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<document ID-CLB-Dataset="59958" ID-DOI="http://dx.doi.org/10.3897/phytokeys.230.103882" ID-GBIF-Dataset="58b283e2-bf49-4656-8668-6086b2b4e003" ID-Pensoft-Pub="1314-2003-230-1" ID-Pensoft-UUID="365B97AE96F45DB295043BF02F4AF4BE" ModsDocID="1314-2003-230-1" checkinTime="1691127498364" checkinUser="pensoft" docAuthor="Peterson, Paul M., Herrera Arrieta, Yolanda, Lobo Cabezas, Silvia &amp; Romaschenko, Konstantin" docDate="2023" docId="6FEE910D34BF5089B95E138DD3861427" docLanguage="en" docName="PhytoKeys 230: 1-106" docOrigin="PhytoKeys 230" docPubDate="2023-08-03" docSource="http://dx.doi.org/10.3897/phytokeys.230.103882" docTitle="Muhlenbergia setarioides E. Fourn., Mexic. Pl. 2: 84. 1886." docType="treatment" docVersion="2" id="365B97AE96F45DB295043BF02F4AF4BE" lastPageNumber="1" masterDocId="365B97AE96F45DB295043BF02F4AF4BE" masterDocTitle="Taxonomic revision of Muhlenbergia (Poaceae, Chloridoideae, Cynodonteae, Muhlenbergiinae) in Central America: phylogeny and classification" masterLastPageNumber="106" masterPageNumber="1" pageNumber="1" updateTime="1691128203638" updateUser="ExternalLinkService">
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<mods:titleInfo id="26268E9243B6828FC4AB0473C8A8240B">
<mods:title id="85E4DE7FAC3A636F456ACC7FC1AF5C80">Taxonomic revision of Muhlenbergia (Poaceae, Chloridoideae, Cynodonteae, Muhlenbergiinae) in Central America: phylogeny and classification</mods:title>
</mods:titleInfo>
<mods:name id="8DF0F23A7AB080745789462FDA1F0686" type="personal">
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</mods:role>
<mods:namePart id="3601CD0A62F8BE7BE4ED3AF4948D65F7">Peterson, Paul M.</mods:namePart>
<mods:nameIdentifier id="6CE39FF2DDC816797EE265231EAAB54F" type="ORCID">https://orcid.org/0000-0001-9405-5528</mods:nameIdentifier>
<mods:affiliation id="797BB28FA36C26B65BC025E6F2AAA4B3">Department of Botany MRC- 166, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier id="294771107B2CE240EC5F2927509808F8" type="email">peterson@si.edu</mods:nameIdentifier>
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<mods:roleTerm id="34297AA17E35599153905E0219FE4BDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFFDA57A731CD03F1B3A0B97924B5A12">Herrera Arrieta, Yolanda</mods:namePart>
<mods:nameIdentifier id="5926D17D05A97A3ABD037A00609D79FB" type="ORCID">https://orcid.org/0000-0003-3814-6518</mods:nameIdentifier>
<mods:affiliation id="02E0E6F6FB5B474913DFEC1B41A5E20C">Instituto Politecnico Nacional, CIIDIR Unidad-Durango-COFAA, Durango, C. P. 34220, Mexico</mods:affiliation>
</mods:name>
<mods:name id="762E359387ACB70FF66761929F4163A6" type="personal">
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<mods:namePart id="85E71C7D377A6C6E2EA03B7F1E8C86C3">Lobo Cabezas, Silvia</mods:namePart>
<mods:nameIdentifier id="AAF22926829B2E0B330122E8435E659A" type="ORCID">https://orcid.org/0009-0007-8100-2559</mods:nameIdentifier>
<mods:affiliation id="CA1E3F05F8718747DFAD12A4E42A1DFB">Herbario Nacional, Museo Nacional de Costa Rica, Apartado Postal 749 - 1000, San Jose, Costa Rica</mods:affiliation>
</mods:name>
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<mods:namePart id="46E871EAEC6907A6E60DE0116D7CEDAD">Romaschenko, Konstantin</mods:namePart>
<mods:nameIdentifier id="B85EA5BDB9828273AB014B2ED0B93501" type="ORCID">https://orcid.org/0000-0002-7248-4193</mods:nameIdentifier>
<mods:affiliation id="493540D8CD95CCF264DCC7C2EBB54F30">Department of Botany MRC- 166, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013 - 7012, USA</mods:affiliation>
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<mods:title id="BD6AD922BC88517367C9F29A1B9346CA">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="E07C814EFE0CF97247AEE27AC327E4A9">
<mods:date id="04D2798253F4F4E8A646CD5B206036EB">2023</mods:date>
<mods:detail id="23381D2E68CA80792899578C421896E2" type="pubDate">
<mods:number id="AB49404761AB9410C5A1F22E45817EB0">2023-08-03</mods:number>
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<mods:detail id="DB9429435C295CCBC2B7E225DD95E604" type="volume">
<mods:number id="D15EC2489A18544A0CF874D1FD885AFE">230</mods:number>
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<mods:start id="83DDAD16BB0C12BA143D787A7E1AD248">1</mods:start>
<mods:end id="EF2C1BC754986D573EFFE7BEA451EA06">106</mods:end>
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<mods:url id="B3D4DACAAE136E696DB1B2623BC2DE81">http://dx.doi.org/10.3897/phytokeys.230.103882</mods:url>
</mods:location>
<mods:classification id="F5AF6B167FBC45AD6A8029F9B797F5B5">journal article</mods:classification>
<mods:identifier id="7F974F03D8C6735C2EFAB70D3AA6D934" type="DOI">http://dx.doi.org/10.3897/phytokeys.230.103882</mods:identifier>
<mods:identifier id="86470BA438234F195888A55AEA69AEA4" type="Pensoft-Pub">1314-2003-230-1</mods:identifier>
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<treatment id="6FEE910D34BF5089B95E138DD3861427" LSID="urn:lsid:plazi:treatment:6FEE910D34BF5089B95E138DD3861427" httpUri="http://treatment.plazi.org/id/6FEE910D34BF5089B95E138DD3861427" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="7A5ECF82184CD0941DC20B27511DE828" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="9ADD2324C90D558DC280EBCA9C5C7D58" pageId="0" pageNumber="1">
30.
<taxonomicName id="244D66B7C5434E932BFEA66722240C1E" ID-CoL="6S39X" ID-ENA="58091" LSID="6FEE910D-34BF-5089-B95E-138DD3861427" authority="E. Fourn., Mexic. Pl. 2: 84. 1886." authorityName="E. Fourn., Mexic. Pl. 2: 84. 1886." class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia setarioides" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="setarioides">Muhlenbergia setarioides E. Fourn., Mexic. Pl. 2: 84. 1886.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="AF83408ADFFC58CF4507BFD3D1D45A47" pageId="0" pageNumber="1" type="description">
<paragraph id="4F86C7DD8C19CE1B2551FAB0C4DF70DA" pageId="0" pageNumber="1">
<figureCitation id="B87F0AD7E9AC3263C4B1C1697F9C290A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. A-E Muhlenbergia spiciformis Trin. A habit B ligule C glumes D floret E stamens, pistil, and lodicules F-J Muhlenbergia setarioides E. Fourn. F habit G ligule H glumes I floret J stamens, pistil, and lodicules. A-E drawn from P. M. Peterson &amp; C. R. Annable 8361 (US) F-J drawn from D. E. Breedlove 9350 (US)." figureDoi="10.3897/phytokeys.230.103882.figure21" httpUri="https://binary.pensoft.net/fig/887889" pageId="0" pageNumber="1">Fig. 21F-J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F791A1CC784988E56DAF1AA1FB3A2707" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="F48C6062B65D2CDD5C22A22D09D87239" pageId="0" pageNumber="1">
<taxonomicName id="FA43017B4A477AA917CC89E510FCACCB" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia polypogonoides" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="polypogonoides">= Muhlenbergia polypogonoides</taxonomicName>
Hack., Ann. K. K. Naturhist. Hofmus. 17: 255. 1902. Type:
<normalizedToken id="7363433CADEA91198CF7DDCEA9C7AEBF" originalValue="México">Mexico</normalizedToken>
,
<emphasis id="512C5DE9311E3B394A5AB38F926A92EB" italics="true" pageId="0" pageNumber="1">A. Schmitz 862</emphasis>
(holotype: W-18890124080!; isotypes: US-3412356! fragm. ex W, W-19160029073!
</paragraph>
</subSubSection>
<subSubSection id="ECB810352FCEDA1BC01A213B96F7CD69" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="D29D12CE9277CA5D2F118F488C906841" pageId="0" pageNumber="1">
<typeStatus id="26132872DDFDCF5B1AB1FF5E934C79BF">Type</typeStatus>
.
</paragraph>
<paragraph id="0DD9239AEF3D56641D37DF7B95262A61" pageId="0" pageNumber="1">
<materialsCitation id="B43969E0C2BC360855EA7E54FA7EAD39" collectingDate="1865-11-14" collectorName="M. Bourgeau, E. Fourn., Beal, Grass. N. Amer." country="Mexico" location="Orizaba" municipality="Borrego" specimenCount="1" typeStatus="lectotype">
<normalizedToken id="63FFA87FB193EE15CE9FC9CE630C26D4" originalValue="México">
<collectingCountry id="61A3D4DB4B4914786C7FC111144DD6F4" name="Mexico">Mexico</collectingCountry>
</normalizedToken>
,
<location id="B34F007CEE650C2F0FFAD1E8106F6C86" LSID="urn:lsid:plazi:treatment:6FEE910D34BF5089B95E138DD3861427:B34F007CEE650C2F0FFAD1E8106F6C86" country="Mexico" municipality="Borrego" name="Orizaba">Orizaba</location>
,
<collectingMunicipality id="DDFFEF9F0EC228A9FA51F7BEA98D8AD2">Borrego</collectingMunicipality>
,
<collectingDate id="1F72742FF419D17ECFA7990258BDB622" value="1865-11-14">14 Nov 1865</collectingDate>
-1866,
<emphasis id="AD6F3C3B89B9344C8BDD9EEFAB1CBCCE" italics="true" pageId="0" pageNumber="1">
<collectorName id="26468BF38D9E5FD369F7B927F015A4A1">M. Bourgeau</collectorName>
3362
</emphasis>
(
<emphasis id="60DE24AF4C8565B4669E301B5C09E962" bold="true" pageId="0" pageNumber="1">
<typeStatus id="69FE04848AA1930A17464A981934F4AB">lectotype</typeStatus>
, designated here
</emphasis>
: MPU-026952 [image!]; isolectotypes: G-00099164 [image!], GH-0024047 [image!] ex P, K-000308906 [image!], MO-2974294!, US-0091990! ex P &amp; ex LE, S-14-29487 [image!]. ≡
<taxonomicName id="EDF6686B4121946AF9306A3A32987F2A" authorityName="Beal" authorityYear="1896" baseAuthorityName="E.Fourn." class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia sylvatica subsp. var. var. setarioides" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="sylvatica" subSpecies="var." variety="setarioides">Muhlenbergia sylvatica var. setarioides</taxonomicName>
(
<collectorName id="6D108A88D8C642082538204750C0FE10">E. Fourn.</collectorName>
)
<collectorName id="CEAF51A3C1B10924AA240CAA51D45CE3">Beal</collectorName>
,
<collectorName id="1329A391B15AE158FC3D8198542C1216">Grass. N. Amer.</collectorName>
2: 249. 1896.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="65907CECCA13B17AA33679E069D5CFA9" pageId="0" pageNumber="1" type="description">
<paragraph id="69001CA4BB69DAA3B7DEAC8805800AE6" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="98B8863222AC222C20443435D7045EFE" pageId="0" pageNumber="1">
Sprawling
<emphasis id="75F4AAC378BC77745A020B1FCE80C324" bold="true" italics="true" pageId="0" pageNumber="1">perennials</emphasis>
, rooting at the lower nodes.
<emphasis id="FA229B7573A5A7D0073A380757B33E0C" bold="true" italics="true" pageId="0" pageNumber="1">Culms</emphasis>
30-70(-100) cm tall, geniculate, usually glabrous, with many branches, about 1.5 mm thick near base.
<emphasis id="BFACE9FD00F6A6705AF082ED24561A54" bold="true" italics="true" pageId="0" pageNumber="1">Leaf sheaths</emphasis>
shorter or longer than internodes, terete, striate, glabrous or scaberulous;
<emphasis id="8E1D8CDCBBEA0035125CFF4C4D64AFFA" bold="true" italics="true" pageId="0" pageNumber="1">ligules</emphasis>
1.5-3.5 mm long, hyaline, becoming yellowish-brown or brownish, erose;
<emphasis id="F09CE6B1BD869642427B8A2C42C1ECAB" bold="true" italics="true" pageId="0" pageNumber="1">blades</emphasis>
4-12(-17) cm long, 3-6(-9) mm wide, flat, thin, dark green, scaberulous above, narrowing to the base, central vein whitish, prominent.
<emphasis id="2B0F6CFA9E04A29D3834000A63B256E8" bold="true" italics="true" pageId="0" pageNumber="1">Panicles</emphasis>
6-13.5(-15) cm long, 1-2 cm wide, yellowish-brown to greenish, densely flowered, interrupted below;
<emphasis id="0337C5BDED52685BF7BDF3DEEBAC70FD" bold="true" italics="true" pageId="0" pageNumber="1">primary branches</emphasis>
1-3.5 cm long, appressed or ascending, flowered to the base;
<emphasis id="42B9DD120FAE784190277843F077F4AB" bold="true" italics="true" pageId="0" pageNumber="1">pedicels</emphasis>
0.2-1.7 mm long, shorter than the spikelets, hispidulous.
<emphasis id="1725B393AB4CB2F8B82BCBD06EA8A0A6" bold="true" italics="true" pageId="0" pageNumber="1">Spikelets</emphasis>
(2-)2.5-3 mm long;
<emphasis id="B4E5E6AD141471BA691EF984A16590CA" bold="true" italics="true" pageId="0" pageNumber="1">glumes</emphasis>
1-2(-2.5) mm long, unequal, 1-veined, green and prominent, apex acute to acuminate;
<emphasis id="E22D33DC62727E8D6D23BE61228A5A1D" bold="true" italics="true" pageId="0" pageNumber="1">lower glumes</emphasis>
1-1.5 mm long;
<emphasis id="4ED277BF2243C4F7158C4528901A638F" bold="true" italics="true" pageId="0" pageNumber="1">upper glumes</emphasis>
1.5-2(-2.5) mm long;
<emphasis id="DE21EA1CC6C4E43E8E72893AB4033485" bold="true" italics="true" pageId="0" pageNumber="1">lemmas</emphasis>
(2-)2.5-3 mm long, lanceolate, pale, awned, mottled with dark green spots, the 3-veins green and prominent, scabrous along the veins, pilose over the central vein and margins on the lower 1/3-1/2, the awns 5-14 mm long, somewhat flexuous, purplish;
<emphasis id="479F97C450AA0C2A847C719B6B3EB21C" bold="true" italics="true" pageId="0" pageNumber="1">paleas</emphasis>
about as long as the lemma, pilose between the veins on lower
<normalizedToken id="6C2258E169FD44E25C9965D6A4DB97DD" originalValue="½">1/2</normalizedToken>
;
<emphasis id="B3C65B54D33ECD4A11E1F51BC9398440" bold="true" italics="true" pageId="0" pageNumber="1">anthers</emphasis>
1-1.2 mm long, yellowish.
<emphasis id="39B2D5E6C594C5BE2797D89F349C1B78" bold="true" italics="true" pageId="0" pageNumber="1">Caryopses</emphasis>
1.3-1.5 mm long, narrowly-ellipsoid, dark reddish-brown. 2
<emphasis id="62C0289F4CD45382E7BDD8957A611080" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 40.
</paragraph>
<caption id="1924701FEC9AB545A6324E87143DD6A4" doi="10.3897/phytokeys.230.103882.figure21" httpUri="https://binary.pensoft.net/fig/887889" pageId="0" pageNumber="1" start="Figure 21" startId="F21">
<paragraph id="466FBEF04307D5F99DEB7AD71F2B0D34" pageId="0" pageNumber="1">
<emphasis id="62D64661C947EE961E958B645DBE7945" bold="true" pageId="0" pageNumber="1">Figure 21.</emphasis>
<emphasis id="A67742DA4A6942B98464A3D01F5EF9DB" bold="true" pageId="0" pageNumber="1">A-E</emphasis>
<taxonomicName id="A72DB356F129A56FDD55227EC08161AD" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia spiciformis" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="spiciformis">
<emphasis id="1EEE502DDC13B930D5E593B67AC2EFBE" italics="true" pageId="0" pageNumber="1">Muhlenbergia spiciformis</emphasis>
</taxonomicName>
Trin.
<emphasis id="5277A3B3BE301D6AE0413AF926361DCD" bold="true" pageId="0" pageNumber="1">A</emphasis>
habit
<emphasis id="E2871489FDF30AF7ACC467C9FB3721BE" bold="true" pageId="0" pageNumber="1">B</emphasis>
ligule
<emphasis id="2D5E482BED82015751185317C8F4480A" bold="true" pageId="0" pageNumber="1">C</emphasis>
glumes
<emphasis id="8D2EB52019B2F5240C0BB7A10745E38B" bold="true" pageId="0" pageNumber="1">D</emphasis>
floret
<emphasis id="C8D3EC86C822C6E6B6A55C24D6194E98" bold="true" pageId="0" pageNumber="1">E</emphasis>
stamens, pistil, and lodicules
<emphasis id="F4E282E7C16B0899110F0808A3BB7430" bold="true" pageId="0" pageNumber="1">F-J</emphasis>
<taxonomicName id="3423EC53C0AE250CFFE9484718ACAD18" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia setarioides" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="setarioides">
<emphasis id="AC8F9E7EBA994CDF37489373FC8F5959" italics="true" pageId="0" pageNumber="1">Muhlenbergia setarioides</emphasis>
</taxonomicName>
E. Fourn.
<emphasis id="60FC5661A936700BDA67F01076D4A8B3" bold="true" pageId="0" pageNumber="1">F</emphasis>
habit
<emphasis id="EC95650DC92A813C3A62274EA9178A53" bold="true" pageId="0" pageNumber="1">G</emphasis>
ligule
<emphasis id="417EE70872C060619616890EB5C8B4C6" bold="true" pageId="0" pageNumber="1">H</emphasis>
glumes
<emphasis id="177D22BCB6DA7BBED1921C2DCEF09BF9" bold="true" pageId="0" pageNumber="1">I</emphasis>
floret
<emphasis id="316295B86A75439EBFC1B7E768FC1C44" bold="true" pageId="0" pageNumber="1">J</emphasis>
stamens, pistil, and lodicules.
<emphasis id="5D8CB0678A6568419A32B02CF9BD904D" bold="true" pageId="0" pageNumber="1">A-E</emphasis>
drawn from
<emphasis id="F9305253BDD384B515CAD5A3549D4996" italics="true" pageId="0" pageNumber="1">P.M. Peterson &amp; C. R. Annable 8361</emphasis>
(US)
<emphasis id="808696CC46CD0E37D726356B6CF9CC93" bold="true" pageId="0" pageNumber="1">F-J</emphasis>
drawn from
<emphasis id="114EDC2D9D48E417D8CFC2CEC8A9B94A" italics="true" pageId="0" pageNumber="1">D.E. Breedlove 9350</emphasis>
(US).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="485352064669A854CC645B75561F99E2" pageId="0" pageNumber="1" type="distribution">
<paragraph id="8DD1C4E336FC1D599F6764D2AAA8E8A2" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="B7F4D08035582196136597CEE6B57F39" pageId="0" pageNumber="1">
<taxonomicName id="9861C2BCD50AE7D130D309A2349B8552" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia setarioides" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="setarioides">
<emphasis id="E5C91B9B3C3285E0C8876DF49F3C4968" italics="true" pageId="0" pageNumber="1">Muhlenbergia setarioides</emphasis>
</taxonomicName>
ranges from Puebla, Oaxaca (
<emphasis id="0702A96970014A8195A301660E3510F7" italics="true" pageId="0" pageNumber="1">Peterson &amp; Annable 9897</emphasis>
, 1.4 mi E of Ayutla on Mex 179), Tlaxcala,Veracruz, and Chiapas in
<normalizedToken id="2A28AA2061C21F4D69E06D4D82CBC4DE" originalValue="México">Mexico</normalizedToken>
to Central America from Guatemala to Panama (
<bibRefCitation id="B0466C2B53DA2EA4CBBCD6D1AC0BA2CD" DOI="https://doi.org/10.21829/abm126.2019.1379" author="Sanchez-Ken, JG" journalOrPublisher="Acta Botanica Mexicana" pageId="0" pageNumber="1" pagination="1 - 115" refId="B53" refString="Sanchez-Ken, JG, 2018. Riqueza de especies, clasificacion y listado de las gramineas (Poaceae) de Mexico. Acta Botanica Mexicana 126 (126): 1 - 115, DOI: https://doi.org/10.21829/abm126.2019.1379" title="Riqueza de especies, clasificacion y listado de las gramineas (Poaceae) de Mexico." url="https://doi.org/10.21829/abm126.2019.1379" volume="126" year="2018">Sanchez-Ken 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D314010A991BF8B3F91FA173C4DEB236" pageId="0" pageNumber="1" type="ecology">
<paragraph id="90C85DAB4364A14BA21391F8B6D2B39A" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="DEF688F3478596D049EBFE9188881657" pageId="0" pageNumber="1">This species is found in tropical-wet forests, shaded banks, near cornfields, and along barrancas; 1500-2400 m.</paragraph>
</subSubSection>
<subSubSection id="73961D655B2710BEC445F78ABE5CD409" pageId="0" pageNumber="1" type="comments">
<paragraph id="1C9B582C8CE1C8E5F5DD9DA07086CD95" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="810DA8B287FDF65F7BAF4A3D3EF39AE4" pageId="0" pageNumber="1">
Morphologically,
<taxonomicName id="8DCD789337F2F988F0051E7EF829DCD1" family="Poaceae" kingdom="Plantae" lsidName="M. setarioides" order="Poales" pageId="0" pageNumber="1" rank="species" species="setarioides">
<emphasis id="251C8EE5D44F6A28FE7ED7C4C0F67A44" italics="true" pageId="0" pageNumber="1">M. setarioides</emphasis>
</taxonomicName>
resembles
<taxonomicName id="763ED990AAEE1850EB93EB48E557649B" family="Poaceae" kingdom="Plantae" lsidName="M. spiciformis" order="Poales" pageId="0" pageNumber="1" rank="species" species="spiciformis">
<emphasis id="370EDDA12C663695D52190AECD6F6122" italics="true" pageId="0" pageNumber="1">M. spiciformis</emphasis>
</taxonomicName>
but differs from the latter in having wider leaf blades 3-6(-9) mm wide (1-3 mm wide in
<taxonomicName id="B0FD3591AC736DC14009E233DAA92914" family="Poaceae" kingdom="Plantae" lsidName="M. spiciformis" order="Poales" pageId="0" pageNumber="1" rank="species" species="spiciformis">
<emphasis id="0E323A9FD106BA095E82C7E7FF62ED58" italics="true" pageId="0" pageNumber="1">M. spiciformis</emphasis>
</taxonomicName>
), shorter spikelets (2-)2.5-3 mm long (versus 3-4 mm long), acute to acuminate (obtuse to acute in
<taxonomicName id="CCC673C3B5817A5015956CFA9462E1BA" family="Poaceae" kingdom="Plantae" lsidName="M. spiciformis" order="Poales" pageId="0" pageNumber="1" rank="species" species="spiciformis">
<emphasis id="5268BCD4FA555890ED88A96DF11DCA9B" italics="true" pageId="0" pageNumber="1">M. spiciformis</emphasis>
</taxonomicName>
) glumes, upper glumes 1.5-2(-2.5) mm long (versus ≤ 1mm long), and lemmas with awns 5-14 mm long [versus (10-)20-40 mm].
</paragraph>
<paragraph id="09AE540B4820777D57994A294826CA3F" pageId="0" pageNumber="1">
<taxonomicName id="F14A8059EFD240DAE65ED8B7366CD24A" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Muhlenbergia setarioides" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="setarioides">
<emphasis id="9DE9BEF20DAF0DB3F300648829DB5465" italics="true" pageId="0" pageNumber="1">Muhlenbergia setarioides</emphasis>
</taxonomicName>
is a member of
<taxonomicName id="363650239E2FB74E0A35A3BED6191AE0" family="Poaceae" kingdom="Plantae" lsidName="M. (Muhlenbergia) subsp. subg." order="Poales" pageId="0" pageNumber="1" rank="subGenus" subGenus="Muhlenbergia">M. subg. Muhlenbergia</taxonomicName>
and is sister to all remaining species in the subgenus (Fig.
<figureCitation id="19E0323EC9C257A891D2233215709979" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A, B Bayesian tree inferred from combined plastid (ndhA intron, rps 16 - trnK, rps 16 intron, rps 3, and rpl 32 - trnL) and ITS sequences. Thick branches indicate posterior probabilities of 0.95 - 1; species in bold occur in Central America; Scale bar: 2 %." figureDoi="10.3897/phytokeys.230.103882.figure1" httpUri="https://binary.pensoft.net/fig/887869" pageId="0" pageNumber="1">1</figureCitation>
;
<bibRefCitation id="9E8D83105A121CE4E990FEBAC2D3CBB5" DOI="https://doi.org/10.1111/jse.12805" author="Peterson, PM" journalOrPublisher="Journal of Systematics and Evolution" pageId="0" pageNumber="1" pagination="621 - 629" refId="B41" refString="Peterson, PM, Roquet, C, Romaschenko, K, Herrera Arrieta, Y, Susanna, A, 2021. A biogeographical analysis of Muhlenbergia (Poaceae: Chloridoideae: Cynodonteae: Muhlenbergiinae). Journal of Systematics and Evolution 60 (3): 621 - 629, DOI: https://doi.org/10.1111/jse.12805" title="A biogeographical analysis of Muhlenbergia (Poaceae: Chloridoideae: Cynodonteae: Muhlenbergiinae)." url="https://doi.org/10.1111/jse.12805" volume="60" year="2021">Peterson et al. 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="85EA4C4FAA646C02EEE7E75EA7FE1D14" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="36C481ACE86D766365A51BF889F4A81A" pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph id="31707EABC9B452DCABAA1DF19815A691" pageId="0" pageNumber="1">
<collectingCountry id="9E10C68E5EB79CBFF6D11575DF931521" name="Costa Rica">Costa Rica</collectingCountry>
.
<emphasis id="8DF5E13EFCD0E9E12FF1BA4B840E0D9E" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="6C2A249307C80ED3FD4F7E6090462B7D" country="Costa Rica" name="Alajuela">Alajuela</collectingRegion>
</emphasis>
: along road below Los Cartagos,
<quantity id="743084080C683775FD45F08867E777B0" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
above Carrizal, wet bank of roadside, in shade of herbs,
<emphasis id="5D0305308195DD979D2CC8D126B7BAB7" italics="true" pageId="0" pageNumber="1">R.W. Pohl &amp; G. Davidse 11735</emphasis>
(CR,
<collectingCountry id="21CDFA6F2AF84471C933E55966353D9C" name="United States of America">US</collectingCountry>
).
<emphasis id="AD1EB67C609955CDE6083793F815668C" bold="true" pageId="0" pageNumber="1">Cartago</emphasis>
: River crossing of
<normalizedToken id="F4B6D35BEC43979E48FD0D01C7827796" originalValue="Río">Rio</normalizedToken>
Reventado between Llano Grande and Tierra Blanca,
<emphasis id="F8A992F60FD7E40C5856BF4389BD6330" italics="true" pageId="0" pageNumber="1">R.W. Pohl &amp; M. Lucas 13097</emphasis>
(MO, CR); Cerro de La Carpintera, in potrero,
<emphasis id="ECB27DEE0F6A48227BACEB2F044CC322" italics="true" pageId="0" pageNumber="1">P.C. Standley 34502</emphasis>
(
<collectingCountry id="F57C29E32ABAC2DF74D3647CB25E4CDA" name="United States of America">US</collectingCountry>
).
<emphasis id="CBB37A9CEE8EB9F3C742F5D561BAE6F8" bold="true" pageId="0" pageNumber="1">
San
<normalizedToken id="0DA18220173764E928EBB1271EF580D0" originalValue="José">Jose</normalizedToken>
</emphasis>
: along Rio Maria Aguilar, near San
<normalizedToken id="F089EE42D56635E5A6D398482F596817" originalValue="José">Jose</normalizedToken>
,
<emphasis id="2AB73C8FF66F816C2A1A595BBFB097C6" italics="true" pageId="0" pageNumber="1">P.C. Standley 38976</emphasis>
(
<collectingCountry id="3F5F68E0271B352D554D4A4AD78DDB83" name="United States of America">US</collectingCountry>
); Vazquez de Coronado, Quebrada Corralillo,
<quantity id="EFE4D8600784FD30B96253E7EE3619C9" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
E of Rancho Redondo, moist mossy canyon walls in shade and along roadsides,
<emphasis id="6A75BF722D7A11D2F7BBD96DAE33C496" italics="true" pageId="0" pageNumber="1">R.W. Pohl &amp; G. Davidse 11701</emphasis>
(CR, MO,
<collectingCountry id="ED4FE59C7692F1B50847748FF8F46D5E" name="United States of America">US</collectingCountry>
); La Palma, wet forest,
<emphasis id="196941CA432B91E00B073A34D03DB17C" italics="true" pageId="0" pageNumber="1">P.C. Standley 33006</emphasis>
(
<collectingCountry id="6BD22135F4979E57E116C08B0AED52FD" name="United States of America">US</collectingCountry>
); San Gabriel, orillas de un arroyo,
<emphasis id="5EFF1093398AB669D7FACE827D28E719" italics="true" pageId="0" pageNumber="1">
O.
<normalizedToken id="754E923C8FE6B81105708D1927A41199" originalValue="Jiménez">Jimenez</normalizedToken>
173
</emphasis>
(
<collectingCountry id="525FDC06298BC091E85AFAF5A0D89828" name="United States of America">US</collectingCountry>
).
<collectingCountry id="B231C7D6E7658B664193B93F9967D020" name="El Salvador">El Salvador</collectingCountry>
.
<emphasis id="AB46ABE04EE41968B457855B52FC79A5" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="35C8F23FF9FB5AB2119B08A6D2FB38DD" country="El Salvador" name="La Libertad">La Libertad</collectingRegion>
</emphasis>
: Volcan San Salvador, near bottom of crater of
<normalizedToken id="99C2C864F844ECFD6A24A7738BAEE94F" originalValue="Volcán">Volcan</normalizedToken>
San Salvador,
<emphasis id="29CAEED7BF66C3B1D694256FD2F54ECF" italics="true" pageId="0" pageNumber="1">N.C. Fassett 28595</emphasis>
(MO, ITIC,
<collectingCountry id="BD0129F8265CF1DFDCF133A082B52F3F" name="United States of America">US</collectingCountry>
); Volcan San Salvador,
<normalizedToken id="0953AF300B55FBFCB707FAF92F5232B5" originalValue="Boquerón">Boqueron</normalizedToken>
de San Salvador,
<emphasis id="E1910FB4739F73C16985F34032F2E1FF" italics="true" pageId="0" pageNumber="1">
L.E.
<normalizedToken id="E487E14DB1AF82B3787CA82D632CF4B0" originalValue="González">Gonzalez</normalizedToken>
1852
</emphasis>
(MO, ITIC).
<emphasis id="FF4F7EB0961D484D9A9D14F28783A6D2" bold="true" pageId="0" pageNumber="1">San Salvador</emphasis>
: San Salvador,
<emphasis id="824CE4F357ABD992B7C0A5134BD991E7" italics="true" pageId="0" pageNumber="1">A.S. Hitchcock 8927</emphasis>
(
<collectingCountry id="84DBDA3ADD67C44F2244F82F09686058" name="United States of America">US</collectingCountry>
).
<emphasis id="679CC6E9DED7788B48FE8BCB15CF8043" bold="true" pageId="0" pageNumber="1">Sonsonate</emphasis>
:
<emphasis id="7A9E97A97CCB3F8C91B8E450F5B62DE6" italics="true" pageId="0" pageNumber="1">A. R. Molina et al. 21747</emphasis>
(LAGU).
<collectingCountry id="53D6C2F40704615E6E2B6B06A6B77F9B" name="Guatemala">Guatemala</collectingCountry>
.
<emphasis id="24DA5D48C6AADBE70BFEEA809AC42C5F" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="67D02853756BA43694BCB5FB0FCA6F30" country="Guatemala" name="Escuintla">Escuintla</collectingRegion>
</emphasis>
: San
<normalizedToken id="DD75C04F4FFF8323E412DAED4EC7996B" originalValue="José">Jose</normalizedToken>
,
<emphasis id="F2BA60361FEB7C9340272AFDD634D27A" italics="true" pageId="0" pageNumber="1">W. A. Kellerman 5115</emphasis>
(MO,
<collectingCountry id="ED4F037DE351C652F42DBD736BE3C931" name="United States of America">US</collectingCountry>
).
<emphasis id="05C1C21697751E0ED201F645806F6E47" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="4D636CD64F92750F1B5DD1685891229C" name="Guatemala">Guatemala</collectingCountry>
</emphasis>
: Near San Rafael,
<emphasis id="582942A3B66A7FDCBC05E57D14E91B28" italics="true" pageId="0" pageNumber="1">W.A. Kellerman 6239</emphasis>
(
<collectingCountry id="C3FAE018410159EFD3AD70E5FA4A64EF" name="United States of America">US</collectingCountry>
).
<emphasis id="890EEB8B04DD6C68787B9608927A7D96" bold="true" pageId="0" pageNumber="1">Huehuetenango</emphasis>
: Thickets and forest in deep canyon of tributary of Rio Blanco, about
<quantity id="BC74DE7881964388A779219C36301CF2" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
W of
<normalizedToken id="79B74B34E6B6723815C4DFAB5559C237" originalValue="Aguacatán">Aguacatan</normalizedToken>
,
<emphasis id="E39CD66595FCA02247789F114F584ED1" italics="true" pageId="0" pageNumber="1">L.O. Williams</emphasis>
,
<emphasis id="76DDC21BE2508928D8182AA3790712B3" italics="true" pageId="0" pageNumber="1">A. Molina R. &amp; T.P. Williams 22329</emphasis>
(
<collectingCountry id="B885C5FCB352513765E5C04E2CC6C7EA" name="United States of America">US</collectingCountry>
).
<emphasis id="5F1CB778941F4CCF5A7289FBE9D98A48" bold="true" pageId="0" pageNumber="1">Quetzaltenango</emphasis>
: Mountains near Santa Maria, just S of Quetzaltenango,
<emphasis id="D5008EAAAF26FCDA6712932CD6DD75F3" italics="true" pageId="0" pageNumber="1">Weatherwax 161 (1679)</emphasis>
(
<collectingCountry id="796BCE5199F45FDE1765A1D5B945FC28" name="United States of America">US</collectingCountry>
); Region of Las Nubes, south of San Martin
<collectingCountry id="DD341337BF2EE3386076C1FBF9F1812E" name="Chile">Chile</collectingCountry>
Verde, densely forested barranco,
<emphasis id="31C4C1EAAA20AA33380594429A4C0234" italics="true" pageId="0" pageNumber="1">P.C. Standley 83711</emphasis>
(
<collectingCountry id="56EB9657E99FFF86C3494367A74DCC65" name="United States of America">US</collectingCountry>
); Ravine below Fuentes Georginas, just above Zunil,
<emphasis id="DB4A9BA4A5D60F7B66C12E7287E1CA6F" italics="true" pageId="0" pageNumber="1">J.A. Steyermark 34480</emphasis>
(
<collectingCountry id="225FF05D75DAE19A48D6C69CDF1CA663" name="United States of America">US</collectingCountry>
); Retalhuleu,
<emphasis id="F5DB78C0EAB9E1BBC16D7A1D499FB52A" italics="true" pageId="0" pageNumber="1">M. De Koninck 226</emphasis>
(
<collectingCountry id="582EF575CF16FC024837D8484546BC08" name="United States of America">US</collectingCountry>
); Mountains above Rio Samala, Sierra Madre Mountains,
<quantity id="8661984B5FA6A77960ECD14FF7198E1F" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
west of Zunil,
<emphasis id="CCCD89F63B357211D1606A95BCEF0DAF" italics="true" pageId="0" pageNumber="1">L. O. Williams</emphasis>
,
<emphasis id="1E733943563F2F2A5BCD8701A92D2716" italics="true" pageId="0" pageNumber="1">A. Molina R. &amp; T.P. Williams 22987</emphasis>
(
<collectingCountry id="0CB80394D72A446D6439A733D8AAC141" name="United States of America">US</collectingCountry>
).
<emphasis id="DB061AEE3AD4C1CC7CFFB3C8F1FC8509" bold="true" pageId="0" pageNumber="1">Sacatepequez</emphasis>
: near Pastores, damp ravine,
<emphasis id="716E1BB45A66F3DEFD9AF65518F2FEE1" italics="true" pageId="0" pageNumber="1">P.C. Standley 59936</emphasis>
(
<collectingCountry id="187DA9B75B539A319A42410FB6E9AB8A" name="United States of America">US</collectingCountry>
); Volcano Agua, near
<collectingCountry id="CB725984287CEB3BD4BE7BA9938A7A79" name="Antigua and Barbuda">Antigua</collectingCountry>
, shady bank,
<emphasis id="780B7000ED6E4DA128B2D335FB9291D2" italics="true" pageId="0" pageNumber="1">A.S. Hitchcock 9134</emphasis>
(
<collectingCountry id="9851DEE22F0360C0B2B31E40180CBCED" name="United States of America">US</collectingCountry>
); Nacimiento del Cangrejal, cuesta de Las
<normalizedToken id="2D64BC0615CD96008FD29ECB2B66FFD8" originalValue="Cañas">Canas</normalizedToken>
,
<emphasis id="5B02A25E0A9C590CB8C624C5BB899F60" italics="true" pageId="0" pageNumber="1">A. Molina R. 15439</emphasis>
(
<collectingCountry id="B7F96B6A6B34D9781D31D501BBC581AB" name="United States of America">US</collectingCountry>
).
<emphasis id="BC9A016BF669C77768EA0F04A8A32546" bold="true" pageId="0" pageNumber="1">San Marcos</emphasis>
: Barrancos
<quantity id="48EB1ABA766E2877596E8EA285573C0F" metricMagnitude="3" metricUnit="m" metricValue="9.656064" unit="mi" value="6.0">6 mi</quantity>
south and west of town of Tajumulco, northwestern slopes of Volcan Tajumulco, moist thickets in quebrada,
<emphasis id="5FAA98115DF14A105697033BD50A1482" italics="true" pageId="0" pageNumber="1">J.A. Steyermark 36594</emphasis>
(
<collectingCountry id="E05E1DE0F4F33B43D2BEF6EEE3ACAA7D" name="United States of America">US</collectingCountry>
); Barranco Eminencia, above San Rafael Pie de la Cuesta, wet meadow,
<emphasis id="C730CCC3D56AC9CBA80C9C88B5A54D41" italics="true" pageId="0" pageNumber="1">P.C. Standley 68464</emphasis>
(
<collectingCountry id="6D9A100AAE50B27847F52CAD4358A929" name="United States of America">US</collectingCountry>
).
<emphasis id="318EBEFFFF97CFB2DA2D7C83D1B5EDC9" bold="true" pageId="0" pageNumber="1">Solola</emphasis>
: Mixed forest area, mountains slopes above Lake Atitlan, about
<quantity id="CE130EA2CD6D9A371AD3AF08FEADA69A" metricMagnitude="3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" unit="km" value="4.0" valueMax="5.0" valueMin="3.0">3-5 km</quantity>
west of Panajachel,
<emphasis id="39047C6239035AE0B03635900E0D92B2" italics="true" pageId="0" pageNumber="1">L.O. Williams</emphasis>
,
<emphasis id="2E5F30FB7EF81579FECCBE889174DEFC" italics="true" pageId="0" pageNumber="1">A. Molina R. &amp; T.P. Williams 25264</emphasis>
(
<collectingCountry id="C71D1F878607C9481BEA950889E085CD" name="United States of America">US</collectingCountry>
).
<collectingCountry id="5E21D891280E5DE8E6251A125E6BAE75" name="Mexico">Mexico</collectingCountry>
.
<emphasis id="35FF88BB8F6CD6717336E1D8991922D5" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="E09F5FAF5A421AD0573BAE578B398E6F" country="Mexico" name="Chiapas">Chiapas</collectingRegion>
. Motozintla
</emphasis>
: along road from Toilman to Niquivil, near Ojo de Agua,
<emphasis id="4AA39221EC5DA1FB06C7983AD52C4027" italics="true" pageId="0" pageNumber="1">D.E. Breedlove 42616</emphasis>
(MO).
<emphasis id="8CA39490825AE6E505BE45A40EEFFC34" bold="true" pageId="0" pageNumber="1">Siltepec</emphasis>
: On the ridge above Siltepec on the road to Huixtla,
<emphasis id="AF7D76DE2F2A8B3C70315D1CEAECC5DF" italics="true" pageId="0" pageNumber="1">D.E. Breedlove &amp; Almeda 58258</emphasis>
(CAS, MO).
<emphasis id="EAEC85104ADE770902C2067C93109868" bold="true" pageId="0" pageNumber="1">Tenejapa</emphasis>
: along trail from Tenejapa center to
<collectingCountry id="5DC023C443CEE276532B4848918F77FC" name="Saint Kitts and Nevis">
San
<normalizedToken id="0E15724F768ED6FF80D37C02F4E417AF" originalValue="Cristóbal">Cristobal</normalizedToken>
</collectingCountry>
de las Casas, in the paraje of Balum
<normalizedToken id="1009023A42A964537AB2CD9F5CC03EB6" originalValue="K">K'</normalizedToken>
anal,
<emphasis id="8FE19E5BA2ACAFFF6432FF6EF5E40ABF" italics="true" pageId="0" pageNumber="1">D.E. Breedlove 9350</emphasis>
(ENCB, DS,
<collectingCountry id="086B9A4F44A2FDCB5BF11F1F31D6688B" name="United States of America">US</collectingCountry>
); La Punta del Cerro Cruz
<normalizedToken id="1AE69A832D080E9F14B5DE1A17CD3B41" originalValue="Chen">Ch'en</normalizedToken>
,
<emphasis id="451E16D00A68C550CC10170A111ECBA1" italics="true" pageId="0" pageNumber="1">
A. Shilom
<normalizedToken id="C6B75595A61A09E369C6B052AE8BAEED" originalValue="Méndez">Mendez</normalizedToken>
Ton 5088
</emphasis>
(MEXU, MO).
<emphasis id="FA42E250C6803F2EF61B52006F3BC337" bold="true" pageId="0" pageNumber="1">
<normalizedToken id="3D067483829BDABE839D8D4DBA58E243" originalValue="Unión">Union</normalizedToken>
<normalizedToken id="8D5DF4AC20FE8FCA8A7CB877FBDE4BE4" originalValue="Juárez">Juarez</normalizedToken>
</emphasis>
: En el
<normalizedToken id="BF6D6B46B7080BFB0099859ED1E2B550" originalValue="volcán">volcan</normalizedToken>
<normalizedToken id="10AA9410664F134367827238A0CB4F99" originalValue="Tacaná">Tacana</normalizedToken>
a
<quantity id="02D1A528298C58AFBF934FA090D132B2" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</quantity>
al E de
<normalizedToken id="162D8AEF138BFEDC6CEDD1A2AC7A429A" originalValue="Talquián">Talquian</normalizedToken>
,
<emphasis id="D38CF0B495CA01905D19FB2098F5D2EC" italics="true" pageId="0" pageNumber="1">
E.M.
<normalizedToken id="EF193A41C2087AFA23E31270BF4B417A" originalValue="Martínez">Martinez</normalizedToken>
S. &amp; A
<normalizedToken id="4EDE221061A273C4DE29A101BA5D778F" originalValue="Reyes-García">Reyes-Garcia</normalizedToken>
20293
</emphasis>
(MO).
<collectingCountry id="C99BAE3076A598AAE3B700C3E3055C43" name="Panama">Panama</collectingCountry>
.
<emphasis id="ABACFD3B0F86163C872AA2ABBCADB497" bold="true" pageId="0" pageNumber="1">
<normalizedToken id="EA063E86930F63E1F153F1DED47AB103" originalValue="Chiriquí">
<collectingRegion id="2B3BA690577C9C67F8A5DF318C23B2BC" country="Panama" name="Chiriqui">Chiriqui</collectingRegion>
</normalizedToken>
</emphasis>
: Rio Caldera,
<quantity id="296742DD339CAF707645490EC7D487B8" metricMagnitude="3" metricUnit="m" metricValue="2.414016" metricValueMax="3.218688" metricValueMin="1.609344" unit="mi" value="1.5" valueMax="2.0" valueMin="1.0">1-2 mi</quantity>
above El Boquete,
<emphasis id="4C5175B35F641E4BA4F154DD3720CA4A" italics="true" pageId="0" pageNumber="1">E.P. Killip 4513</emphasis>
(
<collectingCountry id="35EB265C17099EFE2F211D1770FD54EC" name="United States of America">US</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>