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<document id="DEE63E4C3F51E45B6A5765B0806876E6" ID-CLB-Dataset="307464" ID-DOI="10.3897/zookeys.1223.133267" ID-GBIF-Dataset="da6e1a36-3088-414e-9c26-ecd371657ba6" ID-ZooBank="25B8B14C-2348-4D57-9345-4DA8A2405BC0" ID-publisher-id="133267" URI-arpha="3D6287B4-44A2-5938-B1FE-2B810FFE6288" URI-zoobank="http://zoobank.org/25B8B14C-2348-4D57-9345-4DA8A2405BC0" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1736792558286" checkinUser="pensoft" docAuthor="Borja-Arrieta, Ricardo, López-Orozco, Carlos Mario, Carpio-Díaz, Yesenia M., Gutierrez-Estrada, Miguel, Campos-Filho, Ivanklin Soares, Navas-S., Gabriel R. &amp; Bichuette, Maria Elina" docDate="2025" docId="706611444DB854F3898BA892503DEBD3" docLanguage="en" docName="ZooKeys 1223: 263-288" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/133267/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Diploexochus troglobius Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho 2025, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="288" masterDocId="3D6287B444A25938B1FE2B810FFE6288" masterDocTitle="From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species" masterLastPageNumber="288" masterPageNumber="263" pageNumber="263" updateTime="1736807028442" updateUser="ExternalLinkService">
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<mods:title id="ABF7BA9135A487CF7C705ED9F3B7BEEC">From surface to caves: new species of Diploexochus Brandt, 1833 (Oniscidea, Armadillidae) from Colombia, with the description of the first troglobitic species</mods:title>
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<mods:namePart id="DF0274C8F7B739219D7C4AE0E598B804">Borja-Arrieta, Ricardo</mods:namePart>
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<mods:affiliation id="43E4CCBEE3808DF9E3B0529A61F01C73">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil &amp; Grupo de Investigación en Artrópodos Neotropicales, Universidad de Cartagena, Programa de Biología, Campus San Pablo, Cartagena de Indias, Colombia</mods:affiliation>
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<mods:namePart id="4274B3518E12C27B54C4F2C357C643B5">López-Orozco, Carlos Mario</mods:namePart>
<mods:nameIdentifier id="58FF675E5544F6F2740A93F29F41B8C6" type="ORCID">0000-0002-3251-7739</mods:nameIdentifier>
<mods:affiliation id="954C947DE91D80756EF6A2B0D8E8298B">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil &amp; Grupo de Investigación en Artrópodos Neotropicales, Universidad de Cartagena, Programa de Biología, Campus San Pablo, Cartagena de Indias, Colombia</mods:affiliation>
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<mods:namePart id="0AA41550A1289B558CA652B4C7DEC044">Carpio-Díaz, Yesenia M.</mods:namePart>
<mods:nameIdentifier id="EF9BBF1DA1C54B0B8709AED8044D3648" type="ORCID">0000-0001-5116-9736</mods:nameIdentifier>
<mods:affiliation id="865558A2094279157F63505808E7E378">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil &amp; Grupo de Investigación en Artrópodos Neotropicales, Universidad de Cartagena, Programa de Biología, Campus San Pablo, Cartagena de Indias, Colombia</mods:affiliation>
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<mods:namePart id="5A80C4DEB070862E7B26D0CF46553389">Gutierrez-Estrada, Miguel</mods:namePart>
<mods:nameIdentifier id="949435185CB4BD6FD5F912EF2A2A4041" type="ORCID">0000-0003-2017-4262</mods:nameIdentifier>
<mods:affiliation id="A53AB7C4DF0C642B80C3C62F87481E8E">Grupo de Investigación Biología Descriptiva y Aplicada, Universidad de Cartagena, Programa de Biología, Campus San Pablo, Cartagena de Indias, Colombia</mods:affiliation>
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<mods:namePart id="8A539EB7E1F6393F65B4DF0308B85739">Campos-Filho, Ivanklin Soares</mods:namePart>
<mods:nameIdentifier id="92F8870BAF87C97683A0E7DF625C0B6C" type="ORCID">0000-0001-6139-8241</mods:nameIdentifier>
<mods:affiliation id="05DFF03F84679CBAAEB7986BDAF9439F">Grupo de Investigación Hidrobiología, Universidad de Cartagena, Programa de Biología, Campus San Pablo, Cartagena de Indias, Colombia</mods:affiliation>
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<mods:namePart id="48F0D1F9522925194B62D6765CFBB04E">Navas-S., Gabriel R.</mods:namePart>
<mods:nameIdentifier id="94640DF51F083FE95CCDC332B9FD73F0" type="ORCID">0000-0001-9554-6345</mods:nameIdentifier>
<mods:affiliation id="C547A2492AFF62D079EECC579FAFD6E8">Grupo de Investigación en Artrópodos Neotropicales, Universidad de Cartagena, Programa de Biología, Campus San Pablo, Cartagena de Indias, Colombia</mods:affiliation>
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<mods:namePart id="24C493ECE8293F7D11429448658B39EB">Bichuette, Maria Elina</mods:namePart>
<mods:nameIdentifier id="7A5483AA085D6B585359E5C173E15B9C" type="ORCID">0000-0002-9515-4832</mods:nameIdentifier>
<mods:affiliation id="B5F3D31E62C0CB1C4093DE332AE65767">Laboratório de Estudos Subterrâneos, Universidade Federal de São Carlos, São Carlos, São Paulo, Brazil</mods:affiliation>
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<mods:title id="7FCB2EEBE99AF16C63E08E9B8926B2A7">ZooKeys</mods:title>
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<mods:part id="9AE1F20ECA100A24AECE27EF6B02051A">
<mods:date id="F81C383C08D206B94CD4FC605DA345A8">2025</mods:date>
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<mods:number id="46311C8F069F9B95290711EC2B65ABB9">2025-01-13</mods:number>
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<mods:number id="3162F352C8FB875FC0E85901C170DB58">1223</mods:number>
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<mods:start id="80DD11B41A1F22204C912EDD8DFC610D">263</mods:start>
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<mods:identifier id="5909D190E982C2C15BEBCF313358364D" type="DOI">10.3897/zookeys.1223.133267</mods:identifier>
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<treatment id="706611444DB854F3898BA892503DEBD3" ID-DOI="http://doi.org/10.5281/zenodo.14641053" ID-GBIF-Taxon="252012438" ID-Zenodo-Dep="14641053" ID-arpha="70661144-4DB8-54F3-898B-A892503DEBD3" ID-zoobank="https://zoobank.org/0EB8C9FC-F2AD-4D8F-963F-412AE50B7586" LSID="urn:lsid:plazi:treatment:706611444DB854F3898BA892503DEBD3" LSID-ZBK="urn:lsid:zoobank.org:act:0EB8C9FC-F2AD-4D8F-963F-412AE50B7586" httpUri="http://treatment.plazi.org/id/706611444DB854F3898BA892503DEBD3" scope_family="Armadillidae" scope_subOrder="Oniscidea">
<subSubSection id="084BF4F6D497F68C9F78BC0776DF6E22" type="nomenclature">
<paragraph id="B60D4A58D45C5C30C211572647292B92">
<taxonomicName id="CE457F95E081FD60DC66F9F22C712466" authority="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityName="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityYear="2025" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="91346373FB0E10DE505AFF8DAFB12AB5" italics="true">Diploexochus troglobius</emphasis>
López-Orozco, Borja-Arrieta &amp; Campos-Filho
</taxonomicName>
<taxonomicNameLabel id="0885FA2A776601F04885A66EF2E51757" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2FB1C11AD116BEF9C03DF5C6BC427739" type="description">
<paragraph id="8B3FF53F5A4A1A85D6AA4D90EB452EBF">
<figureCitation id="D49BFCCF37D4CFE399C734D70D4E0846" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Diploexochus species from Colombia. Green areas: Tropical Dry Forest (TDF) from Colombia (García et al. 2014)." figureDoi="10.3897/zookeys.1223.133267.figure1" httpUri="https://binary.pensoft.net/fig/1218786">Figs 1</figureCitation>
,
<figureCitation id="2CE7F3CFD8DC986D6E9B22F000146517" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9</figureCitation>
,
<figureCitation id="90A04BB63333274D844E38A49F86B11E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A left mandible B right mandible C maxillula D maxilla E maxilliped F uropod." figureDoi="10.3897/zookeys.1223.133267.figure10" httpUri="https://binary.pensoft.net/fig/1218795">10</figureCitation>
,
<figureCitation id="800954568045399C33407C3F6890BDA0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♂ paratype, CBUDC-CRU 396) A pereopod 1 B pereopod 7 C genital papilla D pleopod 1 E pleopod 2 F pleopod 3 exopod G pleopod 4 exopod H pleopod 5 exopod." figureDoi="10.3897/zookeys.1223.133267.figure11" httpUri="https://binary.pensoft.net/fig/1218796">11</figureCitation>
,
<figureCitation id="EE6467D1151A7BF4A1F009ABF7008E45" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Tropical Dry Forest around Roca Madre Adventure Park, Sucre B livestock around Roca Madre Adventure Park C Roca Madre Cave Gallery D Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. under limestone rocks." figureDoi="10.3897/zookeys.1223.133267.figure12" httpUri="https://binary.pensoft.net/fig/1218797">12</figureCitation>
,
<figureCitation id="F2119ACCDB68BA52D8F969268A7C4EA5" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of the species of the genus Diploexochus Brandt, 1833 of Colombia: A Diploexochus brevispinis (Pearse, 1915), comb. nov. B Diploexochus cacique López-Orozco, Carpio-Díaz &amp; Campos-Filho, sp. nov. C Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1223.133267.figure13" httpUri="https://binary.pensoft.net/fig/1218798">13 C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENKAG" type="type material">
<paragraph id="9168D6F76520AC452276ACE5C3C78CBE">
<heading id="2BBF716BB9CFADB1BC233B423E9BA5D9" reason="title">Type material.</heading>
</paragraph>
<paragraph id="31DFA2D07FD76AB8A95282FBEC8F3CA9">
<materialsCitation id="38DF1447BD6BFF02E5F3054A2402FDAA" ID-GBIF-Occurrence="5006664306" collectingDate="2018-07-12" collectionCode="CM, R" collectorName="CM Lopez-Orozco &amp; R Borja-Arrieta" country="Colombia" county="Toluviejo" latitude="9.513945" location="Roca madre Cave" longLatPrecision="1" longitude="-75.3935" municipality="Campo Aventura Roca Madre" specimenCode="CBUDC-CRU 393" specimenCount="1" specimenCount-male="1" stateProvince="Sucre" typeStatus="holotype">
<collectingCountry id="5F0BE49608781E9E5F2EDB709A68532A" name="Colombia">Colombia</collectingCountry>
•
<specimenCount id="6955445C2A6F69A1E814B1089FD90D29" count="1" type="male">1 ♂</specimenCount>
,
<emphasis id="6714CBFF245533E88B9442E98C64608F" bold="true" italics="true">
<typeStatus id="C7ED0FECC7EE891CCB0B1A9B6CADD7AE" type="holotype">holotype</typeStatus>
</emphasis>
,
<location id="E5E022EC28D04D659DA218C35861E6C8" LSID="urn:lsid:plazi:treatment:706611444DB854F3898BA892503DEBD3:E5E022EC28D04D659DA218C35861E6C8" country="Colombia" county="Toluviejo" latitude="9.513945" longLatPrecision="1" longitude="-75.3935" municipality="Campo Aventura Roca Madre" name="Roca madre Cave" stateProvince="Sucre">Roca madre Cave</location>
,
<collectingMunicipality id="E6830FD700694AA71218010E716D3987">Campo Aventura Roca Madre</collectingMunicipality>
,
<collectingMunicipality id="FA28ECE0251F3D9E61C7F090485D8225">La Piche</collectingMunicipality>
,
<collectingCounty id="D524D43CFD46ECDA2D34DF8A09DAB36A">Toluviejo</collectingCounty>
,
<collectingRegion id="ED8BA7ADB36B31B83A162CF6322AEEFB">Sucre</collectingRegion>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZKAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-75.393500,9.513944]}">
<geoCoordinate id="DD0731ADD7130D5FFBA76B4E742349F5" degrees="9" direction="north" minutes="30" orientation="latitude" precision="1" seconds="50.2" value="9.513945">9°30'50.2&quot;N</geoCoordinate>
,
<geoCoordinate id="633E5B005776804E808623D330470086" degrees="75" direction="west" minutes="23" orientation="longitude" precision="1" seconds="36.6" value="-75.3935">75°23'36.6&quot;W</geoCoordinate>
</named-content>
,
<date id="1FF55424A45C25C7CBCA14568D3283AA" value="2018-07-12">
<collectingDate id="DE9B546A7E73638C62D00FD3689AC96B" value="2018-07-12">12. VII. 2018</collectingDate>
</date>
, leg.
<collectorName id="FD06CED15B53471D42E4DCC24A9AB8B5">CM López-Orozco</collectorName>
,
<collectorName id="547E3C55A9D4716ED643A96D7147974C">R Borja-Arrieta</collectorName>
,
<specimenCode id="B45C319B33FC50CCE00D54584B91202D" collectionCode="CBUDC-CRU">
<abbrev content-type="institution" id="ABBRID0E5KAG" xlink_title="University of Cartagena, Cartagena, Colombia">CBUDC-CRU</abbrev>
393
</specimenCode>
</materialsCitation>
•
<materialsCitation id="58F70ED0889257337A4A7FFA2B1E138E" ID-GBIF-Occurrence="5006664301" collectingDate="2018-07-12" collectionCode="CM, R" collectorName="CM Lopez-Orozco &amp; R Borja-Arrieta" country="Colombia" county="Toluviejo" latitude="9.513945" location="Roca madre Cave" longLatPrecision="1" longitude="-75.3935" municipality="Campo Aventura Roca Madre" specimenCode="CBUDC-CRU 394" specimenCount="1" specimenCount-male="1" stateProvince="Sucre" typeStatus="paratype">
<specimenCount id="7DB53D57009AB726BD2476BBB9B9AE3F" count="1" type="male">1 ♂</specimenCount>
,
<emphasis id="967DADE7B3498BFE559F1F9B4229D932" bold="true" italics="true">
<typeStatus id="38C2D0C8486A4A8FB3AEDD40EDD98161" type="paratype">paratypes</typeStatus>
</emphasis>
, same data as holotype,
<specimenCode id="C670C52C0629AB367ADBBB82D6A218C2" collectionCode="CBUDC-CRU">
<abbrev content-type="institution" id="ABBRID0EGLAG" xlink_title="University of Cartagena, Cartagena, Colombia">CBUDC-CRU</abbrev>
394
</specimenCode>
</materialsCitation>
•
<materialsCitation id="46D15030E135528DDAEEDBCFAF0E284C" ID-GBIF-Occurrence="5006664302" collectingDate="2018-07-12" collectionCode="CM, R" collectorName="CM Lopez-Orozco &amp; R Borja-Arrieta" country="Colombia" county="Toluviejo" latitude="9.513945" location="Roca madre Cave" longLatPrecision="1" longitude="-75.3935" municipality="Campo Aventura Roca Madre" specimenCode="CBUDC-CRU 396" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sucre" typeStatus="paratype">
<specimenCount id="0D1EA95C02D3E23A3A1E04169C621C73" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="17F81629A03105D5100F0465085B6514" count="1" type="female">1 ♀</specimenCount>
(parts in micro-preparations),
<emphasis id="D7BB6C880C18F04D1DCAC60AE24847AF" bold="true" italics="true">
<typeStatus id="FCCC73ECC060F69A3734B041A314BE15" type="paratype">paratypes</typeStatus>
</emphasis>
, same data as holotype,
<specimenCode id="43EBD5EB8BD4ED2E2E84A50D54D6CC19" collectionCode="CBUDC-CRU">
<abbrev content-type="institution" id="ABBRID0EOLAG" xlink_title="University of Cartagena, Cartagena, Colombia">CBUDC-CRU</abbrev>
396
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETLAG" type="description">
<paragraph id="82520A6BB7C92BF8DD87166EDA3FD793">
<heading id="9136639BE6B06F6C169688DB8CE2D284" reason="title">Description.</heading>
</paragraph>
<paragraph id="F1F361124B99DDF7FE71F44A54AFFAC0">
Maximum body length: male
<quantity id="F0AE842F17B35C387F88E024BCC11D5C" metricMagnitude="-3" metricUnit="m" metricValue="2.7" unit="mm" value="2.7">2.7 mm</quantity>
, female
<quantity id="0E41F652D0D86EB663475BC7877D22BB" metricMagnitude="-3" metricUnit="m" metricValue="4.2" unit="mm" value="4.2">4.2 mm</quantity>
. Body outline as in Fig.
<figureCitation id="387966DD834D9D9EF444DA5473DB139A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 A</figureCitation>
. Color faintly brown; body pigments not discernible in vivo (Fig.
<figureCitation id="3F1A0B9D380370C8250CC57DB36FCB61" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Tropical Dry Forest around Roca Madre Adventure Park, Sucre B livestock around Roca Madre Adventure Park C Roca Madre Cave Gallery D Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. under limestone rocks." figureDoi="10.3897/zookeys.1223.133267.figure12" httpUri="https://binary.pensoft.net/fig/1218797">12 D</figureCitation>
). Endoantennal conglobation (Figs
<figureCitation id="9FAD6394CC931366C3A356D18696FD36" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 A</figureCitation>
,
<figureCitation id="2A4C907732F790323210821A4D67AE16" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Tropical Dry Forest around Roca Madre Adventure Park, Sucre B livestock around Roca Madre Adventure Park C Roca Madre Cave Gallery D Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. under limestone rocks." figureDoi="10.3897/zookeys.1223.133267.figure12" httpUri="https://binary.pensoft.net/fig/1218797">12 D</figureCitation>
,
<figureCitation id="B4A8A9ADE575C5263E485EA7F980029D" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Habitus of the species of the genus Diploexochus Brandt, 1833 of Colombia: A Diploexochus brevispinis (Pearse, 1915), comb. nov. B Diploexochus cacique López-Orozco, Carpio-Díaz &amp; Campos-Filho, sp. nov. C Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1223.133267.figure13" httpUri="https://binary.pensoft.net/fig/1218798">13 C</figureCitation>
). Dorsum covered with large triangular tubercles, arranged as follows (Fig.
<figureCitation id="2A93C5752329E21172E9047F815EBEF6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 A, B</figureCitation>
): vertex of cephalon with 10 tubercles in two rows; pereonite 1 with 19 tubercles; pereonites 2–6 with 13 tubercles; pereonite 7 with 11 tubercles; pleonites 3, 4, and telson with two median tubercles. Pereonites 1–7 epimera with one line of
<emphasis id="47BD0E26D5B7276EB5C35D6E95A06CBD" bold="true" italics="true">noduli laterales</emphasis>
per side inserted on outer surface of second tubercle of posterior row (Fig.
<figureCitation id="26C940AC99A2E79CDE69770CC8AEEB96" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 A</figureCitation>
). Dorsal surface densely covered with elongated semi-circular scale-setae, conferring pilous aspect (Fig.
<figureCitation id="3B13EC7F3E81548428CEA33C1587A596" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 C</figureCitation>
). Cephalon (Fig.
<figureCitation id="5AEA78F6BC68B98989EA6DEE66CF4848" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 D – F</figureCitation>
) with prominent frontal shield, distinctly protruding above vertex; eyes with four ommatidia. Pereonites 1–7 epimera flattened and slightly directed outwards; pereonite 1 grooved on posterior lateral margin, inner lobe of schisma rounded and extending beyond posterior margin of outer lobe. Pereonite 2 with triangular ventral lobe rounded at apex, extending beyond posterior margin of epimera; pereonites 3 and 5–7 with well-marked ventral ridge (Fig.
<figureCitation id="1C1CCE1F17F35C7C7EADECFE0C23DF1B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 A, F, G</figureCitation>
). Pleonites 3–5 epimera (Fig.
<figureCitation id="0E992CA2DA88F50BCA48A293F30B68EB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 H, I</figureCitation>
) well developed, sub-rectangular, and directed outwards, distal margins rounded. Telson (Fig.
<figureCitation id="E59CFA2B21E8CC2E50CE3146F5790A6F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 H</figureCitation>
) with proximal part broader than distal part, dorsum slightly depressed, distal margin sinuous. Antennula (Fig.
<figureCitation id="EB6B56E68DECD7F0A2A66ABE04B211DB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 J</figureCitation>
) of three articles, proximal and distal subequal in length, distal article with five stout aesthetascs inserted sub-apically. Antenna (Fig.
<figureCitation id="7454B934C7181F3051497ECABF4931E3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A habitus, lateral view B dorsal tubercles scheme C dorsal scale-seta D cephalon and pereonite 1, dorsal view E cephalon and pereonite 1, posterior view F cephalon and pereonites 1–4, frontal view G pereonites 1–7 epimera, ventral view H pleotelson and uropods, dorsal view I pereonite 7, pleonites 3–5, telson, and uropods, ventral view J antennula K antenna." figureDoi="10.3897/zookeys.1223.133267.figure9" httpUri="https://binary.pensoft.net/fig/1218794">9 K</figureCitation>
) short, not surpassing posterior margin of pereonite 1 when extended backward; flagellum of two articles, the distal about three times as long as first, bearing one row of two lateral aesthetascs. Mandibles with molar penicil semi-dichotomized; left mandible (Fig.
<figureCitation id="1E5B472FEB3338CF49BEB31F5227E852" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A left mandible B right mandible C maxillula D maxilla E maxilliped F uropod." figureDoi="10.3897/zookeys.1223.133267.figure10" httpUri="https://binary.pensoft.net/fig/1218795">10 A</figureCitation>
) with 2 + 1 penicils, right mandible (Fig.
<figureCitation id="97341D61C3DB6C471DD066C92328414D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A left mandible B right mandible C maxillula D maxilla E maxilliped F uropod." figureDoi="10.3897/zookeys.1223.133267.figure10" httpUri="https://binary.pensoft.net/fig/1218795">10 B</figureCitation>
) with 1 + 1 penicils. Maxillula (Fig.
<figureCitation id="6D62BD63F548A7454D545D85C3D16CB6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A left mandible B right mandible C maxillula D maxilla E maxilliped F uropod." figureDoi="10.3897/zookeys.1223.133267.figure10" httpUri="https://binary.pensoft.net/fig/1218795">10 C</figureCitation>
) inner endite with two stout penicils, distal margin bearing thin setae; outer endite with 4 + 6 teeth simple. Maxilla (Fig.
<figureCitation id="85229B1BE5F81D4B302BA65838D225A7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A left mandible B right mandible C maxillula D maxilla E maxilliped F uropod." figureDoi="10.3897/zookeys.1223.133267.figure10" httpUri="https://binary.pensoft.net/fig/1218795">10 D</figureCitation>
) inner lobe rounded covered with thick setae; outer lobe rounded three times as wide as inner lobe covered with thin setae. Maxilliped basis (Fig.
<figureCitation id="11E6B8207D9B92BDBF0180820103B6C5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A left mandible B right mandible C maxillula D maxilla E maxilliped F uropod." figureDoi="10.3897/zookeys.1223.133267.figure10" httpUri="https://binary.pensoft.net/fig/1218795">10 E</figureCitation>
) rectangular, bearing sparse setae; palp with two distinct setae on basal article; endite subrectangular, medial seta surpassing distal margin, distal margin bearing one seta. Pereopods 1–7 merus and carpus with sparse setae on sternal margin; carpus 1 with distal seta cleft at apex; ungual seta and dactylar organ simple not surpassing outer claw. Uropod (Fig.
<figureCitation id="CCA39B512E839B420A8B7BE284308098" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♀ paratype, CBUDC-CRU 396) A left mandible B right mandible C maxillula D maxilla E maxilliped F uropod." figureDoi="10.3897/zookeys.1223.133267.figure10" httpUri="https://binary.pensoft.net/fig/1218795">10 F</figureCitation>
) protopod flattened, enlarged on basal part, distal part elongated and sub-rectangular, distal margin rounded, medial margin concave with L-shaped appearance; exopod as long as endopod inserted dorsally near medial margin bellow distinct lobe, lobe not extending beyond medial margin; endopod short bearing many short setae. Pleopods 1–5 exopods with monospiracular respiratory structures.
</paragraph>
<caption id="F44D97B0E5FA794D9E6A26879015EBA2" ID-DOI="10.3897/zookeys.1223.133267.figure10" ID-arpha="7375334A-8248-50F0-B99F-D92E75C5CBB7" httpUri="https://binary.pensoft.net/fig/1218795" startId="F10">
<paragraph id="FD6B5B3DDFC41538B6738F130B090DDE">
<label id="00005A590E3D3E8BC28A8C31C6122883">Figure 10.</label>
</paragraph>
<paragraph id="B6A16ED64B86BDFAA7FED35A0C6E5A2C">
<taxonomicName id="D84B424154FD538CC1F977AA781BC4F0" authority="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityName="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityYear="2025" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="10E6DDABB14AB5D02E5F21B238DDBE48" italics="true">Diploexochus troglobius</emphasis>
López-Orozco, Borja-Arrieta &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="8DDD679157322EBFA85133B91B6CD7CE" rank="species">sp. nov.</taxonomicNameLabel>
(♀ paratype,
<abbrev content-type="institution" id="ABBRID0E6OAG" xlink_title="University of Cartagena, Cartagena, Colombia">CBUDC-CRU</abbrev>
396)
<emphasis id="F5230480170A2505D7D045F4EF1E2EEF" bold="true">A</emphasis>
left mandible
<emphasis id="E8DEC999C8E407B769275AB297EFBBD9" bold="true">B</emphasis>
right mandible
<emphasis id="2045ADB43AAF137C793F07D46309DA69" bold="true">C</emphasis>
maxillula
<emphasis id="7248D5DB5DB80695D3F41A60A26339A4" bold="true">D</emphasis>
maxilla
<emphasis id="1E37B1989691CA1C98350F2907BD5334" bold="true">E</emphasis>
maxilliped
<emphasis id="AC9252793BA8593FAE63ACF8BC7AD2A9" bold="true">F</emphasis>
uropod.
</paragraph>
</caption>
<paragraph id="2364FC815A47B1DC31DF6B8B7FB77E7C">
<emphasis id="462DCDCFBFC2E57879C13AABF2EE3A5F" bold="true">Male.</emphasis>
Pereopods 1–7 (Fig.
<figureCitation id="B6DFA304067937DEF4DE13BC92811DF5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♂ paratype, CBUDC-CRU 396) A pereopod 1 B pereopod 7 C genital papilla D pleopod 1 E pleopod 2 F pleopod 3 exopod G pleopod 4 exopod H pleopod 5 exopod." figureDoi="10.3897/zookeys.1223.133267.figure11" httpUri="https://binary.pensoft.net/fig/1218796">11 A, B</figureCitation>
) without particular modifications. Genital papilla as in Fig.
<figureCitation id="AA64F5C3B5EFC9DD086725FBA90D5278" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♂ paratype, CBUDC-CRU 396) A pereopod 1 B pereopod 7 C genital papilla D pleopod 1 E pleopod 2 F pleopod 3 exopod G pleopod 4 exopod H pleopod 5 exopod." figureDoi="10.3897/zookeys.1223.133267.figure11" httpUri="https://binary.pensoft.net/fig/1218796">11 C</figureCitation>
. Pleopod 1 (Fig.
<figureCitation id="BB4CA79711EB7C982481E89FB63E3DC1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♂ paratype, CBUDC-CRU 396) A pereopod 1 B pereopod 7 C genital papilla D pleopod 1 E pleopod 2 F pleopod 3 exopod G pleopod 4 exopod H pleopod 5 exopod." figureDoi="10.3897/zookeys.1223.133267.figure11" httpUri="https://binary.pensoft.net/fig/1218796">11 D</figureCitation>
) exopod hour-glass shaped, twice as wide as long, inner portion rounded, outer portion triangular, distal and proximal margins narrow on middle; endopod about three times as long as exopod. Pleopod 2 (Fig.
<figureCitation id="68475274001412DE0CAED0DA532923C0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♂ paratype, CBUDC-CRU 396) A pereopod 1 B pereopod 7 C genital papilla D pleopod 1 E pleopod 2 F pleopod 3 exopod G pleopod 4 exopod H pleopod 5 exopod." figureDoi="10.3897/zookeys.1223.133267.figure11" httpUri="https://binary.pensoft.net/fig/1218796">11 E</figureCitation>
) exopod triangular, outer margin strongly concave; endopod slightly longer than exopod. Pleopod 3–5 exopods as in Fig.
<figureCitation id="94A5E4BB838E9494170B8BB91A1427AE" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. (♂ paratype, CBUDC-CRU 396) A pereopod 1 B pereopod 7 C genital papilla D pleopod 1 E pleopod 2 F pleopod 3 exopod G pleopod 4 exopod H pleopod 5 exopod." figureDoi="10.3897/zookeys.1223.133267.figure11" httpUri="https://binary.pensoft.net/fig/1218796">11 F – H</figureCitation>
.
</paragraph>
<caption id="0CB3CF8FC125996EBE539B210532CD78" ID-DOI="10.3897/zookeys.1223.133267.figure11" ID-arpha="38AAD98C-A85B-5A27-9467-9B8BD3FAC29A" httpUri="https://binary.pensoft.net/fig/1218796" startId="F11">
<paragraph id="8AD3A08BE33954973CE303AF27DDDC22">
<label id="7B1A6A58A6B64BA64E023631CD9F8E25">Figure 11.</label>
</paragraph>
<paragraph id="90630E8564832159088DE600048093A1">
<taxonomicName id="FFD65CE56ED36A3F61BDC393352E6F03" authority="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityName="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityYear="2025" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="DAB84882CDDC071790FECDDD6EEB0868" italics="true">Diploexochus troglobius</emphasis>
López-Orozco, Borja-Arrieta &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="A967082466FA04B3BA9D4616225E14F8" rank="species">sp. nov.</taxonomicNameLabel>
(♂ paratype,
<abbrev content-type="institution" id="ABBRID0EARAG" xlink_title="University of Cartagena, Cartagena, Colombia">CBUDC-CRU</abbrev>
396)
<emphasis id="4F52B203308DDB7D4C756369EF3E832A" bold="true">A</emphasis>
pereopod 1
<emphasis id="7873D617C8E1F1EA08201B428DD28B0D" bold="true">B</emphasis>
pereopod 7
<emphasis id="BA409619DB262BF480F226390494813E" bold="true">C</emphasis>
genital papilla
<emphasis id="8DDDCC1D4BD176427386E7FD66C79E23" bold="true">D</emphasis>
pleopod 1
<emphasis id="A046F205E6353775FA86DC2259D1B9FB" bold="true">E</emphasis>
pleopod 2
<emphasis id="4FE1B0C62A3C5D9DF7B88A81C82FF841" bold="true">F</emphasis>
pleopod 3 exopod
<emphasis id="8D66450FECE568C3C3B4B76FCFE0031F" bold="true">G</emphasis>
pleopod 4 exopod
<emphasis id="5E3AC87F36E99436146030661EF235D2" bold="true">H</emphasis>
pleopod 5 exopod.
</paragraph>
</caption>
<caption id="1FEF480838D5BCCB558A7E25C469FE77" ID-DOI="10.3897/zookeys.1223.133267.figure12" ID-arpha="1B498088-3694-5AAE-BDAF-1D63F42BFAD5" httpUri="https://binary.pensoft.net/fig/1218797" startId="F12">
<paragraph id="B2992A27E01A95339F312033E6BDD617">
<label id="329B3A98CD4B2492AD02946E1ACBE332">Figure 12.</label>
</paragraph>
<paragraph id="C1D4D6076E8A6F8B1ABE768494ED6493">
<emphasis id="A8946ACC32BE77E14E541FFCA93DC383" bold="true">A</emphasis>
Tropical Dry Forest around Roca Madre Adventure Park, Sucre
<emphasis id="5E1F7B19AEDB36AFA3B4994FC29BD846" bold="true">B</emphasis>
livestock around Roca Madre Adventure Park
<emphasis id="537BFC831F486F12B0EEDC54E2BA8FF2" bold="true">C</emphasis>
Roca Madre Cave Gallery
<emphasis id="36DF3FCD10E77760D7C92B52D679FEAF" bold="true">D</emphasis>
<taxonomicName id="012241F4A1A9807513185E0AA11FA728" authority="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityName="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityYear="2025" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="17FE1DC5DEB9D2AAB2CCD2D5C5DAB739" italics="true">Diploexochus troglobius</emphasis>
López-Orozco, Borja-Arrieta &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="C7BBB845D86E75F700B85604F9625ABB" rank="species">sp. nov.</taxonomicNameLabel>
under limestone rocks.
</paragraph>
</caption>
<caption id="CEA609CF18E91BC00CE92A44D4E21AF0" ID-DOI="10.3897/zookeys.1223.133267.figure13" ID-arpha="DE94BD87-F27B-5EA9-BD0C-4610E8F2AE1F" httpUri="https://binary.pensoft.net/fig/1218798" startId="F13">
<paragraph id="8D01A4A463B80A09053C34C711AB82B2">
<label id="D9D10A97F55C99CD67F7CDE9D0CCAE79">Figure 13.</label>
</paragraph>
<paragraph id="1A00A2445EE036C3F4EED3B465E2E864">
Habitus of the species of the genus
<taxonomicName id="206BD1FE8C258778219FFDFC64E7D5B8" authority="Brandt, 1833" authorityName="Brandt" authorityYear="1833" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="genus">
<emphasis id="5E547C8DD65F305E673877BFD5CC4B52" italics="true">Diploexochus</emphasis>
Brandt, 1833
</taxonomicName>
of Colombia:
<emphasis id="812678702DC925EFCBB67B4FCC200BB8" bold="true">A</emphasis>
<taxonomicName id="13D2E158832CDFD49F22EDB18A9F078C" authority="(Pearse, 1915)" authorityName="Borja-Arrieta &amp; López-Orozco &amp; Carpio-Díaz &amp; Gutierrez-Estrada &amp; Campos-Filho &amp; Navas-S. &amp; Bichuette" authorityYear="2025" baseAuthorityName="Pearse" baseAuthorityYear="1915" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="brevispinis" status="comb. nov.">
<emphasis id="0AFAA197D120C0A4841224D8237A7EBC" italics="true">Diploexochus brevispinis</emphasis>
(
<bibRefCitation id="7DC6E933E5E7A5408BED85D2DC4544FD" DOI="10.5479/si.00963801.49-2123.531" author="Pearse A" journalOrPublisher="Proceedings of the United States National Museum" pagination="531-556" refId="B49" refString="Pearse A (1915) An account of the Crustacea collected by the Walker Expedition to Santa Marta, Colombia. Proceedings of the United States National Museum 49: 531 - 556. https: // doi. org / 10.5479 / si. 00963801.49 - 2123.531" title="An account of the Crustacea collected by the Walker Expedition to Santa Marta, Colombia." volume="49" year="1915">Pearse, 1915</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="B08D030B4BDB630925B22950E9895A8A" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="A2F57C319D82F6CED9F1E34D90CA3EA7" bold="true">B</emphasis>
<taxonomicName id="6A4EBD19DD3ABFD7EFEC6BD7227509D4" authority="Lopez-Orozco, Carpio-Diaz &amp; Campos-Filho" authorityName="Lopez-Orozco, Carpio-Diaz &amp; Campos-Filho" authorityYear="2025" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="cacique" status="sp. nov.">
<emphasis id="132E0073C31BF7AEDA1B604B796AC693" italics="true">Diploexochus cacique</emphasis>
López-Orozco, Carpio-Díaz &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="CDE293F392795A02809B8AB0F76EE63C" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="A24CA172A9C82D866351D51251E762FB" bold="true">C</emphasis>
<taxonomicName id="C37815BB87EA7667A5C7BAB05081A472" authority="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityName="Lopez-Orozco, Borja-Arrieta &amp; Campos-Filho" authorityYear="2025" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="B2219039C8893C7B582AAC371A3DAFEF" italics="true">Diploexochus troglobius</emphasis>
López-Orozco, Borja-Arrieta &amp; Campos-Filho
</taxonomicName>
,
<taxonomicNameLabel id="E22961F0E2C717992E36ADE7196DDBA0" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EVUAG" type="etymology">
<paragraph id="F79CD0D1CE9F73803F0540A599B8A0A0">
<heading id="A04F0EF7C20131573ACF140A8D159FA6" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="004F895B427B3D582EB82603F8064C89">
Latin:
<emphasis id="8CDFC3DC1DEB217EA4A0509FFE77F9B9" italics="true">troglo</emphasis>
+
<emphasis id="F86B53C6C7D6AD2A590884A5003D5497" italics="true">bio</emphasis>
= cave-dwelling. The new name of the species is an adjective that refers to the troglobitic category of the species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6UAG" type="remarks">
<paragraph id="2AA37064B534D2C3BC132C789AA61454">
<heading id="EBC7792D4857E7AF83E012C007939E05" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="D2633FC7E6B97C18D581D56C14FBB4E6">
<taxonomicName id="84F59E63262EE1ADDB6B68FAC955ABC9" authorityName="Borja-Arrieta &amp; López-Orozco &amp; Carpio-Díaz &amp; Gutierrez-Estrada &amp; Campos-Filho &amp; Navas-S. &amp; Bichuette" authorityYear="2025" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="FC343C6E4D1B445FD4548CD549653AC4" italics="true">Diploexochus troglobius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="818A6216516C71D804100707A8AF9906" rank="species">sp. nov.</taxonomicNameLabel>
is easily distinguishable from the congeners in the arrangement of the dorsal tubercles of the pleon, dorsal surface with pilose aspect, eyes of four ommatidia, and the shape of the male pleopod 1 exopod.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQVAG" type="natural history">
<paragraph id="CB7A3E1FBA98DFFC5F8BCD5D77A006D9">
<heading id="CC50FCC512175143D88918AFD2546068" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="03AFD18318148A1FC8D1EA86D6461676">
Specimens of
<taxonomicName id="061BB67DFB6DDD48CC5F524A045D2511" authorityName="Borja-Arrieta &amp; López-Orozco &amp; Carpio-Díaz &amp; Gutierrez-Estrada &amp; Campos-Filho &amp; Navas-S. &amp; Bichuette" authorityYear="2025" class="Malacostraca" family="Armadillidae" genus="Diploexochus" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="troglobius" status="sp. nov.">
<emphasis id="8EFAAEFE758A539BB9E110241040A778" italics="true">D. troglobius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0E9889594CCC28ADE28FF5DF919E1B1F" rank="species">sp. nov.</taxonomicNameLabel>
were collected in the aphotic zone of the Roca Madre Cave, beneath limestone rocks (Fig.
<figureCitation id="E6FE19578B4331C8BFDCE3CC06630320" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A Tropical Dry Forest around Roca Madre Adventure Park, Sucre B livestock around Roca Madre Adventure Park C Roca Madre Cave Gallery D Diploexochus troglobius López-Orozco, Borja-Arrieta &amp; Campos-Filho, sp. nov. under limestone rocks." figureDoi="10.3897/zookeys.1223.133267.figure12" httpUri="https://binary.pensoft.net/fig/1218797">12 D</figureCitation>
). This species is considered troglobitic due to the reduction of body pigments and reduction in the number of ommatidia. In addition, several surveys were conducted outside the cave and other subterranean ecosystems to confirm its restricted distribution. This species is considered endemic to the study area.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFWAG" type="distribution">
<paragraph id="D67D1E490E61325B17F196D046C2A45A">
<heading id="8195F58C606E8D7558F7CA0AE2131D6B" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="F5CFA1A5686A9EF6859CCC72142BA53C">
This species is known only from the type locality at Roca Madre Cave,
<collectingRegion id="50586B4BC7CDBA70AB996AB60EA756C1" country="Venezuela" name="Sucre">Sucre</collectingRegion>
, inserted on
<abbrev id="ABBRID0ELWAG" xlink_title="Tropical Dry Forest">TDF</abbrev>
area (Fig.
<figureCitation id="5B1B0262BAD60491AD0FAC49FCD39DC6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Diploexochus species from Colombia. Green areas: Tropical Dry Forest (TDF) from Colombia (García et al. 2014)." figureDoi="10.3897/zookeys.1223.133267.figure1" httpUri="https://binary.pensoft.net/fig/1218786">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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