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<mods:title id="538A6FB3BADA1B190EBB3432F4AFAAB1">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<mods:namePart id="C5A8CD24DA165B13F56F05011BFDBEAB">Magalhães, Célio</mods:namePart>
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<mods:namePart id="FED090D45113D40C4A6DD332825CC915">Wehrtmann, Ingo S.</mods:namePart>
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<mods:namePart id="86EF896C40304EDE4CE52E5DED8F3169">Lara, Luis Rólier</mods:namePart>
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<mods:namePart id="845EE05062338AF2EAD9C04980987BAA">Mantelatto, Fernando L.</mods:namePart>
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<treatment id="710D87BB291A1605D6BDFB44FD98FECD" ID-DOI="http://doi.org/10.5281/zenodo.5491520" ID-GBIF-Taxon="119590160" ID-Zenodo-Dep="5491520" LSID="urn:lsid:plazi:treatment:710D87BB291A1605D6BDFB44FD98FECD" httpUri="http://treatment.plazi.org/id/710D87BB291A1605D6BDFB44FD98FECD" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313">
<subSubSection id="B1BE6526291A1604D6BDFB44FE89FAF3" pageId="12" pageNumber="313" type="nomenclature">
<paragraph id="F91B36AD291A1604D6BDFB44FD75FAD2" blockId="12.[151,676,1255,1315]" box="[151,676,1255,1282]" pageId="12" pageNumber="313">
<heading id="A25381C1291A1604D6BDFB44FD75FAD2" bold="true" box="[151,676,1255,1282]" fontSize="11" level="1" pageId="12" pageNumber="313" reason="1">
<taxonomicName id="3EA44D2E291A1604D6BDFB44FD75FAD2" ID-CoL="78N6V" authority="Pretzmann, 1978" authorityName="Pretzmann" authorityYear="1978" box="[151,676,1255,1282]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="lavallensis">
<emphasis id="CBD0EABF291A1604D6BDFB44FD75FAD2" bold="true" box="[151,676,1255,1282]" pageId="12" pageNumber="313">
<emphasis id="CBD0EABF291A1604D6BDFB44FE1BFAD1" bold="true" box="[151,458,1255,1281]" italics="true" pageId="12" pageNumber="313">Ptychophallus lavallensis</emphasis>
<bibRefCitation id="9D354B5C291A1604D7FBFB4BFD75FAD2" author="Pretzmann" box="[465,676,1256,1282]" pageId="12" pageNumber="313" refString="Pretzmann, G. (1978) Neue Potamocarcinini, Poglayen-Neuwall leg. 1975 (vorlaufige Mitteilung). Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch Naturwissenchaftliche Klasse, Series 1, 1978 (2), 51 - 54." type="journal article" year="1978">Pretzmann, 1978</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F91B36AD291A1604D6BDFAA9FE89FAF3" blockId="12.[151,676,1255,1315]" box="[151,344,1290,1315]" pageId="12" pageNumber="313">
(
<figureCitation id="619F2A28291A1604D6B5FAA9FEF7FAF3" box="[159,294,1290,1315]" captionStart="FIGURES 22 32" captionStartId="11.[151,264,1746,1769]" captionTargetBox="[156,1427,204,1718]" captionTargetId="figure@11.[156,1431,195,1725]" captionTargetPageId="11" captionText="FIGURES 22 32. Ptychophallus exilipes (Rathbun, 1898): 22, left first gonopod, USNM 39089, caudo-mesial view; 23, same data, latero-cephalic view of distal part; 24 26, outline of right first gonopod from three males of UCR-MZ 741, caudo-mesial view; 27, outline of right first gonopod, USNM 39096, caudo-mesial view; 28, same data, INPA 1857, caudo-mesial view; 29, outline of left first gonopod, SMF 36265, caudo-mesial view; 30, same data, SMF 36264, caudo-mesial view. Ptychophallus lavallensis Pretzmann, 1978: 31, left first gonopod, holotype, NHMW 4002, caudo-mesial view; 32, latero-cephalic view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/235130/files/figure.png" pageId="12" pageNumber="313">Figs. 31, 32</figureCitation>
,
<figureCitation id="619F2A28291A1604D718FAA8FE9EFAF3" box="[306,335,1291,1315]" captionStart="FIGURE 78" captionStartId="32.[151,250,1218,1240]" captionTargetBox="[156,1432,201,1194]" captionTargetId="figure@32.[151,1436,193,1196]" captionTargetPageId="32" captionText="FIGURE 78. Distribution map of Ptychophallus cocleensis Pretzmann, 1965, P. lavallensis Pretzmann, 1978, and P. micracanthus Rodríguez, 1994." httpUri="https://zenodo.org/record/235136/files/figure.png" pageId="12" pageNumber="313">78</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291A1604D6BDFAF1FBB7FA76" pageId="12" pageNumber="313" type="reference_group">
<paragraph id="F91B36AD291A1604D6BDFAF1FBB7FA76" blockId="12.[151,1436,1362,1446]" pageId="12" pageNumber="313">
<treatmentCitationGroup id="D9B41183291A1604D6BDFAF1FBB7FA76" pageId="12" pageNumber="313">
<taxonomicName id="3EA44D2E291A1604D6BDFAF1FC8CFAB8" ID-CoL="78N6V" authority="Pretzmann, 1978: 1" authorityName="Pretzmann" authorityPageNumber="1" authorityYear="1978" box="[151,861,1362,1384]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="subSpecies" species="exilipes" subGenus="Ptychophallus" subSpecies="lavallensis">
<emphasis id="CBD0EABF291A1604D6BDFAF1FEF9FAB8" box="[151,296,1362,1384]" italics="true" pageId="12" pageNumber="313">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF291A1604D71CFAF1FE16FAB8" box="[310,455,1362,1384]" italics="true" pageId="12" pageNumber="313">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF291A1604D7FCFAF1FD47FAB8" box="[470,662,1362,1384]" italics="true" pageId="12" pageNumber="313">exilipes lavallensis</emphasis>
<treatmentCitation id="780510BC291A1604D4B6FAF0FC8CFAB8" author="Pretzmann" box="[668,861,1362,1384]" page="1" pageId="12" pageNumber="313" year="1978">
<bibRefCitation id="9D354B5C291A1604D4B6FAF0FC9AFAB8" author="Pretzmann" box="[668,843,1362,1384]" pageId="12" pageNumber="313" refString="Pretzmann, G. (1978) Neue Potamocarcinini, Poglayen-Neuwall leg. 1975 (vorlaufige Mitteilung). Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch Naturwissenchaftliche Klasse, Series 1, 1978 (2), 51 - 54." type="journal article" year="1978">Pretzmann, 1978</bibRefCitation>
: 1
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="780510BC291A1604D551FAF1FB8DFAB8" author="Pretzmann" box="[891,1116,1362,1384]" page="651" pageId="12" pageNumber="313" year="1980">
<bibRefCitation id="9D354B5C291A1604D551FAF1FBFAFAB8" author="Pretzmann" box="[891,1067,1362,1384]" pageId="12" pageNumber="313" refString="Pretzmann, G. (1980) Von Dr. Ivo Poglayen-Neuwall 1975 in Mittelamerika gesammelte Krabben. Annalen des Naturhistorischen Museums, 83, 651 - 666, pls. 1 - 18. [Wien]" type="journal article" year="1980">Pretzmann 1980</bibRefCitation>
: 651
</treatmentCitation>
, pl. 1, figs. 15.
<taxonomicName id="3EA44D2E291A1604D6BDFAD2FE4BFA57" ID-CoL="78N6V" box="[151,410,1393,1415]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="lavallensis">
<emphasis id="CBD0EABF291A1604D6BDFAD2FE4BFA57" box="[151,410,1393,1415]" italics="true" pageId="12" pageNumber="313">Ptychophallus lavallensis</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC291A1604D79FFAD2FD5CFA57" author="Rodriguez" box="[437,653,1393,1415]" page="304" pageId="12" pageNumber="313" year="1994">
<bibRefCitation id="9D354B5C291A1604D79FFAD2FD8BFA57" author="Rodriguez" box="[437,602,1393,1415]" pageId="12" pageNumber="313" refString="Rodriguez, G. (1994) A revision of the type material of some species of Hypolobocera and Ptychophallus (Crustacea: Decapoda: Pseudothelphusidae) in the National Museum of Natural History, Washington, D. C., with descriptions of a new species and a new subspecies. Proceedings of the Biological Society of Washington, 107 (2), 296 - 307." type="journal article" year="1994">Rodríguez 1994</bibRefCitation>
: 304
</treatmentCitation>
, fig.
<date id="8D1A106D291A1604D4E8FAD2FD05FA57" box="[706,724,1393,1415]" pageId="12" pageNumber="313">3I</date>
J.—
<treatmentCitation id="780510BC291A1604D520FAD2FB9BFA57" author="Campos" box="[778,1098,1393,1415]" page="560" pageId="12" pageNumber="313" year="1999">
<bibRefCitation id="9D354B5C291A1604D520FAD2FBC8FA57" author="Campos" box="[778,1049,1393,1415]" pageId="12" pageNumber="313" refString="Campos, M. R. &amp; Lemaitre, R. (1999) Two new freshwater crabs of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) from Panama, with notes on the distribution of the genus. Proceedings of the Biological Society of Washington, 112 (3), 553 - 561." type="journal article" year="1999">Campos &amp; Lemaitre 1999</bibRefCitation>
: 560
</treatmentCitation>
(map).—
<treatmentCitation id="780510BC291A1604D283FAD2FEFFFA76" author="Rodriguez" page="424" pageId="12" pageNumber="313" year="2000">
<bibRefCitation id="9D354B5C291A1604D283FAD2FF2FFA76" author="Rodriguez" pageId="12" pageNumber="313" refString="Rodriguez, G. &amp; Hedstrom, I. (2000) The freswater crabs of the Barbilla National Park, Costa Rica (Crustacea: Brachyura: Pseudothelphusidae), with notes on the evolution of structures for spermatophore retention. Proceedings of the Biological Society of Washington, 113 (2), 420 - 425." type="journal article" year="2000">Rodríguez &amp; Hedström 2000</bibRefCitation>
: 424
</treatmentCitation>
.—
<treatmentCitation id="780510BC291A1604D766FA33FDC0FA76" author="Ng" box="[332,529,1424,1446]" page="176" pageId="12" pageNumber="313" year="2008">
<bibRefCitation id="9D354B5C291A1604D766FA33FE30FA76" author="Ng" box="[332,481,1424,1446]" pageId="12" pageNumber="313" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF291A1604D758FA32FE74FA76" box="[370,421,1424,1446]" italics="true" pageId="12" pageNumber="313">et al.</emphasis>
2008
</bibRefCitation>
: 176
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC291A1604D457FA33FBDEFA76" author="Villalobos" box="[637,1039,1424,1446]" page="297" pageId="12" pageNumber="313" year="2008">
<bibRefCitation id="9D354B5C291A1604D457FA33FC0EFA76" author="Villalobos" box="[637,991,1424,1446]" pageId="12" pageNumber="313" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 297
</treatmentCitation>
(in list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291A1604D6BDFA71FC68F94F" pageId="12" pageNumber="313" type="materials_examined">
<paragraph id="F91B36AD291A1604D6BDFA71FC68F94F" blockId="12.[151,1437,1490,2019]" pageId="12" pageNumber="313">
<emphasis id="CBD0EABF291A1604D6BDFA71FE50FA3B" bold="true" box="[151,385,1490,1515]" pageId="12" pageNumber="313">Material examined</emphasis>
.
<collectingCountry id="81B3763D291A1604D7BDFA70FDC3FA3B" box="[407,530,1491,1515]" name="Panama" pageId="12" pageNumber="313">PANAMA</collectingCountry>
. Colón Province, Atlantic drainage:
<specimenCount id="EFA2FD24291A1604D5FBFA70FBF6FA3B" box="[977,1063,1490,1515]" pageId="12" pageNumber="313" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE68291A1604D210FA70FB5AFA3B" LSID="urn:lsid:biocol.org:col:34871" box="[1082,1163,1491,1515]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="313">USNM</collectionCode>
184338, El Aguacate,
<date id="8D1A106D291A1604D6BDFA55FEDBF9DF" box="[151,266,1526,1551]" pageId="12" pageNumber="313" value="1973-02-22">22.ii.1973</date>
, leg. unknown;
<specimenCount id="EFA2FD24291A1604D7E3FA54FDCAF9DF" box="[457,539,1526,1551]" pageId="12" pageNumber="313" type="male">1 male</specimenCount>
,
<collectionCode id="9FB5AE68291A1604D400FA54FDAAF9DF" LSID="urn:lsid:biocol.org:col:34871" box="[554,635,1527,1551]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="313">USNM</collectionCode>
184339, El Aguacate,
<date id="8D1A106D291A1604D5A1FA55FC2EF9DF" box="[907,1023,1526,1551]" pageId="12" pageNumber="313" value="1973-02-22">22.ii.1973</date>
, leg. unknown.—
<collectingCountry id="81B3763D291A1604D2E6FA55FAF6F9DF" box="[1228,1319,1526,1551]" name="Panama" pageId="12" pageNumber="313">Panamá</collectingCountry>
Province, Pacific drainage:
<specimenCount id="EFA2FD24291A1604D776F9B8FE64F9E3" box="[348,437,1562,1587]" pageId="12" pageNumber="313" type="male">2 males</specimenCount>
,
<collectionCode id="9FB5AE68291A1604D7EAF9B8FE2EF9E3" box="[448,511,1563,1587]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="12" pageNumber="313">INPA</collectionCode>
1711,
<specimenCount id="EFA2FD24291A1604D466F9B8FD5AF9E3" box="[588,651,1562,1587]" pageId="12" pageNumber="313" type="generic">1 juv.</specimenCount>
male,
<collectionCode id="9FB5AE68291A1604D4FDF9B8FCF2F9E3" box="[727,803,1563,1587]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="12" pageNumber="313">CCDB</collectionCode>
2660, Capira River,
<date id="8D1A106D291A1604D224F9B9FB5BF9E3" box="[1038,1162,1562,1587]" pageId="12" pageNumber="313" value="2008-08-02">2.viii.2008</date>
, L.S. Torati leg.—Coclé Province, Pacific drainage:
<specimenCount id="EFA2FD24291A1604D7CEF99CFD94F987" box="[484,581,1598,1623]" pageId="12" pageNumber="313" type="male">3 males</specimenCount>
,
<collectionCode id="9FB5AE68291A1604D473F99CFD7BF987" LSID="urn:lsid:biocol.org:col:34871" box="[601,682,1599,1623]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="313">USNM</collectionCode>
<date id="8D1A106D291A1604D492F99CFCDEF987" box="[696,783,1599,1623]" pageId="12" pageNumber="313" value="00-01-24">240100</date>
, creek at headwaters of Indio River, above El Valle,
<date id="8D1A106D291A1604D6BDF9C1FEC0F9AB" box="[151,273,1634,1659]" pageId="12" pageNumber="313" value="1962-09-14">14.ix.1962</date>
, H. Loftin
<emphasis id="CBD0EABF291A1604D7B5F9C7FE0CF9AB" box="[415,477,1634,1659]" italics="true" pageId="12" pageNumber="313">et al.</emphasis>
leg.; male,
<typeStatus id="261F880F291A1604D45EF9C1FD08F9AB" box="[628,729,1634,1659]" pageId="12" pageNumber="313" type="holotype">holotype</typeStatus>
,
<collectionCode id="9FB5AE68291A1604D4C0F9C0FC97F9AA" box="[746,838,1635,1658]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="313">NHMW</collectionCode>
4002, brook above El Valle,
<quantity id="3E5C9B48291A1604D298F9C1FAD3F9AA" box="[1202,1282,1634,1659]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="12" pageNumber="313" unit="m" value="670.0">670 m</quantity>
,
<date id="8D1A106D291A1604D33EF9C1FAABF9AB" box="[1300,1402,1634,1659]" pageId="12" pageNumber="313" value="1975-02-02">2.ii.1975</date>
, I. Poglayen-Neuwall leg.;
<specimenCount id="EFA2FD24291A1604D78CF924FDC4F94F" box="[422,533,1670,1695]" pageId="12" pageNumber="313" type="female">4 females</specimenCount>
,
<typeStatus id="261F880F291A1604D40AF924FD5FF94F" box="[544,654,1671,1695]" pageId="12" pageNumber="313" type="paratype">paratypes</typeStatus>
,
<collectionCode id="9FB5AE68291A1604D4B0F924FD27F94E" box="[666,758,1671,1694]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="313">NHMW</collectionCode>
4002, same data.
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291A1604D6EDF909FA5AF84B" pageId="12" pageNumber="313" type="description">
<paragraph id="F91B36AD291A1604D6EDF909FA5AF84B" blockId="12.[151,1437,1490,2019]" pageId="12" pageNumber="313">
<emphasis id="CBD0EABF291A1604D6EDF909FE0DF913" bold="true" box="[199,476,1706,1731]" pageId="12" pageNumber="313">Description of gonopod</emphasis>
. Straight in mesiocaudal view, apex strongly bent over 90° in laterocephalic direction. Marginal suture on mesial side, straight; marginal process rounded, not produced beyond caudal border of apex. Mesial process small, subtriangular, downturned. Lateral process well developed, broadly subtriangular, undivided, approximately 70% of stem length (from proximal opening to caudal border of apex), much wider than lateral end of apex; lateral border distinctly convex, proximal, distal lobes indistinct. Distocaudal ridge short, weakly developed, not well separated from the distal border of lateral process. Apex oblong, narrow; caudal border nearly straight, with small notch at or near lateral end. Field of apical spines well developed, facing towards lateral side.
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291A1604D6EDF805FEB7F833" pageId="12" pageNumber="313" type="materials_examined">
<paragraph id="F91B36AD291A1604D6EDF805FEB7F833" blockId="12.[151,1437,1490,2019]" pageId="12" pageNumber="313">
<emphasis id="CBD0EABF291A1604D6EDF805FDD6F86F" bold="true" box="[199,519,1958,1983]" pageId="12" pageNumber="313">
<typeStatus id="261F880F291A1604D6EDF805FEE5F86F" box="[199,308,1958,1983]" pageId="12" pageNumber="313" type="holotype">Holotype</typeStatus>
and
<typeStatus id="261F880F291A1604D75EF805FE76F86F" box="[372,423,1958,1983]" pageId="12" pageNumber="313">type</typeStatus>
locality
</emphasis>
. Male, cw 31.2, cl 18.7,
<collectionCode id="9FB5AE68291A1604D504F804FC5BF86E" box="[814,906,1959,1982]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="313">NHMW</collectionCode>
4002.
<collectingCountry id="81B3763D291A1604D5F7F804FBE9F86E" box="[989,1080,1959,1982]" name="Panama" pageId="12" pageNumber="313">Panama</collectingCountry>
, Coclé Province, near El Valle (Pacific drainage).
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1605D6EDFF34FD89FF04" pageId="13" pageNumber="314" type="distribution">
<paragraph id="F91B36AD291B1605D6EDFF34FD89FF04" blockId="13.[151,1436,151,285]" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6EDFF34FE88FF60" bold="true" box="[199,345,151,176]" pageId="13" pageNumber="314">Distribution</emphasis>
. The few records available suggest that the species is distributed in central
<collectingCountry id="81B3763D291B1605D2F4FF3BFAEBFF7F" box="[1246,1338,152,175]" name="Panama" pageId="13" pageNumber="314">Panama</collectingCountry>
, in both Atlantic and Pacific drainages (
<figureCitation id="619F2A28291B1605D7DDFF1FFD9AFF04" box="[503,587,188,213]" captionStart="FIGURE 78" captionStartId="32.[151,250,1218,1240]" captionTargetBox="[156,1432,201,1194]" captionTargetId="figure@32.[151,1436,193,1196]" captionTargetPageId="32" captionText="FIGURE 78. Distribution map of Ptychophallus cocleensis Pretzmann, 1965, P. lavallensis Pretzmann, 1978, and P. micracanthus Rodríguez, 1994." httpUri="https://zenodo.org/record/235136/files/figure.png" pageId="13" pageNumber="314">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291B1605D6EDFF7CFD98FECD" pageId="13" pageNumber="314" type="discussion">
<paragraph id="F91B36AD291B1605D6EDFF7CFD98FECD" blockId="13.[151,1436,151,285]" pageId="13" pageNumber="314">
<emphasis id="CBD0EABF291B1605D6EDFF7CFEE5FF28" bold="true" box="[199,308,223,248]" pageId="13" pageNumber="314">Remarks</emphasis>
. This species is very closely related to
<taxonomicName id="3EA44D2E291B1605D520FF42FCAFFF28" box="[778,894,223,248]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="exilipes">
<emphasis id="CBD0EABF291B1605D520FF42FCAFFF28" box="[778,894,223,248]" italics="true" pageId="13" pageNumber="314">P. exilipes</emphasis>
</taxonomicName>
, and their possible differences were discussed under the remarks of the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>