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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1111.77586" ID-GBIF-Dataset="4f820466-2eff-42bb-9c93-16143363528f" ID-PMC="PMC9848911" ID-Pensoft-Pub="1313-2970-1111-43" ID-Pensoft-UUID="6E23DAFA45395554A61E23AD4B06AD68" ID-PubMed="36760855" ID-ZooBank="3FAAEA839E8141A99B868576F8A1F33A" ModsDocID="1313-2970-1111-43" checkinTime="1657577097621" checkinUser="pensoft" docAuthor="Blahnik, Roger &amp; Andersen, Trond" docDate="2022" docId="701827E8CB355A258418943E8DC98B0D" docLanguage="en" docName="ZooKeys 1111: 43-198" docOrigin="ZooKeys 1111" docPubDate="2022-07-11" docSource="http://dx.doi.org/10.3897/zookeys.1111.77586" docTitle="Chimarra amakyei Blahnik &amp; Andersen 2022, sp. nov." docType="treatment" docUuid="7E3AB7DA-C910-461E-9057-754E790215CE" docUuidSource="ZooBank" docVersion="4" id="6E23DAFA45395554A61E23AD4B06AD68" lastPageNumber="43" masterDocId="6E23DAFA45395554A61E23AD4B06AD68" masterDocTitle="New species of the genus Chimarra Stephens from Africa (Trichoptera, Philopotamidae) and characterization of the African groups and subgroups of the genus" masterLastPageNumber="198" masterPageNumber="43" pageNumber="43" updateTime="1732488290164" updateUser="ExternalLinkService">
<mods:mods id="93490BF05BF2A8FFECE6E7741882A37C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3C522B26B4FBAFA9A9A1E13B1E248EB8">
<mods:title id="36AEB22AFF677701B7F31827AD5AEC8C">New species of the genus Chimarra Stephens from Africa (Trichoptera, Philopotamidae) and characterization of the African groups and subgroups of the genus</mods:title>
</mods:titleInfo>
<mods:name id="7E1ED481388EFFFCA7EA3588C908B96A" type="personal">
<mods:role id="C2ADDE3E325B6FD42D26DE43922C3F3B">
<mods:roleTerm id="BBE13C0B788083FE4CDB4C68B1A39B53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31AFB4310E025BA4C828124F1C490DCD">Blahnik, Roger</mods:namePart>
<mods:affiliation id="F1779D967EC1C697CC3C78D8853F07C7">Department of Entomology, University of Minnesota, 1980 Folwell Ave., 219 Hodson Hall, St. Paul, Minnesota, 55108, USA</mods:affiliation>
</mods:name>
<mods:name id="9DA0D4D4D2DC8EFBEE02D192C8574776" type="personal">
<mods:role id="C6A64ACD0C19E024BB34A4D314ED921C">
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</mods:role>
<mods:namePart id="D8DC06B248F520766D5BDFC16FD30999">Andersen, Trond</mods:namePart>
<mods:nameIdentifier id="F63D7879244AB1CA619F447213A2EC19" type="ORCID">https://orcid.org/0000-0003-2201-1870</mods:nameIdentifier>
<mods:affiliation id="31B1889B7A946F5F06DB97926417012F">Department of Natural History, University Museum of Bergen, University of Bergen, PO Box 7800, NO- 5020 Bergen, Norway</mods:affiliation>
<mods:nameIdentifier id="C1471DE9D97B71B99597FCCC788E81A4" type="email">trond.andersen@uib.no</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A32DEB979D9530D38C8D918B57740009">text</mods:typeOfResource>
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<mods:title id="839B63A10D8486132583E65365B17509">ZooKeys</mods:title>
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<mods:part id="7C7A2F6C6C44A1D8AE2E10920931DE8F">
<mods:date id="3C8260BCF853E27CAFA8E5139BD4FDFC">2022</mods:date>
<mods:detail id="75C33632F4AD8169A08E740B8502866A" type="pubDate">
<mods:number id="478166BEDD5C8C59723A3EFDEDF8A524">2022-07-11</mods:number>
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<mods:detail id="BB918FE932C8E4DD4E5DD611217114D4" type="volume">
<mods:number id="69751EDA3708D8427D2F4FFCD5831619">1111</mods:number>
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<mods:url id="7248BB10268E2C91F4037E12FDB39CDA">http://dx.doi.org/10.3897/zookeys.1111.77586</mods:url>
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<mods:classification id="19345272A36D1A8BD8B9CD0A34822BC0">journal article</mods:classification>
<mods:identifier id="0489608C713D0E8BD4E4CD8408FBA248" type="DOI">http://dx.doi.org/10.3897/zookeys.1111.77586</mods:identifier>
<mods:identifier id="B12506EBD0B4BA531573200606321ADB" type="Pensoft-Pub">1313-2970-1111-43</mods:identifier>
<mods:identifier id="DC118A47DDF48A8799710740AF212830" type="ZooBank">3FAAEA839E8141A99B868576F8A1F33A</mods:identifier>
<mods:identifier id="EB6EFECCC04F2866A2C11B9788C0EBE2" type="Pensoft-UUID">6E23DAFA45395554A61E23AD4B06AD68</mods:identifier>
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<treatment id="701827E8CB355A258418943E8DC98B0D" ID-GBIF-Taxon="197225421" LSID="urn:lsid:zoobank.org:act:7E3AB7DA-C910-461E-9057-754E790215CE" httpUri="http://treatment.plazi.org/id/701827E8CB355A258418943E8DC98B0D" lastPageNumber="43" pageId="0" pageNumber="43" scope_family="Philopotamidae" scope_order="Trichoptera">
<subSubSection id="6C15ADB897E3DFAED65783729CDD546D" pageId="0" pageNumber="43" type="nomenclature">
<paragraph id="62C11733BFE6B1F651933964CDC4CED7" pageId="0" pageNumber="43">
<taxonomicName id="2DA3CC6E7E21631D58A3BD5052CAE320" LSID="http://zoobank.org/7E3AB7DA-C910-461E-9057-754E790215CE" authority="Blahnik &amp; Andersen, 2022" authorityName="Blahnik &amp; Andersen" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra amakyei" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="amakyei" status="sp. nov.">Chimarra amakyei</taxonomicName>
<taxonomicNameLabel id="D4F29688E5F2D273956C1E2F7325D00E" pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="60AA4B1ED4024A46108470E45D1B1030" pageId="0" pageNumber="43" type="description">
<paragraph id="AA1F211561EF9BCEFBDBB26185607A97" pageId="0" pageNumber="43">
<figureCitation id="848868E3F594C2EB9A948DF18584630A" captionStart="Figure 24" captionStartId="F23" captionText="Figure 24. Chimarra amakyei sp. nov., ♂ genitalia A lateral B dorsal, segments IX and X C inferior appendage, dorsal D phallus, lateral E phallus, dorsal." figureDoi="10.3897/zookeys.1111.77586.figure24" httpUri="https://binary.pensoft.net/fig/711490" pageId="0" pageNumber="43">Fig. 24A-E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="17BD1578538593B2FF2CB7000062BFC2" pageId="0" pageNumber="43" type="type material">
<paragraph id="76A3FABB67CB0C2D98369837085A3027" pageId="0" pageNumber="43">Type material.</paragraph>
<paragraph id="41D6E7BE41A1F881EF0E1D9FE76D4D62" pageId="0" pageNumber="43">
<emphasis id="0AA85E98736B013B8E6B7289BCBDCB10" bold="true" pageId="0" pageNumber="43">
<emphasis id="99DF1E262DA375BE4F1FE09434D83784" italics="true" pageId="0" pageNumber="43">Holotype</emphasis>
.
</emphasis>
Ghana -
<emphasis id="69EECDF4504019ADDA068F49D4177518" bold="true" pageId="0" pageNumber="43">Volta Reg.</emphasis>
● ♂ (in alcohol); Wli, Agumatsa waterfall, station # 3;
<geoCoordinate id="C7E55CE41218117BA7BD19717C0009A2" degrees="7" direction="north" minutes="07" orientation="latitude" precision="15" seconds="29" value="7.1247225">7°07'29&quot;N</geoCoordinate>
,
<geoCoordinate id="4985E7327904132892DD633357978544" degrees="0" direction="east" minutes="35" orientation="longitude" precision="15" seconds="31" value="0.5919444">0°35'31&quot;E</geoCoordinate>
; 10 Mar. 1993; JS Amakye &amp; J
<normalizedToken id="51C11033EF323E9369AF0A68F50D8E07" originalValue="Kjærandsen">Kjaerandsen</normalizedToken>
leg.; sweep net; UMSP 000550018.
<emphasis id="5C56B295165F4EDB16B18062B6B0F03B" bold="true" pageId="0" pageNumber="43">
<emphasis id="C37015B71E16A791615EE59CFA0357B5" italics="true" pageId="0" pageNumber="43">Paratypes</emphasis>
.
</emphasis>
Ghana -
<emphasis id="A45DFB1B5AA25956FCC9246BB59163BF" bold="true" pageId="0" pageNumber="43">Volta Reg.</emphasis>
● 3♂♂; same data as for holotype except station # 2A; 8-11 Mar. 1993; Malaise trap; ZMBN.
</paragraph>
</subSubSection>
<subSubSection id="60740F7DCF9B640EC60CC0F03BE751E6" pageId="0" pageNumber="43" type="materials_examined">
<paragraph id="CBB3EFC3193F8BC3B402A34048D6DF60" pageId="0" pageNumber="43">Additional material.</paragraph>
<paragraph id="65D0656562A73DAE136C9589AD4E2A80" pageId="0" pageNumber="43">
<materialsCitation id="29B092C08B372E75710A5D0B471B3650" collectingDate="1993-03-05" collectingDateMax="1993-11-21" collectingDateMin="1993-03-05" collectingMethod="Malaise trap" collectorName="J. Kjaerandsen, JS Amakye, J Kjaerandsen" country="Ghana" latitude="7.1247225" location="Agumatsa" longLatPrecision="21" longitude="0.5919444" municipality="Wli" specimenCount="7" specimenCount-female="7" stateProvince="Volta">
<collectingCountry id="81203883D6EB9C3DEB9106EDC0F9F4DB" name="Ghana">Ghana</collectingCountry>
-
<emphasis id="895EFC5EBF8D47FD53CDFF0ABB5E4A19" bold="true" pageId="0" pageNumber="43">
<collectingRegion id="27EB830D1EE74E3CF2D301937EE1DB46" country="Ghana" name="Volta">Volta</collectingRegion>
Reg.
</emphasis>
<specimenCount id="6DB4B284F2B2C79D09DF432A39E2B4C1" type="female">2♀♀</specimenCount>
;
<collectingMunicipality id="09FC3F2E7C5691AD316EF51E4054AACB">Wli</collectingMunicipality>
,
<location id="02C50029852F81D519CD27BFBAB8D840" LSID="urn:lsid:plazi:treatment:701827E8CB355A258418943E8DC98B0D:02C50029852F81D519CD27BFBAB8D840" country="Ghana" latitude="7.1247225" longLatPrecision="21" longitude="0.5919444" municipality="Wli" name="Agumatsa" stateProvince="Volta">Agumatsa</location>
waterfall, station # 1A;
<geoCoordinate id="F55BCC9DF1B279F402F37B23CFD4196A" degrees="7" direction="north" minutes="07" orientation="latitude" precision="15" seconds="29" value="7.1247225">7°07'29&quot;N</geoCoordinate>
,
<geoCoordinate id="C328CB4A34267E5836491E4817730300" degrees="0" direction="east" minutes="35" orientation="longitude" precision="15" seconds="31" value="0.5919444">0°35'31&quot;E</geoCoordinate>
;
<collectingDate id="BF1EC70F188D80560CFE2F139E74986B" value="1993-11-12" valueMax="1993-11-21" valueMin="1993-11-12">12-21 Nov. 1993</collectingDate>
,
<collectorName id="2E4E9D627A391EFFF6C327CA662C1BC1">
J.
<normalizedToken id="6DDAA21082FF00C844508B9A5DFB44C6" originalValue="Kjærandsen">Kjaerandsen</normalizedToken>
</collectorName>
leg.;
<collectingMethod id="55261B5B5F5E3DFC5781DBC5004C9308">Malaise trap</collectingMethod>
; ZMBN ●
<specimenCount id="50F418CCA02BFC0E1085A90866AB5898" type="female">1♀</specimenCount>
; same collection data as for previous except station # 2B;
<collectingDate id="3BA7F45723F0FE255069856D1A7A449A" value="1993-03-05" valueMax="1993-03-14" valueMin="1993-03-05">5-14 Mar. 1993</collectingDate>
;
<collectorName id="757BC0998294864AC8CCC71278266FAC">JS Amakye</collectorName>
&amp;
<collectorName id="FCEE598BC5C22B4486F51FDF69946DF8">
J.
<normalizedToken id="4A46B47AD6C9EECC3B913FEF8F18C96B" originalValue="Kjærandsen">Kjaerandsen</normalizedToken>
</collectorName>
leg.; ZMBN ●
<specimenCount id="5AFDB57F4176B524C84F94CE22B4B5C1" type="female">2♀♀</specimenCount>
; same collection data as for previous except station # 5A;
<collectingDate id="A5A9274D9B91716D633923720987BB59" value="1993-03-10" valueMax="1993-03-13" valueMin="1993-03-10">10-13 Mar. 1993</collectingDate>
; ZMBN ●
<specimenCount id="FDC9FC89F92BF929311C0AE2C4F47B44" type="female">1♀</specimenCount>
; same collection data as for previous except station # 5D;
<collectingDate id="D8D9C4EAB40EFBC6D6D59B301A25F6E6" value="1993-03-09" valueMax="1993-03-12" valueMin="1993-03-09">9-12 Mar. 1993</collectingDate>
; ZMBN ●
<specimenCount id="F3DAE1344C55E4553B2B0C5B54DD9087" type="female">1♀</specimenCount>
; same collection data as for previous except station # 5C;
<collectingDate id="D67A2630CB53B4E26A033875466BCB87" value="1993-03-12" valueMax="1993-03-15" valueMin="1993-03-12">12-15 Mar. 1993</collectingDate>
,
<collectorName id="7547317ED001BE7FB2F97C5731AB32BA">
J
<normalizedToken id="2E535B3945EA9FF33FFA1053F299D199" originalValue="Kjærandsen">Kjaerandsen</normalizedToken>
</collectorName>
&amp;
<collectorName id="9E124DB711A006B18B45CEA80865B781">JS Amakye</collectorName>
leg.; UMSP
</materialsCitation>
.
</paragraph>
<caption id="62BCF62542DF63CB51059E1746175C23" doi="10.3897/zookeys.1111.77586.figure24" httpUri="https://binary.pensoft.net/fig/711490" pageId="0" pageNumber="43" start="Figure 24" startId="F23">
<paragraph id="2A59247B65E34B8F8A853EC264DEC975" pageId="0" pageNumber="43">
<emphasis id="A5B816C28AA45086A4597720D35BEE7F" bold="true" pageId="0" pageNumber="43">Figure 24.</emphasis>
<taxonomicName id="4553148DDF6BBF926B02370885083545" authorityName="Blahnik &amp; Andersen" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra amakyei" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="amakyei">
<emphasis id="41FDBA61A4D95CF67AA77F856C78B7F0" italics="true" pageId="0" pageNumber="43">Chimarra amakyei</emphasis>
</taxonomicName>
sp. nov.,
<specimenCount id="74B2D40E936A6E9CAACDC22D2BB8AF62"></specimenCount>
genitalia
<emphasis id="1A22C73574DC25478A0F47B1A7F0DF92" bold="true" pageId="0" pageNumber="43">A</emphasis>
lateral
<emphasis id="420F384BF587B5F7CCF3965FF18D9630" bold="true" pageId="0" pageNumber="43">B</emphasis>
dorsal, segments IX and X
<emphasis id="F74C61FBF13F2BDE8DD83C00082F5633" bold="true" pageId="0" pageNumber="43">C</emphasis>
inferior appendage, dorsal
<emphasis id="D1E3615262A731B25B591A5A1BEC02FE" bold="true" pageId="0" pageNumber="43">D</emphasis>
phallus, lateral
<emphasis id="6C9BB2F5544866363E18A3AD3FAA7DDD" bold="true" pageId="0" pageNumber="43">E</emphasis>
phallus, dorsal.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5C4B089300B52DD739944FED5F00C7E2" pageId="0" pageNumber="43" type="diagnosis">
<paragraph id="B318222C20D39055A48F88870AB0A151" pageId="0" pageNumber="43">Diagnosis.</paragraph>
<paragraph id="2EF0DF7C27DF78CEF94CF9B2117EC89B" pageId="0" pageNumber="43">
<taxonomicName id="850C17DAE7840054A7F79F5865ADEF0C" authorityName="Blahnik &amp; Andersen" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra amakyei" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="amakyei">
<emphasis id="26973D10DADB5FD3B63E6B632D278018" italics="true" pageId="0" pageNumber="43">Chimarra amakyei</emphasis>
</taxonomicName>
sp. nov. is closely related to
<taxonomicName id="628E78338CD51EB44A17857152003F21" authorityName="Mosely" authorityYear="1936" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra leta" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="leta">
<emphasis id="5A68E0DDB73ED9D3200907021A915868" italics="true" pageId="0" pageNumber="43">Chimarra leta</emphasis>
</taxonomicName>
Mosely. Like that species it has an elongate, narrow tergum X, with preanal appendages flattened and fused basally. It also has inferior appendages projecting approximately midlaterally, above a small basally located ventral process. It differs diagnostically in that the apices of inferior appendages are tapering and acute, rather than subtruncate.
</paragraph>
</subSubSection>
<subSubSection id="FC33195E0806D5674CDFE56E09E3B342" pageId="0" pageNumber="43" type="description">
<paragraph id="3138EE5C871FE5774AE6E1E0CDE1250E" pageId="0" pageNumber="43">Description.</paragraph>
<paragraph id="12EC91E41E3197DBA581A5741A5219E5" pageId="0" pageNumber="43">
<emphasis id="0FD7C313C2A2F1BB3C03AE550D044393" bold="true" italics="true" pageId="0" pageNumber="43">Adult.</emphasis>
Overall color (in alcohol) yellowish brown; vertex of head dark brown, setal warts very pale. Head relatively short (postocular parietal sclerite short). Palps moderately elongate; maxillary palp with 1st segment very short (approximately as long as wide), 2nd segment relatively short (distinctly shorter than 3rd), apex with small cluster of stiff setae, 3rd segment elongate, 4th segment very short (shorter than 2nd), 5th segment elongate (slightly longer than 3rd). Forewing length: male, 4.8-5.3 mm; female, 4.5-5.3 mm. Fore- and hind wings with forks I, II, III, and V present. Forewing with R1 sinuous, stem of Rs distinctly inflected at past midlength (without distinct node at inflection), basal fork of discoidal cell moderately enlarged, length of cell slightly greater than 2
<normalizedToken id="B263EF535913DD696EE8D8B220B9424E" originalValue="×">x</normalizedToken>
width, forks I and II sessile,
<emphasis id="058971C7A0AF0479FD63A9C9E31C56F3" italics="true" pageId="0" pageNumber="43">r</emphasis>
crossvein strongly diagonal, intersecting discoidal cell at just past midlength,
<emphasis id="CFAC77F12D259D1928D0ADCEB8C5DE47" italics="true" pageId="0" pageNumber="43">s</emphasis>
,
<emphasis id="F6FCF418754E2F5BBD896F174FFB4DA8" italics="true" pageId="0" pageNumber="43">r-m</emphasis>
, and
<emphasis id="76A704BCE5C4D3ADA27DF934A70169EE" italics="true" pageId="0" pageNumber="43">m</emphasis>
crossveins linear, s crossvein pigmented (like wing),
<emphasis id="EF0F79A5A0EC7C04FCABBB7DD6BD48DF" italics="true" pageId="0" pageNumber="43">r-m</emphasis>
and
<emphasis id="EC1FA14B52315EDA66261F027BE14D99" italics="true" pageId="0" pageNumber="43">m</emphasis>
crossveins hyaline (
<emphasis id="B33396AF659E3A29D7DE06CE257F0A9E" italics="true" pageId="0" pageNumber="43">m</emphasis>
very indistinct), 2A with crossvein (apparently forked apically to 1A and 3A). Hind wing with R1 reduced, narrowly parallel to subcosta, fork I sessile, fork II subsessile, fork III distal and relatively wide, anal loop small. Forelegs with apical tibial spur short; male without enlarged tarsal claws, apical segments of tarsi narrow, claws very small and symmetrical.
</paragraph>
<paragraph id="D10684293E949EE753C32D8D0DFC699B" pageId="0" pageNumber="43">
<emphasis id="7C5896ECDE962EECBD0ACF1C5D798689" bold="true" pageId="0" pageNumber="43">
<emphasis id="063F4E61CB83584E74B3AF97370E20C9" italics="true" pageId="0" pageNumber="43">Male genitalia</emphasis>
.
</emphasis>
Segment VIII with sternum very short, tergum moderately expanded dorsally (~ 2
<normalizedToken id="8629C5A05F771118A64131AA2E59CD58" originalValue="×">x</normalizedToken>
length of sternum at base). Segment IX, in lateral view, relatively short, anterior margin concave, dorsally with broadly rounded apodeme, anteroventrally with short angular projection at approximately basal
<normalizedToken id="4226CA94D28F2F6E64D8829807A2D299" originalValue="¼">1/4</normalizedToken>
; posterior margin moderately produced midlaterally, basoventrally with very short, subtriangular, posteriorly projected ventral process, segment expanded and sloping dorsal to process, with inferior appendages inserted high above process, approximately midlaterally; as viewed dorsally, with tergum very narrow, but continuous, sternum subtruncate, slightly concave mesally. Lateral lobes of tergum X elongate, rugose and club-like, with small rounded, mesally directed process bearing two sensilla at past midlength; dorsum of tergum X short, membranous, continuous with paired, sclerotized, apically rounded, periphallic processes surrounding phallic apparatus laterally and ventrally. Preanal appendages narrow and flattened, fused basally to lateral lobes. Inferior appendage without pronounced basal inflection, appendage narrow basally, dorsal margin with rounded expansion, tapering to acute, projecting apex; base of dorsal expansion with very short, sclerotized, mesally curved projection, continuous on mesal surface as sclerotized ridge. Phallic apparatus with relatively small, tubular phallobase with usual basodorsal expansion, apicodorsal margin with deep membranous invagination, ventral margin projecting, but without apicomesal projection; endotheca with two very small spines; phallotremal sclerite complex composed of reclinate, ring-like structure, with short rounded apicolateral projections.
</paragraph>
</subSubSection>
<subSubSection id="FD556989EAAAA29A409E5CD4DC886235" pageId="0" pageNumber="43" type="etymology">
<paragraph id="5D31F3FF7481137ABBB37A834685017A" pageId="0" pageNumber="43">Etymology.</paragraph>
<paragraph id="1FE2DB671B0D89AD0E0C0448A90A720D" pageId="0" pageNumber="43">
<taxonomicName id="2104148AF72546352C0C29B8E12A7FD0" authorityName="Blahnik &amp; Andersen" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra amakyei" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="amakyei">
<emphasis id="CB5A0543FB232E5DF417B4837EFEE99A" italics="true" pageId="0" pageNumber="43">Chimarra amakyei</emphasis>
</taxonomicName>
, named for Joseph S. Amakye, who helped collect much of the material represented in this paper, in recognition of his efforts.
</paragraph>
</subSubSection>
</treatment>
</document>