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<document id="C85B7F2D5D8EFDF5DBBF4169CBE6F787" ID-CLB-Dataset="42408" ID-DOI="10.5281/zenodo.235127" ID-GBIF-Dataset="16b434af-bed4-43fd-bdfc-0f29bcaec7bb" ID-ISSN="1175-5326" ID-Zenodo-Dep="235127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460931586988" checkinUser="plazi" docAuthor="Magalhães, Célio, Wehrtmann, Ingo S., Lara, Luis Rólier &amp; Mantelatto, Fernando L." docDate="2015" docId="710D87BB291B1605D6BDFECCFCEBFAB6" docLanguage="en" docName="zt03905p344.pdf" docOrigin="Zootaxa 3905 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptychophallus micracanthus Rodriguez 1994" docType="treatment" docVersion="14" lastPageNumber="314" masterDocId="8D34FFC329161608D62AFFA3FFD1FFD0" masterDocTitle="Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)" masterLastPageNumber="344" masterPageNumber="301" pageNumber="314" updateTime="1698582371579" updateUser="ExternalLinkService">
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<mods:title id="89AFC36D1A2481FEDB6A39DD8EE50BCC">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<mods:namePart id="B298A853D86ABD97570A2BB5A98D7FFE">Magalhães, Célio</mods:namePart>
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<mods:namePart id="257A4110089C1F227A9723470E96A8CE">Wehrtmann, Ingo S.</mods:namePart>
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<mods:namePart id="1768DE9B72F77AE4253E13A5382F10B4">Lara, Luis Rólier</mods:namePart>
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<mods:namePart id="AC38320BC430DE1FF52ECE706769A4E5">Mantelatto, Fernando L.</mods:namePart>
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<mods:title id="6A2847FAA2287E4A0BDA13570A9A259D">Zootaxa</mods:title>
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<mods:date id="A958F37815D7E99331C6EBA7375E81BD">2015</mods:date>
<mods:detail id="8F8D20DA7F01076D6D42B8BA662614B1" type="volume">
<mods:number id="CA87DCE8A5A8293EBF4B88A058AE2775">3905</mods:number>
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<treatment id="710D87BB291B1605D6BDFECCFCEBFAB6" ID-DOI="http://doi.org/10.5281/zenodo.5491522" ID-GBIF-Taxon="119590157" ID-Zenodo-Dep="5491522" LSID="urn:lsid:plazi:treatment:710D87BB291B1605D6BDFECCFCEBFAB6" httpUri="http://treatment.plazi.org/id/710D87BB291B1605D6BDFECCFCEBFAB6" lastPageNumber="314" pageId="13" pageNumber="314">
<subSubSection id="B1BE6526291B1605D6BDFECCFE89FE7B" pageId="13" pageNumber="314" type="nomenclature">
<paragraph id="F91B36AD291B1605D6BDFECCFD15FE5A" blockId="13.[151,708,367,427]" box="[151,708,367,394]" pageId="13" pageNumber="314">
<heading id="A25381C1291B1605D6BDFECCFD15FE5A" bold="true" box="[151,708,367,394]" fontSize="11" level="1" pageId="13" pageNumber="314" reason="1">
<taxonomicName id="3EA44D2E291B1605D6BDFECCFD15FE5A" ID-CoL="78N77" authority="Rodriguez, 1994" authorityName="Rodriguez" authorityYear="1994" box="[151,708,367,394]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="micracanthus">
<emphasis id="CBD0EABF291B1605D6BDFECCFD15FE5A" bold="true" box="[151,708,367,394]" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6BDFECCFE20FE59" bold="true" box="[151,497,367,393]" italics="true" pageId="13" pageNumber="314">Ptychophallus micracanthus</emphasis>
<bibRefCitation id="9D354B5C291B1605D7D2FED3FD15FE5A" author="Rodriguez" box="[504,708,368,394]" pageId="13" pageNumber="314" refString="Rodriguez, G. (1994) A revision of the type material of some species of Hypolobocera and Ptychophallus (Crustacea: Decapoda: Pseudothelphusidae) in the National Museum of Natural History, Washington, D. C., with descriptions of a new species and a new subspecies. Proceedings of the Biological Society of Washington, 107 (2), 296 - 307." type="journal article" year="1994">Rodríguez, 1994</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F91B36AD291B1605D6BDFE31FE89FE7B" blockId="13.[151,708,367,427]" box="[151,344,402,427]" pageId="13" pageNumber="314">
(
<figureCitation id="619F2A28291B1605D6B5FE31FEF7FE7B" box="[159,294,402,427]" captionStart="FIGURES 33 – 43" captionStartId="15.[151,264,1763,1786]" captionTargetBox="[178,1412,197,1737]" captionTargetId="figure@15.[174,1412,193,1742]" captionTargetPageId="15" captionText="FIGURES 33 – 43. Ptychophallus micracanthus Rodríguez, 1994: 33, left first gonopod, holotype, USNM 240106, caudomesial view; 34, same data, latero-cephalic view of distal part. Ptychophallus montanus (Rathbun, 1898): 35, left first gonopod, lectotype, USNM 19486, caudo-mesial view; 36, same data, latero-cephalic view; 37, outline of left first gonopod, INPA 1843, caudo-mesial view; 38, same data, INPA 1847, caudo-mesial view. Ptychophallus tristani (Rathbun, 1896): 39, left first gonopod (partially damaged on proximal part), holotype, USNM 19047, caudo-mesial view; 40, same data, latero-cephalic view of distal part; 41, left first gonopod, holotype of Ptychophallus (Ptychophallus) campylus Pretzmann, 1968, NHMW 21849, caudo-mesial view; 42, outline of right gonopod, INPA 1836, caudo-mesial view; 43, outline of left first gonopod, INPA 1851, caudo-mesial view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235131/files/figure.png" pageId="13" pageNumber="314">Figs. 33, 34</figureCitation>
,
<figureCitation id="619F2A28291B1605D718FE30FE9EFE7B" box="[306,335,403,427]" captionStart="FIGURE 78" captionStartId="32.[151,250,1218,1240]" captionTargetBox="[156,1432,201,1194]" captionTargetId="figure@32.[151,1436,193,1196]" captionTargetPageId="32" captionText="FIGURE 78. Distribution map of Ptychophallus cocleensis Pretzmann, 1965, P. lavallensis Pretzmann, 1978, and P. micracanthus Rodríguez, 1994." httpUri="https://zenodo.org/record/235136/files/figure.png" pageId="13" pageNumber="314">78</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1605D6BDFE79FE0BFDFE" pageId="13" pageNumber="314" type="reference_group">
<paragraph id="F91B36AD291B1605D6BDFE79FE0BFDFE" blockId="13.[151,1436,474,558]" pageId="13" pageNumber="314">
<treatmentCitationGroup id="D9B41183291B1605D6BDFE79FE0BFDFE" pageId="13" pageNumber="314">
<taxonomicName id="3EA44D2E291B1605D6BDFE79FD67FE20" ID-CoL="78N77" authority="Rodriguez, 1994: 305" authorityName="Rodriguez" authorityPageNumber="305" authorityYear="1994" box="[151,694,474,496]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="micracanthus">
<emphasis id="CBD0EABF291B1605D6BDFE79FE6EFE20" box="[151,447,474,496]" italics="true" pageId="13" pageNumber="314">Ptychophallus micracanthus</emphasis>
<treatmentCitation id="780510BC291B1605D7E1FE79FD67FE20" author="Rodriguez" box="[459,694,474,496]" page="305" pageId="13" pageNumber="314" year="1994">
<bibRefCitation id="9D354B5C291B1605D7E1FE79FDAFFE20" author="Rodriguez" box="[459,638,474,496]" pageId="13" pageNumber="314" refString="Rodriguez, G. (1994) A revision of the type material of some species of Hypolobocera and Ptychophallus (Crustacea: Decapoda: Pseudothelphusidae) in the National Museum of Natural History, Washington, D. C., with descriptions of a new species and a new subspecies. Proceedings of the Biological Society of Washington, 107 (2), 296 - 307." type="journal article" year="1994">Rodríguez, 1994</bibRefCitation>
: 305
</treatmentCitation>
</taxonomicName>
, fig. 4A–D.—
<treatmentCitation id="780510BC291B1605D578FE79FB78FE20" author="Campos" box="[850,1193,474,496]" page="560" pageId="13" pageNumber="314" year="1999">
<bibRefCitation id="9D354B5C291B1605D578FE79FBA3FE20" author="Campos" box="[850,1138,474,496]" pageId="13" pageNumber="314" refString="Campos, M. R. &amp; Lemaitre, R. (1999) Two new freshwater crabs of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) from Panama, with notes on the distribution of the genus. Proceedings of the Biological Society of Washington, 112 (3), 553 - 561." type="journal article" year="1999">Campos &amp; Lemaitre 1999</bibRefCitation>
: 560
</treatmentCitation>
(map).—
<treatmentCitation id="780510BC291B1605D324FE79FE49FDDF" author="Rodriguez" page="423" pageId="13" pageNumber="314" year="2000">
<bibRefCitation id="9D354B5C291B1605D324FE79FEB8FDDF" author="Rodriguez" pageId="13" pageNumber="314" refString="Rodriguez, G. &amp; Hedstrom, I. (2000) The freswater crabs of the Barbilla National Park, Costa Rica (Crustacea: Brachyura: Pseudothelphusidae), with notes on the evolution of structures for spermatophore retention. Proceedings of the Biological Society of Washington, 113 (2), 420 - 425." type="journal article" year="2000">Rodríguez &amp; Hedström 2000</bibRefCitation>
: 423
</treatmentCitation>
.—
<treatmentCitation id="780510BC291B1605D79FFE5AFCCFFDDF" author="Rodriguez" box="[437,798,505,527]" page="356" pageId="13" pageNumber="314" year="2005">
<bibRefCitation id="9D354B5C291B1605D79FFE5AFD3FFDDF" author="Rodriguez" box="[437,750,505,527]" pageId="13" pageNumber="314" refString="Rodriguez, G. &amp; Magalhaes, C. (2005) Recent advances in the biology of the Neotropical freshwater crab family of Pseudothelphusidae (Crustacea, Decapoda, Brachyura). Revista Brasileira de Zoologia, 22 (2), 354 - 365. http: // dx. doi. org / 10.1590 / s 0101 - 81752005000200009" type="journal article" year="2005">Rodríguez &amp; Magalhães 2005</bibRefCitation>
: 356
</treatmentCitation>
(table I).—
<treatmentCitation id="780510BC291B1605D5BFFE59FB8CFDDF" author="Ng" box="[917,1117,505,527]" page="176" pageId="13" pageNumber="314" year="2008">
<bibRefCitation id="9D354B5C291B1605D5BFFE59FBFCFDDF" author="Ng" box="[917,1069,505,527]" pageId="13" pageNumber="314" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF291B1605D596FE59FC20FDDF" box="[956,1009,505,527]" italics="true" pageId="13" pageNumber="314">et al.</emphasis>
2008
</bibRefCitation>
: 176
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC291B1605D2E6FE5AFE52FDFE" author="Villalobos" page="297" pageId="13" pageNumber="314" year="2008">
<bibRefCitation id="9D354B5C291B1605D2E6FE5AFE85FDFE" author="Villalobos" pageId="13" pageNumber="314" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 297
</treatmentCitation>
(in list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1605D6BDFDF9FB14FC43" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="F91B36AD291B1605D6BDFDF9FB14FC43" blockId="13.[151,1436,602,1383]" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6BDFDF9FEAEFDA3" bold="true" box="[151,383,602,627]" pageId="13" pageNumber="314">Material examined</emphasis>
.
<collectingCountry id="81B3763D291B1605D7B9FDF8FDDFFDA3" box="[403,526,603,627]" name="Panama" pageId="13" pageNumber="314">PANAMA</collectingCountry>
.
<taxonomicName id="3EA44D2E291B1605D434FDF8FD8BFDA2" box="[542,602,603,626]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="338" phylum="Arthropoda" rank="species" species="kuna">Kuna</taxonomicName>
Yala Comarca, Atlantic drainage:
<specimenCount id="EFA2FD24291B1605D5D7FDF8FB8DFDA3" box="[1021,1116,602,627]" pageId="13" pageNumber="314" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD24291B1605D247FDF8FB33FDA3" box="[1133,1250,602,627]" pageId="13" pageNumber="314" type="female">2 females</specimenCount>
,
<specimenCount id="EFA2FD24291B1605D2D9FDF8FAEAFDA3" box="[1267,1339,602,627]" pageId="13" pageNumber="314" type="generic">4 juv.</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D361FDF8FA4DFDA3" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,603,627]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
<date id="8D1A106D291B1605D6BDFDDDFF3FFD47" box="[151,238,638,663]" pageId="13" pageNumber="314" value="05-01-24">240105</date>
, tributary of Carti Grande River, at trail NW from Nuragandi [Nusagandi Natural Reserve?],
<date id="8D1A106D291B1605D300FDDDFA49FD47" box="[1322,1432,638,663]" pageId="13" pageNumber="314" value="1985-03-05">5.iii.1985</date>
, R.W. Bouchard leg.;
<specimenCount id="EFA2FD24291B1605D7A9FD00FE0CFD6B" box="[387,477,674,699]" pageId="13" pageNumber="314" type="male">3 males</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D7C2FD00FDE8FD6B" LSID="urn:lsid:biocol.org:col:34871" box="[488,569,675,699]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
<date id="8D1A106D291B1605D46BFD00FD46FD6B" box="[577,663,675,699]" pageId="13" pageNumber="314" value="10-01-24">240110</date>
, Pingandi Creek at Llano-Carti road,
<date id="8D1A106D291B1605D217FD01FB7AFD6B" box="[1085,1195,674,699]" pageId="13" pageNumber="314" value="1985-03-04">4.iii.1985</date>
, R.W. Bouchard leg.;
<specimenCount id="EFA2FD24291B1605D6BDFD64FF36FD0F" box="[151,231,710,735]" pageId="13" pageNumber="314" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D6DEFD64FE94FD0F" LSID="urn:lsid:biocol.org:col:34871" box="[244,325,711,735]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
<date id="8D1A106D291B1605D764FD64FE72FD0F" box="[334,419,711,735]" pageId="13" pageNumber="314" value="11-01-24">240111</date>
, Nuragandi [Nusagandi?] off Llando-Carti Rd,
<date id="8D1A106D291B1605D5E1FD65FBE8FD0F" box="[971,1081,710,735]" pageId="13" pageNumber="314" value="1985-03-01">1.iii.1985</date>
, R.W. Bouchard leg.—
<collectingCountry id="81B3763D291B1605D36AFD65FA4DFD0F" box="[1344,1436,710,735]" name="Panama" pageId="13" pageNumber="314">Panama</collectingCountry>
Province, Pacific drainage: male,
<typeStatus id="261F880F291B1605D403FD49FD5FFCD3" box="[553,654,746,771]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
,
<collectionCode id="9FB5AE68291B1605D4B5FD48FD21FCD3" LSID="urn:lsid:biocol.org:col:34871" box="[671,752,747,771]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
<date id="8D1A106D291B1605D4D7FD49FC85FCD3" box="[765,852,746,771]" pageId="13" pageNumber="314" value="06-01-24">240106</date>
, 16 km from Cerro Azul, approximately
<quantity id="3E5C9B48291B1605D362FD48FA46FCD2" box="[1352,1431,747,771]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="314" unit="m" value="330.0">330 m</quantity>
,
<date id="8D1A106D291B1605D6BDFCADFED2FCF7" box="[151,259,782,807]" pageId="13" pageNumber="314" value="1971-01-29">29.i.1971</date>
, L.G. Abele &amp; R. Dressler leg.;
<specimenCount id="EFA2FD24291B1605D44FFCACFD14FCF7" box="[613,709,782,807]" pageId="13" pageNumber="314" type="female">1 female</specimenCount>
<typeStatus id="261F880F291B1605D4E1FCACFCFFFCF7" box="[715,814,783,807]" pageId="13" pageNumber="314" type="paratype">paratype</typeStatus>
,
<specimenCount id="EFA2FD24291B1605D513FCADFC59FCF7" box="[825,904,782,807]" pageId="13" pageNumber="314" type="generic">36 juv.</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D5B8FCACFC32FCF7" LSID="urn:lsid:biocol.org:col:34871" box="[914,995,783,807]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
<date id="8D1A106D291B1605D5C3FCACFB91FCF7" box="[1001,1088,783,807]" pageId="13" pageNumber="314" value="07-01-24">240107</date>
, same data;
<specimenCount id="EFA2FD24291B1605D2EDFCACFACEFCF7" box="[1223,1311,782,807]" pageId="13" pageNumber="314" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD24291B1605D300FCACFA49FCF7" box="[1322,1432,782,807]" pageId="13" pageNumber="314" type="female">4 females</specimenCount>
,
<specimenCount id="EFA2FD24291B1605D6BDFC90FF36FC9B" box="[151,231,818,843]" pageId="13" pageNumber="314" type="generic">22 juv.</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D6D9FC90FE95FC9B" LSID="urn:lsid:biocol.org:col:34871" box="[243,324,819,843]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
<date id="8D1A106D291B1605D766FC90FE72FC9B" box="[332,419,819,843]" pageId="13" pageNumber="314" value="08-01-24">240108</date>
, same data;
<specimenCount id="EFA2FD24291B1605D41AFC90FD5BFC9B" box="[560,650,818,843]" pageId="13" pageNumber="314" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD24291B1605D4BDFC90FD2DFC9B" box="[663,764,818,843]" pageId="13" pageNumber="314" type="female">1 female</specimenCount>
,
<specimenCount id="EFA2FD24291B1605D522FC90FC9AFC9B" box="[776,843,818,843]" pageId="13" pageNumber="314" type="generic">3 juv.</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D57CFC90FC76FC9B" LSID="urn:lsid:biocol.org:col:34871" box="[854,935,819,843]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
<date id="8D1A106D291B1605D59AFC90FBD6FC9B" box="[944,1031,819,843]" pageId="13" pageNumber="314" value="09-01-24">240109</date>
, tributary of Pacora River, creek by road, about
<quantity id="3E5C9B48291B1605D736FCF4FE9FFCBF" box="[284,334,854,879]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="13" pageNumber="314" unit="mi" value="8.0">8 mi</quantity>
N of Cerro Azul,
<date id="8D1A106D291B1605D430FCF5FD75FCBF" box="[538,676,854,879]" pageId="13" pageNumber="314" value="1962-08-27">27.viii.1962</date>
, H. Lofkin &amp; Kosan leg.;
<specimenCount id="EFA2FD24291B1605D5E2FCF4FBC7FCBF" box="[968,1046,854,879]" pageId="13" pageNumber="314" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D20BFCF4FBB1FCBF" box="[1057,1120,855,879]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="13" pageNumber="314">INPA</collectionCode>
1525, same data.—Chiriquí Province, Pacific drainage:
<specimenCount id="EFA2FD24291B1605D7E6FCD8FDCBFC43" box="[460,538,890,915]" pageId="13" pageNumber="314" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE68291B1605D40CFCD8FDA6FC43" LSID="urn:lsid:biocol.org:col:34871" box="[550,631,891,915]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
1089237, Cerro Punta,
<date id="8D1A106D291B1605D5A8FCD9FBDDFC43" box="[898,1036,890,915]" pageId="13" pageNumber="314" value="1963-03-01" valueMax="1963-03-03" valueMin="1963-03-01">1–3.iii.1963</date>
, E.J. Morris leg.
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1605D6EDFC3DFE01FB07" pageId="13" pageNumber="314" type="description">
<paragraph id="F91B36AD291B1605D6EDFC3DFE01FB07" blockId="13.[151,1436,602,1383]" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6EDFC3DFE3CFC67" bold="true" box="[199,493,926,951]" pageId="13" pageNumber="314">Description of gonopod</emphasis>
. Straight in mesiocaudal view, apex bent approximately 90° in laterocephalic direction. Marginal suture on mesial side, straight; marginal process subretangular, not produced beyond distal border of apex. Mesial process small, subtriangular, slightly downturned, with one short spine on laterocephalic side. Lateral process lengthy, strongly bilobed, approximately 80% of stem length (from proximal opening to caudal border of apex), with deep notch in middle; lateral border with deep median incision, proximal lobe narrower, wider than the distal one, surpassing in width the lateral end of apex; distal lobe as wide as lateral end of apex. Distocaudal ridge short, narrow, weakly separated from distal border of lateral process by shallow depression. Apex oblong, narrow, with narrow notch on lateral end. Field of apical spines well developed, facing towards cephalolateral side.
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1605D6EDFB41FE75FACF" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="F91B36AD291B1605D6EDFB41FE75FACF" blockId="13.[151,1436,602,1383]" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6EDFB41FDD5FB2B" bold="true" box="[199,516,1250,1275]" pageId="13" pageNumber="314">
<typeStatus id="261F880F291B1605D6EDFB41FEE5FB2B" box="[199,308,1250,1275]" pageId="13" pageNumber="314" type="holotype">Holotype</typeStatus>
and
<typeStatus id="261F880F291B1605D759FB41FE77FB2B" box="[371,422,1250,1275]" pageId="13" pageNumber="314">type</typeStatus>
locality
</emphasis>
. Male, cw 42.4, cl 25.5,
<collectionCode id="9FB5AE68291B1605D50DFB40FCA9FB2B" LSID="urn:lsid:biocol.org:col:34871" box="[807,888,1251,1275]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="314">USNM</collectionCode>
<date id="8D1A106D291B1605D5ABFB41FC08FB2B" box="[897,985,1250,1275]" pageId="13" pageNumber="314" value="06-01-24">240106</date>
.
<collectingCountry id="81B3763D291B1605D5CFFB40FBEFFB2A" box="[997,1086,1251,1274]" name="Panama" pageId="13" pageNumber="314">Panama</collectingCountry>
Province,
<collectingCountry id="81B3763D291B1605D290FB40FAC7FB2A" box="[1210,1302,1251,1274]" name="Panama" pageId="13" pageNumber="314">Panama</collectingCountry>
, near Cerro Azul (Pacific drainage).
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1605D6EDFA89FB18FA93" box="[199,1225,1322,1347]" pageId="13" pageNumber="314" type="distribution">
<paragraph id="F91B36AD291B1605D6EDFA89FB18FA93" blockId="13.[151,1436,602,1383]" box="[199,1225,1322,1347]" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6EDFA89FE88FA93" bold="true" box="[199,345,1322,1347]" pageId="13" pageNumber="314">Distribution</emphasis>
. Western and central
<collectingCountry id="81B3763D291B1605D466FA88FD79FA92" box="[588,680,1323,1346]" name="Panama" pageId="13" pageNumber="314">Panama</collectingCountry>
, in both Pacific and Atlantic drainages (
<figureCitation id="619F2A28291B1605D242FA89FB6DFA93" box="[1128,1212,1322,1347]" captionStart="FIGURE 78" captionStartId="32.[151,250,1218,1240]" captionTargetBox="[156,1432,201,1194]" captionTargetId="figure@32.[151,1436,193,1196]" captionTargetPageId="32" captionText="FIGURE 78. Distribution map of Ptychophallus cocleensis Pretzmann, 1965, P. lavallensis Pretzmann, 1978, and P. micracanthus Rodríguez, 1994." httpUri="https://zenodo.org/record/235136/files/figure.png" pageId="13" pageNumber="314">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1605D6EDFAEDFCEBFAB6" box="[199,826,1358,1383]" pageId="13" pageNumber="314" type="discussion">
<paragraph id="F91B36AD291B1605D6EDFAEDFCEBFAB6" blockId="13.[151,1436,602,1383]" box="[199,826,1358,1383]" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6EDFAEDFEEAFAB7" bold="true" box="[199,315,1358,1383]" pageId="13" pageNumber="314">Remarks.</emphasis>
See remarks for
<taxonomicName id="3EA44D2E291B1605D7D0FAEDFCE2FAB6" box="[506,819,1358,1383]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="CBD0EABF291B1605D7D0FAEDFCE2FAB6" box="[506,819,1358,1383]" italics="true" pageId="13" pageNumber="314">Ptychophallus costaricensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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