276 lines
31 KiB
XML
276 lines
31 KiB
XML
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<mods:title id="CC9F59B33CB515651A06A4905745E4F8">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<mods:namePart id="FF0DF4DFEAFEC9430FF858CDDE8AF993">Magalhães, Célio</mods:namePart>
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<mods:namePart id="75351F7C3ED6ECDFCE71D3EA28815E11">Lara, Luis Rólier</mods:namePart>
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<mods:namePart id="C55CFC919F10B1ED6CE632D832BC17D9">Mantelatto, Fernando L.</mods:namePart>
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<treatment id="710D87BB2915160DD6BDFA9CFBC6FDEC" ID-DOI="http://doi.org/10.5281/zenodo.5491504" ID-GBIF-Taxon="119590152" ID-Zenodo-Dep="5491504" LSID="urn:lsid:plazi:treatment:710D87BB2915160DD6BDFA9CFBC6FDEC" httpUri="http://treatment.plazi.org/id/710D87BB2915160DD6BDFA9CFBC6FDEC" lastPageId="5" lastPageNumber="306" pageId="3" pageNumber="304">
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<subSubSection id="B1BE65262915160BD6BDFA9CFEEDFAAB" pageId="3" pageNumber="304" type="nomenclature">
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<paragraph id="F91B36AD2915160BD6BDFA9CFD9BFA8A" blockId="3.[151,586,1343,1403]" box="[151,586,1343,1370]" pageId="3" pageNumber="304">
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<heading id="A25381C12915160BD6BDFA9CFD9BFA8A" bold="true" box="[151,586,1343,1370]" fontSize="11" level="1" pageId="3" pageNumber="304" reason="1">
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<taxonomicName id="3EA44D2E2915160BD6BDFA9CFD9BFA8A" ID-CoL="92SC6" authority="Hobbs, 1991" authorityName="Hobbs" authorityYear="1991" box="[151,586,1343,1370]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="puntarenas">
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<emphasis id="CBD0EABF2915160BD6BDFA9CFD9BFA8A" bold="true" box="[151,586,1343,1370]" pageId="3" pageNumber="304">
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<emphasis id="CBD0EABF2915160BD6BDFA9CFE44FA89" bold="true" box="[151,405,1343,1369]" italics="true" pageId="3" pageNumber="304">Achlidon puntarenas</emphasis>
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(
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<bibRefCitation id="9D354B5C2915160BD78FFAE3FD93FA8A" author="Hobbs" box="[421,578,1344,1370]" pageId="3" pageNumber="304" refString="Hobbs, H. H. III (1991) A new pseudothelphusid crab from a cave in southern Costa Rica (Decapoda: Brachyura). Proceedings of the Biological Society of Washington, 104 (2), 295 - 298." type="journal article" year="1991">Hobbs, 1991</bibRefCitation>
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)
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||
</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="F91B36AD2915160BD6BDFAC1FEEDFAAB" blockId="3.[151,586,1343,1403]" box="[151,316,1378,1403]" pageId="3" pageNumber="304">
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(
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||
<figureCitation id="619F2A282915160BD6B5FAC1FEDBFAAA" box="[159,266,1378,1403]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,264,1764,1787]" captionTargetBox="[152,1431,193,1735]" captionTargetId="figure@4.[152,1436,193,1743]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Achlidon agrestis (Rathbun, 1898): 1, left first gonopod (partially damaged), holotype, USNM 19487, mesial view; 2, same data, lateral view of distal part; 3, left first gonopod, NHMW 21827, cephalic view of distal part; 4, same data, apical view of apex; 5, same data, latero-cephalic view. Achlidon puntarenas (Hobbs, 1991): 6, left first gonopod, holotype, USNM 250555, caudo-mesial view; 7, same data, latero-cephalic view of distal part. Ptychophallus cocleensis Pretzmann, 1965: 8, left first gonopod, holotype, USNM 119869, lateral view of distal part; 9, same data, mesial view (dr, disto-caudal ridge; fs, field of apical spines; lp, lateral process; me, mesial process; mp, marginal process; ms, marginal suture). Scale bars = 1 mm." httpUri="https://zenodo.org/record/235128/files/figure.png" pageId="3" pageNumber="304">Figs. 6, 7</figureCitation>
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,
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<figureCitation id="619F2A282915160BD73CFAC0FEE2FAAB" box="[278,307,1379,1403]" captionStart="FIGURE 77" captionStartId="31.[151,250,2011,2033]" captionTargetBox="[157,1433,886,1986]" captionTargetId="figure@31.[150,1436,879,1989]" captionTargetPageId="31" captionText="FIGURE 77. Distribution map of Achlidon agrestis (Rathbun, 1898), and A. puntarenas (Hobbs, 1991)." httpUri="https://zenodo.org/record/235135/files/figure.png" pageId="3" pageNumber="304">77</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="B1BE65262915160BD6BDFA09FCFFFA2E" pageId="3" pageNumber="304" type="reference_group">
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<paragraph id="F91B36AD2915160BD6BDFA09FCFFFA2E" blockId="3.[151,1437,1450,1534]" pageId="3" pageNumber="304">
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<treatmentCitationGroup id="D9B411832915160BD6BDFA09FCFFFA2E" pageId="3" pageNumber="304">
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<taxonomicName id="3EA44D2E2915160BD6BDFA09FD74FA10" ID-CoL="935XX" authority="Hobbs, 1991: 295" authorityName="Hobbs" authorityPageNumber="295" authorityYear="1991" box="[151,677,1450,1472]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="puntarenas">
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<emphasis id="CBD0EABF2915160BD6BDFA09FE17FA10" box="[151,454,1450,1472]" italics="true" pageId="3" pageNumber="304">Pseudothelphusa puntarenas</emphasis>
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<treatmentCitation id="780510BC2915160BD7F2FA09FD74FA10" author="Hobbs" box="[472,677,1450,1472]" page="295" pageId="3" pageNumber="304" year="1991">
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<bibRefCitation id="9D354B5C2915160BD7F2FA09FDB9FA10" author="Hobbs" box="[472,616,1450,1472]" pageId="3" pageNumber="304" refString="Hobbs, H. H. III (1991) A new pseudothelphusid crab from a cave in southern Costa Rica (Decapoda: Brachyura). Proceedings of the Biological Society of Washington, 104 (2), 295 - 298." type="journal article" year="1991">Hobbs, 1991</bibRefCitation>
|
||
: 295
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||
</treatmentCitation>
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||
</taxonomicName>
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||
, fig. 1.—
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||
<treatmentCitation id="780510BC2915160BD53FFA09FBD7FA10" author="Rodriguez" box="[789,1030,1450,1472]" page="184" pageId="3" pageNumber="304" year="1992">
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<bibRefCitation id="9D354B5C2915160BD53FFA09FC1DFA10" author="Rodriguez" box="[789,972,1450,1472]" pageId="3" pageNumber="304" refString="Rodriguez, G. (1992) The freshwater crabs of America. Family Trichodactylidae and supplement to the family Pseudothelphusidae. ORSTOM, Paris, 189 pp. [Faune Tropicale 31]" type="book" year="1992">Rodríguez 1992</bibRefCitation>
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||
: 184
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||
</treatmentCitation>
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||
.—
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||
<treatmentCitation id="780510BC2915160BD20EFA09FB38FA10" author="Hobbs" box="[1060,1257,1450,1472]" page="101" pageId="3" pageNumber="304" year="1994">
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||
<bibRefCitation id="9D354B5C2915160BD20EFA09FB60FA10" author="Hobbs" box="[1060,1201,1450,1472]" pageId="3" pageNumber="304" refString="Hobbs, H. H. III (1994) Biogeography of subterranean decapods in North and Central America and the Caribbean region (Caridea, Astacidea, Brachyura). Hydrobiologia, 287, 95 - 104. http: // dx. doi. org / 10.1007 / bf 00006899" type="journal article" year="1994">Hobbs 1994</bibRefCitation>
|
||
: 101
|
||
</treatmentCitation>
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||
.—
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||
<treatmentCitation id="780510BC2915160BD322FA09FF3DFA0F" author="Suarez" page="156" pageId="3" pageNumber="304" year="2005">
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||
<bibRefCitation id="9D354B5C2915160BD322FA09FA48FA10" author="Suarez" box="[1288,1433,1450,1472]" pageId="3" pageNumber="304" refString="Suarez, H. (2005) A catalog of the type material in the IVIC Reference Collection of Decapoda, Venezuela. Journal of Crustacean Biology, 25 (1), 154 - 158. http: // dx. doi. org / 10.1651 / c- 2557" type="journal article" year="2005">Suárez 2005</bibRefCitation>
|
||
: 156
|
||
</treatmentCitation>
|
||
.—
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||
<treatmentCitation id="780510BC2915160BD723FA6AFD4AFA0F" author="Villalobos" box="[265,667,1481,1503]" page="294" pageId="3" pageNumber="304" year="2008">
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<bibRefCitation id="9D354B5C2915160BD723FA6AFDBDFA0F" author="Villalobos" box="[265,620,1481,1503]" pageId="3" pageNumber="304" refString="Villalobos Hiriart, J. L. & Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. & Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart & Álvarez 2008</bibRefCitation>
|
||
: 294
|
||
</treatmentCitation>
|
||
(in list).—
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||
<treatmentCitation id="780510BC2915160BD522FA6AFB83FA0F" author="Villalobos" box="[776,1106,1481,1503]" httpUri="http://treatment.plazi.org/id/038087A436283C68FC017895B9FAF92B" page="472" pageId="3" pageNumber="304" year="2010">
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<bibRefCitation id="9D354B5C2915160BD522FA6AFBF1FA0F" author="Villalobos" box="[776,1056,1481,1503]" pageId="3" pageNumber="304" refString="Villalobos, J. L. & Alvarez, F. (2010) Phylogenetic analysis of the Mexican freshwater crabs of the tribe Pseudothelphusini (Decapoda: Brachyura: Pseudothelphusidae). Zoological Journal of the Linnean Society, 160, 457 - 481. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2009.00606. x" type="journal article" year="2010">Villalobos & Alvarez 2010</bibRefCitation>
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||
: 472
|
||
</treatmentCitation>
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||
.
|
||
<taxonomicName id="3EA44D2E2915160BD6BDFA4BFE17FA2E" box="[151,454,1512,1534]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="punctarenas">
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||
<emphasis id="CBD0EABF2915160BD6BDFA4BFE17FA2E" box="[151,454,1512,1534]" italics="true" pageId="3" pageNumber="304">Pseudothelphusa punctarenas</emphasis>
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</taxonomicName>
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||
[sic]—
|
||
<treatmentCitation id="780510BC2915160BD43BFA4BFD06FA2E" author="Ng" box="[529,727,1512,1534]" page="176" pageId="3" pageNumber="304" year="2008">
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<bibRefCitation id="9D354B5C2915160BD43BFA4BFD79FA2E" author="Ng" box="[529,680,1512,1534]" pageId="3" pageNumber="304" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
|
||
Ng
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||
<emphasis id="CBD0EABF2915160BD412FA4AFDBDFA2E" box="[568,620,1512,1534]" italics="true" pageId="3" pageNumber="304">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
: 176
|
||
</treatmentCitation>
|
||
(in list).
|
||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="B1BE65262915160BD6BDF989FDE9F903" pageId="3" pageNumber="304" type="materials_examined">
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<paragraph id="F91B36AD2915160BD6BDF989FDE9F903" blockId="3.[151,1437,1578,2035]" pageId="3" pageNumber="304">
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<emphasis id="CBD0EABF2915160BD6BDF989FEACF993" bold="true" box="[151,381,1578,1603]" pageId="3" pageNumber="304">Material examined</emphasis>
|
||
.
|
||
<collectingCountry id="81B3763D2915160BD7A5F988FDEAF993" box="[399,571,1579,1603]" name="Costa Rica" pageId="3" pageNumber="304">COSTA RICA</collectingCountry>
|
||
.
|
||
<taxonomicName id="3EA44D2E2915160BD462F988FD12F992" box="[584,707,1579,1602]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
|
||
Province, Pacific drainage:
|
||
<specimenCount id="EFA2FD242915160BD23AF988FBB3F993" box="[1040,1122,1578,1603]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
|
||
,
|
||
<typeStatus id="261F880F2915160BD25BF989FB07F993" box="[1137,1238,1578,1603]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
|
||
,
|
||
<collectionCode id="9FB5AE682915160BD2CFF988FAE7F993" LSID="urn:lsid:biocol.org:col:34871" box="[1253,1334,1579,1603]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="304">USNM</collectionCode>
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||
<date id="8D1A106D2915160BD36BF989FA49F993" box="[1345,1432,1578,1603]" pageId="3" pageNumber="304" value="55-05-25">250555</date>
|
||
, small stream in Emus Cave,
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||
<date id="8D1A106D2915160BD7CCF9EDFD95F9B7" box="[486,580,1614,1639]" pageId="3" pageNumber="304" value="1989-01-08">8.i.1989</date>
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||
, F. Grady leg.;
|
||
<specimenCount id="EFA2FD242915160BD4D6F9ECFC89F9B7" box="[764,856,1614,1639]" pageId="3" pageNumber="304" type="male">2 males</specimenCount>
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||
,
|
||
<typeStatus id="261F880F2915160BD54CF9ECFC05F9B7" box="[870,980,1615,1639]" pageId="3" pageNumber="304" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<collectionCode id="9FB5AE682915160BD5C8F9ECFBE2F9B7" LSID="urn:lsid:biocol.org:col:34871" box="[994,1075,1615,1639]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="304">USNM</collectionCode>
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<date id="8D1A106D2915160BD217F9EDFB45F9B7" box="[1085,1172,1614,1639]" pageId="3" pageNumber="304" value="56-05-25">250556</date>
|
||
, same data;
|
||
<specimenCount id="EFA2FD242915160BD30CF9ECFA49F9B7" box="[1318,1432,1614,1639]" pageId="3" pageNumber="304" type="female">7 females</specimenCount>
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||
,
|
||
<typeStatus id="261F880F2915160BD6BDF9D0FED4F95B" box="[151,261,1651,1675]" pageId="3" pageNumber="304" type="paratype">paratypes</typeStatus>
|
||
,
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||
<collectionCode id="9FB5AE682915160BD73BF9D0FEB3F95B" LSID="urn:lsid:biocol.org:col:34871" box="[273,354,1651,1675]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="304">USNM</collectionCode>
|
||
<date id="8D1A106D2915160BD743F9D1FE11F95B" box="[361,448,1650,1675]" pageId="3" pageNumber="304" value="57-05-25">250557</date>
|
||
,
|
||
<date id="8D1A106D2915160BD7E6F9D1FDFBF95B" box="[460,554,1650,1675]" pageId="3" pageNumber="304" value="1989-01-01">1.i.1989</date>
|
||
, F. Grady leg.;
|
||
<specimenCount id="EFA2FD242915160BD4F3F9D0FCF6F95B" box="[729,807,1650,1675]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="EFA2FD242915160BD519F9D0FCA2F95B" box="[819,883,1650,1675]" pageId="3" pageNumber="304" type="generic">3 juv.</specimenCount>
|
||
males,
|
||
<specimenCount id="EFA2FD242915160BD5E0F9D0FBFEF95B" box="[970,1071,1650,1675]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
|
||
,
|
||
<specimenCount id="EFA2FD242915160BD210F9D1FBABF95B" box="[1082,1146,1650,1675]" pageId="3" pageNumber="304" type="generic">5 juv.</specimenCount>
|
||
females, UCR-MZ 3215- 0 1, village Río Claro, cascade Las Cavernitas,
|
||
<geoCoordinate id="9C90506A2915160BD4B4F934FCECF97E" box="[670,829,1687,1711]" direction="north" orientation="latitude" pageId="3" pageNumber="304" precision="1" value="8.6968">8°41'48.48" N</geoCoordinate>
|
||
<geoCoordinate id="9C90506A2915160BD56EF934FC20F97F" box="[836,1009,1687,1711]" direction="west" orientation="longitude" pageId="3" pageNumber="304" precision="1" value="-83.04287">83°02'34.34"W</geoCoordinate>
|
||
,
|
||
<quantity id="3E5C9B482915160BD5D1F935FB95F97E" box="[1019,1092,1686,1711]" metricMagnitude="2" metricUnit="m" metricValue="1.06" pageId="3" pageNumber="304" unit="m" value="106.0">106 m</quantity>
|
||
,
|
||
<date id="8D1A106D2915160BD265F935FB6AF97F" box="[1103,1211,1686,1711]" pageId="3" pageNumber="304" value="2014-01-27">27.i.2014</date>
|
||
, I.S. Wehrtmann, C. Magalhães & E. Souza-Carvalho leg.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE65262915160BD6EDF97DFAB4F823" pageId="3" pageNumber="304" type="description">
|
||
<paragraph id="F91B36AD2915160BD6EDF97DFAB4F823" blockId="3.[151,1437,1578,2035]" pageId="3" pageNumber="304">
|
||
<emphasis id="CBD0EABF2915160BD6EDF97DFE30F927" bold="true" box="[199,481,1758,1783]" pageId="3" pageNumber="304">Description of gonopod</emphasis>
|
||
. Nearly straight, only slightly curved towards latero-cephalic side on caudo-mesial view, narrower at end of proximal third. Marginal suture on mesial surface, not twisted to other surfaces. Distal portion of caudo-marginal projection slightly bilobate in mesial view. Marginal process (distal lobe of caudomarginal projection,
|
||
<emphasis id="CBD0EABF2915160BD7A0F8EFFE19F8B3" box="[394,456,1868,1891]" italics="true" pageId="3" pageNumber="304">sensu</emphasis>
|
||
<bibRefCitation id="9D354B5C2915160BD7FEF8E9FCC5F8B3" author="Villalobos" box="[468,788,1866,1891]" pageId="3" pageNumber="304" refString="Villalobos, J. L. & Alvarez, F. (2010) Phylogenetic analysis of the Mexican freshwater crabs of the tribe Pseudothelphusini (Decapoda: Brachyura: Pseudothelphusidae). Zoological Journal of the Linnean Society, 160, 457 - 481. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2009.00606. x" type="journal article" year="2010">Villalobos & Alvarez 2010</bibRefCitation>
|
||
) triangular, produced beyond the mesial crest of apex, directed cephalodistally; proximal lobe broadly rounded, border smooth. Mesial process strongly developed, reniform. Distocaudal ridge elongated, extending distally from the beginning of second third of stem length. Apex elongated, narrow; both mesial, lateral crests nearly straight. Apex cavity situated distally, with spermatic channel opening positioned on cephalic end; field of apical spines weakly developed situated on middle, cephalic end.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="ADDB66252912160CD6BDF947FF08F862" httpUri="https://zenodo.org/record/235128/files/figure.png" pageId="4" pageNumber="305" targetBox="[152,1431,193,1735]" targetPageId="4">
|
||
<paragraph id="F91B36AD2912160CD6BDF947FF08F862" blockId="4.[151,1436,1764,1970]" pageId="4" pageNumber="305">
|
||
<emphasis id="CBD0EABF2912160CD6BDF947FEEAF92A" bold="true" box="[151,315,1764,1787]" pageId="4" pageNumber="305">FIGURES 1–9.</emphasis>
|
||
<taxonomicName id="3EA44D2E2912160CD768F947FD74F92A" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" box="[322,677,1764,1787]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="agrestis">
|
||
<emphasis id="CBD0EABF2912160CD768F947FE22F92B" box="[322,499,1764,1787]" italics="true" pageId="4" pageNumber="305">Achlidon agrestis</emphasis>
|
||
(Rathbun, 1898)
|
||
</taxonomicName>
|
||
: 1, left first gonopod (partially damaged), holotype, USNM 19487, mesial view; 2, same data, lateral view of distal part; 3, left first gonopod, NHMW 21827, cephalic view of distal part; 4, same data, apical view of apex; 5, same data, latero-cephalic view.
|
||
<taxonomicName id="3EA44D2E2912160CD4F5F881FB85F8E8" authority="Hobbs, 1991" authorityName="Hobbs" authorityYear="1991" box="[735,1108,1826,1848]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="puntarenas">
|
||
<emphasis id="CBD0EABF2912160CD4F5F881FC65F8E8" box="[735,948,1826,1848]" italics="true" pageId="4" pageNumber="305">Achlidon puntarenas</emphasis>
|
||
(Hobbs, 1991)
|
||
</taxonomicName>
|
||
: 6, left first gonopod, holotype, USNM 250555, caudo-mesial view; 7, same data, latero-cephalic view of distal part.
|
||
<taxonomicName id="3EA44D2E2912160CD237F8E3FF36F8A5" authority="Pretzmann, 1965: 8" authorityName="Pretzmann" authorityPageNumber="8" authorityYear="1965" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="cocleensis">
|
||
<emphasis id="CBD0EABF2912160CD237F8E3FAF1F886" box="[1053,1312,1856,1878]" italics="true" pageId="4" pageNumber="305">Ptychophallus cocleensis</emphasis>
|
||
Pretzmann, 1965: 8
|
||
</taxonomicName>
|
||
, left first gonopod, holotype, USNM 119869, lateral view of distal part; 9, same data, mesial view (dr, disto-caudal ridge; fs, field of apical spines; lp, lateral process; me, mesial process; mp, marginal process; ms, marginal suture). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="B1BE65262913160DD6EDFF34FE67FF28" pageId="5" pageNumber="306" type="materials_examined">
|
||
<paragraph id="F91B36AD2913160DD6EDFF34FE67FF28" blockId="5.[151,1436,151,573]" pageId="5" pageNumber="306">
|
||
<emphasis id="CBD0EABF2913160DD6EDFF34FDD8FF60" bold="true" box="[199,521,151,176]" pageId="5" pageNumber="306">
|
||
<typeStatus id="261F880F2913160DD6EDFF34FEE5FF60" box="[199,308,151,176]" pageId="5" pageNumber="306" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="261F880F2913160DD75CFF34FE78FF60" box="[374,425,151,176]" pageId="5" pageNumber="306">type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
. Male, cw 18.0, cl 11.5 (chelipeds dearticulated),
|
||
<collectionCode id="9FB5AE682913160DD27CFF3BFB76FF60" LSID="urn:lsid:biocol.org:col:34871" box="[1110,1191,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="306">USNM</collectionCode>
|
||
<date id="8D1A106D2913160DD298FF34FADBFF60" box="[1202,1290,151,176]" pageId="5" pageNumber="306" value="55-05-25">250555</date>
|
||
.
|
||
<collectingCountry id="81B3763D2913160DD33DFF3BFA49FF60" box="[1303,1432,151,176]" name="Costa Rica" pageId="5" pageNumber="306">Costa Rica</collectingCountry>
|
||
,
|
||
<taxonomicName id="3EA44D2E2913160DD6BDFF1EFEC3FF04" box="[151,274,189,212]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="338" phylum="Arthropoda" rank="species" species="puntarenas">Puntarenas</taxonomicName>
|
||
Province, small stream tributary of Claro River, in Emus Cave, near Río Claro village [
|
||
<geoCoordinate id="9C90506A2913160DD322FF1FFABDFF04" box="[1288,1388,188,212]" direction="north" orientation="latitude" pageId="5" pageNumber="306" precision="925" value="8.666667">08°40’N</geoCoordinate>
|
||
<geoCoordinate id="9C90506A2913160DD35FFF1FFF08FF28" direction="west" orientation="longitude" pageId="5" pageNumber="306" precision="925" value="-83.066666">83° 04’W</geoCoordinate>
|
||
] (Pacific drainage).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE65262913160DD6EDFEA7FC50FECC" box="[199,897,260,285]" pageId="5" pageNumber="306" type="distribution">
|
||
<paragraph id="F91B36AD2913160DD6EDFEA7FC50FECC" blockId="5.[151,1436,151,573]" box="[199,897,260,285]" pageId="5" pageNumber="306">
|
||
<emphasis id="CBD0EABF2913160DD6EDFEA7FE88FECD" bold="true" box="[199,345,260,285]" pageId="5" pageNumber="306">Distribution</emphasis>
|
||
. Southern
|
||
<collectingCountry id="81B3763D2913160DD7F9FEA7FD81FECD" box="[467,592,260,285]" name="Costa Rica" pageId="5" pageNumber="306">Costa Rica</collectingCountry>
|
||
, Pacific drainage (
|
||
<figureCitation id="619F2A282913160DD50AFEA7FCA5FECC" box="[800,884,260,285]" captionStart="FIGURE 77" captionStartId="31.[151,250,2011,2033]" captionTargetBox="[157,1433,886,1986]" captionTargetId="figure@31.[150,1436,879,1989]" captionTargetPageId="31" captionText="FIGURE 77. Distribution map of Achlidon agrestis (Rathbun, 1898), and A. puntarenas (Hobbs, 1991)." httpUri="https://zenodo.org/record/235135/files/figure.png" pageId="5" pageNumber="306">Fig. 77</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B1BE65262913160DD6EDFE84FBC6FDEC" pageId="5" pageNumber="306" type="discussion">
|
||
<paragraph id="F91B36AD2913160DD6EDFE84FBC6FDEC" blockId="5.[151,1436,151,573]" pageId="5" pageNumber="306">
|
||
<emphasis id="CBD0EABF2913160DD6EDFE84FEEAFE90" bold="true" box="[199,315,295,320]" pageId="5" pageNumber="306">Remarks.</emphasis>
|
||
The G1 morphology of the present taxon clearly differs from that of the species of the genus
|
||
<taxonomicName id="3EA44D2E2913160DD6BDFEEFFE87FEB5" box="[151,342,332,357]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBD0EABF2913160DD6BDFEEFFE87FEB5" box="[151,342,332,357]" italics="true" pageId="5" pageNumber="306">Pseudothelphusa</emphasis>
|
||
</taxonomicName>
|
||
de Saussure, 1857.
|
||
<bibRefCitation id="9D354B5C2913160DD47EFEEFFBD4FEB4" box="[596,1029,332,357]" pageId="5" pageNumber="306" refString="Villalobos Hiriart, J. L. & Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. & Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter">Villalobos Hiriart & Álvarez (2008)</bibRefCitation>
|
||
excluded
|
||
<taxonomicName id="3EA44D2E2913160DD2ACFEEEFAF7FEB4" box="[1158,1318,333,356]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="puntarenas">
|
||
<emphasis id="CBD0EABF2913160DD2ACFEEEFAF7FEB4" box="[1158,1318,333,356]" italics="true" pageId="5" pageNumber="306">P. puntarenas</emphasis>
|
||
</taxonomicName>
|
||
from the Pseudothelphusini
|
||
<bibRefCitation id="9D354B5C2913160DD747FED3FDC9FE58" author="Ortmann" box="[365,536,368,392]" pageId="5" pageNumber="306" refString="Ortmann, A. E. (1897) Carcinologische Studien. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Thiere, 10 (3), 258 - 372, pl. 17." type="journal article" year="1897">Ortmann, 1897</bibRefCitation>
|
||
and proposed that it should be placed in
|
||
<taxonomicName id="3EA44D2E2913160DD5DFFECCFB8BFE58" box="[1013,1114,367,392]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBD0EABF2913160DD5DFFECCFB8BFE58" box="[1013,1114,367,392]" italics="true" pageId="5" pageNumber="306">Achlidon</emphasis>
|
||
</taxonomicName>
|
||
. Such recommendation was based on the results of their phylogenetic analysis of the tribe Pseudothelphusini, which was subsequently published (
|
||
<bibRefCitation id="9D354B5C2913160DD73EFE14FD9DFE00" author="Villalobos" box="[276,588,439,464]" pageId="5" pageNumber="306" refString="Villalobos, J. L. & Alvarez, F. (2010) Phylogenetic analysis of the Mexican freshwater crabs of the tribe Pseudothelphusini (Decapoda: Brachyura: Pseudothelphusidae). Zoological Journal of the Linnean Society, 160, 457 - 481. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2009.00606. x" type="journal article" year="2010">Villalobos & Alvarez 2010</bibRefCitation>
|
||
) when they confirmed the new combination.
|
||
<bibRefCitation id="9D354B5C2913160DD27FFE14FA4DFE00" author="Villalobos" box="[1109,1436,439,464]" pageId="5" pageNumber="306" refString="Villalobos, J. L. & Alvarez, F. (2010) Phylogenetic analysis of the Mexican freshwater crabs of the tribe Pseudothelphusini (Decapoda: Brachyura: Pseudothelphusidae). Zoological Journal of the Linnean Society, 160, 457 - 481. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2009.00606. x" type="journal article" year="2010">Villalobos & Alvarez (2010)</bibRefCitation>
|
||
used somatic and sex characters to argue that four autapomorphies seen in the G1 of
|
||
<taxonomicName id="3EA44D2E2913160DD278FE7EFB3AFE24" box="[1106,1259,477,500]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="puntarenas">
|
||
<emphasis id="CBD0EABF2913160DD278FE7EFB3AFE24" box="[1106,1259,477,500]" italics="true" pageId="5" pageNumber="306">P. puntarenas</emphasis>
|
||
</taxonomicName>
|
||
supported such exclusion and that the apical position of the spermatic channel opening and the lack of torsion of the G1 would “indicate that the species belongs to the Hypolobocerini in the genus
|
||
<taxonomicName id="3EA44D2E2913160DD5B4FD87FBD2FDED" box="[926,1027,548,573]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CBD0EABF2913160DD5B4FD87FBD2FDED" box="[926,1027,548,573]" italics="true" pageId="5" pageNumber="306">Achlidon</emphasis>
|
||
</taxonomicName>
|
||
”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |