454 lines
55 KiB
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454 lines
55 KiB
XML
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<mods:title id="9E4E87B89478EB8BD00791C369CF9F7D">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<treatment id="710D87BB291F1602D6BDFECCFE9BFCE7" ID-DOI="http://doi.org/10.5281/zenodo.5491514" ID-GBIF-Taxon="119590154" ID-Zenodo-Dep="5491514" LSID="urn:lsid:plazi:treatment:710D87BB291F1602D6BDFECCFE9BFCE7" httpUri="http://treatment.plazi.org/id/710D87BB291F1602D6BDFECCFE9BFCE7" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
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<subSubSection id="B1BE6526291F1601D6BDFECCFE89FE7B" pageId="9" pageNumber="310" type="nomenclature">
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<paragraph id="F91B36AD291F1601D6BDFECCFCB8FE59" blockId="9.[151,873,367,427]" box="[151,873,367,394]" pageId="9" pageNumber="310">
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<heading id="A25381C1291F1601D6BDFECCFCB8FE59" bold="true" box="[151,873,367,394]" fontSize="11" level="1" pageId="9" pageNumber="310" reason="1">
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<emphasis id="CBD0EABF291F1601D6BDFECCFCB8FE59" bold="true" box="[151,873,367,394]" pageId="9" pageNumber="310">
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<taxonomicName id="3EA44D2E291F1601D6BDFECCFDABFE5A" ID-CoL="6WN6V" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" box="[151,634,367,394]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="exilipes">
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<emphasis id="CBD0EABF291F1601D6BDFECCFE77FE59" bold="true" box="[151,422,367,393]" italics="true" pageId="9" pageNumber="310">Ptychophallus exilipes</emphasis>
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(
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<bibRefCitation id="9D354B5C291F1601D79FFED3FDA3FE5A" author="Rathbun" box="[437,626,368,394]" pageId="9" pageNumber="310" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
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)
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</taxonomicName>
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,
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<emphasis id="CBD0EABF291F1601D4ADFECCFCB8FE59" bold="true" box="[647,873,367,393]" italics="true" pageId="9" pageNumber="310">species inquirenda</emphasis>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="F91B36AD291F1601D6BDFE31FE89FE7B" blockId="9.[151,873,367,427]" box="[151,344,402,427]" pageId="9" pageNumber="310">
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(
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<figureCitation id="619F2A28291F1601D6B5FE31FEF6FE7B" box="[159,295,402,427]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="9" pageNumber="310">Figs. 22–30</figureCitation>
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,
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<figureCitation id="619F2A28291F1601D718FE30FE9EFE7B" box="[306,335,403,427]" captionStart="FIGURE 80" captionStartId="34.[151,250,1534,1556]" captionTargetBox="[163,1432,524,1513]" captionTargetId="figure@34.[150,1436,519,1513]" captionTargetPageId="34" captionText="FIGURE 80. Distribution map of Ptychophallus costaricensis Villalobos, 1974, and P. exilipes (Rathbun, 1898)." httpUri="https://zenodo.org/record/235138/files/figure.png" pageId="9" pageNumber="310">80</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B1BE6526291F1601D6BDFE79FBF7FD78" pageId="9" pageNumber="310" type="reference_group">
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<paragraph id="F91B36AD291F1601D6BDFE79FDE4FDDF" blockId="9.[151,1436,474,680]" pageId="9" pageNumber="310">
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<treatmentCitationGroup id="D9B41183291F1601D6BDFE79FDE4FDDF" pageId="9" pageNumber="310">
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<taxonomicName id="3EA44D2E291F1601D6BDFE79FDBDFE20" ID-CoL="4PD8Z" authority="Rathbun, 1898: 514" authorityName="Rathbun" authorityPageNumber="514" authorityYear="1898" box="[151,620,474,496]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="exilipes">
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<emphasis id="CBD0EABF291F1601D6BDFE79FE49FE20" box="[151,408,474,496]" italics="true" pageId="9" pageNumber="310">Pseudothelphusa exilipes</emphasis>
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<treatmentCitation id="780510BC291F1601D7B5FE79FDBDFE20" author="Rathbun" box="[415,620,474,496]" page="514" pageId="9" pageNumber="310" year="1898">
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<bibRefCitation id="9D354B5C291F1601D7B5FE79FDE8FE20" author="Rathbun" box="[415,569,474,496]" pageId="9" pageNumber="310" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
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: 514
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</treatmentCitation>
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</taxonomicName>
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, 534, fig. 1.—
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||
<treatmentCitation id="780510BC291F1601D4D1FE79FC7EFE20" author="Young" box="[763,943,474,496]" page="211" pageId="9" pageNumber="310" year="1900">
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||
<bibRefCitation id="9D354B5C291F1601D4D1FE79FC51FE20" author="Young" box="[763,896,474,496]" pageId="9" pageNumber="310" refString="Young, C. G. (1900) The stalk-eyed Crustacea of British Guiana, West Indies, and Bermuda. J. M. Watkins, London, 514 pp., 4 pls." type="book" year="1900">Young 1900</bibRefCitation>
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: 211
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</treatmentCitation>
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||
.—
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||
<treatmentCitation id="780510BC291F1601D5E7FE79FB45FE20" author="Rathbun" box="[973,1172,474,496]" page="242" pageId="9" pageNumber="310" year="1904">
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<bibRefCitation id="9D354B5C291F1601D5E7FE79FBB2FE20" author="Rathbun" box="[973,1123,474,496]" pageId="9" pageNumber="310" refString="Rathbun, M. J. (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle Paris, 6 (4), 225 - 312, text-figs. 1 - 37, volume pls. 9 - 18 (pls. I - X for Rathbun's article)." type="journal article" year="1904">Rathbun 1904</bibRefCitation>
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: 242
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||
</treatmentCitation>
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||
(in list).—
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||
<treatmentCitation id="780510BC291F1601D328FE79FF3CFDDF" author="Rathbun" page="279" pageId="9" pageNumber="310" year="1905">
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||
<bibRefCitation id="9D354B5C291F1601D328FE79FA49FE20" author="Rathbun" box="[1282,1432,474,496]" pageId="9" pageNumber="310" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905">Rathbun 1905</bibRefCitation>
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||
: 279
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||
</treatmentCitation>
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||
.—
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||
<treatmentCitation id="780510BC291F1601D720FE5AFE0FFDDF" author="Coifmann" box="[266,478,505,527]" page="107" pageId="9" pageNumber="310" year="1939">
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||
<bibRefCitation id="9D354B5C291F1601D720FE5AFE7FFDDF" author="Coifmann" box="[266,430,505,527]" pageId="9" pageNumber="310" refString="Coifmann, I. (1939) Potamonidi della Guiana inglese raccolti dal Prof. Nello Beccari. Archivio Zoologico Italiano, 27, 93 - 116, pl. 3." type="journal article" year="1939">Coifmann 1939</bibRefCitation>
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: 107
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</treatmentCitation>
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(in list).
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="F91B36AD291F1601D6BDFDBBFC4DFDFE" blockId="9.[151,1436,474,680]" box="[151,924,536,558]" pageId="9" pageNumber="310">
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<treatmentCitationGroup id="D9B41183291F1601D6BDFDBBFC4DFDFE" box="[151,924,536,558]" pageId="9" pageNumber="310">
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<taxonomicName id="3EA44D2E291F1601D6BDFDBBFDEEFDFE" ID-CoL="4PD8Z" box="[151,575,536,558]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="exilipes" subGenus="Ptychophallus">
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<emphasis id="CBD0EABF291F1601D6BDFDBBFE95FDFE" box="[151,324,536,558]" italics="true" pageId="9" pageNumber="310">Pseudothelphusa</emphasis>
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(
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<emphasis id="CBD0EABF291F1601D779FDBBFE35FDFE" box="[339,484,536,558]" italics="true" pageId="9" pageNumber="310">Ptychophallus</emphasis>
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)
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<emphasis id="CBD0EABF291F1601D7D8FDBBFDEEFDFE" box="[498,575,536,558]" italics="true" pageId="9" pageNumber="310">exilipes</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="780510BC291F1601D470FDBBFCC8FDFE" author="Smalley" box="[602,793,536,558]" page="11" pageId="9" pageNumber="310" year="1964">
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<bibRefCitation id="9D354B5C291F1601D470FDBBFD24FDFE" author="Smalley" box="[602,757,536,558]" pageId="9" pageNumber="310" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley 1964b</bibRefCitation>
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: 11
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</treatmentCitation>
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, figs. 13, 14.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="F91B36AD291F1601D6BDFD95FEF6FDBB" blockId="9.[151,1436,474,680]" pageId="9" pageNumber="310">
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<treatmentCitationGroup id="D9B41183291F1601D6BDFD95FEF6FDBB" pageId="9" pageNumber="310">
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<taxonomicName id="3EA44D2E291F1601D6BDFD95FDFBFD9C" ID-CoL="6WN6V" box="[151,554,566,588]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="exilipes" subGenus="Ptychophallus">
|
||
<emphasis id="CBD0EABF291F1601D6BDFD95FEF9FD9C" box="[151,296,566,588]" italics="true" pageId="9" pageNumber="310">Ptychophallus</emphasis>
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||
(
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<emphasis id="CBD0EABF291F1601D710FD95FE1AFD9C" box="[314,459,566,588]" italics="true" pageId="9" pageNumber="310">Ptychophallus</emphasis>
|
||
)
|
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<emphasis id="CBD0EABF291F1601D7F7FD95FDFBFD9C" box="[477,554,566,588]" italics="true" pageId="9" pageNumber="310">exilipes</emphasis>
|
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</taxonomicName>
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||
—
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||
<treatmentCitation id="780510BC291F1601D46FFD94FCDEFD9C" author="Pretzmann" box="[581,783,566,588]" page="5" pageId="9" pageNumber="310" year="1965">
|
||
<bibRefCitation id="9D354B5C291F1601D46FFD94FD26FD9C" author="Pretzmann" box="[581,759,566,588]" pageId="9" pageNumber="310" refString="Pretzmann, G. (1965) Vorlaufiger Bericht uber die Familie Pseudothelphusidae. Sitzungsberichte der Mathematisch- Naturwissenschaftlichen Klasse der Osterreichischen Akademie der Wissenschaften, Wien, 1, 1 - 11." type="journal article" year="1965">Pretzmann, 1965</bibRefCitation>
|
||
: 5
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||
</treatmentCitation>
|
||
.—
|
||
<treatmentCitation id="780510BC291F1601D506FD95FBD0FD9C" author="Pretzmann" box="[812,1025,566,588]" page="21" pageId="9" pageNumber="310" year="1971">
|
||
<bibRefCitation id="9D354B5C291F1601D506FD95FC0CFD9C" author="Pretzmann" box="[812,989,566,588]" pageId="9" pageNumber="310" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche Klasse der Osterreichischen Akademie der Wissenschaften, Series 1, 179 (1 - 4), 12 - 24." type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
|
||
: 21
|
||
</treatmentCitation>
|
||
.—
|
||
<treatmentCitation id="780510BC291F1601D235FD95FB29FD9C" author="Pretzmann" box="[1055,1272,566,588]" page="87" pageId="9" pageNumber="310" year="1972">
|
||
<bibRefCitation id="9D354B5C291F1601D235FD95FB01FD9C" author="Pretzmann" box="[1055,1232,566,588]" pageId="9" pageNumber="310" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
|
||
: 87
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||
</treatmentCitation>
|
||
, figs. 508–510, 522–524.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="F91B36AD291F1601D6BDFDD7FBF7FD78" blockId="9.[151,1436,474,680]" pageId="9" pageNumber="310">
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<treatmentCitationGroup id="D9B41183291F1601D6BDFDD7FBF7FD78" pageId="9" pageNumber="310">
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<taxonomicName id="3EA44D2E291F1601D6BDFDD7FE53FD5A" ID-CoL="6WN6V" box="[151,386,628,650]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="exilipes">
|
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<emphasis id="CBD0EABF291F1601D6BDFDD7FE53FD5A" box="[151,386,628,650]" italics="true" pageId="9" pageNumber="310">Ptychophallus exilipes</emphasis>
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</taxonomicName>
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||
—
|
||
<treatmentCitation id="780510BC291F1601D7B7FDD7FDA2FD5A" author="Rodriguez" box="[413,627,628,650]" page="85" pageId="9" pageNumber="310" year="1982">
|
||
<bibRefCitation id="9D354B5C291F1601D7B7FDD7FD98FD5A" author="Rodriguez" box="[413,585,628,650]" pageId="9" pageNumber="310" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez 1982</bibRefCitation>
|
||
: 85
|
||
</treatmentCitation>
|
||
.—
|
||
<treatmentCitation id="780510BC291F1601D4BBFDD7FC3CFD5A" author="Campos" box="[657,1005,628,650]" page="560" pageId="9" pageNumber="310" year="1999">
|
||
<bibRefCitation id="9D354B5C291F1601D4BBFDD7FC67FD5A" author="Campos" box="[657,950,628,650]" pageId="9" pageNumber="310" refString="Campos, M. R. & Lemaitre, R. (1999) Two new freshwater crabs of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) from Panama, with notes on the distribution of the genus. Proceedings of the Biological Society of Washington, 112 (3), 553 - 561." type="journal article" year="1999">Campos & Lemaitre 1999</bibRefCitation>
|
||
: 560
|
||
</treatmentCitation>
|
||
(map).—
|
||
<treatmentCitation id="780510BC291F1601D27EFDD7FF3FFD78" author="Rodriguez" page="423" pageId="9" pageNumber="310" year="2000">
|
||
<bibRefCitation id="9D354B5C291F1601D27EFDD7FA48FD5A" author="Rodriguez" box="[1108,1433,628,650]" pageId="9" pageNumber="310" refString="Rodriguez, G. & Hedstrom, I. (2000) The freswater crabs of the Barbilla National Park, Costa Rica (Crustacea: Brachyura: Pseudothelphusidae), with notes on the evolution of structures for spermatophore retention. Proceedings of the Biological Society of Washington, 113 (2), 420 - 425." type="journal article" year="2000">Rodríguez & Hedström 2000</bibRefCitation>
|
||
: 423
|
||
</treatmentCitation>
|
||
.—
|
||
<treatmentCitation id="780510BC291F1601D721FD31FE00FD78" author="Ng" box="[267,465,658,680]" page="176" pageId="9" pageNumber="310" year="2008">
|
||
<bibRefCitation id="9D354B5C291F1601D721FD31FE70FD78" author="Ng" box="[267,417,658,680]" pageId="9" pageNumber="310" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
|
||
Ng
|
||
<emphasis id="CBD0EABF291F1601D718FD30FEB4FD78" box="[306,357,658,680]" italics="true" pageId="9" pageNumber="310">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
: 176
|
||
</treatmentCitation>
|
||
(in list).—
|
||
<treatmentCitation id="780510BC291F1601D417FD31FC1EFD78" author="Villalobos" box="[573,975,658,680]" page="297" pageId="9" pageNumber="310" year="2008">
|
||
<bibRefCitation id="9D354B5C291F1601D417FD31FC4EFD78" author="Villalobos" box="[573,927,658,680]" pageId="9" pageNumber="310" refString="Villalobos Hiriart, J. L. & Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. & Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart & Álvarez 2008</bibRefCitation>
|
||
: 297
|
||
</treatmentCitation>
|
||
(in list).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291F1601D6BDFD76FB4CFB36" pageId="9" pageNumber="310" type="materials_examined">
|
||
<paragraph id="F91B36AD291F1601D6BDFD76FB4CFB36" blockId="9.[151,1437,725,2010]" pageId="9" pageNumber="310">
|
||
<emphasis id="CBD0EABF291F1601D6BDFD76FEABFD3E" bold="true" box="[151,378,725,750]" pageId="9" pageNumber="310">Material examined</emphasis>
|
||
.
|
||
<collectingCountry id="81B3763D291F1601D7A0FD75FDE2FD3E" box="[394,563,726,750]" name="Costa Rica" pageId="9" pageNumber="310">COSTA RICA</collectingCountry>
|
||
. San José Province, Atlantic drainage:
|
||
<specimenCount id="EFA2FD24291F1601D5DBFD75FB87FD3E" box="[1009,1110,725,750]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<typeStatus id="261F880F291F1601D249FD76FB19FD3E" box="[1123,1224,725,750]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D2FEFD75FAF4FD3E" LSID="urn:lsid:biocol.org:col:34871" box="[1236,1317,726,750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="310">USNM</collectionCode>
|
||
19488, El Coronel,
|
||
<quantity id="3E5C9B48291F1601D728FD59FE9CFCC1" box="[258,333,762,786]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="310" unit="m" value="700.0">700 m</quantity>
|
||
, undated, P. Biolley & E. Fernández leg.—San José Province, Pacific drainage:
|
||
<specimenCount id="EFA2FD24291F1601D2C7FD59FAECFCC2" box="[1261,1341,761,786]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D361FD59FA4DFCC2" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,762,786]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="310">USNM</collectionCode>
|
||
39096, Cangrejal de Aserrí, Pacific drainage,
|
||
<quantity id="3E5C9B48291F1601D4BFFCBDFD0FFCE5" box="[661,734,798,822]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="310" unit="m" value="800.0">800 m</quantity>
|
||
,
|
||
<date id="8D1A106D291F1601D4C3FCBEFCEEFCE6" box="[745,831,797,822]" pageId="9" pageNumber="310" value="1906-04">iv.1906</date>
|
||
, J.F.
|
||
<collectingCountry id="81B3763D291F1601D55DFCBEFC14FCE6" box="[887,965,797,822]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="9" pageNumber="310">Tristan</collectingCountry>
|
||
leg.;
|
||
<specimenCount id="EFA2FD24291F1601D229FCBDFB80FCE6" box="[1027,1105,797,822]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D276FCBDFB7CFCE6" LSID="urn:lsid:biocol.org:col:34871" box="[1116,1197,798,822]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="310">USNM</collectionCode>
|
||
39089, Dota Canton, San José, Santa María de Dota,
|
||
<quantity id="3E5C9B48291F1601D7D3FCE2FD81FC89" box="[505,592,833,858]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="310" unit="m" value="1600.0">1600 m</quantity>
|
||
,
|
||
<date id="8D1A106D291F1601D471FCE2FD75FC8A" box="[603,676,833,858]" pageId="9" pageNumber="310" value="1907-01">i.1907</date>
|
||
, P. Biolley leg.;
|
||
<specimenCount id="EFA2FD24291F1601D571FCE1FC78FC8A" box="[859,937,833,858]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="EFA2FD24291F1601D59EFCE1FBF3FC8A" box="[948,1058,833,858]" pageId="9" pageNumber="310" type="female">4 females</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D207FCE1FBAFFC8A" LSID="urn:lsid:biocol.org:col:34871" box="[1069,1150,834,858]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="310">USNM</collectionCode>
|
||
39089 (sic), same data;
|
||
<specimenCount id="EFA2FD24291F1601D3A4FCE1FF01FCAE" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
, UCR-MZ 742-02, Los Tubos Creek,
|
||
<quantity id="3E5C9B48291F1601D4A8FCC5FD6DFCAE" box="[642,700,869,894]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="310" unit="km" value="1.0">1 km</quantity>
|
||
W Boquete, Adan Bonilla’s farm (San Isidro del El General), V. Juárez leg.;
|
||
<specimenCount id="EFA2FD24291F1601D734FC29FEBCFC72" box="[286,365,905,930]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D753FC29FE69FC72" box="[377,440,906,930]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="310">INPA</collectionCode>
|
||
1857, same data;
|
||
<specimenCount id="EFA2FD24291F1601D4AFFC29FD0FFC72" box="[645,734,905,930]" pageId="9" pageNumber="310" type="male">3 males</specimenCount>
|
||
, UCR-MZ 737, Pérez Zeledón Canton, Painer, creek
|
||
<quantity id="3E5C9B48291F1601D313FC29FA50FC72" box="[1337,1409,905,930]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="310" unit="km" value="14.0">14 km</quantity>
|
||
N San Isidro,
|
||
<date id="8D1A106D291F1601D732FC0EFE5DFC16" box="[280,396,941,966]" pageId="9" pageNumber="310" value="1972-02-26">26.ii.1972</date>
|
||
, V. Suarez leg.;
|
||
<specimenCount id="EFA2FD24291F1601D46EFC0DFD42FC16" box="[580,659,941,966]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="EFA2FD24291F1601D4B5FC0DFCD5FC16" box="[671,772,941,966]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
, UCR-MZ 739, Talari River, Los Ángeles de Canaan, road to Chirripó,
|
||
<date id="8D1A106D291F1601D70DFC72FE4FFC3A" box="[295,414,977,1002]" pageId="9" pageNumber="310" value="1972-04-11">11.iv.1972</date>
|
||
, V. Juárez leg.;
|
||
<specimenCount id="EFA2FD24291F1601D471FC71FD66FC3A" box="[603,695,977,1002]" pageId="9" pageNumber="310" type="male">3 males</specimenCount>
|
||
, UCR-MZ 741, Perez Zeledón Canton, creek
|
||
<quantity id="3E5C9B48291F1601D2E4FC71FADBFC3A" box="[1230,1290,977,1002]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="310" unit="km" value="4.0">4 km</quantity>
|
||
N (La Ese),
|
||
<date id="8D1A106D291F1601D6BDFC56FED0FBDE" box="[151,257,1013,1038]" pageId="9" pageNumber="310" value="1972-04-07">7.iv.1972</date>
|
||
, V. Juárez leg.; 1 right G1,
|
||
<specimenCount id="EFA2FD24291F1601D414FC55FD72FBDE" box="[574,675,1013,1038]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
, UCR-MZ 2784, grassland between Peje River and road, El Joroz, approximately
|
||
<quantity id="3E5C9B48291F1601D715FBBAFEA8FBE2" box="[319,377,1049,1074]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="310" unit="km" value="6.0">6 km</quantity>
|
||
S San Isidro del General,
|
||
<date id="8D1A106D291F1601D4B5FBB9FCC1FBE2" box="[671,784,1050,1074]" pageId="9" pageNumber="310" value="1972-05-14">14.v.1972</date>
|
||
, J.E. Baldrige leg.;
|
||
<specimenCount id="EFA2FD24291F1601D5C1FBB9FBEBFBE2" box="[1003,1082,1049,1074]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="EFA2FD24291F1601D26FFBB9FB78FBE2" box="[1093,1193,1049,1074]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D29EFBB9FB22FBE2" box="[1204,1267,1050,1074]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="310">INPA</collectionCode>
|
||
1846, forest in Frank Weber’s farm, approximately
|
||
<quantity id="3E5C9B48291F1601D406FB9DFDBBFB86" box="[556,618,1085,1110]" metricMagnitude="4" metricUnit="m" metricValue="1.7702784" pageId="9" pageNumber="310" unit="mi" value="11.0">11 mi</quantity>
|
||
S San Isidro del General,
|
||
<date id="8D1A106D291F1601D5A4FB9EFC35FB86" box="[910,996,1085,1110]" pageId="9" pageNumber="310" value="1972-06">vi.1972</date>
|
||
, J.E. Baldrige leg.—Alajuela Province, Atlantic drainage:
|
||
<specimenCount id="EFA2FD24291F1601D758FBC1FE14FBAA" box="[370,453,1121,1146]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="EFA2FD24291F1601D7FCFBC1FDEFFBAA" box="[470,574,1121,1146]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D465FBC1FD4AFBAA" box="[591,667,1122,1146]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="9" pageNumber="310">CCDB</collectionCode>
|
||
4538, Paradero 1, unnamed creek, Balsa River tributary,
|
||
<collectionCode id="9FB5AE68291F1601D367FBC1FA4DFBAA" box="[1357,1436,1122,1146]" pageId="9" pageNumber="310">CTRM</collectionCode>
|
||
480153/
|
||
<date id="8D1A106D291F1601D6DEFB26FE98FB4E" box="[244,329,1157,1182]" pageId="9" pageNumber="310" value="28-12-25">251228</date>
|
||
,
|
||
<date id="8D1A106D291F1601D777FB25FE10FB4E" box="[349,449,1158,1182]" pageId="9" pageNumber="310" value="2010-10-07">7.x.2010</date>
|
||
, L.R. Lara leg.—
|
||
<taxonomicName id="3EA44D2E291F1601D4B3FB26FCC6FB4E" box="[665,791,1157,1182]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
|
||
Province, Pacific drainage:
|
||
<specimenCount id="EFA2FD24291F1601D259FB25FB18FB4E" box="[1139,1225,1157,1182]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D2F6FB25FAC5FB4E" LSID="urn:lsid:biocol.org:col:34838" box="[1244,1300,1158,1182]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="310">SMF</collectionCode>
|
||
36264, La Cusinga, near Uvita,
|
||
<geoCoordinate id="9C90506A291F1601D7A1FB09FDC7FB12" box="[395,534,1194,1218]" direction="north" orientation="latitude" pageId="9" pageNumber="310" precision="1" value="9.135917">9°08'09.3"N</geoCoordinate>
|
||
<geoCoordinate id="9C90506A291F1601D435FB09FD46FB12" box="[543,663,1194,1218]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="92" value="-83.72">83°43.2'W</geoCoordinate>
|
||
,
|
||
<date id="8D1A106D291F1601D48EFB0AFCC0FB12" box="[676,785,1193,1218]" pageId="9" pageNumber="310" value="2008-01-18">18.i.2008</date>
|
||
, G. Köhler leg.;
|
||
<specimenCount id="EFA2FD24291F1601D5FBFB09FBF0FB12" box="[977,1057,1193,1218]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="9FB5AE68291F1601D205FB09FBB6FB12" LSID="urn:lsid:biocol.org:col:34838" box="[1071,1127,1194,1218]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="9" pageNumber="310">SMF</collectionCode>
|
||
36265, Coto Brus Canton, Estación Biológica Las Cruces,
|
||
<geoCoordinate id="9C90506A291F1601D7D7FB6DFD50FB36" box="[509,641,1230,1254]" direction="north" orientation="latitude" pageId="9" pageNumber="310" precision="15" value="8.785833">08°47'09"N</geoCoordinate>
|
||
<geoCoordinate id="9C90506A291F1601D4A2FB6EFCC2FB36" box="[648,787,1229,1254]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="15" value="-82.96027">82°57'37"W</geoCoordinate>
|
||
,
|
||
<quantity id="3E5C9B48291F1601D534FB6DFCA4FB35" box="[798,885,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="9" pageNumber="310" unit="m" value="1220.0">1220 m</quantity>
|
||
,
|
||
<date id="8D1A106D291F1601D5AAFB6EFC24FB36" box="[896,1013,1229,1254]" pageId="9" pageNumber="310" value="2007-02-15">15.ii.2007</date>
|
||
, G. Köhler leg.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291F1601D6EDFB52FDCDF9FA" pageId="9" pageNumber="310" type="description">
|
||
<paragraph id="F91B36AD291F1601D6EDFB52FDCDF9FA" blockId="9.[151,1437,725,2010]" pageId="9" pageNumber="310">
|
||
<emphasis id="CBD0EABF291F1601D6EDFB52FE3CFADA" bold="true" box="[199,493,1265,1290]" pageId="9" pageNumber="310">Description of gonopod</emphasis>
|
||
. Straight in mesiocaudal view, apex bent approximately 90° in laterocephalic direction. Marginal suture on mesial side, straight; marginal process subtriangular, not produced beyond caudal border of apex. Mesial process small, subtriangular, usually strongly downturned. Lateral process well developed, slightly bilobed, approximately 50–60% of stem length (from proximal opening to caudal border of apex), with shallow depression around middle, usually distinctly wider than lateral end of apex; lateral border nearly straight or slightly bilobed, with proximal lobe rounded, usually larger, wider, distal lobe thicker distally. Distocaudal ridge short, narrow, separated from distal border of lateral process by distinct depression. Apex oblong, slightly oblique, narrow; caudal border slightly convex, with small notch near lateral end. Field of apical spines well developed, facing towards cephalolateral side.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291F1601D6EDF996FCF8F946" pageId="9" pageNumber="310" type="materials_examined">
|
||
<paragraph id="F91B36AD291F1601D6EDF996FCF8F946" blockId="9.[151,1437,725,2010]" pageId="9" pageNumber="310">
|
||
<emphasis id="CBD0EABF291F1601D6EDF996FDDEF99E" bold="true" box="[199,527,1589,1614]" pageId="9" pageNumber="310">
|
||
<typeStatus id="261F880F291F1601D6EDF996FEE5F99E" box="[199,308,1589,1614]" pageId="9" pageNumber="310" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="261F880F291F1601D750F996FE7CF99E" box="[378,429,1589,1614]" pageId="9" pageNumber="310">type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
. Female, cw 24.0, cl 14.3,
|
||
<collectionCode id="9FB5AE68291F1601D54BF995FC63F99E" LSID="urn:lsid:biocol.org:col:34871" box="[865,946,1590,1614]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="310">USNM</collectionCode>
|
||
19488.
|
||
<collectingCountry id="81B3763D291F1601D233F995FB4AF99E" box="[1049,1179,1589,1614]" name="Costa Rica" pageId="9" pageNumber="310">Costa Rica</collectingCountry>
|
||
, El Coronel (Atlantic drainage). According to
|
||
<bibRefCitation id="9D354B5C291F1601D79AF9FAFDA1F9A2" author="Smalley" box="[432,624,1625,1650]" pageId="9" pageNumber="310" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley (1964b)</bibRefCitation>
|
||
, this locality is probably the one situated on the Sucio River,
|
||
<geoCoordinate id="9C90506A291F1601D311F9F9FA4DF9A2" box="[1339,1436,1626,1650]" direction="north" orientation="latitude" pageId="9" pageNumber="310" precision="925" value="10.116667">10°07’N</geoCoordinate>
|
||
<geoCoordinate id="9C90506A291F1601D6BDF9DEFF2CF946" box="[151,253,1661,1686]" direction="west" orientation="longitude" pageId="9" pageNumber="310" precision="925" value="-83.916664">83°55’W</geoCoordinate>
|
||
, at the border of San José and Cartago Provinces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291F1601D6EDF902FF12F8D2" pageId="9" pageNumber="310" type="distribution">
|
||
<paragraph id="F91B36AD291F1601D6EDF902FF12F8D2" blockId="9.[151,1437,725,2010]" pageId="9" pageNumber="310">
|
||
<emphasis id="CBD0EABF291F1601D6EDF902FE88F96A" bold="true" box="[199,345,1697,1722]" pageId="9" pageNumber="310">Distribution</emphasis>
|
||
. Except for two records (including the
|
||
<typeStatus id="261F880F291F1601D56EF901FCA5F96A" box="[836,884,1698,1722]" pageId="9" pageNumber="310">type</typeStatus>
|
||
locality), all the available records point to a distribution on the Pacific drainage of central and southern
|
||
<collectingCountry id="81B3763D291F1601D56BF965FC11F90E" box="[833,960,1733,1758]" name="Costa Rica" pageId="9" pageNumber="310">Costa Rica</collectingCountry>
|
||
, San José and
|
||
<taxonomicName id="3EA44D2E291F1601D241F965FB37F90D" box="[1131,1254,1734,1757]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
|
||
Provinces (
|
||
<figureCitation id="619F2A28291F1601D345F966FF66F8D2" captionStart="FIGURE 80" captionStartId="34.[151,250,1534,1556]" captionTargetBox="[163,1432,524,1513]" captionTargetId="figure@34.[150,1436,519,1513]" captionTargetPageId="34" captionText="FIGURE 80. Distribution map of Ptychophallus costaricensis Villalobos, 1974, and P. exilipes (Rathbun, 1898)." httpUri="https://zenodo.org/record/235138/files/figure.png" pageId="9" pageNumber="310">Fig. 80</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE6526291F1602D6EDF8AEFE9BFCE7" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310" type="discussion">
|
||
<paragraph id="F91B36AD291F1602D6EDF8AEFA91FF28" blockId="9.[151,1437,725,2010]" lastBlockId="10.[151,1437,151,824]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
|
||
<emphasis id="CBD0EABF291F1601D6EDF8AEFEE5F8F6" bold="true" box="[199,308,1805,1830]" pageId="9" pageNumber="310">Remarks</emphasis>
|
||
. The USNM has two jars with the same catalogue number (39089) from Santa María de Dota, one with a single male (cw 23.8, cl 14.3) and the other with one male and four females.
|
||
<bibRefCitation id="9D354B5C291F1601D24AF892FACCF89A" author="Smalley" box="[1120,1309,1841,1866]" pageId="9" pageNumber="310" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley (1964b)</bibRefCitation>
|
||
assigned a male from this lot to the taxon, which was identified with the
|
||
<typeStatus id="261F880F291F1601D58CF8F5FC30F8BE" box="[934,993,1878,1902]" pageId="9" pageNumber="310">types</typeStatus>
|
||
by M.-J. Rathbun.
|
||
<bibRefCitation id="9D354B5C291F1601D2F3F8F6FA4DF8BE" author="Smalley" box="[1241,1436,1877,1902]" pageId="9" pageNumber="310" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley (1964b)</bibRefCitation>
|
||
nevertheless remarked that collections made only six miles from the
|
||
<typeStatus id="261F880F291F1601D5BDF8D9FC16F842" box="[919,967,1914,1938]" pageId="9" pageNumber="310">type</typeStatus>
|
||
locality consisted only of specimens of
|
||
<taxonomicName id="3EA44D2E291F1601D3ADF8D9FEC3F865" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="tumimanus">
|
||
<emphasis id="CBD0EABF291F1601D3ADF8D9FEC3F865" italics="true" pageId="9" pageNumber="310">P. tumimanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3EA44D2E291F1601D77FF83DFE0FF865" box="[341,478,1950,1973]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="montatus">
|
||
<emphasis id="CBD0EABF291F1601D77FF83DFE0FF865" box="[341,478,1950,1973]" italics="true" pageId="9" pageNumber="310">P. montatus</emphasis>
|
||
</taxonomicName>
|
||
and, therefore, males from the
|
||
<typeStatus id="261F880F291F1601D54FF83DFC44F866" box="[869,917,1950,1974]" pageId="9" pageNumber="310">type</typeStatus>
|
||
locality would be very valuable. We were unfortunately not able to obtain new collections from the
|
||
<typeStatus id="261F880F291F1601D51EF861FCB5F80A" box="[820,868,1986,2010]" pageId="9" pageNumber="310">type</typeStatus>
|
||
locality. Specimens collected at nearby locations from El Coronel examined in the present study belong to
|
||
<taxonomicName id="3EA44D2E291C1602D51AFF3AFC4EFF60" box="[816,927,153,176]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="tristani">
|
||
<emphasis id="CBD0EABF291C1602D51AFF3AFC4EFF60" box="[816,927,153,176]" italics="true" pageId="10" pageNumber="311">P. tristani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3EA44D2E291C1602D5F0FF3AFBB5FF60" box="[986,1124,153,176]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="CBD0EABF291C1602D5F0FF3AFBB5FF60" box="[986,1124,153,176]" italics="true" pageId="10" pageNumber="311">P. montanus</emphasis>
|
||
</taxonomicName>
|
||
. A male (cw 23.5, cl 14.3; USNM 39096) from Cangrejal de Aserrí (possibly Acosta Canton instead of Aserrí Canton), a locality about
|
||
<quantity id="3E5C9B48291C1602D37CFF1FFA4DFF05" box="[1366,1436,188,213]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="311" unit="km" value="50.0">50 km</quantity>
|
||
SW from the
|
||
<typeStatus id="261F880F291C1602D704FF43FE8FFF28" box="[302,350,224,248]" pageId="10" pageNumber="311">type</typeStatus>
|
||
locality, also shows a G1 similar to that of the male from Santa María de Dota (
|
||
<figureCitation id="619F2A28291C1602D2F5FF7CFAE2FF28" box="[1247,1331,223,248]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="10" pageNumber="311">Fig. 27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F91B36AD291C1602D6EDFEA7FA49FEB5" blockId="10.[151,1437,151,824]" pageId="10" pageNumber="311">
|
||
The Sucio River, the presumptive
|
||
<typeStatus id="261F880F291C1602D47DFEA6FD56FECD" box="[599,647,261,285]" pageId="10" pageNumber="311">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="3EA44D2E291C1602D538FEA6FC56FECD" box="[786,903,260,285]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="exilipes">
|
||
<emphasis id="CBD0EABF291C1602D538FEA6FC56FECD" box="[786,903,260,285]" italics="true" pageId="10" pageNumber="311">P. exilipes</emphasis>
|
||
</taxonomicName>
|
||
, is on the Atlantic drainage. Almost all other specimens examined in this study, however, came from rivers that drain into the Pacific. We consider this taxon a
|
||
<emphasis id="CBD0EABF291C1602D6BDFEEEFEB8FEB5" box="[151,361,332,357]" italics="true" pageId="10" pageNumber="311">species inquirenda</emphasis>
|
||
rather than a
|
||
<emphasis id="CBD0EABF291C1602D428FEEEFD79FEB5" box="[514,680,332,357]" italics="true" pageId="10" pageNumber="311">nomen dubium</emphasis>
|
||
status until further investigation helps establish its correct identity.
|
||
</paragraph>
|
||
<paragraph id="F91B36AD291C1602D6EDFECCFCC4FDC8" blockId="10.[151,1437,151,824]" pageId="10" pageNumber="311">
|
||
The shape of the lateral process of the G1 can vary to some extent. It is usually broadly subtriangular, with the proximal lobe larger and wider, which gives the lateral border a gentle asymmetric concave outline (e.g. SMF 36264,
|
||
<figureCitation id="619F2A28291C1602D6C1FE14FEEAFE00" box="[235,315,439,464]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="10" pageNumber="311">Fig. 30</figureCitation>
|
||
) or even this border be more or less convex: e.g. USNM 39096 (
|
||
<figureCitation id="619F2A28291C1602D225FE14FBB3FE00" box="[1039,1122,439,464]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="10" pageNumber="311">Fig. 27</figureCitation>
|
||
), SMF 36265 (
|
||
<figureCitation id="619F2A28291C1602D320FE14FA8FFE00" box="[1290,1374,439,464]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="10" pageNumber="311">Fig. 29</figureCitation>
|
||
). In a few specimens this concavity can be a little more pronounced, e.g. INPA 1857,
|
||
<figureCitation id="619F2A28291C1602D23AFE7FFBB2FE24" box="[1040,1123,476,501]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="10" pageNumber="311">Fig. 28</figureCitation>
|
||
). All of these variations can be found in the males of lot UCR-MZ 741 (
|
||
<figureCitation id="619F2A28291C1602D455FE5CFCDBFDC8" box="[639,778,511,536]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="10" pageNumber="311">Figs. 24–26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F91B36AD291C1602D6EDFD86FE9BFCE7" blockId="10.[151,1437,151,824]" pageId="10" pageNumber="311">
|
||
In spite of the great similarities between the gonopods of
|
||
<taxonomicName id="3EA44D2E291C1602D56EFD86FC65FDED" box="[836,948,548,573]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="exilipes">
|
||
<emphasis id="CBD0EABF291C1602D56EFD86FC65FDED" box="[836,948,548,573]" italics="true" pageId="10" pageNumber="311">P. exilipes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="CBD0EABF291C1602D5C3FD86FBA9FDEC" box="[1001,1144,548,573]" italics="true" pageId="10" pageNumber="311">P. l av a l l en s i s</emphasis>
|
||
we prefer to treat them as separate species based on their distribution (central-southern
|
||
<collectingCountry id="81B3763D291C1602D577FDEBFC0BFDB0" box="[861,986,583,608]" name="Costa Rica" pageId="10" pageNumber="311">Costa Rica</collectingCountry>
|
||
and central
|
||
<collectingCountry id="81B3763D291C1602D244FDEBFB1BFD8F" box="[1134,1226,584,607]" name="Panama" pageId="10" pageNumber="311">Panama</collectingCountry>
|
||
, respectively) and because of some of the gonopod characters, as already mentioned by
|
||
<bibRefCitation id="9D354B5C291C1602D583FDCFFBA2FD54" author="Rodriguez" box="[937,1139,620,645]" pageId="10" pageNumber="311" refString="Rodriguez, G. (1994) A revision of the type material of some species of Hypolobocera and Ptychophallus (Crustacea: Decapoda: Pseudothelphusidae) in the National Museum of Natural History, Washington, D. C., with descriptions of a new species and a new subspecies. Proceedings of the Biological Society of Washington, 107 (2), 296 - 307." type="journal article" year="1994">Rodríguez (1994)</bibRefCitation>
|
||
. Although the presence of long setae in the G1 of
|
||
<taxonomicName id="3EA44D2E291C1602D7B3FD32FDFDFD78" box="[409,556,655,680]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lavallensis">
|
||
<emphasis id="CBD0EABF291C1602D7B3FD32FDFDFD78" box="[409,556,655,680]" italics="true" pageId="10" pageNumber="311">P. lavallensis</emphasis>
|
||
</taxonomicName>
|
||
varies (the
|
||
<typeStatus id="261F880F291C1602D484FD2CFCC1FD78" box="[686,784,655,680]" pageId="10" pageNumber="311" type="holotype">holotype</typeStatus>
|
||
bears a few, see
|
||
<figureCitation id="619F2A28291C1602D5E6FD2CFBCCFD78" box="[972,1053,655,680]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="10" pageNumber="311">Fig. 31</figureCitation>
|
||
), in
|
||
<taxonomicName id="3EA44D2E291C1602D267FD32FB6FFD78" box="[1101,1214,655,680]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="exilipes">
|
||
<emphasis id="CBD0EABF291C1602D267FD32FB6FFD78" box="[1101,1214,655,680]" italics="true" pageId="10" pageNumber="311">P. exilipes</emphasis>
|
||
</taxonomicName>
|
||
the apical peduncle is a little longer and the apex is not so strongly bent towards the cephalic side; in addition, the caudal ridge is more clearly distinct and longer than in
|
||
<taxonomicName id="3EA44D2E291C1602D433FD7AFD7CFD20" box="[537,685,727,752]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lavallensis">
|
||
<emphasis id="CBD0EABF291C1602D433FD7AFD7CFD20" box="[537,685,727,752]" italics="true" pageId="10" pageNumber="311">P. lavallensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="619F2A28291C1602D497FD74FCA0FD20" box="[701,881,727,752]" captionStart="FIGURES 22 – 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 – 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 – 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="10" pageNumber="311">Figs. 22, 30, 31</figureCitation>
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). Whether these differences are consistent or due to variability must be further investigated when new collections become available, in particular from western and central
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<collectingCountry id="81B3763D291C1602D6C0FC83FE97FCE7" box="[234,326,800,823]" name="Panama" pageId="10" pageNumber="311">Panama</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |