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<mods:title id="E14055DF997740835D53416C00F6A581">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<treatment id="710D87BB2914160BD6BDF95AFEA0FB27" ID-DOI="http://doi.org/10.5281/zenodo.5491502" ID-GBIF-Taxon="119590151" ID-Zenodo-Dep="5491502" LSID="urn:lsid:plazi:treatment:710D87BB2914160BD6BDF95AFEA0FB27" httpUri="http://treatment.plazi.org/id/710D87BB2914160BD6BDF95AFEA0FB27" lastPageId="3" lastPageNumber="304" pageId="2" pageNumber="303">
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<paragraph id="F91B36AD2914160AD6BDF95AFDEDF8C3" blockId="2.[151,572,1785,1845]" box="[151,572,1785,1811]" pageId="2" pageNumber="303">
<heading id="A25381C12914160AD6BDF95AFDEDF8C3" bold="true" box="[151,572,1785,1811]" fontSize="11" level="1" pageId="2" pageNumber="303" reason="1">
<taxonomicName id="3EA44D2E2914160AD6BDF95AFDEDF8C3" ID-CoL="97ZR" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" box="[151,572,1785,1811]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="agrestis">
<emphasis id="CBD0EABF2914160AD6BDF95AFDEDF8C3" bold="true" box="[151,572,1785,1811]" pageId="2" pageNumber="303">
<emphasis id="CBD0EABF2914160AD6BDF95AFEBBF8C3" bold="true" box="[151,362,1785,1811]" italics="true" pageId="2" pageNumber="303">Achlidon agrestis</emphasis>
(
<bibRefCitation id="9D354B5C2914160AD753F95AFDE5F8C3" author="Rathbun" box="[377,564,1785,1811]" pageId="2" pageNumber="303" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
)
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<paragraph id="F91B36AD2914160AD6BDF8BFFEEDF8E4" blockId="2.[151,572,1785,1845]" box="[151,316,1820,1845]" pageId="2" pageNumber="303">
(
<figureCitation id="619F2A282914160AD6B5F8BFFEDBF8E5" box="[159,266,1820,1845]" captionStart="FIGURES 1 9" captionStartId="4.[151,264,1764,1787]" captionTargetBox="[152,1431,193,1735]" captionTargetId="figure@4.[152,1436,193,1743]" captionTargetPageId="4" captionText="FIGURES 1 9. Achlidon agrestis (Rathbun, 1898): 1, left first gonopod (partially damaged), holotype, USNM 19487, mesial view; 2, same data, lateral view of distal part; 3, left first gonopod, NHMW 21827, cephalic view of distal part; 4, same data, apical view of apex; 5, same data, latero-cephalic view. Achlidon puntarenas (Hobbs, 1991): 6, left first gonopod, holotype, USNM 250555, caudo-mesial view; 7, same data, latero-cephalic view of distal part. Ptychophallus cocleensis Pretzmann, 1965: 8, left first gonopod, holotype, USNM 119869, lateral view of distal part; 9, same data, mesial view (dr, disto-caudal ridge; fs, field of apical spines; lp, lateral process; me, mesial process; mp, marginal process; ms, marginal suture). Scale bars = 1 mm." httpUri="https://zenodo.org/record/235128/files/figure.png" pageId="2" pageNumber="303">Figs. 15</figureCitation>
,
<figureCitation id="619F2A282914160AD73CF8BFFEE2F8E4" box="[278,307,1820,1844]" captionStart="FIGURE 77" captionStartId="31.[151,250,2011,2033]" captionTargetBox="[157,1433,886,1986]" captionTargetId="figure@31.[150,1436,879,1989]" captionTargetPageId="31" captionText="FIGURE 77. Distribution map of Achlidon agrestis (Rathbun, 1898), and A. puntarenas (Hobbs, 1991)." httpUri="https://zenodo.org/record/235135/files/figure.png" pageId="2" pageNumber="303">77</figureCitation>
)
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<taxonomicName id="3EA44D2E2914160AD6BDF8C7FDBCF8AA" ID-CoL="84G7D" authority="Rathbun, 1898: 514" authorityName="Rathbun" authorityPageNumber="514" authorityYear="1898" box="[151,621,1892,1914]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="agrestis">
<emphasis id="CBD0EABF2914160AD6BDF8C7FE4BF8AA" box="[151,410,1892,1914]" italics="true" pageId="2" pageNumber="303">Pseudothelphusa agrestis</emphasis>
<treatmentCitation id="780510BC2914160AD78BF8C7FDBCF8AA" author="Rathbun" box="[417,621,1892,1914]" page="514" pageId="2" pageNumber="303" year="1898">
<bibRefCitation id="9D354B5C2914160AD78BF8C7FDEBF8AA" author="Rathbun" box="[417,570,1892,1914]" pageId="2" pageNumber="303" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537. http: // dx. doi. org / 10.5479 / si. 00963801.21 - 1158.507" type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
: 514
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, 532, fig. 2.—
<treatmentCitation id="780510BC2914160AD4D4F8C7FC60F8AA" author="Young" box="[766,945,1892,1914]" page="212" pageId="2" pageNumber="303" year="1900">
<bibRefCitation id="9D354B5C2914160AD4D4F8C7FC51F8AA" author="Young" box="[766,896,1892,1914]" pageId="2" pageNumber="303" refString="Young, C. G. (1900) The stalk-eyed Crustacea of British Guiana, West Indies, and Bermuda. J. M. Watkins, London, 514 pp., 4 pls." type="book" year="1900">Young 1900</bibRefCitation>
: 212
</treatmentCitation>
.—
<treatmentCitation id="780510BC2914160AD5E5F8C7FB45F8AA" author="Rathbun" box="[975,1172,1892,1914]" page="242" pageId="2" pageNumber="303" year="1904">
<bibRefCitation id="9D354B5C2914160AD5E5F8C7FBB5F8AA" author="Rathbun" box="[975,1124,1892,1914]" pageId="2" pageNumber="303" refString="Rathbun, M. J. (1904) Les crabes d'eau douce (Potamonidae). Nouvelles Archives du Museum d'Histoire naturelle Paris, 6 (4), 225 - 312, text-figs. 1 - 37, volume pls. 9 - 18 (pls. I - X for Rathbun's article)." type="journal article" year="1904">Rathbun 1904</bibRefCitation>
: 242
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC2914160AD328F8C7FF3DF848" author="Rathbun" page="285" pageId="2" pageNumber="303" year="1905">
<bibRefCitation id="9D354B5C2914160AD328F8C7FA49F8AA" author="Rathbun" box="[1282,1432,1892,1914]" pageId="2" pageNumber="303" refString="Rathbun, M. J. (1905) Les crabes d'eau douce. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 7 (4), 159 - 321, textfigs. 38 - 105, volume pls. 13 - 22. [pls. XI - X for Rathbun's article]" type="journal article" year="1905">Rathbun 1905</bibRefCitation>
: 285
</treatmentCitation>
.—
<treatmentCitation id="780510BC2914160AD723F821FE0FF848" author="Coifmann" box="[265,478,1922,1944]" page="106" pageId="2" pageNumber="303" year="1939">
<bibRefCitation id="9D354B5C2914160AD723F821FE7FF848" author="Coifmann" box="[265,430,1922,1944]" pageId="2" pageNumber="303" refString="Coifmann, I. (1939) Potamonidi della Guiana inglese raccolti dal Prof. Nello Beccari. Archivio Zoologico Italiano, 27, 93 - 116, pl. 3." type="journal article" year="1939">Coifmann 1939</bibRefCitation>
: 106
</treatmentCitation>
(in list).
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</paragraph>
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<taxonomicName id="3EA44D2E2914160AD6BDF802FDDAF867" ID-CoL="84G7D" box="[151,523,1953,1975]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="agrestis" subGenus="Achlidon">
<emphasis id="CBD0EABF2914160AD6BDF802FE95F867" box="[151,324,1953,1975]" italics="true" pageId="2" pageNumber="303">Pseudothelphusa</emphasis>
(
<emphasis id="CBD0EABF2914160AD779F802FE7FF867" box="[339,430,1953,1975]" italics="true" pageId="2" pageNumber="303">Achlidon</emphasis>
)
<emphasis id="CBD0EABF2914160AD796F801FDDAF867" box="[444,523,1954,1975]" italics="true" pageId="2" pageNumber="303">agrestis</emphasis>
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<treatmentCitation id="780510BC2914160AD40CF802FD04F867" author="Smalley" box="[550,725,1953,1975]" page="8" pageId="2" pageNumber="303" year="1964">
<bibRefCitation id="9D354B5C2914160AD40CF802FD10F867" author="Smalley" box="[550,705,1953,1975]" pageId="2" pageNumber="303" refString="Smalley, A. E. (1964 b) The river crabs of Costa Rica, and the subfamilies of the Pseudothelphusidae. Tulane Studies in Zoology, 12 (1), 5 - 13." type="journal article" year="1964" yearSuffix="b">Smalley 1964b</bibRefCitation>
: 8
</treatmentCitation>
.—
<treatmentCitation id="780510BC2914160AD4DEF802FC65F867" author="Schmitt" box="[756,948,1953,1975]" page="105" pageId="2" pageNumber="303" year="1969">
<bibRefCitation id="9D354B5C2914160AD4DEF802FC53F867" author="Schmitt" box="[756,898,1953,1975]" pageId="2" pageNumber="303" refString="Schmitt, W. L. (1969) Colombian freshwater crab notes. Proceedings of the Biological Society of Washington, 82, 93 - 112." type="journal article" year="1969">Schmitt 1969</bibRefCitation>
: 105
</treatmentCitation>
, fig. 6.
<emphasis id="CBD0EABF2914160AD6BDF863FEB1F806" box="[151,352,1984,2006]" italics="true" pageId="2" pageNumber="303">
<taxonomicName id="3EA44D2E2914160AD6BDF863FF22F806" ID-CoL="7NMS6" box="[151,243,1984,2006]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">Achlidon</taxonomicName>
agrestis—
</emphasis>
<treatmentCitation id="780510BC2914160AD74AF863FDF4F806" author="Smalley" box="[352,549,1984,2006]" page="105" pageId="2" pageNumber="303" year="1970">
<bibRefCitation id="9D354B5C2914160AD74AF863FE20F806" author="Smalley" box="[352,497,1984,2006]" pageId="2" pageNumber="303" refString="Smalley, A. E. (1970) A new genus of freshwater crabs from Guatemala, with a key to the middle American genera (Crustacea, Decapoda, Pseudothelphusidae). The American Midland Naturalist, 83 (1), 96 - 106. http: // dx. doi. org / 10.2307 / 2424009" type="journal article" year="1970">Smalley 1970</bibRefCitation>
: 105
</treatmentCitation>
, fig. 3.—
<treatmentCitation id="780510BC2914160AD4AEF863FCB1F806" author="Rodriguez" box="[644,864,1984,2006]" page="106" pageId="2" pageNumber="303" year="1982">
<bibRefCitation id="9D354B5C2914160AD4AEF863FCE1F806" author="Rodriguez" box="[644,816,1984,2006]" pageId="2" pageNumber="303" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez 1982</bibRefCitation>
: 106
</treatmentCitation>
.—
<treatmentCitation id="780510BC2914160AD554F863FB88F806" author="Villalobos" box="[894,1113,1984,2006]" page="220" pageId="2" pageNumber="303" year="1982">
<bibRefCitation id="9D354B5C2914160AD554F863FBF6F806" author="Villalobos" box="[894,1063,1984,2006]" pageId="2" pageNumber="303" refString="Villalobos, A. (1982) Decapoda. In: Hurlbert, S. H. &amp; Villalobos, A. (Eds.), Aquatic Biota of Mexico, Central America and the West Indies. San Diego State University, San Diego, pp. 215 - 239." type="book chapter" year="1982">Villalobos 1982</bibRefCitation>
: 220
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC2914160AD2E0F863FE52F824" author="Villalobos" page="295" pageId="2" pageNumber="303" year="2008">
<bibRefCitation id="9D354B5C2914160AD2E0F863FE85F824" author="Villalobos" pageId="2" pageNumber="303" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 295
</treatmentCitation>
(in list).
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</paragraph>
<paragraph id="F91B36AD2915160BD6BDFF3AFD9EFF3B" blockId="3.[151,932,152,266]" pageId="3" pageNumber="304">
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<taxonomicName id="3EA44D2E2915160BD6BDFF3AFDD2FF7E" box="[151,515,152,174]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="agrestis" subGenus="Achlidon">
<emphasis id="CBD0EABF2915160BD6BDFF3AFEEDFF7E" box="[151,316,153,174]" italics="true" pageId="3" pageNumber="304">Potamocarcinus</emphasis>
(
<emphasis id="CBD0EABF2915160BD761FF3BFE77FF7E" box="[331,422,152,174]" italics="true" pageId="3" pageNumber="304">Achlidon</emphasis>
)
<emphasis id="CBD0EABF2915160BD79EFF3AFDD2FF7E" box="[436,515,153,174]" italics="true" pageId="3" pageNumber="304">agrestis</emphasis>
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<treatmentCitation id="780510BC2915160BD437FF3AFD3DFF7E" author="Pretzmann" box="[541,748,152,174]" page="20" pageId="3" pageNumber="304" year="1971">
<bibRefCitation id="9D354B5C2915160BD437FF3AFD17FF7E" author="Pretzmann" box="[541,710,152,174]" pageId="3" pageNumber="304" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche Klasse der Osterreichischen Akademie der Wissenschaften, Series 1, 179 (1 - 4), 12 - 24." type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: 20
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.
<taxonomicName id="3EA44D2E2915160BD6BDFF15FE6BFF1D" ID-CoL="97ZR" box="[151,442,182,205]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="agrestis" subGenus="Achlidon">
<emphasis id="CBD0EABF2915160BD6BDFF15FF22FF1C" box="[151,243,182,204]" italics="true" pageId="3" pageNumber="304">Achlidon</emphasis>
(
<emphasis id="CBD0EABF2915160BD72BFF15FE8DFF1C" box="[257,348,182,204]" italics="true" pageId="3" pageNumber="304">Achlidon</emphasis>
)
<emphasis id="CBD0EABF2915160BD741FF1BFE6BFF1D" box="[363,442,184,205]" italics="true" pageId="3" pageNumber="304">agrestis</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC2915160BD7FEFF14FD73FF1C" author="Pretzmann" box="[468,674,182,204]" page="82" pageId="3" pageNumber="304" year="1972">
<bibRefCitation id="9D354B5C2915160BD7FEFF14FDACFF1C" author="Pretzmann" box="[468,637,182,204]" pageId="3" pageNumber="304" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
: 82
</treatmentCitation>
, figs. 429432, 435437.?
<emphasis id="CBD0EABF2915160BD682FF76FEA1FF3B" box="[168,368,213,235]" italics="true" pageId="3" pageNumber="304">
<taxonomicName id="3EA44D2E2915160BD682FF76FED5FF3B" ID-CoL="7NMS6" box="[168,260,213,235]" class="Malacostraca" family="Pseudothelphusidae" genus="Achlidon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">Achlidon</taxonomicName>
agrestis—
</emphasis>
<treatmentCitation id="780510BC2915160BD75AFF75FD9AFF3B" author="Pretzmann" box="[368,587,213,235]" page="653" pageId="3" pageNumber="304" year="1980">
<bibRefCitation id="9D354B5C2915160BD75AFF75FDC8FF3B" author="Pretzmann" box="[368,537,213,235]" pageId="3" pageNumber="304" refString="Pretzmann, G. (1980) Von Dr. Ivo Poglayen-Neuwall 1975 in Mittelamerika gesammelte Krabben. Annalen des Naturhistorischen Museums, 83, 651 - 666, pls. 1 - 18. [Wien]" type="journal article" year="1980">Pretzmann 1980</bibRefCitation>
: 653
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD2915160BD6BDFF57FD6CFEDA" blockId="3.[151,932,152,266]" box="[151,701,244,266]" pageId="3" pageNumber="304">
<treatmentCitationGroup id="D9B411832915160BD6BDFF57FD6CFEDA" box="[151,701,244,266]" pageId="3" pageNumber="304">
<taxonomicName id="3EA44D2E2915160BD6BDFF57FE85FEDA" box="[151,340,244,266]" class="Malacostraca" family="Pseudothelphusidae" genus="Anchlidon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="agrestis">
<emphasis id="CBD0EABF2915160BD6BDFF57FE85FEDA" box="[151,340,244,266]" italics="true" pageId="3" pageNumber="304">Anchlidon agrestis</emphasis>
</taxonomicName>
[sic]—
<treatmentCitation id="780510BC2915160BD7B5FF57FDB4FEDA" author="Ng" box="[415,613,244,266]" page="173" pageId="3" pageNumber="304" year="2008">
<bibRefCitation id="9D354B5C2915160BD7B5FF57FDE4FEDA" author="Ng" box="[415,565,244,266]" pageId="3" pageNumber="304" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF2915160BD7ECFF56FE28FEDA" box="[454,505,244,266]" italics="true" pageId="3" pageNumber="304">et al.</emphasis>
2008
</bibRefCitation>
: 173
</treatmentCitation>
(in list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262915160BD6BDFE95FCC5FD0B" pageId="3" pageNumber="304" type="materials_examined">
<paragraph id="F91B36AD2915160BD6BDFE95FCC5FD0B" blockId="3.[151,1437,310,1271]" pageId="3" pageNumber="304">
<emphasis id="CBD0EABF2915160BD6BDFE95FEAAFE9F" bold="true" box="[151,379,310,335]" pageId="3" pageNumber="304">Material examined</emphasis>
.
<collectingCountry id="81B3763D2915160BD7A6FE94FDE4FE9F" box="[396,565,311,335]" name="Costa Rica" pageId="3" pageNumber="304">COSTA RICA</collectingCountry>
. Limón Province, Atlantic drainage:
<specimenCount id="EFA2FD242915160BD5F4FE94FBFFFE9F" box="[990,1070,310,335]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
, 2 young, UCR-MZ, Road 232, Gonzalez creek,
<geoCoordinate id="9C90506A2915160BD77EFEF9FE23FEA3" box="[340,498,346,371]" direction="north" orientation="latitude" pageId="3" pageNumber="304" precision="1" value="10.162499">10°0945.0”N</geoCoordinate>
<geoCoordinate id="9C90506A2915160BD7D3FEF9FD4FFEA3" box="[505,670,346,371]" direction="west" orientation="longitude" pageId="3" pageNumber="304" precision="1" value="-83.9375">83°5615.0”W</geoCoordinate>
,
<quantity id="3E5C9B482915160BD480FEF8FD22FEA2" box="[682,755,347,371]" metricMagnitude="2" metricUnit="m" metricValue="4.7299999999999995" pageId="3" pageNumber="304" unit="m" value="473.0">473 m</quantity>
,
<date id="8D1A106D2915160BD52AFEF9FCBDFEA3" box="[768,876,346,371]" pageId="3" pageNumber="304" value="2013-01-29">29.i.2013</date>
, I.S. Wehrtmann
<emphasis id="CBD0EABF2915160BD21AFEFFFBB3FEA3" box="[1072,1122,346,371]" italics="true" pageId="3" pageNumber="304">et al</emphasis>
.;
<specimenCount id="EFA2FD242915160BD252FEF9FB03FEA3" box="[1144,1234,346,371]" pageId="3" pageNumber="304" type="male">5 males</specimenCount>
,
<specimenCount id="EFA2FD242915160BD2F4FEF8FA9CFEA3" box="[1246,1357,346,371]" pageId="3" pageNumber="304" type="female">2 females</specimenCount>
, UCR- MZ, Braulio Carrillo National Park, Road 232,
<geoCoordinate id="9C90506A2915160BD487FEDCFC9AFE47" box="[685,843,383,407]" direction="north" orientation="latitude" pageId="3" pageNumber="304" precision="1" value="10.150472">10°0901.7”N</geoCoordinate>
<geoCoordinate id="9C90506A2915160BD579FEDDFC26FE47" box="[851,1015,382,407]" direction="west" orientation="longitude" pageId="3" pageNumber="304" precision="1" value="-83.95372">83°5713.4”W</geoCoordinate>
,
<quantity id="3E5C9B482915160BD229FEDDFB9DFE46" box="[1027,1100,382,407]" metricMagnitude="2" metricUnit="m" metricValue="6.75" pageId="3" pageNumber="304" unit="m" value="675.0">675 m</quantity>
,
<date id="8D1A106D2915160BD272FEDDFB14FE47" box="[1112,1221,382,407]" pageId="3" pageNumber="304" value="2013-01-29">29.i.2013</date>
, I.S. Wehrtmann
<emphasis id="CBD0EABF2915160BD3A2FEDCFF7CFE6B" italics="true" pageId="3" pageNumber="304">et al</emphasis>
. leg.;
<specimenCount id="EFA2FD242915160BD6D9FE00FE9DFE6B" box="[243,332,418,443]" pageId="3" pageNumber="304" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD242915160BD77DFE00FE6AFE6B" box="[343,443,418,443]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE682915160BD7ECFE00FDD4FE6B" box="[454,517,419,443]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="304">INPA</collectionCode>
2083, same data;
<specimenCount id="EFA2FD242915160BD4FAFE00FCF8FE6B" box="[720,809,418,443]" pageId="3" pageNumber="304" type="male">2 males</specimenCount>
,
<specimenCount id="EFA2FD242915160BD51EFE00FC49FE6B" box="[820,920,418,443]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE682915160BD58EFE00FC21FE6B" box="[932,1008,419,443]" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="3" pageNumber="304">CCDB</collectionCode>
4556, same data;
<specimenCount id="EFA2FD242915160BD291FE00FAD8FE6B" box="[1211,1289,418,443]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD242915160BD33EFE00FA52FE6B" box="[1300,1411,418,443]" pageId="3" pageNumber="304" type="female">4 females</specimenCount>
,
<specimenCount id="EFA2FD242915160BD3A4FE00FF32FE0F" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
with young, UCR-MZ, Hondo River, Road 232,
<geoCoordinate id="9C90506A2915160BD56AFE65FC0FFE0F" box="[832,990,454,479]" direction="north" orientation="latitude" pageId="3" pageNumber="304" precision="1" value="10.092278">10°0532.2”N</geoCoordinate>
<geoCoordinate id="9C90506A2915160BD5C6FE65FB40FE0F" box="[1004,1169,454,479]" direction="west" orientation="longitude" pageId="3" pageNumber="304" precision="1" value="-83.41747">83°2502.9”W</geoCoordinate>
,
<quantity id="3E5C9B482915160BD288FE65FB35FE0E" box="[1186,1252,454,479]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="3" pageNumber="304" unit="m" value="64.0">64 m</quantity>
,
<date id="8D1A106D2915160BD2DDFE65FAB2FE0F" box="[1271,1379,454,479]" pageId="3" pageNumber="304" value="2013-01-29">29.i.2013</date>
, I.S. Wehrtmann
<emphasis id="CBD0EABF2915160BD70EFE4FFE88FDD3" box="[292,345,490,515]" italics="true" pageId="3" pageNumber="304">et al</emphasis>
. leg.;
<specimenCount id="EFA2FD242915160BD78DFE48FDD5FDD3" box="[423,516,490,515]" pageId="3" pageNumber="304" type="male">2 males</specimenCount>
,
<collectionCode id="9FB5AE682915160BD43EFE48FDA1FDD2" box="[532,624,491,514]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="304">NHMW</collectionCode>
21827, Siquirres, Guayacán Farm,
<quantity id="3E5C9B482915160BD238FE48FB8FFDD3" box="[1042,1118,490,515]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="304" unit="km" value="40.0">40 km</quantity>
N Turrialba,
<date id="8D1A106D2915160BD32BFE49FAADFDD3" box="[1281,1404,490,515]" pageId="3" pageNumber="304" value="1975-03-11">11.iii.1975</date>
, I. Poglayen-Neuwall leg.;
<specimenCount id="EFA2FD242915160BD782FDACFE26FDF7" box="[424,503,526,551]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD242915160BD42EFDACFDA5FDF7" box="[516,628,526,551]" pageId="3" pageNumber="304" type="female">2 females</specimenCount>
, UCR-MZ 757, Finca Guayacán,
<date id="8D1A106D2915160BD5D9FDADFBBCFDF7" box="[1011,1133,526,551]" pageId="3" pageNumber="304" value="1976-09-11">11.ix.1976</date>
, R.A. Cruz leg.—San José Province:
<specimenCount id="EFA2FD242915160BD720FD90FEB4FD9B" box="[266,357,562,587]" pageId="3" pageNumber="304" type="male">2 males</specimenCount>
, UCR-MZ 716, Vásquez de Coronado Canton, Bajo de La Hondura, Laguna Farm,
<date id="8D1A106D2915160BD335FD91FA49FD9B" box="[1311,1432,562,587]" pageId="3" pageNumber="304" value="1972-04-22">22.iv.1972</date>
, H. Camacho leg.—Cartago Province:
<specimenCount id="EFA2FD242915160BD46EFDF4FD42FDBF" box="[580,659,598,623]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
,
<typeStatus id="261F880F2915160BD48AFDF5FCD4FDBF" box="[672,773,598,623]" pageId="3" pageNumber="304" type="holotype">holotype</typeStatus>
,
<collectionCode id="9FB5AE682915160BD53BFDF4FCB3FDBF" LSID="urn:lsid:biocol.org:col:34871" box="[785,866,599,623]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="304">USNM</collectionCode>
19487, La Flor, farm near Torito,
<quantity id="3E5C9B482915160BD2C2FDF4FAA6FDBE" box="[1256,1399,599,623]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="3" pageNumber="304" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">9001000 m</quantity>
, E. Fernández leg.;
<specimenCount id="EFA2FD242915160BD760FDD8FE46FD43" box="[330,407,634,659]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD242915160BD788FDD8FDC0FD43" box="[418,529,634,659]" pageId="3" pageNumber="304" type="female">4 females</specimenCount>
, UCR-MZ 714, La
<collectingCountry id="81B3763D2915160BD4C1FDD9FCFDFD43" box="[747,812,634,659]" name="Switzerland" pageId="3" pageNumber="304">Suiza</collectingCountry>
, El Alto de
<collectingCountry id="81B3763D2915160BD59AFDD9FBCCFD43" box="[944,1053,634,659]" name="Germany" pageId="3" pageNumber="304">Alemania</collectingCountry>
Farm, km 4 NE road to Pacayitas, C. Villalobos, R.A. Cruz &amp; Marín leg.;
<specimenCount id="EFA2FD242915160BD47BFD3CFD4EFD67" box="[593,671,670,695]" pageId="3" pageNumber="304" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD242915160BD480FD3CFCDFFD67" box="[682,782,670,695]" pageId="3" pageNumber="304" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE682915160BD533FD3CFC89FD67" box="[793,856,671,695]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="3" pageNumber="304">INPA</collectionCode>
1870, same data;
<specimenCount id="EFA2FD242915160BD209FD3CFBADFD67" box="[1059,1148,670,695]" pageId="3" pageNumber="304" type="male">2 males</specimenCount>
, UCR-MZ 715, La
<collectingCountry id="81B3763D2915160BD37DFD3DFA49FD67" box="[1367,1432,670,695]" name="Switzerland" pageId="3" pageNumber="304">Suiza</collectingCountry>
,
<quantity id="3E5C9B482915160BD6BDFD60FF01FD0B" box="[151,208,706,731]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="304" unit="km" value="1.0">1 km</quantity>
NE on road to Pacayitas,
<date id="8D1A106D2915160BD7D9FD61FD95FD0B" box="[499,580,706,731]" pageId="3" pageNumber="304" value="1971-02">ii.1971</date>
, C. Villalobos leg.
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262915160BD6EDFD45FEB8FBCF" pageId="3" pageNumber="304" type="description">
<paragraph id="F91B36AD2915160BD6EDFD45FEB8FBCF" blockId="3.[151,1437,310,1271]" pageId="3" pageNumber="304">
<emphasis id="CBD0EABF2915160BD6EDFD45FE34FD2F" bold="true" box="[199,485,742,767]" pageId="3" pageNumber="304">Description of gonopod</emphasis>
. Slightly curved towards cephalic side on caudo-mesial view, narrower at end of proximal third. Marginal suture on mesial surface, not distorted to other surfaces. Distal portion of caudo-marginal projection unilobate, with slightly convex outline in mesial view. Marginal process (cephalic end of caudomarginal projection,
<emphasis id="CBD0EABF2915160BD7AFFCF7FE12FCBB" box="[389,451,852,875]" italics="true" pageId="3" pageNumber="304">sensu</emphasis>
<bibRefCitation id="9D354B5C2915160BD7E6FCF1FCDAFCBB" author="Villalobos" box="[460,779,850,875]" pageId="3" pageNumber="304" refString="Villalobos, J. L. &amp; Alvarez, F. (2010) Phylogenetic analysis of the Mexican freshwater crabs of the tribe Pseudothelphusini (Decapoda: Brachyura: Pseudothelphusidae). Zoological Journal of the Linnean Society, 160, 457 - 481. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2009.00606. x" type="journal article" year="2010">Villalobos &amp; Alvarez, 2010</bibRefCitation>
) rounded, not produced beyond the mesial crest of apex. Mesial process weakly developed, as short, rounded protuberance. Distocaudal ridge elongated, extending distally from beginning of the second third of stem length. Apex elongated, narrow; mesial crest nearly straight, lateral crest slightly concave in apical view, both crests slightly convex in mesial view. Apex cavity situated distally, with both spermatic channel opening, field of apical spines positioned towards cephalic end; field of apical spines weakly developed.
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262915160BD6EDFB89FD73FBB7" pageId="3" pageNumber="304" type="materials_examined">
<paragraph id="F91B36AD2915160BD6EDFB89FD73FBB7" blockId="3.[151,1437,310,1271]" pageId="3" pageNumber="304">
<emphasis id="CBD0EABF2915160BD6EDFB89FDDFFB93" bold="true" box="[199,526,1066,1091]" pageId="3" pageNumber="304">
<typeStatus id="261F880F2915160BD6EDFB89FEE5FB93" box="[199,308,1066,1091]" pageId="3" pageNumber="304" type="holotype">Holotype</typeStatus>
and
<typeStatus id="261F880F2915160BD750FB89FE7CFB93" box="[378,429,1066,1091]" pageId="3" pageNumber="304">type</typeStatus>
locality
</emphasis>
. Male, cw 26.5, cl 15.8 (partially dearticulated),
<collectionCode id="9FB5AE682915160BD275FB88FB61FB93" LSID="urn:lsid:biocol.org:col:34871" box="[1119,1200,1067,1091]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="304">USNM</collectionCode>
19487.
<collectingCountry id="81B3763D2915160BD33CFB88FA49FB93" box="[1302,1432,1066,1091]" name="Costa Rica" pageId="3" pageNumber="304">Costa Rica</collectingCountry>
, Cartago Province, between La Flor and Torito.
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262915160BD6EDFBD1FCAAFB5B" box="[199,891,1138,1163]" pageId="3" pageNumber="304" type="distribution">
<paragraph id="F91B36AD2915160BD6EDFBD1FCAAFB5B" blockId="3.[151,1437,310,1271]" box="[199,891,1138,1163]" pageId="3" pageNumber="304">
<emphasis id="CBD0EABF2915160BD6EDFBD1FE88FB5B" bold="true" box="[199,345,1138,1163]" pageId="3" pageNumber="304">Distribution</emphasis>
. Central
<collectingCountry id="81B3763D2915160BD7EAFBD0FDECFB5B" box="[448,573,1138,1163]" name="Costa Rica" pageId="3" pageNumber="304">Costa Rica</collectingCountry>
, Atlantic drainage (
<figureCitation id="619F2A282915160BD533FBD1FCBFFB5B" box="[793,878,1138,1163]" captionStart="FIGURE 77" captionStartId="31.[151,250,2011,2033]" captionTargetBox="[157,1433,886,1986]" captionTargetId="figure@31.[150,1436,879,1989]" captionTargetPageId="31" captionText="FIGURE 77. Distribution map of Achlidon agrestis (Rathbun, 1898), and A. puntarenas (Hobbs, 1991)." httpUri="https://zenodo.org/record/235135/files/figure.png" pageId="3" pageNumber="304">Fig. 77</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262915160BD6EDFB35FEA0FB27" pageId="3" pageNumber="304" type="discussion">
<paragraph id="F91B36AD2915160BD6EDFB35FEA0FB27" blockId="3.[151,1437,310,1271]" pageId="3" pageNumber="304">
<emphasis id="CBD0EABF2915160BD6EDFB35FEE5FB7F" bold="true" box="[199,308,1174,1199]" pageId="3" pageNumber="304">Remarks</emphasis>
.
<bibRefCitation id="9D354B5C2915160BD76FFB34FD9FFB7F" author="Pretzmann" box="[325,590,1174,1199]" pageId="3" pageNumber="304" refString="Pretzmann, G. (1980) Von Dr. Ivo Poglayen-Neuwall 1975 in Mittelamerika gesammelte Krabben. Annalen des Naturhistorischen Museums, 83, 651 - 666, pls. 1 - 18. [Wien]" type="journal article" year="1980">Pretzmann (1980: 653)</bibRefCitation>
recorded four females from Agua Buena River (NHMW 4017) but their identification is doubtful because other pseudothelphusid species are recorded from the same area (e.g.
<taxonomicName id="3EA44D2E2915160BD33CFB19FF32FB26" class="Malacostraca" family="Pseudothelphusidae" genus="Allacanthos" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="pittieri">
<emphasis id="CBD0EABF2915160BD33CFB19FF32FB26" italics="true" pageId="3" pageNumber="304">Allacanthos pittieri</emphasis>
</taxonomicName>
, see below).
</paragraph>
</subSubSection>
</treatment>
</document>