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<mods:title id="38925E2F820ADE5E82C99C65082365DE">The external larval morphology of aquatic and terrestrial Luciolinae fireflies (Coleoptera: Lampyridae)</mods:title>
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<mods:namePart id="42812A6C6F1D0D8899DE91979D26F07D">Fu, Xinhua</mods:namePart>
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<paragraph id="F931254E0B62C030FF3FFC8583DBFC82" blockId="15.[151,483,864,924]" box="[151,483,864,891]" pageId="15" pageNumber="16">
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<taxonomicName id="3E8E5ECD0B62C030FF3FFC8583DBFC82" ID-CoL="8MYPS" authority="Gorham" authorityName="Gorham" box="[151,483,864,891]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="substriata">
<emphasis id="CBFAF95C0B62C030FF3FFC8583DBFC82" bold="true" box="[151,483,864,891]" pageId="15" pageNumber="16">
<emphasis id="CBFAF95C0B62C030FF3FFC858349FC83" bold="true" box="[151,369,865,891]" italics="true" pageId="15" pageNumber="16">Luciola substriata</emphasis>
Gorham
</emphasis>
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</heading>
</paragraph>
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<figureCitation id="61B539CB0B62C030FF3FFC6082D1FC64" box="[151,233,900,924]" captionStart="FIGURES 31 36" captionStartId="16.[151,264,1671,1694]" captionTargetBox="[151,1435,193,1649]" captionTargetId="figure@16.[151,1436,193,1650]" captionTargetPageId="16" captionText="FIGURES 31 36. Luciola substriata first instar larva, SEM. 31, 32 body, dorsal (31) and ventral; 33 ventral abdominal segments 3 7; 34 anterior portion of prothorax and retracted head, ventral; 35 left antenna, ventral; 36 apical maxillary palpomere, ventral." httpUri="https://zenodo.org/record/211304/files/figure.png" pageId="15" pageNumber="16">Figs 31</figureCitation>
42, 70, 83, 84
</paragraph>
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<treatmentCitationGroup id="D99E02600B62C030FF3FFC288126FBF8" pageId="15" pageNumber="16">
<taxonomicName id="3E8E5ECD0B62C030FF3FFC288028FC19" ID-CoL="8MYPS" authority="Gorham 1880:100" authorityName="Gorham" authorityPageNumber="100" authorityYear="1880" box="[151,528,971,994]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="substriata">
<emphasis id="CBFAF95C0B62C030FF3FFC288377FC1A" box="[151,335,972,994]" italics="true" pageId="15" pageNumber="16">Luciola substriata</emphasis>
<treatmentCitation id="782F035F0B62C030FEFDFC2F8028FC19" author="Gorham" box="[341,528,971,993]" page="100" pageId="15" pageNumber="16" year="1880">
<bibRefCitation id="9D1F58BF0B62C030FEFDFC2F83DCFC19" author="Gorham" box="[341,484,971,993]" pageId="15" pageNumber="16" refString="Gorham, H. S. (1880) Materials for a revision of the Lampyridae. Transactions of the Royal Entomological Society of London, 1880, 1 - 36, 83 - 112." type="journal article" year="1880">Gorham 1880</bibRefCitation>
:100
</treatmentCitation>
</taxonomicName>
; 1895:305.
<treatmentCitation id="782F035F0B62C030FD2DFC2F810CFC19" author="Olivier" box="[645,820,971,993]" page="86" pageId="15" pageNumber="16" year="1902">
<bibRefCitation id="9D1F58BF0B62C030FD2DFC2F812EFC19" author="Olivier" box="[645,790,971,993]" pageId="15" pageNumber="16" refString="Olivier, E. (1902 a) Catalogue des especes de ' Luciola'et genres voisins decrits jusqu'ace jour. Revue Scientifique du Bourbonnais, 15, 69 - 88." type="journal article" year="1902" yearSuffix="a">Olivier 1902a</bibRefCitation>
:86
</treatmentCitation>
; 1913a:271; 1913b:59.
<treatmentCitation id="782F035F0B62C030FB8AFC2986E2FC19" author="Ho" box="[1058,1242,972,994]" page="47" pageId="15" pageNumber="16" year="1997">
<bibRefCitation id="9D1F58BF0B62C030FB8AFC29868CFC19" author="Ho" box="[1058,1204,972,994]" pageId="15" pageNumber="16" refString="Ho, J. Z. &amp; Jiang, B. H. (1997) Two firefly species with aquatic larvae in Taiwan. Conservation Quarterly, 17, 42 - 46." type="journal article" year="1997">
Ho
<emphasis id="CBFAF95C0B62C030FBEEFC29864AFC1A" box="[1094,1138,972,994]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. 1997
</bibRefCitation>
: 47
</treatmentCitation>
.
<treatmentCitation id="782F035F0B62C030FB4BFC29875CFC19" author="Ho" box="[1251,1380,971,994]" page="50" pageId="15" pageNumber="16" year="2002">
<bibRefCitation id="9D1F58BF0B62C030FB4BFC29877EFC19" author="Ho" box="[1251,1350,971,994]" pageId="15" pageNumber="16" refString="Ho, J. Z (2002) Larval morphology of twenty-one species and bionomics of fireflies (Coleoptera: Lampyridae) in Taiwan. Doctoral dissertation in the Department of Entomology, National Chung Hsing University Taiwan." type="book" year="2002">Ho, 2002</bibRefCitation>
:50
</treatmentCitation>
.
<treatmentCitation id="782F035F0B62C030FAC5FC29835BFBF8" author="Jeng" page="539" pageId="15" pageNumber="16" year="2003">
<bibRefCitation id="9D1F58BF0B62C030FAC5FC29830FFBF8" author="Jeng" pageId="15" pageNumber="16" refString="Jeng, M. L., Lai, J. &amp; Yang, P. S. (2003) Lampyridae: A synopsis of aquatic fireflies with description of a new species. In: Jach, M. A. &amp; Ji, L. (eds), Water Beetles of China Vol. III, Zoologisch-Botanische Gesellschaft in Osterreich, Wiener Coleopterologenverein pp. 539 - 562." type="book chapter" year="2003">
Jeng
<emphasis id="CBFAF95C0B62C030FF6FFC0F82C2FBF8" box="[199,250,1002,1024]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2003
</bibRefCitation>
:539
</treatmentCitation>
.
<treatmentCitation id="782F035F0B62C030FEC4FC0F8015FBF8" author="Fu" box="[364,557,1002,1024]" page="83" pageId="15" pageNumber="16" year="2005">
<bibRefCitation id="9D1F58BF0B62C030FEC4FC0F8030FBF8" author="Fu" box="[364,520,1002,1024]" pageId="15" pageNumber="16" refString="Fu, X. H., Ohba, N., Vencl, F. V., &amp; Lei, C. L. (2005 a) Structure, behavior and the life cycle of an aquatic firefly, Luciola substriata (Coleoptera: Lampyridae), in China. The Canadian Entomologist, 137, 83 - 90." type="journal article" year="2005" yearSuffix="a">
Fu
<emphasis id="CBFAF95C0B62C030FE25FC0F83F8FBF8" box="[397,448,1002,1024]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2005a
</bibRefCitation>
: 83
</treatmentCitation>
; 2005b:501; 2009: 243.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B19476C50B62C030FF3FFBC98317FB75" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="F931254E0B62C030FF3FFBC98317FB75" blockId="15.[151,1436,1069,1381]" pageId="15" pageNumber="16">
<emphasis id="CBFAF95C0B62C030FF3FFBC983DEFBBE" bold="true" box="[151,486,1069,1094]" pageId="15" pageNumber="16">
Material examined.
<collectingCountry id="819965DE0B62C030FE21FBC983DEFBBE" box="[393,486,1069,1094]" name="China" pageId="15" pageNumber="16">CHINA</collectingCountry>
</emphasis>
: Eleven first and six sixth instars bred from an original population of
<specimenCount id="EF88EEC70B62C030FAAEFBC98754FBBD" box="[1286,1388,1069,1093]" pageId="15" pageNumber="16" type="male">12 males</specimenCount>
and four females collected from Xianjian village, Hongshan District, Wuhan City, Hubei Province.
<date id="8D30038E0B62C030FB53FBB587A4FB91" box="[1275,1436,1105,1129]" pageId="15" pageNumber="16" value="2008-07-23">July 23, 2008</date>
(NHMHAU).
</paragraph>
</subSubSection>
<subSubSection id="B19476C50B62C030FF6FFB7D801EFB01" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="F931254E0B62C030FF6FFB7D801EFB01" blockId="15.[151,1436,1069,1381]" pageId="15" pageNumber="16">
<emphasis id="CBFAF95C0B62C030FF6FFB7D8379FB4A" bold="true" box="[199,321,1177,1202]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
Dark brown well sclerotised metapneustic larvae with all body plates closely adpressed; mandibles lacking teeth; apical labial and maxillary palpomeres with sense organs along ventral surfaces; not possible to distinguish from larvae of
<taxonomicName id="3E8E5ECD0B62C030FE37FB068027FB01" box="[415,543,1249,1273]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="aquatilis">
<emphasis id="CBFAF95C0B62C030FE37FB068027FB01" box="[415,543,1249,1273]" italics="true" pageId="15" pageNumber="16">L. aquatilis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B19476C50B62C030FF6FFAE181A8FA9D" pageId="15" pageNumber="16" type="discussion">
<paragraph id="F931254E0B62C030FF6FFAE181A8FA9D" blockId="15.[151,1436,1069,1381]" pageId="15" pageNumber="16">
<emphasis id="CBFAF95C0B62C030FF6FFAE18303FAE5" bold="true" box="[199,315,1285,1309]" pageId="15" pageNumber="16">Remarks.</emphasis>
<bibRefCitation id="9D1F58BF0B62C030FEEAFAE18090FAE5" author="Ballantyne" box="[322,680,1285,1309]" pageId="15" pageNumber="16" refString="Ballantyne, L. A. &amp; Lambkin, C. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne and Lambkin (2009)</bibRefCitation>
scored two different populations of males of this species from both mainland
<collectingCountry id="819965DE0B62C030FEADFACD8370FAB9" box="[261,328,1321,1345]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
and offshore
<collectingCountry id="819965DE0B62C030FE49FACD801CFAB9" box="[481,548,1321,1345]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
(
<collectingCountry id="819965DE0B62C030FD9AFACD80BFFAB9" box="[562,647,1321,1345]" name="Taiwan" pageId="15" pageNumber="16">Taiwan</collectingCountry>
) with results indicating morphological distinctiveness and two distinct species in an unresolved species complex within
<taxonomicName id="3E8E5ECD0B62C030FD14FAA981B1FA9D" box="[700,905,1357,1381]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="substriata">
<emphasis id="CBFAF95C0B62C030FD14FAA981B1FA9D" box="[700,905,1357,1381]" italics="true" pageId="15" pageNumber="16">Luciola substriata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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