:date>
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<mods:number id="6C2D62078409D258A281F47FA10D9EE0">2024-05-27</mods:number>
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<mods:number id="4263EB0DA3154B0702B7BC2D75810CA3">1202</mods:number>
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<mods:identifier id="900D84DA65A5BC97E69D5381B8482CB6" type="ZooBank">F2DE6CAC-842E-4275-B3C1-C84D7CEDF5B7</mods:identifier>
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<treatment id="715AB4B4D68E591BBC30B7284CB2823C" ID-DOI="http://doi.org/10.5281/zenodo.11356052" ID-Zenodo-Dep="11356052" ID-arpha="715AB4B4-D68E-591B-BC30-B7284CB2823C" ID-zoobank="https://zoobank.org/D21705D9-7D42-4F9A-A4E9-27720466DF1D" LSID="urn:lsid:plazi:treatment:715AB4B4D68E591BBC30B7284CB2823C" LSID-ZBK="urn:lsid:zoobank.org:act:D21705D9-7D42-4F9A-A4E9-27720466DF1D" httpUri="http://treatment.plazi.org/id/715AB4B4D68E591BBC30B7284CB2823C">
<subSubSection id="F8CDAF8FF2CAB8F5C2487D4462F6252F" type="nomenclature">
<paragraph id="1803EBF5A4F8C08FFABD608D9783CCC2">
<taxonomicName id="E9C76E9B808065A7524670DF61156459" authority="Zhang, Li &amp; Yao" authorityName="Zhang, Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tengchong" status="sp. nov.">
<emphasis id="ED14961E82818C173234AF9A00731125" italics="true">Belisana tengchong</emphasis>
Zhang, Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="2FAB9CFCF0D3D52FABC287EAD334BB3A" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FA24E80C8571F5160A7354A788F95770" type="description">
<paragraph id="827CAEC22FC3AE6823D800FECB099D78">
<figureCitation id="55208BB3E84F52961C873E5EB747958B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">Figs 10</figureCitation>
,
<figureCitation id="880C58D581395C6BDB7A5D8847877966" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11</figureCitation>
,
<figureCitation id="A75BF44A20F5AEC2F7626C3FDDC2DB33" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 I, J</figureCitation>
,
<figureCitation id="2B029E98600F803823AF113457B9BF7D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGRBG" type="type material">
<paragraph id="C096992012815160F018067B2EAB6E3A">
<heading id="2308F579A6B8523E61AF0DFB2D4C1AE1" reason="title">Type material.</heading>
</paragraph>
<paragraph id="11EFE50667F0BCC2643749A750E3EC4B">
<materialsCitation id="6052AEE2B8066CF41AD74F31BB50395D" collectingDate="2010-08-18" collectionCode="IZCAS" collectorName="C Wang &amp; Q Zhao &amp; L Lin" country="China" county="Tengchong County" elevation="1802" latitude="25.340784" location="Kongming Cave" longLatPrecision="1" longitude="98.538284" municipality="Baoshan" specimenCode="Ar 44969, Ar 44970-71, Ar 44972-74" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="BBB4DDAB0F6640908582CEC3D11563F7" bold="true" italics="true">
<typeStatus id="2E7D5653D3295D914139A4AF2E6E20FA" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="51083B3F04C3CF50B1AE626170559481" count="1" type="male">♂</specimenCount>
(
<named-content id="C79329961EDED7D4D79BAD23C6B758D6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="BA5649DC230FB47863452D3C22577097" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="22868968CDE12BFE521C78627A6D8EBA">Ar 44969</specimenCode>
) and
<emphasis id="E9439905C52BB2911E8C7C1A4AA98D6C" bold="true" italics="true">
<typeStatus id="EE165C0EB27012767FADB959D17D74B2" type="paratype">paratypes</typeStatus>
</emphasis>
<specimenCount id="058C48A215BBED00B844BF50F8196A17" count="2" type="male">2 ♂</specimenCount>
(
<named-content id="89E0C4B1ADDBA06DEB605DC43213EBE6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="BEAEE9C9DFD60347782B79E34C042099" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="C1C39618C1F2B60D63EF8DE86E37837C">Ar 44970–71</specimenCode>
)
<specimenCount id="0122017474C398F051C57DE29D297308" count="3" type="female">3 ♀</specimenCount>
(
<named-content id="58FA252E6E5C40D37D9611EF5B633082" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="4D5A18A1C97E7E0B53D8FF1C25CF8878" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="9C9884D92EBCA7FC4F9674A4888DF97C">Ar 44972–74</specimenCode>
),
<location id="AB8A1D1D7D246AB961234F8B579C04DB" LSID="urn:lsid:plazi:treatment:715AB4B4D68E591BBC30B7284CB2823C:AB8A1D1D7D246AB961234F8B579C04DB" country="China" county="Tengchong County" latitude="25.340784" longLatPrecision="1" longitude="98.538284" municipality="Baoshan" name="Kongming Cave" stateProvince="Yunnan">Kongming Cave</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDSBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[98.538283,25.340783]}">
<geoCoordinate id="DE0BCDF0E0EC1080C15DE9CD77CFD7C6" degrees="25" direction="north" minutes="20.447" orientation="latitude" precision="1" value="25.340784">25 ° 20.447 ' N</geoCoordinate>
,
<geoCoordinate id="04521530B06E5ADD659F9893CE3BBDC2" degrees="98" direction="east" minutes="32.297" orientation="longitude" precision="1" value="98.538284">98 ° 32.297 ' E</geoCoordinate>
</named-content>
,
<quantity id="44600B2B0460824BA6D657F834D9730D" metricMagnitude="3" metricUnit="m" metricValue="1.802" unit="m" value="1802.0">
<elevation id="D75F084446DCE7A940E29551BE76DE66" metricMagnitude="3" metricUnit="m" metricValue="1.802" unit="m" value="1802.0">1802 m</elevation>
</quantity>
),
<location id="11C4E83397B5C34FB50A8BA5A58E7F04" LSID="urn:lsid:plazi:treatment:715AB4B4D68E591BBC30B7284CB2823C:11C4E83397B5C34FB50A8BA5A58E7F04" country="China" county="Tengchong County" latitude="25.340784" longLatPrecision="1" longitude="98.538284" municipality="Baoshan" name="Jiangdong Village" stateProvince="Yunnan">Jiangdong Village</location>
,
<location id="41C1328A8A9F639D656E86BB06E37312" LSID="urn:lsid:plazi:treatment:715AB4B4D68E591BBC30B7284CB2823C:41C1328A8A9F639D656E86BB06E37312" country="China" county="Tengchong County" latitude="25.340784" longLatPrecision="1" longitude="98.538284" municipality="Baoshan" name="Gudong Town" stateProvince="Yunnan">Gudong Town</location>
,
<collectingCounty id="8755D135148FE9AB2693A7FADF4C0D0D">Tengchong County</collectingCounty>
,
<collectingMunicipality id="FE364060ECB4EABDEAA1214AED0842C9">Baoshan</collectingMunicipality>
,
<emphasis id="717F427D541AEF2DB1644C8989D0EA33" bold="true">
<collectingRegion id="CEF386BFD8756ED6F7CA2AF1B649614F" country="China" name="Yunnan">Yunnan</collectingRegion>
</emphasis>
,
<emphasis id="47FB4EEF3AA8F1ADD44A5EBE7C51F170" bold="true">
<collectingCountry id="B00343D5B38739398C98FC2A12E15879" name="China">China</collectingCountry>
</emphasis>
,
<date id="E673C505816A4C0826299B1313959B17" value="2010-08-18">
<collectingDate id="D0C4522F7E92DE1B2BF35082B8F7277D" value="2010-08-18">18 / 08 / 2010</collectingDate>
</date>
,
<collectorName id="5D2D90C654B3EA0F863DFF6F536E0BBA">C Wang</collectorName>
,
<collectorName id="AB7D773EA6F3E8CF61C9F8707CC19011">Q Zhao</collectorName>
&amp;
<collectorName id="10E0DF2DC60EDF1DAF756E939204D30E">L Lin</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMSBG" type="etymology">
<paragraph id="F38568D40CC3E923551CA90905CBB2F2">
<heading id="C621EBD60CB6067A149B9711E83245C6" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="0680AF2B2FEF10CE9A9198EC64110D9A">
The specific name refers to the
<typeStatus id="361B2975E6738803C7AC1AD09806BCEB">type</typeStatus>
locality, which is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERSBG" type="diagnosis">
<paragraph id="8BE6E10FCB679D7C5F8BE496CC02E6A1">
<heading id="2C49C81739FB658DBA02A67F61A70F39" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="A22C73E5BA8BE5C1DCF51C2FFC72310C">
The new species resembles
<taxonomicName id="A3795A304B98D67B8C4B61EAA1679299" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="pianma">
<emphasis id="F039A7A4AB364519D1A0B6E17087D141" italics="true">B. pianma</emphasis>
<bibRefCitation id="7D3484E7836D1E003F9D08ADB300B3EF" author="Huber BA" journalOrPublisher="Zoologica" pagination="1-126" refId="B1" refString="Huber BA (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: The case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica 155: 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: The case of Belisana Thorell 1898 (Araneae, Pholcidae)." volume="155" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="BB366828E56688A780F5CB9DC6DFD041" author="Huber BA" firstAuthor="Huber" journalOrPublisher="Zoologica" page="14" pagination="1-126" refId="B1" refString="Huber BA (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: The case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica 155: 1–126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: The case of Belisana Thorell 1898 (Araneae, Pholcidae)." volume="155" year="2005">Huber 2005: 14</bibRefCitation>
, figs 5, 6, 56, 88–92, 112–130) by having similar bulbal apophyses and epigyne (Figs
<figureCitation id="A996ECC97052F75D96B8BC7DD01C006D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 A, C</figureCitation>
,
<figureCitation id="D3C04CE645EBD4FC4B04E01475148367" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 A</figureCitation>
), but can be distinguished by differences in males: clypeus unmodified (Fig.
<figureCitation id="1B0F1F657B6217040DB3820B19EE28D5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 E</figureCitation>
vs with a pair of front apophyses), cheliceral distal apophyses on median part of chelicerae (Fig.
<figureCitation id="4A8D524527F838EA1066492A82018564" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 D</figureCitation>
vs distal part), procursus without distal spine and dorso-distal sclerite (Figs
<figureCitation id="7C693FD8E5F8F64EDB0244AFE4BCEC6F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">10 C</figureCitation>
,
<figureCitation id="17CC8EDDBE9FDA430EB9B84D89B47F73" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 I</figureCitation>
vs present), and with angular distal membranous process (arrow
<quantity id="748AD7334D2DFA41AFEF7777A939D8F1" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="87B3BB8656939661FB7442A00BD116BF" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">10 C</figureCitation>
,
<figureCitation id="C6A1EB4D53ECF361C4ACB1E1F65AF217" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 I</figureCitation>
vs absent); differences in females: pore plates wide and curved (2 × longer than wide, Figs
<figureCitation id="D86C49999C303ED22A23FA0D00732F49" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 B</figureCitation>
,
<figureCitation id="17EB91BA15DBAB2355FE23D88313588E" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 B</figureCitation>
vs narrow, 6 × longer than wide).
</paragraph>
<caption id="6E89E7524CA3CEEFFFB73FE44643ACAB" ID-DOI="10.3897/zookeys.1202.121633.figure10" ID-arpha="BCF2B87E-D255-5FE3-89CF-524EBD33E703" httpUri="https://binary.pensoft.net/fig/1055875" startId="F10">
<paragraph id="8D1B4B91D10C49558E775A1CF45AE626">
<label id="70EB06179272F3419203A59B0E4D8328">Figure 10.</label>
</paragraph>
<paragraph id="6CB7A6EC92D39CEB9E15F4EB3BB6739F">
<taxonomicName id="9F30458AE474268233FAF51DB1CE89EC" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tengchong" status="sp. nov.">
<emphasis id="4040BF433AEBE0A66F1F7C38527295BE" italics="true">Belisana tengchong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F15840D615CB9685338AFC4BCAF3DBAF" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="68C8ADBCC1880B57A16B076698A6AED4" bold="true">A, B</emphasis>
palp (
<emphasis id="7ADBF404322706C2D186B02342E0FF82" bold="true">A</emphasis>
prolateral view
<emphasis id="30741ABB8BF46CB0BE17F63258FF1B6C" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="40641B573045C6408C85D7C8EBB8A8D3" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="5DEC359FB3898F75E126B4658090959F" bold="true">C</emphasis>
prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process
<emphasis id="8185B3CFBAEC971D1A0A34FFBD804C27" bold="true">D</emphasis>
retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation:
<abbrev id="ABBRID0ENVBG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="53F8B7A8A5957C5356882888489252FD" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="80DDA1DE2EB2FA6B77861E0ACDD676F9" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="165954EC3DFB45477CA4DF93F213019F" ID-DOI="10.3897/zookeys.1202.121633.figure11" ID-arpha="087C27FE-48CE-5108-803A-55D42E7A9D81" httpUri="https://binary.pensoft.net/fig/1055876" startId="F11">
<paragraph id="F5D5F61D220EB0246FD97638138583B5">
<label id="C5BB6B963385DD6107DDC44F7E85D667">Figure 11.</label>
</paragraph>
<paragraph id="EDD97E5D5F2FC07488DA3D39E442E6F0">
<taxonomicName id="F8C59D2FEB79862AD371CEDC93B4C4DD" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tengchong" status="sp. nov.">
<emphasis id="613A6953F932D40819EEC6F6FFE884EB" italics="true">Belisana tengchong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A007EA0FD56894BEDBB844D0EB425209" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="BC028870F15038624D4D1538FCD96815" bold="true">C – F</emphasis>
) and paratype female (
<emphasis id="9317382E26C39A1D4922721A5BF75032" bold="true">A, B, G, H</emphasis>
)
<emphasis id="D93469B66D34E5E22BDD90DA1BAA92E9" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="27FA0E1B4B5442561FAA9EF0D5F1928D" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="2C2453EBED1A14104D029FFF5ACB8802" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="C476E68577748FEA42D6E51D85976822" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="C15431C72090DDC14B10364E4A16B23E" bold="true">E – H</emphasis>
habitus (
<emphasis id="291EA2DF8D3769B3805845DC55C69FCB" bold="true">E, G</emphasis>
dorsal view
<emphasis id="7FE4501C71352920FFFF73809E73F190" bold="true">F</emphasis>
lateral view
<emphasis id="776AACC1E88296E7F02010681B64C716" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EBXBG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EFXBG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EJXBG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0ENXBG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0ERXBG" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0EVXBG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.05 (
<emphasis id="4FA1E84779172ED361A82E1D01F57FE5" bold="true">A, B</emphasis>
); 0.10 (
<emphasis id="CBB2ED1AB91E3AE6C6BBA02A2237CE88" bold="true">C, D</emphasis>
); 0.30 (
<emphasis id="3BC80DBEA11EFA7834A6C2D115D64172" bold="true">E – H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EEYBG" type="description">
<paragraph id="D9315A59F13AB40DD08F8C7AD2E252A5">
<heading id="1DD034FDF56817A8E5CF720B2F99EA96" reason="title">Description.</heading>
</paragraph>
<paragraph id="D40C5D4344AAD77E574DA0002D2484F5">
<emphasis id="BA68EAB7B7676762A69931BC3852C44F" bold="true">Male</emphasis>
(
<emphasis id="84FA2F3F0A47A2E5E32EA2354E53162C" bold="true" italics="true">
<typeStatus id="5DF3CF6D1B9371F62DDCFF8878FEE9A6" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.95 (2.08 with clypeus), carapace 0.71 long, 0.85 wide, opisthosoma 1.24 long, 0.93 wide. Leg I: 18.34 (4.75, 0.32, 4.90, 6.67, 1.70), leg II: 12.82 (3.56, 0.26, 3.40, 4.55, 1.05), legs III and IV missing; tibia I
<abbrev id="ABBRID0EPYBG" xlink_title="length/diameter">L / d</abbrev>
: 65. Eye inter-distances and diameters:
<abbrev id="ABBRID0ETYBG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EXYBG" xlink_title="posterior median eye">PME</abbrev>
0.14,
<abbrev id="ABBRID0E2YBG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0E6YBG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EDZBG" xlink_title="anterior lateral eye">ALE</abbrev>
0.03,
<abbrev id="ABBRID0EHZBG" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum width / length: 0.56 / 0.50. Habitus as in Fig.
<figureCitation id="D52C0DE3442A44E8248C6EFE7AB65D77" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 E, F</figureCitation>
. Carapace and sternum yellowish, without marks. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae (Fig.
<figureCitation id="A7D7D689C492024DAB627FDB26B34B92" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 D</figureCitation>
) with a pair of proximo-lateral apophyses and a pair of curved distal apophyses (distance between tips: 0.28). Palp as in Fig.
<figureCitation id="42F99E3CACE0C39B93EB1A4EBDA261D7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">10 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="D414AB73B13D1B214E444AB4C3E8A645" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="C5D48DCB6A532C62A3E74F8294958014" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">10 B</figureCitation>
); femur with tiny retrolatero-proximal protrusion (arrow
<quantity id="7E945CC397A33BDB42600B0C656D955C" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="E7D1F520409EF83665991B3707FDC491" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">10 B</figureCitation>
); procursus simple proximally but complex distally, with curved, sclerotized prolatero-distal apophysis (arrow
<quantity id="B8BFFB61F5C83045D4C75721FC6259EB" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="ECF4BE2667B139261B17BA269529E42F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">10 C</figureCitation>
,
<figureCitation id="AA67A39A7BFC83B48D1AAB6775159E8A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 I</figureCitation>
), angular distal membranous process (arrow
<quantity id="397447475472F2499AEE74DA0848D05D" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="87DD4FE6D3AA033163536450416BFFF1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">10 C</figureCitation>
,
<figureCitation id="67FD5648334A7DEED4DCA3279AF80B2D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 I</figureCitation>
), and narrow, curved retrolatero-subdistal membranous process (arrow in Figs
<figureCitation id="5E1CEF1CE90C65DFAAEC6EEF3A5E7CCE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Belisana tengchong sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at sclerotized prolatero-distal apophysis, arrow 2 points at distal membranous process D retrolateral view, arrow points at retrolatero-subdistal membranous process). Abbreviation: pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure10" httpUri="https://binary.pensoft.net/fig/1055875">10 D</figureCitation>
,
<figureCitation id="972DB50518239734DB8898EE3547DD91" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 J</figureCitation>
); bulb (Fig.
<figureCitation id="0A44234E30EFDD59A26E5E5E036B9E99" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 C</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria on tibia I on 17 % proximally; legs with short vertical setae on metatarsi; tarsus I with 17 distinct pseudosegments.
</paragraph>
<paragraph id="227E013B7DBD128D3877904E92DB1B37">
<emphasis id="FFA25A023C3A89E45A044732B8244D89" bold="true">Female</emphasis>
(
<emphasis id="59766100051DEFA4FE878C196DB91295" bold="true" italics="true">
<typeStatus id="B2CE56035B3BC8A574F837D5A42D19F4" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="0C3779AD1ED95A77750E90635FED029C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="027B0AA30717141F1DBB82C716442331" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="75F5BDD993304D8BBD88293B8255996D">Ar 44972</specimenCode>
): Similar to male, habitus as in Fig.
<figureCitation id="A88A5E886C6B60E9A5FE3E99410CE160" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 G, H</figureCitation>
. Total length 2.14 (2.26 with clypeus), carapace 0.66 long, 0.76 wide, opisthosoma 1.48 long, 1.34 wide; tibia I: 3.36; tibia I
<abbrev id="ABBRID0EK2BG" xlink_title="length/diameter">L / d</abbrev>
: 47. Eye inter-distances and diameters:
<abbrev id="ABBRID0EO2BG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0ES2BG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0EW2BG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0E12BG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0E52BG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0EC3BG" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum width / length: 0.54 / 0.50. Epigyne (Figs
<figureCitation id="5AD4861DC22B6CA1C833061D4AE17AA5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 A</figureCitation>
,
<figureCitation id="EE6600B6F2367BD7C002CB7D966B04D9" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 A</figureCitation>
) simple and flat, marginally slightly sclerotized, with a pair of lateral pockets 0.30 apart. Vulva (Figs
<figureCitation id="14F8ABAD7016A19FF80AF83FB0B4E94F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Belisana tengchong sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A, B); 0.10 (C, D); 0.30 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure11" httpUri="https://binary.pensoft.net/fig/1055876">11 B</figureCitation>
,
<figureCitation id="8F354ADD0D448D1A71B20FEA8D4AFB9C" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 B</figureCitation>
) with curved anterior arch and a pair of wide, curved pore plates (2 × longer than wide) bearing indistinct teeth.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EW3BG" type="variation">
<paragraph id="AD300FEF7F2DE0E4D711409BE57B5CF1">
<heading id="1F951310F6EE9FC412191C3870D0BD12" reason="title">Variation.</heading>
</paragraph>
<paragraph id="A732DFD2CE43CD32BC59EAAC5A526737">
Tibia I in
<specimenCount id="1424D21184B82EB93D5EE15AD917DBC2" count="2" type="male">two male</specimenCount>
<typeStatus id="F7AB93067073DA552397E4DB2E573BA8" type="paratype">paratypes</typeStatus>
(
<named-content id="C88595CA1FF33DFFF41B1A159AAD57FF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="E4C9635CE87402E33849224236380441" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="07119F070F1E02FDED5FD3122615DD95">
<specimenCode id="DEB93B432B6924F9B8DDA611C51EBA68">Ar 44970</specimenCode>
–71
</specimenCode>
): 5.45, 5.83. Tibia I in the other
<specimenCount id="386772B106BE9C800F5BA8250C82B510" count="2" type="female">two female</specimenCount>
<typeStatus id="8DF4266B4845ECF280B6ED8F065A706D" type="paratype">paratypes</typeStatus>
(
<named-content id="BFB7C6EAF70901026DA13FBC00817A34" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="EF53E1E5FF78DA2E6FC5B455DEE1E6D9" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="6CD6E9D079359E31545C4C3830588B99">
<specimenCode id="7279B19F6A6967DE338C8EB28532E6E2">Ar 44973</specimenCode>
–74
</specimenCode>
): 3.45, 4.05.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE4BG" type="habitat">
<paragraph id="9DD4DD774902A05420C332390E5DDDD6">
<heading id="9D6CEDE004DB8031AA23B2C787B1C28E" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="E0102FD0CF52B53809EC0DDD64B3F336">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJ4BG" type="distribution">
<paragraph id="171195DD9F935E8C52D386F67B8D3C74">
<heading id="D73AD0699A595924BBE2BBDF4F1BC5D5" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="74BF16FACF2B39C593FABFF6C91AE937">
<collectingCountry id="CEA41DD4A818BA22ACB66E9311AE3C57" name="China">China</collectingCountry>
(
<collectingRegion id="21198B5E58EB6DC8230A28270155297F" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<typeStatus id="4868F30F381333A25E2926E0ACF60570">type</typeStatus>
locality; Fig.
<figureCitation id="4DB7F292D1BC675986B3E7748671DFFB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the new Belisana species from Yunnan, China 1 Belisana honghe sp. nov. 2 B. jiuxiang sp. nov. 3 B. lincang sp. nov. 4 B. luxi sp. nov. 5 B. tengchong sp. nov. 6 B. tongi sp. nov. 7 B. yongsheng sp. nov. 8 B. yunnan sp. nov." figureDoi="10.3897/zookeys.1202.121633.figure1" httpUri="https://binary.pensoft.net/fig/1055866">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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