342 lines
39 KiB
XML
342 lines
39 KiB
XML
<document id="F1F00B8E986777E39183A4A472D85B49" ID-CLB-Dataset="156939" ID-DOI="10.11646/zootaxa.5190.3.3" ID-GBIF-Dataset="0feedd02-1573-4a52-bce0-f390e7a3bfcd" ID-ISSN="1175-5326" ID-Zenodo-Dep="7138200" ID-ZooBank="AEE72ABB-4358-4BB2-9394-DB5FD8105871" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1664792091404" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A., Semenchenko, Alexander A. & Palatov, Dmitry M." docDate="2022" docId="714487C0FFC6FFF402E7FEB6D001FEBF" docLanguage="en" docName="zootaxa.5190.3.3.pdf" docOrigin="Zootaxa 5190 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diamesa leoniella Hansen 1976" docType="treatment" docVersion="7" lastPageNumber="375" masterDocId="8D7DFFB8FFCBFFFA0270FFF6D360FFFE" masterDocTitle="Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of known species. II. Subgroups davisi, leona and loeffleri" masterLastPageNumber="392" masterPageNumber="361" pageNumber="374" updateTime="1699386040637" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E6045DDFF76C78D542E8375C5EB80897">Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of known species. II. Subgroups davisi, leona and loeffleri</mods:title>
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<mods:name id="548B68734E5F272B339871711C25A18C" type="personal">
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<mods:roleTerm id="407A3773CBA2062779973C6FB66F61CA">Author</mods:roleTerm>
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<mods:namePart id="FC26C642305D4470B6E522E5A21B92C7">Makarchenko, Eugenyi A.</mods:namePart>
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<mods:affiliation id="5C4D65662C69155220D2471674DFBFF7">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia</mods:affiliation>
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<mods:name id="FB1EF817FDAA18BC3E71C21E83C4196D" type="personal">
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<mods:namePart id="C6DA88009DFDC978F34BD244218EA6C3">Semenchenko, Alexander A.</mods:namePart>
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<mods:nameIdentifier id="2D65130589E3586D18F68F3371BC8304" type="ORCID">0000-0001-7207-9529</mods:nameIdentifier>
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<mods:affiliation id="928BD2C9B8949ECDD8F95E451C3D555D">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia & semenchenko _ alexander @ mail. ru; https: // orcid. org / 0000 - 0001 - 7207 - 9529</mods:affiliation>
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<mods:nameIdentifier id="A3AE6FC98D42648F6736677FDD57CFA0" type="email">semenchenko_alexander@mail.ru</mods:nameIdentifier>
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<mods:namePart id="B1502B5C1B3F700B07F251058EB569F3">Palatov, Dmitry M.</mods:namePart>
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<mods:affiliation id="ADBCFF6AEA47501798FC321FB936AE34">A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninskij prosp. 33, 119071 Moscow, Russia triops @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8826 - 9316</mods:affiliation>
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<mods:nameIdentifier id="AEA7844D3BF154BB7F4CBA8559E8EDB1" type="email">triops@yandex.ru</mods:nameIdentifier>
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<mods:title id="6B6598AEAB6C6C18D45CC6DF69038956">Zootaxa</mods:title>
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<mods:date id="29203CA3DDD7EF20925D0DC0E4C73A12">2022</mods:date>
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<mods:number id="0DBB9CF3CFF09A5D0244ED41F83B1A9E">2022-09-29</mods:number>
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<mods:number id="44A5641D84749195EE1AB236E9C1F97C">5190</mods:number>
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<treatment id="714487C0FFC6FFF402E7FEB6D001FEBF" ID-DOI="http://doi.org/10.5281/zenodo.7138238" ID-GBIF-Taxon="201582226" ID-Zenodo-Dep="7138238" LSID="urn:lsid:plazi:treatment:714487C0FFC6FFF402E7FEB6D001FEBF" httpUri="http://treatment.plazi.org/id/714487C0FFC6FFF402E7FEB6D001FEBF" lastPageId="14" lastPageNumber="375" pageId="13" pageNumber="374">
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<heading id="A21A81BAFFC6FFF702E7FEB6D2B7FEA4" bold="true" box="[151,471,319,346]" fontSize="11" level="1" pageId="13" pageNumber="374" reason="1">
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<taxonomicName id="3EED4D55FFC6FFF702E7FEB6D2B7FEA4" ID-CoL="359FB" authority="Hansen" authorityName="Hansen" authorityYear="1976" box="[151,471,319,346]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="374" phylum="Arthropoda" rank="species" species="leoniella">
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<emphasis id="CB99EAC4FFC6FFF702E7FEB6D2B7FEA4" bold="true" box="[151,471,319,346]" pageId="13" pageNumber="374">
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<emphasis id="CB99EAC4FFC6FFF702E7FEB6D210FEA4" bold="true" box="[151,368,319,346]" italics="true" pageId="13" pageNumber="374">Diamesa leoniella</emphasis>
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Hansen
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="B1F7655DFFC6FFF702E7FE92D399FE80" box="[151,249,356,382]" pageId="13" pageNumber="374" type="description">
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<paragraph id="F95236D6FFC6FFF702E7FE92D399FE80" blockId="13.[151,471,319,382]" box="[151,249,356,382]" pageId="13" pageNumber="374">
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(
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<figureCitation id="61D62A53FFC6FFF702EFFE92D391FE80" box="[159,241,356,382]" captionStart="FIGURES 23–28" captionStartId="14.[152,270,1923,1947]" captionTargetBox="[164,1418,345,1898]" captionTargetId="figure-79@14.[164,1422,345,1899]" captionTargetPageId="14" captionText="FIGURES 23–28.Adult males of Diamesa japonica Tokunaga from Honshu, Japan (23, 27), Hokkaido, Japan (24) and Kunashir Island (25–26), Diamesa leoniella Hansen from Alaska (28). 23, flagellum of antenna; 24, tergite IX (bent out on slide); 25, anal point in lateral view; 26–28, hypopygium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7138214" httpUri="https://zenodo.org/record/7138214/files/figure.png" pageId="13" pageNumber="374">Fig. 28</figureCitation>
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)
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<subSubSection id="B1F7655DFFC6FFF702E7FE52D01EFE24" pageId="13" pageNumber="374" type="reference_group">
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<paragraph id="F95236D6FFC6FFF702E7FE52D0BDFE45" blockId="13.[151,989,419,474]" box="[151,989,419,443]" pageId="13" pageNumber="374">
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<taxonomicName id="3EED4D55FFC6FFF702E7FE52D2C3FE45" ID-CoL="359FB" authority="Hansen" authorityName="Hansen" authorityYear="1976" box="[151,419,419,443]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="374" phylum="Arthropoda" rank="species" species="leoniella">
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<emphasis id="CB99EAC4FFC6FFF702E7FE52D231FE45" box="[151,337,419,443]" italics="true" pageId="13" pageNumber="374">Diamesa leoniella</emphasis>
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Hansen
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</taxonomicName>
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in
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<treatmentCitation id="784C10C7FFC6FFF703B3FE52D1B4FE45" author="Hansen, D. C. & Cook, E. F." box="[451,724,419,443]" page="111" pageId="13" pageNumber="374" year="1976">
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<bibRefCitation id="9D7C4B27FFC6FFF703B3FE52D1B4FE45" author="Hansen, D. C. & Cook, E. F." box="[451,724,419,443]" pageId="13" pageNumber="374" pagination="1 - 203" refId="ref17978" refString="Hansen, D. C. & Cook, E. F. (1976) The systematics and morphology of the Nearctic species of Diamesa Meigen, 1835 (Diptera: Chironomidae). Memoirs of the American Entomological Society, 30, 1 - 203." type="journal article" year="1976">Hansen & Cook 1976: 111</bibRefCitation>
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;
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<treatmentCitation id="784C10C7FFC6FFF700AEFE55D0B9FE45" author="Makarchenko, E. A." box="[734,985,419,443]" page="439" pageId="13" pageNumber="374" year="2021">
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<bibRefCitation id="9D7C4B27FFC6FFF700AEFE55D0B9FE45" author="Makarchenko, E. A." box="[734,985,419,443]" pageId="13" pageNumber="374" pagination="439 - 442" refId="ref18845" refString="Makarchenko, E. A. (2021) Confirmation of a species status of Diamesa leoniella Hansen, 1976, stat. resurr. (Diptera, Chironomidae) from North America. Zootaxa, 4985 (3), 439 - 442. https: // doi. org / 10.11646 / zootaxa. 4985.3.11" type="journal article" year="2021">Makarchenko 2021: 439</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="F95236D6FFC6FFF702E7FE34D01EFE24" blockId="13.[151,989,419,474]" box="[151,894,450,474]" pageId="13" pageNumber="374">
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<taxonomicName id="3EED4D55FFC6FFF702E7FE34D2DDFE24" ID-CoL="6CQN4" authority="Tokunaga" authorityName="Tokunaga" authorityYear="1936" box="[151,445,450,474]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="374" phylum="Arthropoda" rank="species" species="japonica">
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<emphasis id="CB99EAC4FFC6FFF702E7FE34D231FE27" box="[151,337,450,473]" italics="true" pageId="13" pageNumber="374">Diamesa japonica</emphasis>
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Tokunaga
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</taxonomicName>
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;
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<treatmentCitation id="784C10C7FFC6FFF703B7FE34D1A0FE24" author="Makarchenko, E. A." box="[455,704,450,474]" page="110" pageId="13" pageNumber="374" year="1981">
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<bibRefCitation id="9D7C4B27FFC6FFF703B7FE34D1A0FE24" author="Makarchenko, E. A." box="[455,704,450,474]" pageId="13" pageNumber="374" pagination="89 - 113" refId="ref18598" refString="Makarchenko, E. A. (1981) Taxonomy and distribution of some chironomids of subfamily Diamesinae (Diptera, Chironomidae) from the Soviet Far East. In: Invertebrate animals in ecosystems of salmon rivers of the Far East. DVNC AN SSSR Press, Vladivostok, pp. 89 - 113. [in Russian]" type="book chapter" year="1981">Makarchenko 1981: 110</bibRefCitation>
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</treatmentCitation>
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, misidentification.
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</paragraph>
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<emphasis id="CB99EAC4FFC6FFF702E7FDF4D2E7FDE2" bold="true" box="[151,391,514,540]" pageId="13" pageNumber="374">Material examined.</emphasis>
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<collectingCountry id="81FA7646FFC6FFF703E5FDF5D285FDE3" box="[405,485,515,541]" name="United States of America" pageId="13" pageNumber="374">U.S.A.</collectingCountry>
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:
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male,
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,
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,
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,
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58.4217
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,
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<date id="8D531016FFC6FFF702E7FDD1D278FDBF" box="[151,280,551,577]" pageId="13" pageNumber="374" value="2013-03-26">
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<collectingDate id="9D17E9FEFFC6FFF702E7FDD1D278FDBF" box="[151,280,551,577]" pageId="13" pageNumber="374" value="2013-03-26">26.III.2013</collectingDate>
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</date>
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, leg.
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<collectorName id="54185300FFC6FFF70321FDD1D2ACFDBF" box="[337,460,551,577]" pageId="13" pageNumber="374">K. Frangos</collectorName>
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and
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<collectorName id="54185300FFC6FFF70071FDD1D115FDBF" box="[513,629,551,577]" pageId="13" pageNumber="374">P. Hudson</collectorName>
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;
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<materialsCitation id="49853C8BFFC6FFF7000FFDD1D2B8FD9B" ID-GBIF-Occurrence="3920576343" collectingDate="2013-07-17" collectorName="Nugget Falls, N & P. Hudson." country="United States of America" location="Nugget Falls" longitude="58.427" municipality="Juneau City" pageId="13" pageNumber="374" specimenCount="1" specimenCount-adult="1" stateProvince="Alaska">
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<specimenCount id="EFEBFD5FFFC6FFF7000FFDD1D1ABFDBF" box="[639,715,551,577]" pageId="13" pageNumber="374" type="adult">1 adult</specimenCount>
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male, the same data except
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<location id="FC32600DFFC6FFF7018FFDD1D7F0FDBF" LSID="urn:lsid:plazi:treatment:714487C0FFC6FFF402E7FEB6D001FEBF:FC32600DFFC6FFF7018FFDD1D7F0FDBF" box="[1023,1168,551,577]" country="United States of America" longitude="58.427" name="Nugget Falls" pageId="13" pageNumber="374">Nugget Falls</location>
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,
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58.4270
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<geoCoordinate id="9CD95011FFC6FFF70765FDD1D6F8FDBF" box="[1301,1432,550,577]" degrees="58.4270" direction="east" orientation="longitude" pageId="13" pageNumber="374" precision="5" value="58.427">E 134.5373</geoCoordinate>
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,
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<date id="8D531016FFC6FFF702E7FDBDD243FD9B" box="[151,291,587,613]" pageId="13" pageNumber="374" value="2013-07-17">
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<collectingDate id="9D17E9FEFFC6FFF702E7FDBDD243FD9B" box="[151,291,587,613]" pageId="13" pageNumber="374" value="2013-07-17">17.VII.2013</collectingDate>
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</date>
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, leg.
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<collectorName id="54185300FFC6FFF7032EFDBDD2B8FD9B" box="[350,472,587,613]" pageId="13" pageNumber="374">P. Hudson.</collectorName>
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</paragraph>
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|
||
<subSubSection id="B1F7655DFFC6FFF702B7FD98D28CF870" pageId="13" pageNumber="374" type="description">
|
||
<paragraph id="F95236D6FFC6FFF702B7FD98D231FD76" blockId="13.[151,1437,514,1189]" box="[199,337,622,648]" pageId="13" pageNumber="374">
|
||
<emphasis id="CB99EAC4FFC6FFF702B7FD98D231FD76" bold="true" box="[199,337,622,648]" pageId="13" pageNumber="374">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="F95236D6FFC6FFF702B7FD64D6F2FD53" blockId="13.[151,1437,514,1189]" box="[199,1426,658,685]" pageId="13" pageNumber="374">
|
||
<emphasis id="CB99EAC4FFC6FFF702B7FD64D226FD53" bold="true" box="[199,326,658,685]" italics="true" pageId="13" pageNumber="374">Adult male</emphasis>
|
||
(n = 2). Total length
|
||
<quantity id="3E159B33FFC6FFF70044FD65D1EAFD53" box="[564,650,659,685]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="13" pageNumber="374" unit="mm" value="4.7">4.7 mm</quantity>
|
||
(n = 1). Wing length
|
||
<quantity id="3E159B33FFC6FFF7010CFD65D080FD53" box="[892,992,659,685]" metricMagnitude="-3" metricUnit="m" metricValue="3.44" pageId="13" pageNumber="374" unit="mm" value="3.44">3.44 mm</quantity>
|
||
(n = 1). Total length/wing length 1.37.
|
||
</paragraph>
|
||
<paragraph id="F95236D6FFC6FFF702B7FD41D058FD0B" blockId="13.[151,1437,514,1189]" pageId="13" pageNumber="374">Coloration. Head, thorax, legs and hypopygium dark brown; antenna brown; palpomeres light brown; abdomen light brown to brown; wing greyish, veins yellowish brown.</paragraph>
|
||
<paragraph id="F95236D6FFC6FFF702B7FD09D13BFC0F" blockId="13.[151,1437,514,1189]" pageId="13" pageNumber="374">
|
||
Head. Eyes hairy. Frontal tubercles 16–44 μm long, covered with microtrichia. Temporal setae including about 8–12 frontals, 13–15 verticals, 5 postorbitals. Clypeus with 5–6 setae. Antenna with 8 flagellomeres and reduced plume of setae; flagellomeres 1–7 with 2–3 setae, 24–40 μm long; terminal flagellomere in basal part with 4–5 setae, 60–72 μm long and with 2 subapical setae, 16–20 μm long. Flagellomeres 1–8 length (μm): 88–104, 36–40, 40, 32, 32, 26–32, 32–44, 164–248; AR 0.41–0.78. Antennal length/palp length 1.09–1.21. Palpomeres lengths (in μm): 32–40; 56–64; 108; 84–100; 136–156. Palpomere
|
||
<quantity id="3E159B33FFC6FFF7017CFC45D058FC33" box="[780,824,947,973]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="374" unit="in" value="3.0">3 in</quantity>
|
||
distal part with sensilla capitata with diameter 20 μm. Palpomeres 1–5 length/head width 0.92.
|
||
</paragraph>
|
||
<paragraph id="F95236D6FFC6FFF702B7FC0DD645FBEB" blockId="13.[151,1437,514,1189]" box="[199,1317,1018,1045]" pageId="13" pageNumber="374">Thorax. Antepronotum with 9–11 ventrolateral setae. Dorsocentrals 5–7, prealars 7, scutellars 9–11.</paragraph>
|
||
<paragraph id="F95236D6FFC6FFF702B7FBE9D7A0FBA3" blockId="13.[151,1437,514,1189]" pageId="13" pageNumber="374">
|
||
Wing. R and R
|
||
<subScript id="65693493FFC6FFF70301FBC7D219FBC1" attach="left" box="[369,377,1073,1087]" fontSize="6" pageId="13" pageNumber="374">1</subScript>
|
||
with 25–34 setae; R
|
||
<subScript id="65693493FFC6FFF70014FBC7D11EFBC1" attach="left" box="[612,638,1073,1087]" fontSize="6" pageId="13" pageNumber="374">4+5</subScript>
|
||
with 18–19 setae. Costa extension 52–60 μm long. RM/MCu 2.8–3.5. Anal lobe slightly reduced, angularly. Squama with 14–17 setae, 52–68 μm long. VR 0.96–1.0.
|
||
</paragraph>
|
||
<paragraph id="F95236D6FFC6FFF702B7FB91D7ABFB5B" blockId="13.[151,1437,514,1189]" pageId="13" pageNumber="374">
|
||
Legs. Spur of front tibia 40–44 μm long; spurs of mid tibia 44 μm long; of hind tibia 74–80 μm and 44–48 μm long. Hind tibial comb with 15–16 setae. Lengths and proportions of leg segments as in
|
||
<tableCitation id="B46F036DFFC6FFF70602FB7DD7A7FB5B" box="[1138,1223,1163,1189]" captionStart="TABLE 9" captionStartId="13.[151,239,1231,1255]" captionTargetPageId="13" captionText="TABLE 9. Lengths (in µm) and proportions of leg segments of Diamesa leoniella Hansen, male (n=2)" httpUri="http://table.plazi.org/id/AD92665EFFC6FFF702E7FB39D7BCFB19" pageId="13" pageNumber="374" tableUuid="AD92665EFFC6FFF702E7FB39D7BCFB19">Table 9</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="AD92665EFFC6FFF702E7FB39D7BCFB19" ID-Table-UUID="AD92665EFFC6FFF702E7FB39D7BCFB19" box="[151,1244,1231,1256]" httpUri="http://table.plazi.org/id/AD92665EFFC6FFF702E7FB39D7BCFB19" pageId="13" pageNumber="374" startId="13.[151,239,1231,1255]" targetBox="[151,1370,1271,1656]" targetIsTable="true" targetPageId="13">
|
||
<paragraph id="F95236D6FFC6FFF702E7FB39D7BCFB19" blockId="13.[151,1244,1231,1256]" box="[151,1244,1231,1256]" pageId="13" pageNumber="374">
|
||
<emphasis id="CB99EAC4FFC6FFF702E7FB39D26AFB19" bold="true" box="[151,266,1231,1255]" pageId="13" pageNumber="374">TABLE 9.</emphasis>
|
||
Lengths (in µm) and proportions of leg segments of
|
||
<taxonomicName id="3EED4D55FFC6FFF7014FFB26D73CFB19" authority="Hansen" authorityName="Hansen" authorityYear="1976" box="[831,1116,1231,1255]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="374" phylum="Arthropoda" rank="species" species="leoniella">
|
||
<emphasis id="CB99EAC4FFC6FFF7014FFB26D762FB19" box="[831,1026,1231,1255]" italics="true" pageId="13" pageNumber="374">Diamesa leoniella</emphasis>
|
||
Hansen
|
||
</taxonomicName>
|
||
, male (n=2)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F95236D6FFC6FFF7029FFB01D79EF98C" pageId="13" pageNumber="374">
|
||
<table id="8BEDC476FFC6000502E7FB01D63AF986" box="[151,1370,1271,1656]" colsContinueIn="25.[151,1316,1406,1791]" gridcols="9" gridrows="9" pageId="13" pageNumber="374">
|
||
<tr id="47DD3494FFC6000502E7FB01D63AFAE8" box="[151,1370,1271,1302]" gridrow="0" pageId="13" pageNumber="374" rowspan-0="1" rowspan-7="1">
|
||
<th id="040C5DE8FFC60005029FFB01D201FAE8" box="[239,353,1271,1302]" gridcol="1" gridrow="0" pageId="13" pageNumber="374">fe</th>
|
||
<th id="040C5DE8FFC6000503ECFB01D16FFAE8" box="[412,527,1271,1302]" gridcol="2" gridrow="0" pageId="13" pageNumber="374">ti</th>
|
||
<th id="040C5DE8FFC60005003AFB01D1DCFAE8" box="[586,700,1271,1302]" gridcol="3" gridrow="0" pageId="13" pageNumber="374">ta1</th>
|
||
<th id="040C5DE8FFC600050088FB01D031FAE8" box="[760,849,1271,1302]" gridcol="4" gridrow="0" pageId="13" pageNumber="374">ta2</th>
|
||
<th id="040C5DE8FFC6000501F4FB01D09FFAE8" box="[900,1023,1271,1302]" gridcol="5" gridrow="0" pageId="13" pageNumber="374">ta3</th>
|
||
<th id="040C5DE8FFC600050623FB01D70DFAE8" box="[1107,1133,1271,1302]" gridcol="6" gridrow="0" pageId="13" pageNumber="374">ta4</th>
|
||
<th id="040C5DE8FFC600050771FB01D63AFAE8" box="[1281,1370,1271,1302]" gridcol="8" gridrow="0" pageId="13" pageNumber="374">ta 5</th>
|
||
</tr>
|
||
<tr id="47DD3494FFC6000502E7FAE9D63AFAC3" box="[151,1370,1311,1341]" gridrow="1" pageId="13" pageNumber="374">
|
||
<th id="040C5DE8FFC6000502E7FAE9D3DBFAC3" box="[151,187,1311,1341]" gridcol="0" gridrow="1" pageId="13" pageNumber="374">P1</th>
|
||
<td id="040C5DE8FFC60005029FFAE9D201FAC3" box="[239,353,1311,1341]" gridcol="1" gridrow="1" pageId="13" pageNumber="374">1607–2280</td>
|
||
<td id="040C5DE8FFC6000503ECFAE9D16FFAC3" box="[412,527,1311,1341]" gridcol="2" gridrow="1" pageId="13" pageNumber="374">1591–2120</td>
|
||
<td id="040C5DE8FFC60005003AFAE9D1DCFAC3" box="[586,700,1311,1341]" gridcol="3" gridrow="1" pageId="13" pageNumber="374">1082–1400</td>
|
||
<td id="040C5DE8FFC600050088FAE9D031FAC3" box="[760,849,1311,1341]" gridcol="4" gridrow="1" pageId="13" pageNumber="374">541–720</td>
|
||
<td id="040C5DE8FFC6000501F4FAE9D09FFAC3" box="[900,1023,1311,1341]" gridcol="5" gridrow="1" pageId="13" pageNumber="374">295–440</td>
|
||
<td id="040C5DE8FFC600050623FAE9D7B4FAC3" box="[1107,1236,1311,1341]" colspan="2" colspanRight="1" gridcol="6" gridrow="1" pageId="13" pageNumber="374">107–140</td>
|
||
<td id="040C5DE8FFC600050771FAE9D63AFAC3" box="[1281,1370,1311,1341]" gridcol="8" gridrow="1" pageId="13" pageNumber="374">148–180</td>
|
||
</tr>
|
||
<tr id="47DD3494FFC6000502E7FABED63AFA98" box="[151,1370,1352,1382]" gridrow="2" pageId="13" pageNumber="374">
|
||
<th id="040C5DE8FFC6000502E7FABED3DBFA98" box="[151,187,1352,1382]" gridcol="0" gridrow="2" pageId="13" pageNumber="374">P 2</th>
|
||
<td id="040C5DE8FFC60005029FFABED201FA98" box="[239,353,1352,1382]" gridcol="1" gridrow="2" pageId="13" pageNumber="374">1706–2160</td>
|
||
<td id="040C5DE8FFC6000503ECFABED16FFA98" box="[412,527,1352,1382]" gridcol="2" gridrow="2" pageId="13" pageNumber="374">1394–1760</td>
|
||
<td id="040C5DE8FFC60005003AFABED1DCFA98" box="[586,700,1352,1382]" gridcol="3" gridrow="2" pageId="13" pageNumber="374">763–960</td>
|
||
<td id="040C5DE8FFC600050088FABED031FA98" box="[760,849,1352,1382]" gridcol="4" gridrow="2" pageId="13" pageNumber="374">377–440</td>
|
||
<td id="040C5DE8FFC6000501F4FABED09FFA98" box="[900,1023,1352,1382]" gridcol="5" gridrow="2" pageId="13" pageNumber="374">230–360</td>
|
||
<td id="040C5DE8FFC600050623FABED7B4FA98" box="[1107,1236,1352,1382]" colspan="2" colspanRight="1" gridcol="6" gridrow="2" pageId="13" pageNumber="374">98–140</td>
|
||
<td id="040C5DE8FFC600050771FABED63AFA98" box="[1281,1370,1352,1382]" gridcol="8" gridrow="2" pageId="13" pageNumber="374">148–180</td>
|
||
</tr>
|
||
<tr id="47DD3494FFC6000502E7FA86D63AFA70" box="[151,1370,1392,1422]" gridrow="3" pageId="13" pageNumber="374">
|
||
<th id="040C5DE8FFC6000502E7FA86D3DBFA70" box="[151,187,1392,1422]" gridcol="0" gridrow="3" pageId="13" pageNumber="374">P3</th>
|
||
<td id="040C5DE8FFC60005029FFA86D201FA70" box="[239,353,1392,1422]" gridcol="1" gridrow="3" pageId="13" pageNumber="374">1738–2400</td>
|
||
<td id="040C5DE8FFC6000503ECFA86D16FFA70" box="[412,527,1392,1422]" gridcol="2" gridrow="3" pageId="13" pageNumber="374">1624–2160</td>
|
||
<td id="040C5DE8FFC60005003AFA86D1DCFA70" box="[586,700,1392,1422]" gridcol="3" gridrow="3" pageId="13" pageNumber="374">1033–1440</td>
|
||
<td id="040C5DE8FFC600050088FA86D031FA70" box="[760,849,1392,1422]" gridcol="4" gridrow="3" pageId="13" pageNumber="374">607–800</td>
|
||
<td id="040C5DE8FFC6000501F4FA86D09FFA70" box="[900,1023,1392,1422]" gridcol="5" gridrow="3" pageId="13" pageNumber="374">312–400</td>
|
||
<td id="040C5DE8FFC600050623FA86D7B4FA70" box="[1107,1236,1392,1422]" colspan="2" colspanRight="1" gridcol="6" gridrow="3" pageId="13" pageNumber="374">98–160</td>
|
||
<td id="040C5DE8FFC600050771FA86D63AFA70" box="[1281,1370,1392,1422]" gridcol="8" gridrow="3" pageId="13" pageNumber="374">148–200</td>
|
||
</tr>
|
||
<tr id="47DD3494FFC6000502E7FA4FD63AFA2F" box="[151,1370,1465,1489]" gridrow="4" pageId="13" pageNumber="374">
|
||
<th id="040C5DE8FFC6000502E7FA4FD63AFA2F" box="[151,1370,1465,1489]" colspan="9" colspanRight="8" gridcol="0" gridrow="4" pageId="13" pageNumber="374">continued</th>
|
||
</tr>
|
||
<tr id="47DD3494FFC6000502E7FA14D63AFA04" box="[151,1370,1506,1530]" gridrow="5" pageId="13" pageNumber="374" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
|
||
<td id="040C5DE8FFC60005029FFA14D201FA04" box="[239,353,1506,1530]" gridcol="1" gridrow="5" pageId="13" pageNumber="374">LR</td>
|
||
<td id="040C5DE8FFC60005003AFA14D1DCFA04" box="[586,700,1506,1530]" gridcol="3" gridrow="5" pageId="13" pageNumber="374">BV</td>
|
||
<td id="040C5DE8FFC6000501F4FA14D09FFA04" box="[900,1023,1506,1530]" gridcol="5" gridrow="5" pageId="13" pageNumber="374">SV</td>
|
||
<td id="040C5DE8FFC6000506C2FA14D7B4FA04" box="[1202,1236,1506,1530]" gridcol="7" gridrow="5" pageId="13" pageNumber="374">BR</td>
|
||
</tr>
|
||
<tr id="47DD3494FFC6000502E7F9FCD63AF9D6" box="[151,1370,1546,1576]" gridrow="6" pageId="13" pageNumber="374" rowspan-4="1" rowspan-6="1">
|
||
<th id="040C5DE8FFC6000502E7F9FCD3DBF9D6" box="[151,187,1546,1576]" gridcol="0" gridrow="6" pageId="13" pageNumber="374">P 1</th>
|
||
<td id="040C5DE8FFC60005029FF9FCD16FF9D6" box="[239,527,1546,1576]" colspan="2" colspanRight="1" gridcol="1" gridrow="6" pageId="13" pageNumber="374">0.66–0.68</td>
|
||
<td id="040C5DE8FFC60005003AF9FCD1DCF9D6" box="[586,700,1546,1576]" gridcol="3" gridrow="6" pageId="13" pageNumber="374">3.92</td>
|
||
<td id="040C5DE8FFC6000501F4F9FCD09FF9D6" box="[900,1023,1546,1576]" gridcol="5" gridrow="6" pageId="13" pageNumber="374">2.96–3.14</td>
|
||
<td id="040C5DE8FFC6000506C2F9FCD63AF9D6" box="[1202,1370,1546,1576]" colspan="2" colspanRight="1" gridcol="7" gridrow="6" pageId="13" pageNumber="374">0.9–1.1</td>
|
||
</tr>
|
||
<tr id="47DD3494FFC6000502E7F9C4D63AF9AE" box="[151,1370,1586,1616]" gridrow="7" pageId="13" pageNumber="374" rowspan-2="1" rowspan-4="1" rowspan-6="1">
|
||
<th id="040C5DE8FFC6000502E7F9C4D3DBF9AE" box="[151,187,1586,1616]" gridcol="0" gridrow="7" pageId="13" pageNumber="374">P2</th>
|
||
<td id="040C5DE8FFC60005029FF9C4D201F9AE" box="[239,353,1586,1616]" gridcol="1" gridrow="7" pageId="13" pageNumber="374">0.55</td>
|
||
<td id="040C5DE8FFC60005003AF9C4D1DCF9AE" box="[586,700,1586,1616]" gridcol="3" gridrow="7" pageId="13" pageNumber="374">4.36–4.53</td>
|
||
<td id="040C5DE8FFC6000501F4F9C4D09FF9AE" box="[900,1023,1586,1616]" gridcol="5" gridrow="7" pageId="13" pageNumber="374">4.06–4.08</td>
|
||
<td id="040C5DE8FFC6000506C2F9C4D63AF9AE" box="[1202,1370,1586,1616]" colspan="2" colspanRight="1" gridcol="7" gridrow="7" pageId="13" pageNumber="374">0.9–1.0</td>
|
||
</tr>
|
||
<tr id="47DD3494FFC6000502E7F9ACD63AF986" box="[151,1370,1626,1656]" gridrow="8" pageId="13" pageNumber="374" rowspan-4="1" rowspan-6="1">
|
||
<th id="040C5DE8FFC6000502E7F9ACD3DBF986" box="[151,187,1626,1656]" gridcol="0" gridrow="8" pageId="13" pageNumber="374">P3</th>
|
||
<td id="040C5DE8FFC60005029FF9ACD16FF986" box="[239,527,1626,1656]" colspan="2" colspanRight="1" gridcol="1" gridrow="8" pageId="13" pageNumber="374">0.64–0.67</td>
|
||
<td id="040C5DE8FFC60005003AF9ACD1DCF986" box="[586,700,1626,1656]" gridcol="3" gridrow="8" pageId="13" pageNumber="374">3.77–3.85</td>
|
||
<td id="040C5DE8FFC6000501F4F9ACD09FF986" box="[900,1023,1626,1656]" gridcol="5" gridrow="8" pageId="13" pageNumber="374">3.17–3.25</td>
|
||
<td id="040C5DE8FFC6000506C2F9ACD63AF986" box="[1202,1370,1626,1656]" colspan="2" colspanRight="1" gridcol="7" gridrow="8" pageId="13" pageNumber="374">1.0–1.1</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="F95236D6FFC6FFF702B7F96AD794F895" blockId="13.[151,1437,1692,2043]" pageId="13" pageNumber="374">
|
||
Hypopygium (
|
||
<figureCitation id="61D62A53FFC6FFF70318F96AD2DAF949" box="[360,442,1692,1719]" captionStart="FIGURES 23–28" captionStartId="14.[152,270,1923,1947]" captionTargetBox="[164,1418,345,1898]" captionTargetId="figure-79@14.[164,1422,345,1899]" captionTargetPageId="14" captionText="FIGURES 23–28.Adult males of Diamesa japonica Tokunaga from Honshu, Japan (23, 27), Hokkaido, Japan (24) and Kunashir Island (25–26), Diamesa leoniella Hansen from Alaska (28). 23, flagellum of antenna; 24, tergite IX (bent out on slide); 25, anal point in lateral view; 26–28, hypopygium in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7138214" httpUri="https://zenodo.org/record/7138214/files/figure.png" pageId="13" pageNumber="374">Fig. 28</figureCitation>
|
||
). Tergite IX with 11–13 setae (from one side), 16–24 μm long and wedge-shaped anal point 100–140 μm long, covered in basal half with numerous microtrichia forward and laterally directed. Laterosternite IX with 10–11 short setae. Transverse sternapodeme triangular, 128 μm long. Gonocoxite 400–496 μm long. Inferior volsella finger-like, 40–48 μm long and 24 μm width, covered with short setae and microtrichia. Superior volsella in form of collar, covered with microtrichia and short setae.Gonostylus slightly curved, 212–246 μm long, with numerous, proximally directed strong setae, 48–64 μm long; megaseta 6–8 μm long. HR 1.65–1.93.
|
||
</paragraph>
|
||
<paragraph id="F95236D6FFC6FFF702B7F883D28CF870" blockId="13.[151,1437,1692,2043]" box="[199,492,1908,1935]" pageId="13" pageNumber="374">
|
||
<emphasis id="CB99EAC4FFC6FFF702B7F883D264F871" bold="true" box="[199,260,1909,1935]" italics="true" pageId="13" pageNumber="374">Pupa</emphasis>
|
||
and
|
||
<emphasis id="CB99EAC4FFC6FFF7034AF882D215F871" bold="true" box="[314,373,1908,1935]" italics="true" pageId="13" pageNumber="374">larva</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1F7655DFFC6FFF402B7F86FD399FF07" lastPageId="14" lastPageNumber="375" pageId="13" pageNumber="374" type="discussion">
|
||
<paragraph id="F95236D6FFC6FFF402B7F86FD399FF07" blockId="13.[151,1437,1692,2043]" lastBlockId="14.[151,1437,151,321]" lastPageId="14" lastPageNumber="375" pageId="13" pageNumber="374">
|
||
<emphasis id="CB99EAC4FFC6FFF702B7F86FD25BF84D" bold="true" box="[199,315,1945,1971]" pageId="13" pageNumber="374">Remarks.</emphasis>
|
||
Male of
|
||
<taxonomicName id="3EED4D55FFC6FFF703C0F86FD159F84C" authorityName="Hansen" authorityYear="1976" box="[432,569,1944,1970]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="374" phylum="Arthropoda" rank="species" species="leoniella">
|
||
<emphasis id="CB99EAC4FFC6FFF703C0F86FD159F84C" box="[432,569,1944,1970]" italics="true" pageId="13" pageNumber="374">D. leoniella</emphasis>
|
||
</taxonomicName>
|
||
is very closely related to
|
||
<taxonomicName id="3EED4D55FFC6FFF70106F86FD09FF84C" authorityName="Tokunaga" authorityYear="1936" box="[886,1023,1945,1970]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="374" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="CB99EAC4FFC6FFF70106F86FD09FF84C" box="[886,1023,1945,1970]" italics="true" pageId="13" pageNumber="374">D. japonica</emphasis>
|
||
</taxonomicName>
|
||
and many years ago senior author misidentified this species as
|
||
<taxonomicName id="3EED4D55FFC6FFF70390F84BD001F829" authority="(Makarchenko 1981)" baseAuthorityName="Makarchenko" baseAuthorityYear="1981" box="[480,865,1980,2007]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="374" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="CB99EAC4FFC6FFF70390F84BD106F828" box="[480,614,1981,2006]" italics="true" pageId="13" pageNumber="374">D. japonica</emphasis>
|
||
(
|
||
<bibRefCitation id="9D7C4B27FFC6FFF70007F84AD038F829" author="Makarchenko, E. A." box="[631,856,1980,2007]" pageId="13" pageNumber="374" pagination="89 - 113" refId="ref18598" refString="Makarchenko, E. A. (1981) Taxonomy and distribution of some chironomids of subfamily Diamesinae (Diptera, Chironomidae) from the Soviet Far East. In: Invertebrate animals in ecosystems of salmon rivers of the Far East. DVNC AN SSSR Press, Vladivostok, pp. 89 - 113. [in Russian]" type="book chapter" year="1981">Makarchenko 1981</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. After obtaining comparative material from North America, it turned out that these two species can be well distinguished by the structure of the hypopygium and some other features (
|
||
<bibRefCitation id="9D7C4B27FFC5FFF4034EFF61D17CFF4F" author="Makarchenko, E. A." box="[318,540,151,177]" pageId="14" pageNumber="375" pagination="439 - 442" refId="ref18845" refString="Makarchenko, E. A. (2021) Confirmation of a species status of Diamesa leoniella Hansen, 1976, stat. resurr. (Diptera, Chironomidae) from North America. Zootaxa, 4985 (3), 439 - 442. https: // doi. org / 10.11646 / zootaxa. 4985.3.11" type="journal article" year="2021">Makarchenko 2021</bibRefCitation>
|
||
). So, the male of
|
||
<taxonomicName id="3EED4D55FFC5FFF400AEFF61D001FF4F" authorityName="Hansen" authorityYear="1976" box="[734,865,151,177]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="375" phylum="Arthropoda" rank="species" species="leoniella">
|
||
<emphasis id="CB99EAC4FFC5FFF400AEFF61D001FF4F" box="[734,865,151,177]" italics="true" pageId="14" pageNumber="375">D. leoniella</emphasis>
|
||
</taxonomicName>
|
||
has a finger-shaped inferior volsella and a superior volsella in the form of a collar, while
|
||
<taxonomicName id="3EED4D55FFC5FFF4004DFF4DD1A2FF2A" baseAuthorityName="Makarchenko" baseAuthorityYear="1981" box="[573,706,187,212]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="375" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="CB99EAC4FFC5FFF4004DFF4DD1A2FF2A" box="[573,706,187,212]" italics="true" pageId="14" pageNumber="375">D. japonica</emphasis>
|
||
</taxonomicName>
|
||
without inferior volsella and superior volsella in form of angular tubercle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1F7655DFFC5FFF402B7FEF4D001FEBF" pageId="14" pageNumber="375" type="distribution">
|
||
<paragraph id="F95236D6FFC5FFF402B7FEF4D001FEBF" blockId="14.[151,1437,151,321]" pageId="14" pageNumber="375">
|
||
<emphasis id="CB99EAC4FFC5FFF402B7FEF4D201FEE2" bold="true" box="[199,353,258,284]" pageId="14" pageNumber="375">Distribution.</emphasis>
|
||
Nearctic species. Known from
|
||
<collectingCountry id="81FA7646FFC5FFF40093FEF5D047FEE3" box="[739,807,259,285]" name="United States of America" pageId="14" pageNumber="375">U.S.A.</collectingCountry>
|
||
—
|
||
<collectingRegion id="3B29F834FFC5FFF40135FEF5D0A1FEE3" box="[837,961,259,285]" country="United States of America" name="California" pageId="14" pageNumber="375">California</collectingRegion>
|
||
,
|
||
<collectingRegion id="3B29F834FFC5FFF401A2FEF5D758FEE3" box="[978,1080,259,285]" country="United States of America" name="Montana" pageId="14" pageNumber="375">Montana</collectingRegion>
|
||
,
|
||
<collectingRegion id="3B29F834FFC5FFF4063AFEF5D7E3FEE3" box="[1098,1155,259,285]" country="United States of America" name="Utah" pageId="14" pageNumber="375">Utah</collectingRegion>
|
||
,
|
||
<collectingRegion id="3B29F834FFC5FFF406E5FEF5D67DFEE3" box="[1173,1309,259,285]" country="United States of America" name="Washington" pageId="14" pageNumber="375">Washington</collectingRegion>
|
||
,
|
||
<collectingRegion id="3B29F834FFC5FFF4075EFEF5D6FCFEE3" box="[1326,1436,259,285]" country="United States of America" name="Wyoming" pageId="14" pageNumber="375">Wyoming</collectingRegion>
|
||
(
|
||
<bibRefCitation id="9D7C4B27FFC5FFF402EFFED1D2F5FEBF" author="Hansen, D. C. & Cook, E. F." box="[159,405,295,321]" pageId="14" pageNumber="375" pagination="1 - 203" refId="ref17978" refString="Hansen, D. C. & Cook, E. F. (1976) The systematics and morphology of the Nearctic species of Diamesa Meigen, 1835 (Diptera: Chironomidae). Memoirs of the American Entomological Society, 30, 1 - 203." type="journal article" year="1976">Hansen & Cook 1976</bibRefCitation>
|
||
) and
|
||
<collectingRegion id="3B29F834FFC5FFF403A2FED1D140FEBF" box="[466,544,295,321]" country="United States of America" name="Alaska" pageId="14" pageNumber="375">Alaska</collectingRegion>
|
||
(recorded for the first time).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |