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<mods:title id="D4BB9908DA2CBE7977C6036DE95E2829">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<emphasis id="CBD0EABF291B1605D6BDFA19FE12FA04" bold="true" box="[151,451,1466,1492]" italics="true" pageId="13" pageNumber="314">Ptychophallus montanus</emphasis>
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<bibRefCitation id="9D354B5C291B1605D7F8FA18FD5CFA05" author="Rathbun" box="[466,653,1467,1493]" pageId="13" pageNumber="314" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
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<figureCitation id="619F2A28291B1605D6B5FA7EFEF7FA26" box="[159,294,1501,1526]" captionStart="FIGURES 33 43" captionStartId="15.[151,264,1763,1786]" captionTargetBox="[178,1412,197,1737]" captionTargetId="figure@15.[174,1412,193,1742]" captionTargetPageId="15" captionText="FIGURES 33 43. Ptychophallus micracanthus Rodríguez, 1994: 33, left first gonopod, holotype, USNM 240106, caudomesial view; 34, same data, latero-cephalic view of distal part. Ptychophallus montanus (Rathbun, 1898): 35, left first gonopod, lectotype, USNM 19486, caudo-mesial view; 36, same data, latero-cephalic view; 37, outline of left first gonopod, INPA 1843, caudo-mesial view; 38, same data, INPA 1847, caudo-mesial view. Ptychophallus tristani (Rathbun, 1896): 39, left first gonopod (partially damaged on proximal part), holotype, USNM 19047, caudo-mesial view; 40, same data, latero-cephalic view of distal part; 41, left first gonopod, holotype of Ptychophallus (Ptychophallus) campylus Pretzmann, 1968, NHMW 21849, caudo-mesial view; 42, outline of right gonopod, INPA 1836, caudo-mesial view; 43, outline of left first gonopod, INPA 1851, caudo-mesial view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235131/files/figure.png" pageId="13" pageNumber="314">Figs. 3538</figureCitation>
,
<figureCitation id="619F2A28291B1605D718FA7DFE9EFA26" box="[306,335,1502,1526]" captionStart="FIGURE 79" captionStartId="33.[151,250,1676,1698]" captionTargetBox="[161,1436,668,1649]" captionTargetId="figure@33.[150,1436,661,1654]" captionTargetPageId="33" captionText="FIGURE 79. Distribution map of Ptychophallus colombianus (Rathbun, 1896), and P. montanus (Rathbun, 1898)." httpUri="https://zenodo.org/record/235137/files/figure.png" pageId="13" pageNumber="314">79</figureCitation>
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<emphasis id="CBD0EABF291B1605D6BDF986FE77F9EB" box="[151,422,1573,1595]" italics="true" pageId="13" pageNumber="314">Pseudothelphusa montana</emphasis>
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, 535, figs. 5, 6.—
<treatmentCitation id="780510BC291B1605D517F986FC37F9EB" author="Young" box="[829,998,1573,1595]" page="14" pageId="13" pageNumber="314" year="1900">
<bibRefCitation id="9D354B5C291B1605D517F986FC13F9EB" author="Young" box="[829,962,1573,1595]" pageId="13" pageNumber="314" refString="Young, C. G. (1900) The stalk-eyed Crustacea of British Guiana, West Indies, and Bermuda. J. M. Watkins, London, 514 pp., 4 pls." type="book" year="1900">Young 1900</bibRefCitation>
: 14
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.—
<treatmentCitation id="780510BC291B1605D22EF986FB01F9EB" author="Rathbun" box="[1028,1232,1573,1595]" page="242" pageId="13" pageNumber="314" year="1904">
<bibRefCitation id="9D354B5C291B1605D22EF986FB4CF9EB" author="Rathbun" box="[1028,1181,1573,1595]" pageId="13" pageNumber="314" refString="Rathbun, M. J. (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle Paris, 6 (4), 225 - 312, text-figs. 1 - 37, volume pls. 9 - 18 (pls. I - X for Rathbun's article)." type="journal article" year="1904">Rathbun 1904</bibRefCitation>
: 242
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(in list).—
<treatmentCitation id="780510BC291B1605D369F986FEFCF98A" author="Rathbun" page="286" pageId="13" pageNumber="314" year="1905">
<bibRefCitation id="9D354B5C291B1605D369F986FF2CF98A" author="Rathbun" pageId="13" pageNumber="314" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905">Rathbun 1905</bibRefCitation>
: 286
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.—
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<bibRefCitation id="9D354B5C291B1605D760F9E7FE0EF98A" author="Rathbun" box="[330,479,1604,1626]" pageId="13" pageNumber="314" refString="Rathbun, M. J. (1906) Catalogue des Potamonides des collections du Museum d'histoire naturelle de Paris, d'apres les revisions et determinations de Mlle. Mary Rathbun. Seconde et derniere partie. Bulletin du Museum d'histoire naturelle Paris, 12 (7), 500 - 504." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
: 501
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(in list).—
<treatmentCitation id="780510BC291B1605D451F9E7FC81F98A" author="Coifmann" box="[635,848,1604,1626]" page="108" pageId="13" pageNumber="314" year="1939">
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: 108
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(in list).—
<treatmentCitation id="780510BC291B1605D5EBF9E7FB5CF98A" author="Ng" box="[961,1165,1604,1626]" page="176" pageId="13" pageNumber="314" year="2008">
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Ng.
<emphasis id="CBD0EABF291B1605D5C1F9E6FBCFF98A" box="[1003,1054,1604,1626]" italics="true" pageId="13" pageNumber="314">et al.</emphasis>
2008
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: 176
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.
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<emphasis id="CBD0EABF291B1605D6BDF9C1FE71F9A8" box="[151,416,1634,1656]" italics="true" pageId="13" pageNumber="314">Pseudothelphusa convexa</emphasis>
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, fig. 16.—
<treatmentCitation id="780510BC291B1605D4C0F9C1FC75F9A8" author="Young" box="[746,932,1634,1656]" page="218" pageId="13" pageNumber="314" year="1900">
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.—
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(in list).—
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.—
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: 107
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(in list).
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<emphasis id="CBD0EABF291B1605D6BDF903FE95F966" box="[151,324,1696,1718]" italics="true" pageId="13" pageNumber="314">Pseudothelphusa</emphasis>
(
<emphasis id="CBD0EABF291B1605D779F903FE35F966" box="[339,484,1696,1718]" italics="true" pageId="13" pageNumber="314">Ptychophallus</emphasis>
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<emphasis id="CBD0EABF291B1605D7D8F902FD9AF966" box="[498,587,1697,1718]" italics="true" pageId="13" pageNumber="314">montana</emphasis>
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: 10
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, figs. 9, 10.
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<paragraph id="F91B36AD291B1605D6BDF91DFCC6F904" blockId="13.[151,1436,1573,1902]" box="[151,791,1726,1749]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="D9B41183291B1605D6BDF91DFCC6F904" box="[151,791,1726,1749]" pageId="13" pageNumber="314">
<taxonomicName id="3EA44D2E291B1605D6BDF91DFDE9F905" ID-CoL="6WMV6" box="[151,568,1726,1749]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="montanus" subGenus="Ptychophallus">
<emphasis id="CBD0EABF291B1605D6BDF91DFEF9F904" box="[151,296,1726,1748]" italics="true" pageId="13" pageNumber="314">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF291B1605D71CF91DFE16F904" box="[310,455,1726,1748]" italics="true" pageId="13" pageNumber="314">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF291B1605D7FCF963FDE9F905" box="[470,568,1728,1749]" italics="true" pageId="13" pageNumber="314">montanus</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC291B1605D478F91CFCC2F904" author="Pretzmann" box="[594,787,1726,1748]" page="5" pageId="13" pageNumber="314" year="1965">
<bibRefCitation id="9D354B5C291B1605D478F91CFD2AF904" author="Pretzmann" box="[594,763,1726,1748]" pageId="13" pageNumber="314" refString="Pretzmann, G. (1965) Vorlaufiger Bericht uber die Familie Pseudothelphusidae. Sitzungsberichte der Mathematisch- Naturwissenschaftlichen Klasse der Osterreichischen Akademie der Wissenschaften, Wien, 1, 1 - 11." type="journal article" year="1965">Pretzmann 1965</bibRefCitation>
: 5
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD291B1605D6BDF97EFADEF923" blockId="13.[151,1436,1573,1902]" box="[151,1295,1757,1779]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="D9B41183291B1605D6BDF97EFADEF923" box="[151,1295,1757,1779]" pageId="13" pageNumber="314">
<taxonomicName id="3EA44D2E291B1605D6BDF97EFD70F923" ID-CoL="6WMV6" box="[151,673,1757,1779]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="subSpecies" species="montanus" subGenus="Ptychophallus" subSpecies="montanus">
<emphasis id="CBD0EABF291B1605D6BDF97EFEF9F923" box="[151,296,1757,1779]" italics="true" pageId="13" pageNumber="314">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF291B1605D71CF97EFE16F923" box="[310,455,1757,1779]" italics="true" pageId="13" pageNumber="314">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF291B1605D7FCF97DFD70F923" box="[470,673,1758,1779]" italics="true" pageId="13" pageNumber="314">montanus montanus</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC291B1605D490F97DFC57F923" author="Pretzmann" box="[698,902,1757,1779]" page="20" pageId="13" pageNumber="314" year="1971">
<bibRefCitation id="9D354B5C291B1605D490F97DFCB2F923" author="Pretzmann" box="[698,867,1757,1779]" pageId="13" pageNumber="314" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche Klasse der Osterreichischen Akademie der Wissenschaften, Series 1, 179 (1 - 4), 12 - 24." type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: 20
</treatmentCitation>
.—
<treatmentCitation id="780510BC291B1605D589F97EFBA2F923" author="Pretzmann" box="[931,1139,1757,1779]" page="85" pageId="13" pageNumber="314" year="1972">
<bibRefCitation id="9D354B5C291B1605D589F97EFB9FF923" author="Pretzmann" box="[931,1102,1757,1779]" pageId="13" pageNumber="314" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
: 85
</treatmentCitation>
, figs. 511513.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD291B1605D6BDF95FFAA1F8C2" blockId="13.[151,1436,1573,1902]" box="[151,1392,1788,1810]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="D9B41183291B1605D6BDF95FFAA1F8C2" box="[151,1392,1788,1810]" pageId="13" pageNumber="314">
<taxonomicName id="3EA44D2E291B1605D6BDF95FFD4AF8C2" box="[151,667,1788,1810]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="subSpecies" species="montanus" subGenus="Ptychophallus" subSpecies="convexus">
<emphasis id="CBD0EABF291B1605D6BDF95FFEF9F8C2" box="[151,296,1788,1810]" italics="true" pageId="13" pageNumber="314">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF291B1605D71CF95FFE16F8C2" box="[310,455,1788,1810]" italics="true" pageId="13" pageNumber="314">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF291B1605D7FCF95EFD4AF8C2" box="[470,667,1789,1810]" italics="true" pageId="13" pageNumber="314">montanus convexus</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC291B1605D49FF95EFC51F8C2" author="Pretzmann" box="[693,896,1788,1810]" page="20" pageId="13" pageNumber="314" year="1971">
<bibRefCitation id="9D354B5C291B1605D49FF95EFC8CF8C2" author="Pretzmann" box="[693,861,1788,1810]" pageId="13" pageNumber="314" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche Klasse der Osterreichischen Akademie der Wissenschaften, Series 1, 179 (1 - 4), 12 - 24." type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: 20
</treatmentCitation>
.—
<treatmentCitation id="780510BC291B1605D5B7F95FFBBFF8C2" author="Pretzmann" box="[925,1134,1788,1810]" page="86" pageId="13" pageNumber="314" year="1972">
<bibRefCitation id="9D354B5C291B1605D5B7F95FFB98F8C2" author="Pretzmann" box="[925,1097,1788,1810]" pageId="13" pageNumber="314" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
: 86
</treatmentCitation>
, figs. 502504, 532, 533.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD291B1605D6BDF8B9FAA0F89F" blockId="13.[151,1436,1573,1902]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="D9B41183291B1605D6BDF8B9FAA0F89F" pageId="13" pageNumber="314">
<taxonomicName id="3EA44D2E291B1605D6BDF8B9FE42F8E0" ID-CoL="6WMV6" box="[151,403,1818,1840]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="CBD0EABF291B1605D6BDF8B9FE42F8E0" box="[151,403,1818,1840]" italics="true" pageId="13" pageNumber="314">Ptychophallus montanus</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC291B1605D786F8B9FDA9F8E0" author="Rodriguez" box="[428,632,1818,1840]" page="85" pageId="13" pageNumber="314" year="1982">
<bibRefCitation id="9D354B5C291B1605D786F8B9FD82F8E0" author="Rodriguez" box="[428,595,1818,1840]" pageId="13" pageNumber="314" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez 1982</bibRefCitation>
: 85
</treatmentCitation>
.—
<treatmentCitation id="780510BC291B1605D4BCF8B9FC9CF8E0" author="Hobbs" box="[662,845,1818,1840]" page="101" pageId="13" pageNumber="314" year="1994">
<bibRefCitation id="9D354B5C291B1605D4BCF8B9FCCEF8E0" author="Hobbs" box="[662,799,1818,1840]" pageId="13" pageNumber="314" refString="Hobbs, H. H. III (1994) Biogeography of subterranean decapods in North and Central America and the Caribbean region (Caridea, Astacidea, Brachyura). Hydrobiologia, 287, 95 - 104. http: // dx. doi. org / 10.1007 / bf 00006899" type="journal article" year="1994">Hobbs, 1994</bibRefCitation>
: 101
</treatmentCitation>
.—
<treatmentCitation id="780510BC291B1605D546F8B9FB63F8E0" author="Campos" box="[876,1202,1818,1840]" page="560" pageId="13" pageNumber="314" year="1999">
<bibRefCitation id="9D354B5C291B1605D546F8B9FB51F8E0" author="Campos" box="[876,1152,1818,1840]" pageId="13" pageNumber="314" refString="Campos, M. R. &amp; Lemaitre, R. (1999) Two new freshwater crabs of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) from Panama, with notes on the distribution of the genus. Proceedings of the Biological Society of Washington, 112 (3), 553 - 561." type="journal article" year="1999">Campos &amp; Lemaitre 1999</bibRefCitation>
: 560
</treatmentCitation>
(map).—
<treatmentCitation id="780510BC291B1605D339F8B9FE48F89F" author="Rodriguez" page="423" pageId="13" pageNumber="314" year="2000">
<bibRefCitation id="9D354B5C291B1605D339F8B9FEB9F89F" author="Rodriguez" pageId="13" pageNumber="314" refString="Rodriguez, G. &amp; Hedstrom, I. (2000) The freswater crabs of the Barbilla National Park, Costa Rica (Crustacea: Brachyura: Pseudothelphusidae), with notes on the evolution of structures for spermatophore retention. Proceedings of the Biological Society of Washington, 113 (2), 420 - 425." type="journal article" year="2000">Rodríguez &amp; Hedström 2000</bibRefCitation>
: 423
</treatmentCitation>
, 424, fig. 2B, G.—
<treatmentCitation id="780510BC291B1605D472F89AFCCDF89F" author="Ng" box="[600,796,1849,1871]" page="176" pageId="13" pageNumber="314" year="2008">
<bibRefCitation id="9D354B5C291B1605D472F89AFD3CF89F" author="Ng" box="[600,749,1849,1871]" pageId="13" pageNumber="314" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF291B1605D457F899FD60F89F" box="[637,689,1849,1871]" italics="true" pageId="13" pageNumber="314">et al.</emphasis>
2008
</bibRefCitation>
: 176
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC291B1605D5A3F89AFACBF89F" author="Villalobos" box="[905,1306,1849,1871]" page="296" pageId="13" pageNumber="314" year="2008">
<bibRefCitation id="9D354B5C291B1605D5A3F89AFB3BF89F" author="Villalobos" box="[905,1258,1849,1871]" pageId="13" pageNumber="314" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 296
</treatmentCitation>
(in list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD291B1605D6BDF8FBFDA2F8BE" blockId="13.[151,1436,1573,1902]" box="[151,627,1880,1902]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="D9B41183291B1605D6BDF8FBFDA2F8BE" box="[151,627,1880,1902]" pageId="13" pageNumber="314">
<taxonomicName id="3EA44D2E291B1605D6BDF8FBFEEFF8BE" ID-CoL="7W7TX" box="[151,318,1880,1902]" class="Malacostraca" family="Pseudothelphusidae" genus="Fredius" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="CBD0EABF291B1605D6BDF8FBFEEFF8BE" box="[151,318,1880,1902]" italics="true" pageId="13" pageNumber="314">Fredius convexa</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC291B1605D772F8FAFDCAF8BE" author="Ng" box="[344,539,1880,1902]" page="174" pageId="13" pageNumber="314" year="2008">
<bibRefCitation id="9D354B5C291B1605D772F8FAFE3AF8BE" author="Ng" box="[344,491,1880,1902]" pageId="13" pageNumber="314" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF291B1605D756F8FAFE61F8BE" box="[380,432,1880,1902]" italics="true" pageId="13" pageNumber="314">et al.</emphasis>
2008
</bibRefCitation>
: 174
</treatmentCitation>
(in list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1606D6BDF839FD86FA4D" lastPageId="14" lastPageNumber="315" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="F91B36AD291B1606D6BDF839FC0CFA85" blockId="13.[151,1437,1946,2007]" lastBlockId="14.[151,1437,151,2013]" lastPageId="14" lastPageNumber="315" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6BDF839FEA8F863" bold="true" box="[151,377,1946,1971]" pageId="13" pageNumber="314">Material examined</emphasis>
.
<collectingCountry id="81B3763D291B1605D7A2F838FDE3F863" box="[392,562,1947,1971]" name="Costa Rica" pageId="13" pageNumber="314">COSTA RICA</collectingCountry>
. Alajuela Province, Atlantic drainage:
<specimenCount id="EFA2FD24291B1605D5CFF838FBE2F863" box="[997,1075,1946,1971]" pageId="13" pageNumber="314" type="male">1 male</specimenCount>
, 2 ovigerous females, UCR-MZ 3124-01, Macadamia Creek,
<geoCoordinate id="9C90506A291B1605D7CCF81CFD4DF807" box="[486,668,1983,2007]" direction="south" orientation="latitude" pageId="13" pageNumber="314" precision="1" value="-10.283898">10°1702.033”S</geoCoordinate>
<geoCoordinate id="9C90506A291B1605D48CF81DFCB6F807" box="[678,871,1982,2007]" direction="west" orientation="longitude" pageId="13" pageNumber="314" precision="1" value="-84.51526">84°3054.932”W</geoCoordinate>
,
<date id="8D1A106D291B1605D55FF81DFC30F807" box="[885,993,1982,2007]" pageId="13" pageNumber="314" value="2013-01-23">23.i.2013</date>
, L.R. Lara;
<specimenCount id="EFA2FD24291B1605D25BF81CFB13F807" box="[1137,1218,1982,2007]" pageId="13" pageNumber="314" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D2FBF81CFAC1F807" box="[1233,1296,1983,2007]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="13" pageNumber="314">INPA</collectionCode>
2085, same data;
<specimenCount id="EFA2FD2429181606D6FFFF3BFEF5FF60" box="[213,292,151,176]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE6829181606D71BFF3BFEACFF60" box="[305,381,152,176]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="14" pageNumber="315">CCDB</collectionCode>
4540, same data;
<specimenCount id="EFA2FD2429181606D466FF3BFD4AFF60" box="[588,667,151,176]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ 3126-01, creek tributary of Balsa River,
<geoCoordinate id="9C90506A29181606D2CDFF34FA4DFF60" box="[1255,1436,151,176]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.281005">10°16'51.618&quot;N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D6BDFF1FFE82FF05" box="[151,339,188,213]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.48578">84°29'08.825&quot;W</geoCoordinate>
,
<date id="8D1A106D29181606D74AFF1FFE14FF04" box="[352,453,188,212]" pageId="14" pageNumber="315" value="2010-10-07">7.x.2010</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429181606D4ACFF1FFD06FF05" box="[646,727,188,213]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD2429181606D4CFFF1FFC9DFF05" box="[741,844,188,213]" pageId="14" pageNumber="315" type="female">1 female</specimenCount>
, UCR-MZ 3126-02, creek tributary of Balsa River,
<geoCoordinate id="9C90506A29181606D6BDFF7CFE9DFF28" box="[151,332,223,248]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.280423">10°16'49.526&quot;N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D77FFF7CFDD2FF28" box="[341,515,223,248]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.51453">84°30'52.29&quot;W</geoCoordinate>
,
<date id="8D1A106D29181606D43AFF43FDA4FF28" box="[528,629,224,248]" pageId="14" pageNumber="315" value="2010-10-07">7.x.2010</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429181606D51CFF43FC56FF28" box="[822,903,223,248]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE6829181606D5BFFF43FC05FF28" box="[917,980,224,248]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="315">INPA</collectionCode>
2086, same data;
<specimenCount id="EFA2FD2429181606D283FF43FB28FF28" box="[1193,1273,223,248]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE6829181606D32DFF43FA82FF28" box="[1287,1363,224,248]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="14" pageNumber="315">CCDB</collectionCode>
4536, same data;
<specimenCount id="EFA2FD2429181606D733FEA7FEA5FECD" box="[281,372,260,285]" pageId="14" pageNumber="315" type="male">3 males</specimenCount>
,
<specimenCount id="EFA2FD2429181606D7A8FEA7FE39FECD" box="[386,488,260,285]" pageId="14" pageNumber="315" type="female">1 female</specimenCount>
, UCR-MZ 3127-02, spring of Santa Rita,
<geoCoordinate id="9C90506A29181606D5E5FEA7FBA7FECD" box="[975,1142,260,285]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.268711">10°16'07.36&quot;N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D255FEA7FACFFECC" box="[1151,1310,260,284]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.51217">84°30'43.8&quot;W</geoCoordinate>
,
<date id="8D1A106D29181606D301FEA7FA46FECD" box="[1323,1431,260,285]" pageId="14" pageNumber="315" value="2013-01-22">22.i.2013</date>
, L.R. Lara leg.;
<specimenCount id="EFA2FD2429181606D76DFE8BFE73FE90" box="[327,418,295,320]" pageId="14" pageNumber="315" type="male">3 males</specimenCount>
,
<specimenCount id="EFA2FD2429181606D785FE8BFDC5FE90" box="[431,532,295,320]" pageId="14" pageNumber="315" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE6829181606D40BFE8BFDB1FE90" box="[545,608,296,320]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="315">INPA</collectionCode>
2084, same data;
<specimenCount id="EFA2FD2429181606D518FE8BFC5CFE90" box="[818,909,295,320]" pageId="14" pageNumber="315" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD2429181606D5B0FE8BFBD1FE90" box="[922,1024,295,320]" pageId="14" pageNumber="315" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE6829181606D227FE8BFB88FE90" box="[1037,1113,296,320]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="14" pageNumber="315">CCDB</collectionCode>
4546, same data.—Alajuela Province, Pacific drainage:
<specimenCount id="EFA2FD2429181606D7FBFEEFFDCCFEB5" box="[465,541,332,357]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
(cw 25.3, cl 14.8),
<collectionCode id="9FB5AE6829181606D52AFEEFFC9DFEB4" box="[768,844,332,356]" pageId="14" pageNumber="315">UREG</collectionCode>
, Zapotal, San Antonio, Maquina Creek (tributary of Barrancas River),
<geoCoordinate id="9C90506A29181606D744FED3FDD1FE58" box="[366,512,368,392]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.15145">10°09.087N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D427FECCFD74FE58" box="[525,677,367,392]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.61418">84°36.851W</geoCoordinate>
,
<date id="8D1A106D29181606D49CFECCFCC4FE58" box="[694,789,367,392]" pageId="14" pageNumber="315" value="2009-12">xii.2009</date>
, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D22CFED3FB87FE58" box="[1030,1110,367,392]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
(cw 18.4, cl 11.4),
<collectionCode id="9FB5AE6829181606D366FED3FA49FE58" box="[1356,1432,368,392]" pageId="14" pageNumber="315">UREG</collectionCode>
, Valvera Creek (tributary of Barranca River), near Zapotal,
<geoCoordinate id="9C90506A29181606D571FE37FC38FE7D" box="[859,1001,404,429]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.158116">10°09,487N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D5D2FE37FB41FE7D" box="[1016,1168,404,429]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.64923">84°38.954W</geoCoordinate>
,
<quantity id="3E5C9B4829181606D289FE37FAD1FE7C" box="[1187,1280,404,429]" metricMagnitude="3" metricUnit="m" metricValue="1.157" pageId="14" pageNumber="315" unit="m" value="1157.0">1157 m</quantity>
,
<date id="8D1A106D29181606D33EFE37FAA2FE7D" box="[1300,1395,404,429]" pageId="14" pageNumber="315" value="2009-12">xii.2009</date>
, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D779FE1BFE73FE00" box="[339,418,439,464]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
(cw 29.6, cl 11.7),
<collectionCode id="9FB5AE6829181606D4BEFE1BFD31FE00" box="[660,736,440,464]" pageId="14" pageNumber="315">UREG</collectionCode>
, Valvera Creek (tributary of Barranca River), near Zapotal,
<geoCoordinate id="9C90506A29181606D6BDFE7FFEF8FE25" box="[151,297,476,501]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.16095">10°09.657N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D705FE7FFE19FE24" box="[303,456,476,500]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.645134">84°38.708W</geoCoordinate>
,
<date id="8D1A106D29181606D7F8FE7FFDE0FE25" box="[466,561,476,501]" pageId="14" pageNumber="315" value="2009-12">xii.2009</date>
, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D520FE7FFC84FE25" box="[778,853,476,501]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
(cw 20.7, cl 12.2),
<collectionCode id="9FB5AE6829181606D204FE7FFBABFE24" box="[1070,1146,476,500]" pageId="14" pageNumber="315">UREG</collectionCode>
, Palmital Creek (tributary of Barranca River), near San Antonio (near Zapotal),
<geoCoordinate id="9C90506A29181606D52DFE5CFC49FDC8" box="[775,920,511,536]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.158684">10°09.521N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D588FE5CFBEBFDC8" box="[930,1082,511,536]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.61653">84°36.992W</geoCoordinate>
,
<date id="8D1A106D29181606D262FE5CFB79FDC8" box="[1096,1192,511,536]" pageId="14" pageNumber="315" value="2009-12">xii.2009</date>
, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D3A4FDA3FF1CFDED" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
(cw 26.0, cl 14.9),
<collectionCode id="9FB5AE6829181606D79CFD87FDD3FDEC" box="[438,514,548,572]" pageId="14" pageNumber="315">UREG</collectionCode>
, Palmital Creek (tributary of Barranca River), near San Antonio (near Zapotal),
<geoCoordinate id="9C90506A29181606D6BDFDEBFEF8FDB0" box="[151,297,584,608]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.148">10°08.880N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D717FDE4FE04FDB0" box="[317,469,583,608]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.61452">84°36.871W</geoCoordinate>
,
<date id="8D1A106D29181606D7C7FDE4FD9CFDB0" box="[493,589,583,608]" pageId="14" pageNumber="315" value="2009-12">xii.2009</date>
, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D577FDEBFC69FDB0" box="[861,952,583,608]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ, near Zapotal,
<geoCoordinate id="9C90506A29181606D325FDEBFA4CFDB0" box="[1295,1437,584,608]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.158016">10°09.481'N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D6BDFDCFFEFAFD55" box="[151,299,620,645]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.6359">84°38.154'W</geoCoordinate>
, date unknown, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D497FDCFFCDCFD55" box="[701,781,620,645]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ, near Zapotal,
<geoCoordinate id="9C90506A29181606D21CFDCFFB12FD54" box="[1078,1219,620,644]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.148">10°08.880'N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D2E6FDCFFAB1FD55" box="[1228,1376,620,645]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.61462">84°36.877'W</geoCoordinate>
, date unknown, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D7D2FD33FD9CFD78" box="[504,589,655,680]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ, near Zapotal,
<geoCoordinate id="9C90506A29181606D5A0FD2CFBC9FD78" box="[906,1048,655,680]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.152917">10°09.175'N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D20CFD2CFB6BFD78" box="[1062,1210,655,680]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.61977">84°37.186'W</geoCoordinate>
, date unknown, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D764FD17FE4EFD1D" box="[334,415,692,717]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ, near Zapotal,
<geoCoordinate id="9C90506A29181606D4E6FD17FC88FD1D" box="[716,857,692,717]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.16125">10°09.675'N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D549FD17FC26FD1C" box="[867,1015,692,716]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.64515">84°38.709'W</geoCoordinate>
, date unknown, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D3A4FD17FF01FD20" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ, near Zapotal,
<geoCoordinate id="9C90506A29181606D7C5FD74FDAAFD20" box="[495,635,727,752]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.158517">10°09.511'N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D4A8FD74FCC7FD20" box="[642,790,727,752]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.61653">84°36.992'W</geoCoordinate>
, date unknown, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D2B3FD7BFB36FD20" box="[1177,1255,727,752]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ, near Zapotal,
<geoCoordinate id="9C90506A29181606D6D4FD5FFE5DFCC4" box="[254,396,764,788]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.15145">10°09.087'N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D7BDFD5FFDFBFCC5" box="[407,554,764,789]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.61428">84°36.857'W</geoCoordinate>
, date unknown, T. Poettinger leg.;
<specimenCount id="EFA2FD2429181606D5E3FD5FFBCBFCC5" box="[969,1050,764,789]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ, Valverde Creek, near Zapotal,
<geoCoordinate id="9C90506A29181606D72BFCBCFE5FFCE8" box="[257,398,799,824]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="10.161167">10°09.670'N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D7B1FCBCFDFEFCE8" box="[411,559,799,824]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-84.64658">84°38.795'W</geoCoordinate>
,
<quantity id="3E5C9B4829181606D46AFC83FD4FFCE7" box="[576,670,800,824]" metricMagnitude="3" metricUnit="m" metricValue="1.033" pageId="14" pageNumber="315" unit="m" value="1033.0">1033 m</quantity>
, T. Poettinger &amp; N. Thiercelin leg.—Heredia Province, Atlantic drainage:
<specimenCount id="EFA2FD2429181606D722FCE7FEB3FC8D" box="[264,354,836,861]" pageId="14" pageNumber="315" type="male">3 males</specimenCount>
,
<specimenCount id="EFA2FD2429181606D745FCE7FE31FC8D" box="[367,480,836,861]" pageId="14" pageNumber="315" type="female">3 females</specimenCount>
, UCR-MZ 178, Vara Blanca,
<date id="8D1A106D29181606D51FFCE7FC46FC8D" box="[821,919,836,861]" pageId="14" pageNumber="315" value="1967-05-07">7.v.1967</date>
, leg. D.C. Robinson
<emphasis id="CBD0EABF29181606D2A2FCE6FB13FC8D" box="[1160,1218,836,861]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
;
<specimenCount id="EFA2FD2429181606D2F8FCE7FAF3FC8D" box="[1234,1314,836,861]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ 170, Vara Blanca,
<date id="8D1A106D29181606D74EFCC4FE3EFC50" box="[356,495,871,896]" pageId="14" pageNumber="315" value="1967-08-27">27.viii.1967</date>
, D.C. Robinson leg.—Heredia Province, Pacific drainage:
<specimenCount id="EFA2FD2429181606D2ABFCCBFB0BFC50" box="[1153,1242,871,896]" pageId="14" pageNumber="315" type="male">3 males</specimenCount>
,
<specimenCount id="EFA2FD2429181606D2CFFCCBFA85FC50" box="[1253,1364,871,896]" pageId="14" pageNumber="315" type="female">2 females</specimenCount>
,
<specimenCount id="EFA2FD2429181606D374FCCBFA4CFC50" box="[1374,1437,871,896]" pageId="14" pageNumber="315" type="generic">2 juv.</specimenCount>
females, UCR-MZ 733, San José de La
<taxonomicName id="3EA44D2E29181606D4B6FC2EFCD3FC74" box="[668,770,909,932]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="montana">Montaña</taxonomicName>
, Porroz River,
<date id="8D1A106D29181606D5E2FC2FFBE4FC75" box="[968,1077,908,933]" pageId="14" pageNumber="315" value="1971-11-03">3.xi.1971</date>
, C. Villalobos &amp; R.A. Cruz leg.—Cartago Province, Atlantic drainage:
<specimenCount id="EFA2FD2429181606D4ABFC13FD0AFC18" box="[641,731,943,968]" pageId="14" pageNumber="315" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD2429181606D4CDFC0CFC86FC18" box="[743,855,943,968]" pageId="14" pageNumber="315" type="female">5 females</specimenCount>
,
<collectionCode id="9FB5AE6829181606D549FC13FC65FC18" LSID="urn:lsid:biocol.org:col:34871" box="[867,948,944,968]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="315">USNM</collectionCode>
32257, Cachí, Reventazón River,
<date id="8D1A106D29181606D313FC0CFA49FC18" box="[1337,1432,943,968]" pageId="14" pageNumber="315" value="1902-12">xii.1902</date>
, P. Biolley leg.;
<specimenCount id="EFA2FD2429181606D769FC77FE4AFC3D" box="[323,411,980,1005]" pageId="14" pageNumber="315" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD2429181606D78DFC77FDC7FC3D" box="[423,534,980,1005]" pageId="14" pageNumber="315" type="female">6 females</specimenCount>
, UCR-MZ 652, reservoir of Cachí,
<date id="8D1A106D29181606D58EFC77FBC7FC3D" box="[932,1046,980,1005]" pageId="14" pageNumber="315" value="1972-10-15">15.x.1972</date>
, C.E. &amp; C. Villalobos leg.;
<specimenCount id="EFA2FD2429181606D360FC77FA49FC3D" box="[1354,1432,980,1005]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE6829181606D6BDFC5BFF07FBC0" box="[151,214,1016,1040]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="315">INPA</collectionCode>
1843, same data;
<specimenCount id="EFA2FD2429181606D781FC5BFDD7FBC0" box="[427,518,1015,1040]" pageId="14" pageNumber="315" type="male">4 males</specimenCount>
,
<collectionCode id="9FB5AE6829181606D43EFC5BFD82FBC0" box="[532,595,1016,1040]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="14" pageNumber="315">INPA</collectionCode>
2019, Turrialba, road Vargas in Santa Cruz de Turrialba,
<date id="8D1A106D29181606D2DBFC54FAA2FBC0" box="[1265,1395,1015,1040]" pageId="14" pageNumber="315" value="2013-12-24">24.xii.2013</date>
, D. Hernández leg.;
<specimenCount id="EFA2FD2429181606D77AFBBFFE4FFBE5" box="[336,414,1052,1077]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ 3227-01, Buena Vista, Pacayas, Alvarado Canton,
<geoCoordinate id="9C90506A29181606D272FBBFFB23FBE5" box="[1112,1266,1052,1077]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="1" value="9.928734">9°55'43.44&quot;N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D2D3FBBFFA49FBE5" box="[1273,1432,1052,1077]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="1" value="-83.7935">83°47'36.6&quot;W</geoCoordinate>
,
<quantity id="3E5C9B4829181606D6BDFB9CFF3EFB87" box="[151,239,1087,1112]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="315" unit="m" value="1750.0">1750 m</quantity>
,
<date id="8D1A106D29181606D6D6FB9CFEB9FB88" box="[252,360,1087,1112]" pageId="14" pageNumber="315" value="2013-11-09">9.xi.2013</date>
, D. Hernández leg.—San José Province, Atlantic drainage:
<specimenCount id="EFA2FD2429181606D225FBE3FB8FFB88" box="[1039,1118,1087,1112]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
, UCR-MZ 718, Vásquez de Coronado Canton, Claro River, Bajo de La Hondura,
<date id="8D1A106D29181606D530FBC7FC43FBAD" box="[794,914,1124,1149]" pageId="14" pageNumber="315" value="1972-04-29">29.iv.1972</date>
, R.A. Cruz &amp; C. Villalobos leg.—San José Province, Pacific drainage:
<specimenCount id="EFA2FD2429181606D7E6FB2BFDF3FB70" box="[460,546,1159,1184]" pageId="14" pageNumber="315" type="male">3 males</specimenCount>
(cw 28.8, cl 17.8; cw 31.8, cl 18.9; cw 38.0, cl 23.8) 1 immat. female (cw 20.0, cl 12.8), UCR-MZ 171, San Isidro de Coronado,
<date id="8D1A106D29181606D494FB0FFCF2FB15" box="[702,803,1196,1221]" pageId="14" pageNumber="315" value="1968-02-03">3.ii.1968</date>
, col. unknown;
<specimenCount id="EFA2FD2429181606D5FDFB0FFBF3FB15" box="[983,1058,1196,1221]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
<typeStatus id="261F880F29181606D203FB0FFB43FB15" box="[1065,1170,1196,1221]" pageId="14" pageNumber="315" type="lectotype">lectotype</typeStatus>
,
<specimenCount id="EFA2FD2429181606D2B4FB0FFB38FB15" box="[1182,1257,1196,1221]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
paralectoptype,
<specimenCount id="EFA2FD2429181606D6BDFB73FED5FB38" box="[151,260,1231,1256]" pageId="14" pageNumber="315" type="female">2 females</specimenCount>
<typeStatus id="261F880F29181606D727FB6CFE63FB38" box="[269,434,1231,1256]" pageId="14" pageNumber="315" type="paralectotype">paralectotypes</typeStatus>
,
<collectionCode id="9FB5AE6829181606D794FB73FDDEFB38" LSID="urn:lsid:biocol.org:col:34871" box="[446,527,1232,1256]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="315">USNM</collectionCode>
19486, La Palma,
<quantity id="3E5C9B4829181606D4C2FB6CFC91FB38" box="[744,832,1231,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="315" unit="m" value="1500.0">1500 m</quantity>
,
<date id="8D1A106D29181606D564FB6CFC4EFB38" box="[846,927,1231,1256]" pageId="14" pageNumber="315" value="1896-02">ii.1896</date>
, G.F.
<collectingCountry id="81B3763D29181606D5CAFB6CFBFFFB38" box="[992,1070,1231,1256]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="14" pageNumber="315">Tristan</collectingCountry>
leg.;
<specimenCount id="EFA2FD2429181606D25BFB73FB11FB38" box="[1137,1216,1231,1256]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD2429181606D2E7FB6CFAECFB38" box="[1229,1341,1231,1256]" pageId="14" pageNumber="315" type="female">5 females</specimenCount>
,
<collectionCode id="9FB5AE6829181606D360FB72FA4DFB38" box="[1354,1436,1233,1256]" httpUri="http://grbio.org/cool/1h73-pg9e" name="Tianjin Museum of Natural History" pageId="14" pageNumber="315">TMNH</collectionCode>
4442, Grande de Tárcoles River,
<quantity id="3E5C9B4829181606D42DFB57FD9CFADD" box="[519,589,1268,1293]" metricMagnitude="2" metricUnit="m" metricValue="3.218688" pageId="14" pageNumber="315" unit="mi" value="0.2">0.2 mi</quantity>
S Alto La Palma,
<quantity id="3E5C9B4829181606D533FB57FC8EFADD" box="[793,863,1268,1293]" metricMagnitude="3" metricUnit="m" metricValue="7.0811136" pageId="14" pageNumber="315" unit="mi" value="4.4">4.4 mi</quantity>
N San Jerónimo, Moravia Canton,
<date id="8D1A106D29181606D2C0FB57FA8FFADD" box="[1258,1374,1268,1293]" pageId="14" pageNumber="315" value="1962-07-09">9.vii.1962</date>
, A.E. &amp; G. Smalley leg.—
<taxonomicName id="3EA44D2E29181606D75DFAB4FE24FAE0" box="[375,501,1303,1328]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
Province, Pacific drainage:
<specimenCount id="EFA2FD2429181606D51BFABBFCADFAE0" box="[817,892,1303,1328]" pageId="14" pageNumber="315" type="male">1 male</specimenCount>
(cw 39.3, cl 24.9),
<typeStatus id="261F880F29181606D27CFAB4FB69FAE0" box="[1110,1208,1303,1328]" pageId="14" pageNumber="315" type="holotype">holotype</typeStatus>
of
<taxonomicName id="3EA44D2E29181606D2F6FAB4FE70FA84" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="CBD0EABF29181606D2F6FAB4FF22FA84" italics="true" pageId="14" pageNumber="315">Pseudothelphusa convexa</emphasis>
<bibRefCitation id="9D354B5C29181606D6D0FA9FFE70FA84" author="Rathbun" box="[250,417,1340,1365]" pageId="14" pageNumber="315" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
</taxonomicName>
,
<collectionCode id="9FB5AE6829181606D787FA9FFE2FFA84" LSID="urn:lsid:biocol.org:col:34871" box="[429,510,1340,1364]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="315">USNM</collectionCode>
19483, Palmar, undated, G.K. Chemie leg.
</paragraph>
<paragraph id="F91B36AD29181606D6EDFAC3FD86FA4D" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="315">
<collectingCountry id="81B3763D29181606D6EDFAC3FE93FAA8" box="[199,322,1376,1400]" name="Panama" pageId="14" pageNumber="315">PANAMA</collectingCountry>
. Chiriquí Province, Pacific drainage:
<specimenCount id="EFA2FD2429181606D537FAC3FC50FAA8" box="[797,897,1375,1400]" pageId="14" pageNumber="315" type="male">4 males</specimenCount>
,
<specimenCount id="EFA2FD2429181606D5BDFAFCFBC0FAA8" box="[919,1041,1375,1400]" pageId="14" pageNumber="315" type="female">5 females</specimenCount>
,
<collectionCode id="9FB5AE6829181606D202FAC3FBA8FAA8" LSID="urn:lsid:biocol.org:col:34871" box="[1064,1145,1376,1400]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="315">USNM</collectionCode>
1089227, vicinities La Concepción,
<date id="8D1A106D29181606D701FA27FE53FA4D" box="[299,386,1412,1437]" pageId="14" pageNumber="315" value="1964-06">vi.1964</date>
, local children leg.
</paragraph>
</subSubSection>
<subSubSection id="B1BE652629181606D6EDFA04FE91F80D" pageId="14" pageNumber="315" type="description">
<paragraph id="F91B36AD29181606D6EDFA04FC5BF8D5" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="315">
<emphasis id="CBD0EABF29181606D6EDFA04FE35FA10" bold="true" box="[199,484,1447,1472]" pageId="14" pageNumber="315">Description of gonopod</emphasis>
. Straight in mesiocaudal view. Marginal suture on mesial side, straight; marginal process rounded to subtriangular, not produced beyond distal border of apex. Mesial process subtriangular, slightly downturned. Lateral process moderately long, approximately 7075% of stem length (from proximal opening to caudal border of apex), strongly bilobed, with distinct median concavity on caudal surface; lateral border with distinct median incision; proximal lobe rather angled, usually as wide as or sometimes wider than distal lobe, with few to several scattered minute setae on caudal, lateral surfaces; distal lobe broad, usually transversal, moderately swollen distally on cephalic surface, narrower or usually as wide as lateral end of apex. Distocaudal ridge distinct, narrow, well separated from distal border of lateral process by distinct depression. Apex bent approximately 90° towards laterocephalic side; oblong, narrow in distal view; caudal border with narrow notch near lateral end. Field of apical spines well developed, facing towards cephalolateral side.
</paragraph>
<paragraph id="F91B36AD29181606D6EDF8ACFE91F80D" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="315">
<emphasis id="CBD0EABF29181606D6EDF8ACFDD8F8F8" bold="true" box="[199,521,1807,1832]" pageId="14" pageNumber="315">
<typeStatus id="261F880F29181606D6EDF8ACFEECF8F8" box="[199,317,1807,1832]" pageId="14" pageNumber="315" type="lectotype">Lectotype</typeStatus>
and
<typeStatus id="261F880F29181606D753F8ACFE7DF8F8" box="[377,428,1807,1832]" pageId="14" pageNumber="315">type</typeStatus>
locality
</emphasis>
. Male, cw 25.5, cl 15.1, USNM 19486.
<collectingCountry id="81B3763D29181606D5FAF8B3FB9CF8F8" box="[976,1101,1807,1832]" name="Costa Rica" pageId="14" pageNumber="315">Costa Rica</collectingCountry>
, San José, La Palma (Alto La Palma,
<geoCoordinate id="9C90506A29181606D6C6F897FE9DF89C" box="[236,332,1844,1868]" direction="north" orientation="latitude" pageId="14" pageNumber="315" precision="925" value="10.05">10°03N</geoCoordinate>
<geoCoordinate id="9C90506A29181606D77FF897FE6AF89D" box="[341,443,1844,1869]" direction="west" orientation="longitude" pageId="14" pageNumber="315" precision="925" value="-83.96667">83°58W</geoCoordinate>
, according to
<bibRefCitation id="9D354B5C29181606D476F897FCDDF89D" author="Smalley" box="[604,780,1844,1869]" pageId="14" pageNumber="315" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley, 1964b</bibRefCitation>
) (Pacific drainage).
<bibRefCitation id="9D354B5C29181606D5DEF897FB77F89C" author="Rathbun" box="[1012,1190,1844,1869]" pageId="14" pageNumber="315" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun (1898)</bibRefCitation>
based the description of
<taxonomicName id="3EA44D2E29181606D692F8F4FE35F8A0" box="[184,484,1879,1904]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="CBD0EABF29181606D692F8F4FE35F8A0" box="[184,484,1879,1904]" italics="true" pageId="14" pageNumber="315">Pseudothelphusa montana</emphasis>
</taxonomicName>
on two males and two females. The smaller male is designated here as
<typeStatus id="261F880F29181606D305F8F4FA48F8A0" box="[1327,1433,1879,1904]" pageId="14" pageNumber="315" type="lectotype">lectotype</typeStatus>
. Both gonopods were dissected and probably was the specimen used by
<bibRefCitation id="9D354B5C29181606D598F8DFFB8CF844" author="Rathbun" box="[946,1117,1916,1941]" pageId="14" pageNumber="315" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun (1898</bibRefCitation>
: 516, fig. 6b, c) and
<bibRefCitation id="9D354B5C29181606D36AF8DFFE4DF868" author="Smalley" pageId="14" pageNumber="315" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley (1964b: 12, figs. 9, 10)</bibRefCitation>
to illustrate the G1. The larger male (27.3:15.4) and the females (32.5:18.2; 33.2:18.8) are
<typeStatus id="261F880F29181606D6BDF867FEECF80D" box="[151,317,1988,2013]" pageId="14" pageNumber="315" type="paralectotype">paralectotypes</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="ADDB662529191607D6BDF940FDB2F800" httpUri="https://zenodo.org/record/235131/files/figure.png" pageId="15" pageNumber="316" targetBox="[178,1412,197,1737]" targetPageId="15">
<paragraph id="F91B36AD29191607D6BDF940FDB2F800" blockId="15.[151,1436,1763,2000]" pageId="15" pageNumber="316">
<emphasis id="CBD0EABF29191607D6BDF940FE80F929" bold="true" box="[151,337,1763,1786]" pageId="15" pageNumber="316">FIGURES 3343</emphasis>
.
<taxonomicName id="3EA44D2E29191607D74BF947FCB9F92A" authority="Rodriguez, 1994: 33" authorityName="Rodriguez" authorityPageNumber="33" authorityYear="1994" box="[353,872,1764,1786]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="micracanthus">
<emphasis id="CBD0EABF29191607D74BF947FD57F92A" box="[353,646,1764,1786]" italics="true" pageId="15" pageNumber="316">Ptychophallus micracanthus</emphasis>
Rodríguez, 1994: 33
</taxonomicName>
, left first gonopod, holotype, USNM 240106, caudomesial view; 34, same data, latero-cephalic view of distal part.
<taxonomicName id="3EA44D2E29191607D53AF8A1FB69F8C8" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" box="[784,1208,1794,1816]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="CBD0EABF29191607D53AF8A1FBDBF8C8" box="[784,1034,1794,1816]" italics="true" pageId="15" pageNumber="316">Ptychophallus montanus</emphasis>
(Rathbun, 1898)
</taxonomicName>
: 35, left first gonopod, lectotype, USNM 19486, caudo-mesial view; 36, same data, latero-cephalic view; 37, outline of left first gonopod, INPA 1843, caudo-mesial view; 38, same data, INPA 1847, caudo-mesial view.
<taxonomicName id="3EA44D2E29191607D544F89CFADCF885" authority="Rathbun, 1896" authorityName="Rathbun" authorityYear="1896" box="[878,1293,1855,1878]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="tristani">
<emphasis id="CBD0EABF29191607D544F89CFB82F886" box="[878,1107,1855,1878]" italics="true" pageId="15" pageNumber="316">Ptychophallus tristani</emphasis>
(Rathbun, 1896)
</taxonomicName>
: 39, left first gonopod (partially damaged on proximal part), holotype, USNM 19047, caudo-mesial view; 40, same data, latero-cephalic view of distal part; 41, left first gonopod, holotype of
<taxonomicName id="3EA44D2E29191607D4F1F8DEFAEAF843" authority="Pretzmann, 1968" authorityName="Pretzmann" authorityYear="1968" box="[731,1339,1917,1939]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="campylus" subGenus="Ptychophallus">
<emphasis id="CBD0EABF29191607D4F1F8DEFCBDF843" box="[731,876,1917,1939]" italics="true" pageId="15" pageNumber="316">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF29191607D557F8DEFBDFF843" box="[893,1038,1917,1939]" italics="true" pageId="15" pageNumber="316">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF29191607D20BF8DEFB50F843" box="[1057,1153,1917,1939]" italics="true" pageId="15" pageNumber="316">campylus</emphasis>
Pretzmann, 1968
</taxonomicName>
, NHMW 21849, caudo-mesial view; 42, outline of right gonopod, INPA 1836, caudo-mesial view; 43, outline of left first gonopod, INPA 1851, caudo-mesial view. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="B1BE652629061618D6EDFF34FDF2FF04" pageId="16" pageNumber="317" type="distribution">
<paragraph id="F91B36AD29061618D6EDFF34FDF2FF04" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="317">
<emphasis id="CBD0EABF29061618D6EDFF34FE88FF60" bold="true" box="[199,345,151,176]" pageId="16" pageNumber="317">Distribution</emphasis>
. Central and southeastern
<collectingCountry id="81B3763D29061618D4B4FF3BFCF0FF60" box="[670,801,151,176]" name="Costa Rica" pageId="16" pageNumber="317">Costa Rica</collectingCountry>
, in both Atlantic and Pacific drainages, and western
<collectingCountry id="81B3763D29061618D6BDFF1EFF22FF04" box="[151,243,189,212]" name="Panama" pageId="16" pageNumber="317">Panama</collectingCountry>
, Pacific drainage (
<figureCitation id="619F2A2829061618D7E8FF1FFDC7FF04" box="[450,534,188,213]" captionStart="FIGURE 79" captionStartId="33.[151,250,1676,1698]" captionTargetBox="[161,1436,668,1649]" captionTargetId="figure@33.[150,1436,661,1654]" captionTargetPageId="33" captionText="FIGURE 79. Distribution map of Ptychophallus colombianus (Rathbun, 1896), and P. montanus (Rathbun, 1898)." httpUri="https://zenodo.org/record/235137/files/figure.png" pageId="16" pageNumber="317">Fig. 79</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B1BE652629061618D6EDFF7CFE1BFCC4" pageId="16" pageNumber="317" type="discussion">
<paragraph id="F91B36AD29061618D6EDFF7CFABBFD54" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="317">
<emphasis id="CBD0EABF29061618D6EDFF7CFEE5FF28" bold="true" box="[199,308,223,248]" pageId="16" pageNumber="317">Remarks</emphasis>
.
<bibRefCitation id="9D354B5C29061618D769FF7CFE24FF28" author="Rathbun" box="[323,501,223,248]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun (1898)</bibRefCitation>
mentioned that the G1 of
<taxonomicName id="3EA44D2E29061618D509FF42FC6AFF28" box="[803,955,225,248]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="tumimanus">
<emphasis id="CBD0EABF29061618D509FF42FC6AFF28" box="[803,955,225,248]" italics="true" pageId="16" pageNumber="317">P. tumimanus</emphasis>
</taxonomicName>
, although resembling that of
<taxonomicName id="3EA44D2E29061618D321FF42FA44FF28" box="[1291,1429,225,248]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="CBD0EABF29061618D321FF42FA44FF28" box="[1291,1429,225,248]" italics="true" pageId="16" pageNumber="317">P. montanus</emphasis>
</taxonomicName>
, had less thickened distal lobe and the proximal lobe less angular than in the latter species.
<bibRefCitation id="9D354B5C29061618D298FEA7FAB4FECC" author="Rathbun" box="[1202,1381,260,285]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun (1898)</bibRefCitation>
also distinguished them by the presence of a large protuberance on the palm between the fingers in
<taxonomicName id="3EA44D2E29061618D29DFE8AFA9FFE90" box="[1207,1358,296,320]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="tumimanus">
<emphasis id="CBD0EABF29061618D29DFE8AFB14FE90" box="[1207,1221,297,320]" italics="true" pageId="16" pageNumber="317">P</emphasis>
.
<emphasis id="CBD0EABF29061618D2F9FE8AFA9FFE90" box="[1235,1358,297,320]" italics="true" pageId="16" pageNumber="317">tumimanus</emphasis>
</taxonomicName>
. In the keys provided by Rathbun both species are differentiated by the morphology of the lobes of the lateral process.
<bibRefCitation id="9D354B5C29061618D6BDFED3FEB2FE58" author="Pretzmann" box="[151,355,368,392]" pageId="16" pageNumber="317" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann (1972)</bibRefCitation>
considered
<taxonomicName id="3EA44D2E29061618D7C4FED2FD56FE58" box="[494,647,369,392]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="tumimanus">
<emphasis id="CBD0EABF29061618D7C4FED2FD56FE58" box="[494,647,369,392]" italics="true" pageId="16" pageNumber="317">P. tumimanus</emphasis>
</taxonomicName>
having a lateral process bearing a thick section in contrast to a slim section in
<taxonomicName id="3EA44D2E29061618D726FE36FE47FE7C" box="[268,406,405,428]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="CBD0EABF29061618D726FE36FE47FE7C" box="[268,406,405,428]" italics="true" pageId="16" pageNumber="317">P. montanus</emphasis>
</taxonomicName>
. For
<bibRefCitation id="9D354B5C29061618D7FFFE37FD40FE7D" author="Smalley" box="[469,657,404,429]" pageId="16" pageNumber="317" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley (1964b)</bibRefCitation>
and
<bibRefCitation id="9D354B5C29061618D4E1FE37FC43FE7C" author="Rodriguez" box="[715,914,404,429]" pageId="16" pageNumber="317" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez (1982)</bibRefCitation>
<taxonomicName id="3EA44D2E29061618D5B0FE36FBF5FE7C" box="[922,1060,405,428]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="CBD0EABF29061618D5B0FE36FBF5FE7C" box="[922,1060,405,428]" italics="true" pageId="16" pageNumber="317">P. montanus</emphasis>
</taxonomicName>
has a marginal process nearly as long as distal border of apex and a subacute proximal lobe in contrast to a marginal process much shorter and a proximal lobe broadly rounded in
<taxonomicName id="3EA44D2E29061618D41BFE7EFD1CFE24" box="[561,717,477,500]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="tumimanus">
<emphasis id="CBD0EABF29061618D41BFE7EFD1CFE24" box="[561,717,477,500]" italics="true" pageId="16" pageNumber="317">P. tumimanus</emphasis>
</taxonomicName>
.
<bibRefCitation id="9D354B5C29061618D4CAFE7FFC70FE25" author="Smalley" box="[736,929,476,501]" pageId="16" pageNumber="317" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley (1964b)</bibRefCitation>
also distinguished them by size: small (
<taxonomicName id="3EA44D2E29061618D3ADFE7EFED5FDC8" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="CBD0EABF29061618D3ADFE7EFED5FDC8" italics="true" pageId="16" pageNumber="317">P. montanus</emphasis>
</taxonomicName>
) or large (
<taxonomicName id="3EA44D2E29061618D7A2FDA2FDF5FDC8" box="[392,548,513,536]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="tumimanus">
<emphasis id="CBD0EABF29061618D7A2FDA2FDF5FDC8" box="[392,548,513,536]" italics="true" pageId="16" pageNumber="317">P. tumimanus</emphasis>
</taxonomicName>
) species. These diagnostic characters are indeed subject to some degree of variation in our material, particularly the size and shape of the lateral process of the G1 (see
<figureCitation id="619F2A2829061618D283FD87FA8CFDEC" box="[1193,1373,548,573]" captionStart="FIGURES 33 43" captionStartId="15.[151,264,1763,1786]" captionTargetBox="[178,1412,197,1737]" captionTargetId="figure@15.[174,1412,193,1742]" captionTargetPageId="15" captionText="FIGURES 33 43. Ptychophallus micracanthus Rodríguez, 1994: 33, left first gonopod, holotype, USNM 240106, caudomesial view; 34, same data, latero-cephalic view of distal part. Ptychophallus montanus (Rathbun, 1898): 35, left first gonopod, lectotype, USNM 19486, caudo-mesial view; 36, same data, latero-cephalic view; 37, outline of left first gonopod, INPA 1843, caudo-mesial view; 38, same data, INPA 1847, caudo-mesial view. Ptychophallus tristani (Rathbun, 1896): 39, left first gonopod (partially damaged on proximal part), holotype, USNM 19047, caudo-mesial view; 40, same data, latero-cephalic view of distal part; 41, left first gonopod, holotype of Ptychophallus (Ptychophallus) campylus Pretzmann, 1968, NHMW 21849, caudo-mesial view; 42, outline of right gonopod, INPA 1836, caudo-mesial view; 43, outline of left first gonopod, INPA 1851, caudo-mesial view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235131/files/figure.png" pageId="16" pageNumber="317">Figs. 35, 37, 38</figureCitation>
). The cheliped protuberance and the size are also poor characters because some large specimens can have gonopods with subacute proximal border of the lateral process and bear a protuberance in the palm (e.g., UCR-MZ 170, 171).
</paragraph>
<paragraph id="F91B36AD29061618D6EDFD2CFE1BFCC4" blockId="16.[151,1437,151,789]" pageId="16" pageNumber="317">
Such variability as well as the largely sympatric distribution of both species would suggest that they could be synonymized. We nevertheless prefer to keep them separate for the time being until genetic studies are undertaken.
<taxonomicName id="3EA44D2E29061618D6BDFD74FDA3FD20" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" box="[151,626,727,752]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="CBD0EABF29061618D6BDFD74FE6FFD20" box="[151,446,727,752]" italics="true" pageId="16" pageNumber="317">Pseudothelphusa convexa</emphasis>
<bibRefCitation id="9D354B5C29061618D7E3FD74FDA3FD20" author="Rathbun" box="[457,626,727,752]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
</taxonomicName>
was already synonymized by
<bibRefCitation id="9D354B5C29061618D5F0FD74FB74FD20" author="Rodriguez" box="[986,1189,727,752]" pageId="16" pageNumber="317" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez (1982)</bibRefCitation>
under
<taxonomicName id="3EA44D2E29061618D2D6FD74FE14FCC4" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="CBD0EABF29061618D2D6FD74FED5FCC4" italics="true" pageId="16" pageNumber="317">Ptychophallus montanus</emphasis>
(
<bibRefCitation id="9D354B5C29061618D738FD5FFE6CFCC4" author="Rathbun" box="[274,445,764,789]" pageId="16" pageNumber="317" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>